taxonID	type	description	language	source
03C0A869FFD0FFDE838EFC5BFDF3193B.taxon	description	(Fig. 1) Description. Female. Body length 4.5 mm; fore wing length 3.3 mm. Head. Antennae almost filiform, more than 25 - segmented (apical segments missing). Scape 1.4 × as long as its maximum width. First flagellar segment 5.5 × as long as its apical width, as long as second segment. Head width (dorsal view) 1.7 × median length. Head behind eyes (dorsal view) roundly narrowed; transverse diameter of eye 1.9 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.3 × its sides; POL: Od: OOL = 6: 5: 12. Vertex and frons entirely densely aciculate. Eye glabrous, 1.4 × as high as broad. Malar space 0.4 × eye height and equal to basal width of mandible. Temple almost smooth, with very fine striation. Face with very dense, fine and absolutely even transverse aciculation (like surface of gramophone record or CD); its width 1.1 × eye height and 1.1 × height of face and clypeus combined. Clypeal suture absent. Hypoclypeal depression rather small and round, its width 0.6 × distance from edge of depression to eye, 0.3 × width of face. Occipital carina complete dorsally, not joined with hypostomal carina being obliterated upper base of mandible. Mesosoma. Length of mesosoma 2.0 × its height. Pronotal keel fine, its posterior branch widely fused with posterior margin of pronotum, anterior branch situated submedially on pronotum. Mesoscutum highly and roundly elevated above pronotum, its median lobe convex anteriorly and without anterolateral corners. Mesoscutum mostly glabrous, with rather sparse, more or less long and almost erect pale setae arranged narrow along notauli and laterally, densely and small reticulate-areolate with additional very fine granulation, lateral lobes of mesoscutum only granulate; with two distinct and convergent posteriorly carinae medioposteriorly. Notauli anteriorly distinct, wide, crenulate, very shallow in posterior half. Prescutellar depression deep, long, with four carinae, finely rugulose-reticulate, 0.4 × as long as scutellum. Scutellum entirely densely granulate, distinctly convex, with distinct lateral carinae. Mesopleuron weakly rugulose. Subalar depression very shallow, wide, finely reticulate-granulate and with sparse striae. Precoxal sulcus distinct, deep, weakly curved, crenulate, running along anterior half of lower part of mesopleuron. Propodeum with very short and wide lateral tubercles, with distinctly delineated by carinae and granulate basolateral areas, its rest part coarsely and transversely striate with rugosity; areola narrow and short; basal carina present in basal half, 3.0 × as long as anterior fork of areola. Wings. Fore wing 3.7 × as long as wide. Vein r arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 20: 5: 27: 16. Second submarginal cell not narrowed distally, its length 3.5 × maximum width, 1.45 × length of first subdiscal cell. Vein CU 1 a interstitial. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, weakly curved and strongly oblique towards base of wing. Legs. Hind coxa dorsally finely transversely striate, widely and densely granulate at rest part, with basoventral tooth and corner. Hind femur distinctly and densely granulate-coriaceous, smooth ventrally, 3.1 × as long as wide. Hind tibia apically without spines on outer side. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 0.9 – 1.0 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, as longer as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly curved ventrally, dorsally arched in basal third and straight in apical two-thirds, thickened in basal third; widened in apical quarter (dorsal view), with spiracular tubercles in basal third; distinctly reticulate-rugulose and with sparse striae in basal 0.7, finely striate apically. Length of petiole 3.0 × its apical width (dorsal view), 1.9 × length of propodeum. Second tergite with separate laterotergites only basally. Median length of second and third tergites combined 1.25 × basal width of second tergite, 0.7 × as long as their maximum width. Second suture absent. Second and third tergites densely reticulate-granulate, almost smooth narrowly apically. Fourth tergite densely reticulate-granulate, almost smooth narrowly apically. Remaining tergites smooth. Ovipositor straight. Ovipositor sheath 2.3 × as long as petiole, as long as metasoma, 1.5 × as long as mesosoma, 0.8 × as long as fore wing. Colour. Body yellowish brown, precoxal sulcus reddish brown. Antennae yellow brown. Palpi pale yellow. Legs pale yellowish brown, fore and middle coxae and all trochanters white, hind coxa white with infuscation; all tibiae white basally; hind leg brownish yellow. Ovipositor sheath brownish yellow in basal half and black in apical half. Fore wing faintly infuscate, with brown spots near vein 1 - SR, 1 - M and pterostigma. Pterostigma brown, pale in basal third. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFD0FFDE838EFC5BFDF3193B.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Jianfengling, 7. VI. 2007, Liu Jingxian, No. 200702556 (ZJUH). Comparative diagnosis. This new species belongs to the S. vulnificus Wilkinson species group. Spathius aciculatus sp. nov. is similar to S. paracritolaus Belokobylskij, but differs in having the petiole longer, its length 3.0 × apical width and 1.9 × length of propodeum (2.0 × its apical width and 1.5 × length of propodeum in S. paracritolaus); vein CU 1 a interstitial (not interstitial in S. paracritolaus); mesoscutum densely and small reticulate-areolate with additional very fine granulation (finely and densely reticulate in S. paracritolaus); vertex entirely densely aciculate (laterally smooth, finely aciculate only medially in S. paracritolaus); ovipositor long (short in S. paracritolaus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFD0FFDE838EFC5BFDF3193B.taxon	etymology	Etymology. From “ aciculus ” (Greek for “ aciculate ”), because its aciculate sculpture on vertex and frons.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFD0FFDE838EFC5BFDF3193B.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCEFFDF838EFA8DFDF31B46.taxon	description	(Fig. 2) Description. Female. Body length 5.0 mm; fore wing length 3.8 mm. Head. Antennae slender, weakly setiform, 36 - segmented. Scape 1.5 × as long as its maximum width. First flagellar segment 6.0 × as long as its apical width, 1.2 × as long as second segment. Head not depressed, its width (dorsal view) 1.3 × median length. Head behind eyes roundly narrowed. Transverse diameter of eye 1.7 × length of temple. Ocelli arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 5: 5: 16. Vertex mostly smooth, only finely striate medio-laterally. Frons densely and transversely striate. Eye glabrous, 1.2 × as high as broad. Malar space 0.5 × eye height and equal to basal width of mandible. Temple smooth. Face finely rugose-reticulate, with finely granulation between rugosity, smooth narrowly medially; its width 1.2 × eye height and 1.4 × height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.3 × its height. Pronotal keel distinct, its posterior branch situated closely to posterior margin of pronotum, but not fused with it; anterior branch fine and situated submedially. Mesoscutum rather highly and roundly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; densely granulate, with rather long and more or less distinct rugae near notauli and laterally; lateral lobes medially with wide granulate areas, with two convergent posteriorly carinae and narrow rugosity between them in medioposterior half. Notauli complete, wide, deep anteriorly and shallow posteriorly, coarsely crenulate. Prescutellar depression distinct, deep, with three carinae, 0.3 × as long as scutellum. Scutellum finely granulate, weakly convex, with distinct lateral carinae. Mesopleuron weakly coriaceous-granulate. Subalar depression shallow, wide, coarsely rugose-striate. Precoxal sulcus deep, sinuate, distinctly crenulate, running along entire length of lower part of mesopleuron. Propodeum with distinct tubercles, densely and coarsely rugose-reticulate, densely granulate in basal half, with areas more or less distinctly delineated by carinae; basal carina 1.5 × as long as anterior fork of areola; areola large and pentagonal; petiolate area distinctly separated from areola. Wings. Fore wing 4.0 × as long as wide. Vein r distinctly arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 25: 7: 42: 25. Second submarginal cell not narrowed distally, its length 3.5 × maximum width, 1.4 × length of first subdiscal cell. Vein cu-a weakly postfurcal. Vein m-cu postfurcal. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, weakly curved towards base of wing. Legs. Hind coxa transversely striate and granulate dorsally, coriaceous-granulate laterally and ventrally, with basoventral tooth and corner. Hind femur almost smooth, finely striate dorsally, 3.1 × as long as wide. Hind tibia apically with two spines on outer margin. Dorsal surface of hind tibia with rather long, sparse and semi-erect setae, their length 1.9 – 2.2 × maximum width of tibia. Hind tarsus 0.8 × as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Second segment of hind tarsus 0.4 × as long as basitarsus, 0.9 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) straight ventrally and distinct arched dorsally, with spiracular tubercles in basal third; sparsely striate in basal 0.7, densely striate in apically 0.3, with granulation between striae. Length of petiole 2.3 × its apical width (dorsal view), 1.5 × length of propodeum. Second tergite without separated laterotergites. Second suture indistinct. Median length of second and third tergites combined 1.4 × basal width of second tergite, 0.7 × as long as their maximum width. Second tergite densely but finely striate, smooth laterally; third tergite densely but finely striate in basal half, smooth lateral and in apical half; fourth tergite very dense semi-circularly anastomose striate, with very dense rugulosity between striae. Fifth tergite finely and sparsely coriaceous-reticulate basally, finely semi-circularly striate apically. Remaining tergites smooth. Ovipositor sheath 1.8 × as long as petiole, 0.7 × as long as metasoma, as long as mesosoma, 0.5 × as long as fore wing. Colour. Body pale yellow; vertex with brown patch in middle part of temples. Mesosoma dark reddish brown to almost black in lower part (lateral view). Petiole yellowish brown; second and third tergites mostly black with yellow narrow lateral (partly) and wide posterior areas, second tergite without yellow spots. Remaining tergites mainly black, but yellow laterally and narrowly apically. Antenna brownish yellow basally, yellowish brown to brown in median 0.3, five subapical segments pale yellow, last apical segment brown. Palpi pale yellow. Middle femur brown subapically; hind femur dark brown in apical half. Ovipositor sheath brown, but medially widely pale brown. Fore wing faintly infuscate, with faint maculation. Pterostigma brown, yellow in basal 0.25 – 0.30 and apically. Variation. Body length 5.4 mm; fore wing length 3.9 mm. First flagellar segment 5.8 × as long as its apical width. Transverse diameter of eye 1.8 × length of temple. Mesosoma 2.2 × as long as its height. Fore wing 3.9 × as long as maximum width. Hind femur 3.2 × as long as wide. Length of petiole 2.4 × its apical width (dorsal view). Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCEFFDF838EFA8DFDF31B46.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Jianfengling, 6. VI. 2007, Liu Jingxian, No. 200703850 (ZJUH). Paratype: 1 female, China, Hainan Prov., Jianfengling, 4 – 7. VI. 2007, Zeng Jie, No. 200711138 (ZJUH). Comparative diagnosis. This new species belongs to the S. moderabilis Wilkinson species group. Spathius albithorax sp. nov. is similar to S. moderabilis Wilkinson, but differs in having the second and third tergites striate (finely punctate-reticulate basally in S. moderabilis), ovipositor sheath shorter than metasoma (distinctly longer than metasoma and as long as body in S. moderabilis), mesoscutum roundly elevated above pronotum (perpendicularly elevated in S. moderabilis), and pronotal keel posterior branch not fused with posterior margin of pronotum (fused in S. moderabilis).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCEFFDF838EFA8DFDF31B46.taxon	etymology	Etymology. From “ albus ” (Latin for “ white ”) and “ thorax ” (Latin for “ thorax ”) because its mesosoma is white.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCEFFDF838EFA8DFDF31B46.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCAFFDA838EFF45FDCE1DB4.taxon	description	(Fig. 3)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCAFFDA838EFF45FDCE1DB4.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Yunnan Prov., Xishuangbanna, 17. VIII. 1958, Wang Shuyong. Distribution. China (Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCAFFDA838EFE07FC9C1F30.taxon	materials_examined	Material examined. China (ZJUH). Zhejiang Prov.: 1 female, Suichang Jiulongshan, 18. VIII. 1994, He Junhua, No. 944072; 1 female, Kaihua Gutianshan, VII – VIII. 1990, Ma Yun, No. 905982; 1 female, Gutianshan, 18. VII. 1992, Ma Yun, No. 923885; 1 female, Qingyuan Baishanzu, 20. VII. 1994, Wu Hong, No. 946944; Fujian Prov.: 1 female, Guadun, 7. X. 1991, Chen Xuexin, No. 920401.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCAFFDA838EFE07FC9C1F30.taxon	distribution	Distribution. China (Zhejiang, Fujian) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCAFFDB838EFC88FD421AD6.taxon	description	(Fig. 4) Description. Female. Body length 5.9 mm; fore wing length 3.8 mm. Head. Antennae slender, filiform, 43 - segmented. Scape 1.4 × as long as its maximum width. First flagellar segment 5.3 × as long as its apical width, 1.1 × as long as second segment. Head width (dorsal view) 1.4 × median length. Head behind eyes (dorsal view) roundly narrowed; transverse diameter of eye 1.7 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 6: 5: 14. Vertex almost smooth, only finely striate anteriorly. Frons densely and transversely striate. Eye glabrous, 1.3 × as high as broad. Malar space 0.4 × eye height and 0.9 × basal width of mandible. Temple smooth. Face almost entirely transversely rugose-reticulate, clypeus rugose-striate; its width equal to eye height and 1.1 × height of face and clypeus combined. Clypeal suture distinct, but shallow, without additional carina. Clypeus flat, with narrow ventral flange. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.4 × its height. Pronotal keel situated on middle of pronotum, its posterior branch distinctly separated from posterior margin of pronotum. Mesoscutum not highly and roundly elevated above pronotum, its median lobe protruding forwards, convex-rounded anteriorly (dorsal view), without anterolateral corners; with sparse long almost erect setae situated widely along notauli and narrowly marginally; densely granulate with additional dense long rugosity. Notauli deep anteriorly and shallow posteriorly, wide, complete, coarsely crenulate. Prescutellar depression shallow, with three carinae, finely rugulose, 0.4 × as long as scutellum. Scutellum granulate-coriaceous, weakly convex, with lateral carinae. Mesopleuron entirely coarsely and curvedly rugose-striate. Subalar depression shallow, wide, coarsely and sparsely rugose-striate. Precoxal sulcus deep and wide in anterior 0.7, shallow and narrow in posterior 0.3, weakly curved, coarsely crenulate, running along almost entire lower part of mesopleuron. Propodeum without tubercles, mostly coarsely rugulose-reticulate, its basolateral areas granulate-punctate in basal half and coarsely rugose in posterior half; basal carina long, 2.6 × as long as anterior fork of areola; areola unclear delineated by carinae. Wings. Fore wing 4.7 × as long as wide. Vein r arising before middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 12: 3: 20: 10. Second submarginal cell not narrowed distally, its length 3.1 × maximum width, 1.3 × length of first subdiscal cell. Vein m-cu postfurcal. Vein CU 1 a postfurcal. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, strongly oblique towards base of wing. Legs. Hind coxa dorsally transversely and rather finely semi-circularly striate, almost entirely finely granulate in lower half, without basoventral tooth and corner. Hind femur almost smooth, elongate-oval, 3.5 × as long as wide. Hind tibia apically with four spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 0.8 – 1.0 × maximum width of tibia. Hind tarsus 0.8 × as long as hind tibia. Hind basitarsus 0.8 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 0.9 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) almost straight, more or less evenly thick, weakly widened in apical 0.2 (dorsal view), with small spiracular tubercles in basal third; sparsely and finely striate, finely coriaceous and granulate between striae in basal third, only finely granulate-coriaceous on median third, and densely striate in apical third. Length of petiole 8.6 × its apical width (dorsal view), 3.1 × length of propodeum. Second and third tergites with separate laterotergites. Second suture absent. Median length of second and third tergites combined 2.9 × basal width of second tergite, 1.4 their maximum width. Second and third tergites with very fine aciculation. Remaining tergites smooth. Ovipositor sheath 3.1 × as long as petiole, 1.5 × as long as metasoma, 3.5 × as long as mesosoma, 1.6 × as long as fore wing. Colour. Head mostly pale yellowish brown, face pale brown, with dark area on vertex and temple. Mesosoma mostly black, mesoscutum yellowish brown, mesopleuron with a small yellow spot. Petiole black; second and third tergites mostly black, yellow apically; remaining tergites widely dark medially, yellow laterally. Antenna brownish yellow, faintly infuscate towards apex. Palpi pale yellow. Legs pale yellow; fore and middle femora faintly darkened subapically, hind femur widely black; all tibiae faintly infuscate submedially. Ovipositor sheath brown or pale brown, almost black apically. Fore wing faintly infuscate. Pterostigma brown, basally and apically yellow. Variation. Body length 4.2 – 7.0 mm; fore wing length 2.7 – 4.5 mm. First flagellar segment 5.0 – 5.5 × as long as its apical width, 1.2 × as long as second segment. Transverse diameter of eye 1.8 × length of temple. Ocelli arranged in triangle with base 1.3 × its sides. Mesosoma 2.2 × as long as its height. Fore wing 4.5 – 4.8 × as long as maximum width. Length of second submarginal cell 3.0 – 3.2 × maximum width, 1.2 × length of first subdiscal cell. Body paler, mesosoma without dark areas. Male. Body length 3.8 – 4.2 mm; fore wing length 2.5 – 2.7 mm. Hind femur 3.8 × as long as wide. Length of petiole 7.8 – 8.8 × its apical width (dorsal view), 3.0 – 3.2 × length of propodeum. Median length of second and third tergites combined 2.8 – 3.0 × basal width of second tergite, 1.3 – 1.4 × their maximum width. Otherwise similar to female.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCAFFDB838EFC88FD421AD6.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Jianfengling, 6. VI. 2007, Liu Jingxian, No. 200703766 (ZJUH). Paratypes: China: 1 female, Hainan Prov., Diaoluoshan, 29. V – 2. VI. 2007, Xiao Bin, No. 200804562 (ZJUH); 1 female, Hainan Prov., Bawangling, 9 – 10. VI. 2007, Liu Jingxian, No. 200703548 (ZJUH); 1 female, Hainan Prov., Jianfengling Tianchi, 12 – 15. VII. 2006, Chen Tianfei, No. 200803208 (ZJUH); 1 female, Hainan Prov., Jianfengling Tianchi, 12 – 15. VII. 2006, Liu Jingxian, No. 200803659 (ZISP); 1 male, Hainan Prov., Jianfengling Tianchi, 25. XI. 2008, Tan Jiangli, No. 200805087 (ZJUH); 1 female, Hainan Prov., Jianfengling, 22 – 23. XI. 2008, Wang Manman, No. 200805934 (ZJUH); 1 male, Hainan Prov., Jianfengling, 25. XI. 2008, Wang Manman, No. 200806096 (ZJUH); 1 male, Hainan Prov., Yinggeling, 24 – 25. V. 2007, Liu Jingxian, No. 200702624 (ZJUH); 1 female, Guangdong Prov., Shixing Chebaling, 15. VII. 2003, Xu Zaifu, No. 20058750 (ZJUH); 1 female, Guangdong Prov., Shixing Chebaling, 21. VIII. 2003, Xu Zaifu, No. 20052432 (ZJUH). Comparative diagnosis. This new species belongs to the S. imbecillus Enderlein species group. S. angustalatus sp. nov. is similar to S. festus Nixon, but differs in having the mesoscutum widely densely granulate with additional dense long rugosity (fine sculpture at most part in S. festus), areola of propodeum unclear (distinctly delineated by carinae in S. festus), mesopleuron entirely coarsely and curvedly rugose-striate (mostly smooth, only with faintly sculpture at discal of mesopleuron in S. festus), vein m-cu of fore wing postfurcal (almost interstitial in S. festus), hind coxa without basoventral tooth and corner (with basoventral tooth and corner in S. festus), most part of mesosoma, metasoma and hind femur widely black or dark (mainly pale brown in S. festus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCAFFDB838EFC88FD421AD6.taxon	etymology	Etymology. From “ angustus ” (Latin for “ narrow ”) and “ ala ” (Latin for “ wing ”), because its narrow wings.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCAFFDB838EFC88FD421AD6.taxon	distribution	Distribution. China (Hainan, Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCBFFD6838EF8E6FE521DB1.taxon	materials_examined	Material examined. China (ZJUH). Guangdong Prov.: 1 female, Fengkai, 16. V. 1992, Chen Xuexin, No. 921587. Hainan Prov.: 1 female, Bawangling, 26. XI. 2008, Wang Manman, No. 200805631; 1 female, Jianfengling, 6. VI. 2007, Liu Jingxian, No. 200703731; 1 female, Wuzhishan, 16 – 20. V. 2007, Liu Jingxian, No. 200703284; 2 females, Yinggeling, 28. V – 3. VI. 2007, Weng Liqiong, Nos. 200804067, 200804196. Yunnan Prov.: 1 female, Xishuangbanna, 31. VII. 2003, Jiang Qian, No. 20045892.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFCBFFD6838EF8E6FE521DB1.taxon	distribution	Distribution. China (Guangdong, Hainan, Yunnan, Taiwan); India, Malaysia, Papua New Guinea, Philippines, Thailand, Vietnam, Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC6FFD6838EFE0EFF0A1FC9.taxon	materials_examined	Material examined. Female (holotype of Spathius crebristriatus Chao, IZCAS), Fujian Prov., Jianyang Huangkenggongshe Aotou, 20. VII. 1965, She Rijian. Female (holotype of Spathius mundus Chao, IZCAS), Fujian Prov., Meifeng, 7. V. 1966. China (ZJUH). Hainan Prov.: 1 female, Bawangling, 26. XI. 2008, Wang Manman, No. 200805636; 2 females, Yinggeling Hongmaocun, 23 – 25. V. 2007, Zeng Jie, Nos. 200804510 and 200804471.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC6FFD6838EFE0EFF0A1FC9.taxon	distribution	Distribution. China (Hainan); Indonesia, Malaysia, Taiwan, Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC6FFD6838EFE0EFF0A1FC9.taxon	discussion	Remarks. Spathius crebristriatus Chao and S. mundus Chao were synonymized as S. mimeticus (Enderlein) by Chen & Shi (2004), but S. crebristriatus and S. mundus have their mesoscutum highly and vertically elevated above pronotum, median lobe of mesoscutum (dorsal view) almost straight anteriorly and with distinct anterolateral corners and vertex with steep and posteriorly declivous surface which are identical to S. araeceri Nixon.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC6FFD6838EFB80FA951B89.taxon	materials_examined	Material examined. Female (Holotype, IZCAS), Fujian Prov., Dehua, 3 – 4. XI. 1974, Chen Jiahua. China (ZJUH). Guangdong Prov.: 1 female, Longmen Nankunshan, 14 – 15. VII. 2003, Xu Zaifu, No. 20053752; 1 female, 1 male, Meizhou Fengxi, 29. VII. 2003, Chen Jujian, Nos. 20048697 and 20048847; 1 female, Shixing Chebaling, 26. VII. 2008, Wen Min, No. 200809154. Hainan Prov.: 2 females, Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, Nos. 200802158 and 200802268; 1 female, Diaoluoshan, 28 – 31. V. 2007, Liu Jingxian, No. 200702958; 1 male, Diaoluoshan, 16 – 17. VII. 2006, Weng Liqiong, No. 200802468; 1 female, 2 males, Jianfengling, 6. VI. 2007, Liu Jingxian, Nos. 200703765, 200703788 and 200703797; 1 female, 3 males, Jianfengling, 7. VI. 2007, Liu Jingxian, Nos. 200702430, 200702447, 200702475 and 200702561; 1 female, Jianfengling, 5 – 7. VI. 2007, Xiao Bin, No. 200806894; 2 females, 2 males, Jianfengling, 4 – 7. VI. 2007, Zeng Jie, Nos. 200711023, 200711112, 200710909 and 200710925; 1 male, Jianfengling Tianchi, 12 – 15. VII. 2006, Chen Tianfei, No. 200803222; 1 female, Jianfengling Tianchi, 22 – 23. X. 2007, Liu Jingxian, No. 200710713; 1 male, Jianfengling Tianchi, 12 – 15. VII. 2007, Liu Jingxian, No. 200803587; 1 female, 1 male, Jianfengling Tianchi, 24. XI. 2008, Tan Jiangli, Nos. 200806012 and 200806045; 1 female, Jianfengling Tianchi, 25. XI. 2008, Wang Manman, No. 200806074; 1 female, Jianfengling Tianchi, 12 – 15. VII. 2006, Weng Liqiong, No. 200803404; 2 females, 1 male, Jianfengling Tianchi, 12 – 15. VII. 2006, Zhang Wenyong, Nos. 200803416, 200803421 and 200803479; 1 female, Wuzhishan Shuimanxiang, 16 – 18. V. 2007, Zeng Jie, No. 200807543; 1 female, Yinggeling, 17. XI. 2008, Wang Manman, No. 200805405. Yunnan Prov.: 2 males, Xishuangbanna, 31. VII. 2003, Xu Zaifu, Nos. 20053527 and 20053546; 1 male, Baoshan Gaoligongshan, 16 – 17. VII. 2006, Zeng Jie, No. 200802050.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC6FFD6838EFB80FA951B89.taxon	distribution	Distribution. China (Fujian, Guangdong, Hainan, Yunnan); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC6FFD6838EFB80FA951B89.taxon	discussion	Remarks. Record of this species in the fauna of the Russian Far East was erroneous (Belokobylskij, 2009).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC7FFD4838EFF45FD421EB3.taxon	description	(Fig. 5) Description. Female. Body length 4.2 mm; fore wing length 3.3 mm. Head. Antennae slender, weakly setiform, more than 31 - segmented (apical segments missing). Scape 1.6 × as long as its maximum width. First flagellar segment 5.7 × as long as its apical width, 1.4 × as long as second segment. Head not depressed, its width (dorsal view) 1.3 × median length. Head behind eyes roundly narrowed. Transverse diameter of eye 1.8 × length of temple. Ocelli arranged in triangle with base 1.3 × as long as its sides; POL: Od: OOL = 6: 4: 18. Vertex and frons densely coarsely rugose-striate, with rugulosity between striae. Eye glabrous, 1.5 × as high as broad. Malar space 0.3 × eye height and 0.8 × basal width of mandible. Temple distinctly coarsely rugose-striate, with rugulosity between striae. Face densely coarsely rugose-reticulate; its width equal to eye height and 1.2 × height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.4 × its height. Pronotal keel distinct, its posterior branch widely fused with posterior margin of pronotum; anterior branch distinct, situated in posterior third of pronotum. Mesoscutum highly and roundly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; densely irregularly rugose-striate with granulation, with rather long and more or less distinct rugae near notauli and laterally; lateral lobes medially with wide granulate areas, with two convergent posteriorly carinae and narrow rugosity between them in medioposterior half. Notauli complete, wide, shallow, densely crenulate. Prescutellar depression shallow, with five carinae, 0.3 × as long as scutellum. Scutellum densely granulate, flat, with distinct lateral carinae. Mesopleuron entirely granulate and irregularly rugose-striate. Subalar depression shallow, wide, coarsely rugose-striate and granulate. Precoxal sulcus shallow, narrow, straight, distinctly crenulate and granulate, running along anterior 0.6 of lower part of mesopleuron. Propodeum without distinct tubercles, distinctly granulate in basal half, without areas delineated by carinae; basal carina 1.3 × as long as anterior fork of areola; areola narrow and long; petiolate area indistinct. Wings. Fore wing 3.8 × as long as wide. Vein r arising from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 22: 6: 30: 17. Second submarginal cell not narrowed distally, its length 3.0 × maximum width, 1.2 × length of first subdiscal cell. Vein cu-a weakly postfurcal. Vein m-cu postfurcal. Vein CU 1 a interstitial. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, straight, oblique towards base of wing. Legs. Hind coxa densely granulate, finely striate dorsally, without basoventral tooth and corner. Hind femur coriaceous, 2.6 × as long as wide. Hind tibia apically with five spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 0.8 – 1.2 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 0.9 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) almost straight ventrally and distinctly arched in basal third dorsally, with spiracular tubercles in basal third; entirely rugose-striate, with rugulosity and granulation between striae. Length of petiole 3.2 × its apical width (dorsal view), 1.5 × length of propodeum. Second and third tergites with separated laterotergites. Second suture absent. Median length of second and third tergites combined 1.9 × basal width of second tergite, 0.8 × as long as their maximum width. Second and third tergites rather densely and small reticulate-areolate. Remaining tergites reticulate-coriaceous, narrowly smooth apically. Ovipositor sheath 3.1 × as long as petiole, 1.2 × as long as metasoma, 1.7 × as long as mesosoma, 0.8 × as long as fore wing. Colour. Body mainly yellowish brown, but mesosoma reddish brown. Antenna brownish yellow basally, darken toward apex. Palpi pale yellow. Legs yellowish brown, hind leg mostly reddish brown; fore tarsus dark brown. Ovipositor sheath brown, almost black apically. Fore wing faintly infuscate, with distinct maculation. Pterostigma brown, pale in basal 0.3. Variation. Body length 3.8 – 4.6 mm; fore wing length 3.0 – 3.4 mm. Scape 1.5 – 1.8 × as long as its maximum width. First flagellar segment 5.6 – 6.0 × as long as its apical width, 1.3 – 1.5 × as long as second segment. Eye glabrous, 1.3 × as high as broad. Mesosoma 2.2 × as long as its height. Hind femur 2.8 × as long as wide. Median length of second and third tergites combined 1.8 × basal width of second tergite, 0.7 × as long as their maximum width. Male. Body length 4.0 mm; fore wing length 3.2 mm. Scape 1.7 × as long as its maximum width. First flagellar segment 6.0 × as long as its apical width, 1.5 × as long as second segment. Eye glabrous, 1.4 × as high as broad. Prescutellar depression with three carinae. Fore wing 3.6 × as long as wide. Hind femur 3.0 × as long as wide. Length of petiole 3.5 × its apical width (dorsal view), 2.0 × length of propodeum. Otherwise similar to female.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC7FFD4838EFF45FD421EB3.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Jianfengling, 22. XI. 2008, Tan Jiangli, No. 200805315 (ZJUH). Paratypes: 1 female, China, Hainan Prov., Jianfengling, 5 – 7. VI. 2007, Weng Liqiong, No. 200806451 (ZJUH); 1 male, Hainan Prov., Wuzhishan, 16 – 20. V. 2007, Liu Jingxian, No. 200703196 (ZJUH); 1 female, Guangdong Prov., Zijin Qimuzhang, 31. VII. 2003, Song Yanxia, No. 20059059 (ZJUH). Comparative diagnosis. This new species belongs to the S. spilopterus Cameron species group. Spathius aspratiloides sp. nov. is similar to S. aspratilis Chen et Shi, but differs in having the mesoscutum densely and irregularly rugose-striate with granulation and with rather long and more or less distinct rugae near notauli and laterally (only granulate in S. aspratilis), mesopleuron entirely granulate and rugose (only granulate in S. aspratilis), and ovipositor sheath longer than metasoma (shorter in S. aspratilis).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC7FFD4838EFF45FD421EB3.taxon	etymology	Etymology. From “ spiratilus ”, the name of the most similar species, and suffix “ oides ” because this species resemble S. aspiratilus Chen et Shi.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC7FFD4838EFF45FD421EB3.taxon	distribution	Distribution. China (Hainan, Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC4FFD2838EFD05FD421803.taxon	description	(Fig. 6) Description. Female. Body length 4.0 mm; fore wing length 3.0 mm. Head. Antennae slender, weakly setiform, 23 - segmented. Scape 1.3 × as long as its maximum width. First flagellar segment 4.7 × as long as its apical width, 1.2 × as long as second segment. Head not depressed, its width (dorsal view) 1.4 × median length. Head behind eyes roundly narrowed. Transverse diameter of eye 1.7 × length of temple. Ocelli arranged in triangle with base 1.3 × as long as its sides; POL: Od: OOL = 12: 7: 30. Vertex densely striate, finely granulate laterally. Frons densely coarsely striate. Eye glabrous, 1.3 × as high as broad. Malar space 0.4 × eye height and 0.8 × basal width of mandible. Temple finely rugose-striate. Face densely and finely rugose-reticulate, with granulation between rugulosity; its width equal to eye height and 1.2 × height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 1.9 × its height. Pronotal keel distinct, its posterior branch widely fused with posterior margin of pronotum; anterior branch situated submedially on pronotum. Mesoscutum highly and roundly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; densely granulate, with two convergent posteriorly carinae and narrow rugosity between them in medioposterior half. Notauli complete, wide, shallow, densely crenulate. Prescutellar depression shallow, with six carinae, 0.3 × as long as scutellum. Scutellum densely granulate, flat, with distinct lateral carinae. Mesopleuron entirely coriaceous-granulate. Subalar depression shallow, wide, very sparsely rugose-striate. Precoxal sulcus shallow, wide, straight, distinctly crenulate and granulate, running along anterior 0.6 of lower part of mesopleuron. Propodeum with distinct tubercles, distinctly granulate in basal half, with areas distinctly delineated by carinae; basal carina 0.7 × as long as anterior fork of areola; areola narrow and long. Wings. Fore wing 3.8 × as long as wide. Vein r arising weakly before middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 20: 6: 28: 18. Second submarginal cell not narrowed distally, its length 3.5 × maximum width, 1.5 × length of first subdiscal cell. Vein cu-a postfurcal. Vein m-cu postfurcal. Vein CU 1 a not interstitial, arising from anterior 0.3 of distal vein of subdiscal cell. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, weakly curved towards base of wing. Legs. Hind coxa densely granulate, finely striate dorsally, with basoventral tooth and corner. Hind femur coriaceous, 3.0 × as long as wide. Hind tibia apically with five spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 0.8 – 0.9 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 0.8 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) straight ventrally and distinct arched in basal third dorsally, with spiracular tubercles in basal third; densely striate, with rugulosity between striae. Second tergite entirely and third in basal half with separate laterotergites. Length of petiole 1.9 × its apical width (dorsal view), 1.6 × length of propodeum. Second suture absent. Median length of second and third tergites combined 1.3 × basal width of second tergite, 0.8 × as long as their maximum width. Second and third tergites rather densely and small reticulate-areolate. Remaining tergites reticulate-coriaceous, smooth apically on narrow area. Ovipositor sheath 1.8 × as long as petiole, 0.6 × as long as metasoma, as long as mesosoma, 0.4 × as long as fore wing. Colour. Body yellowish brown; scutellum dark brown; mesopleuron reddish brown. Petiole dark reddish brown; remaining tergites dark brown. Antenna brownish yellow basally, darken toward apex. Palpi pale yellow. Legs pale yellow; hind femur dark brown in apical two-thirds; hind tibia in apical third and all fifth segments of tarsi brown. Ovipositor sheath brown, almost black apically. Fore wing faintly infuscate, with distinct maculation. Pterostigma brown, pale in basal 0.3. Variation. Body length 2.9 – 3.8 mm; fore wing length 2.4 – 2.9 mm. First flagellar segment 4.6 – 5.0 × as long as its apical width, 1.3 – 1.5 × as long as second segment. Transverse diameter of eye 1.8 × length of temple. Mesosoma 1.8 – 2.0 × as long as its height. Prescutellar depression with five carinae. Basal carina of propodeum 0.8 × as long as anterior fork of areola. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC4FFD2838EFD05FD421803.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Jianfengling, 7. VI. 2007, Liu Jingxian, No. 200702543 (ZJUH). Paratypes: 1 female, China, Hainan Prov., Jianfengling, 7. VI. 2007, Liu Jingxian, No. 200702566 (ZJUH); 1 female, Hainan Prov., Jianfengling Tianchi, 12 – 15. VII. 2006, Weng Liqiong, No. 200803262 (ZJUH); 1 female, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802192 (ZJUH); 1 female, Guangdong Prov., Yunan Tongledashan, 12 – 13. VIII. 2003, Xu Zaifu, No. 20054691 (ZJUH). Comparative diagnosis. This new species belongs to the S. spilopterus Cameron species group. Spathius basalis sp. nov. is similar to S. decebalus Nixon, but differs in having the hind coxa with basoventral tooth and corner (without corner and tooth in S. decebalus), palpi pale yellow (darkened in S. decebalus), hind tibia in apical third brown and pale yellow on remaining part (entirely dark in S. decebalus), and antenna distinctly darkened toward apex (hardly darken apically in S. decebalus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC4FFD2838EFD05FD421803.taxon	etymology	Etymology. From “ basalis ” (Latin for “ basal ”), because of its hind coxa with basoventral tooth and corner.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC4FFD2838EFD05FD421803.taxon	distribution	Distribution. China (Hainan, Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC2FFD3838EFB55FD6F1B46.taxon	description	(Fig. 7) Description. Female. Body length 3.5 mm; fore wing length 2.4 mm. Head. Antennae filiform, 30 - segmented. Scape 1.7 × as long as its maximum width. First flagellar segment 5.0 × as long as its apical width, 1.2 × as long as second segment. Penultimate segment 4.0 × as long as wide, about as long as pointed apical segment. Head width (dorsal view) 1.5 × its median length, 1.3 × as wide as mesoscutum. Head behind eyes (dorsal view) distinctly and roundly narrowed; transverse diameter of eye 1.8 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 3.0: 2.0: 6.0. Vertex almost entirely smooth; frons entirely, coarsely, densely and curvedly transversely striate. Eye glabrous, 1.2 × as high as broad. Malar space 0.4 × eye height and 0.9 × basal width of mandible. Temple mainly smooth. Face densely, distinctly and weakly curvedly transversely striate with additional very fine reticulation. Face width 1.1 × eye height and 1.1 × as long as height of face and clypeus combined. Hypoclypeal depression medium-sized and round, its width 0.9 × distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, ventrally not joined with hypostomal carina being obliterated upper base of mandible. Mesosoma. Length of mesosoma about 2.0 × its height. Pronotal keel distinct, its posterior branch coarse, distinctly separated from posterior margin of pronotum; anterior branch of carina fine and situated submedially. Mesoscutum highly and roundly elevated above pronotum, its median lobe weakly convex anteriorly and without anterolateral corners. Mesoscutum distinctly and densely granulate, its median lobe anteriorly with granules partly situated in fine transverse lines, without rugae along notauli and laterally, with two distinct and weakly convergent posteriorly straight carinae in medioposterior half; mostly glabrous, with sparse, long and semi-erect pale setae arranged in wide stripe along notauli and in single line laterally. Notauli distinct, wide, coarsely densely crenulate, deep anteriorly and shallow posteriorly. Prescutellar depression rather deep, long, with three carinae, finely rugulose, 0.4 × as long as scutellum. Scutellum entirely densely and distinctly granulate, without rugae, almost flat, with lateral carinae. Mesopleuron finely or partly distinctly striate, smooth in some small areas. Subalar depression shallow, wide, densely and coarsely striate with rugulosity partly. Precoxal sulcus deep, wide and oblique in anterior 0.5, shallow and straight in posterior 0.5, sparsely and distinctly crenulate and partly with granulation, running along entire lower part of mesopleuron. Propodeum with distinct and rather wide lateral tubercles, with basolateral areas distinctly delineated by carinae and densely granulate with long and coarse rugosity, rest part of propodeum densely striate; areola rather narrow and long; basal carina present in basal 0.25, 0.8 × as long as anterior fork of areola. Wings. Fore wing 3.8 × as long as wide. Vein r arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 14: 4: 32: 14. Second submarginal cell not narrowed distally, its length 3.0 × maximum width, 1.5 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior third of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.7 × as long as vein 1 - M. Vein m-cu weakly antefurcal, strongly oblique towards base of wing, almost hyaline. Legs. Hind coxa dorsally transversely striate with dense distinct granulation, distinctly densely granulate laterally, but finer below, with distinct basoventral corner and tooth. Hind femur densely and finely granulate with fine striation dorsally, almost smooth on rest part, 3.3 × as long as wide. Hind tibia apically with two slender spines on outer side. Dorsal surface of hind tibia with long, dense and semi-erect setae, their length 0.8 – 1.3 × maximum width of tibia. Hind tarsus as long as hind tibia. Hind basitarsus 0.5 × as long as second-fifth segments combined. Second segment of hind tarsus 0.6 × as long as basitarsus, 1.2 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) curved ventrally, dorsally rather distinctly arched in basal 0.5 and almost straight in apical 0.5, thickened in basal 0.3; widened in apical fifth (dorsal view), with spiracular tubercles in weakly widened basal third. Length of petiole 2.9 × its apical width, 1.6 × length of propodeum. Second tergite without separate laterotergites. Median length of second and third tergites combined 1.8 × basal width of second tergite, 0.8 × as long as their maximum width. Second suture absent. Petiole distinctly and rather densely longitudinally striate, with additional dense reticulation between striae. Remaining tergites entirely smooth. Ovipositor straight. Ovipositor sheath 1.3 × as long as petiole, 0.6 × as long as metasoma, 0.9 × as long as mesosoma, 0.45 × as long as fore wing. Colour. Head light reddish brown, reddish brown on face and posteriorly. Mesosoma and metasoma reddish brown with several pale spots and stripes. Antennae brown to reddish brown, two basal segments yellow, nine subapical segments whitish yellow, three last apical segment brown. Palpi whitish. Legs brownish yellow, fore and middle coxae and trochanters pale yellow, hind coxa reddish brown, but paler apically, hind femur brown to reddish brown, paler in basal quarter and apically, hind tibia yellow to brownish yellow, pale yellow in subbasal short area. Ovipositor sheath yellow to brownish yellow in basal 0.3 and dark brown in apical 0.7. Fore wing evenly faintly infuscate. Pterostigma brown, pale yellow in basal quarter. Variation. Body length 2.8 mm; fore wing length 2.0 mm. Mesoscutum with fine and short rugae near notauli. Mesopleuron smooth on wide median area. Length of petiole 3.2 × its apical width. Head and mesosoma paler. Otherwise similar to female. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC2FFD3838EFB55FD6F1B46.taxon	materials_examined	Material examined. Holotype. female, China, Fujian Prov., Yongtai Geling, 17. IX. 2002, Wang Yiping, No. 20023760 (ZJUH). Paratype: 1 female, China, Guangxi Prov., Guangxi Botanic Garden, 30. X. 2002, Lin Naiquan, No. 20034969 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius beatoides sp. nov. is similar to S. beatus Chao, but differs in having the mesoscutum with distinct rugae penetrating on the lobes near notauli (not penetrated in S. beatus), scutellum distinctly granulate (very weakly granulate-coriaceous in S. beatus), mesopleuron medially more or less distinctly and widely rugosity-striate (smooth in S. beatus), basal carina of propodeum distinctly shorter than areola fork (same length in S. beatus), hind coxa long (rather short in S. beatus), petiole slender and more straight (wide and distinctly curved in S. beatus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC2FFD3838EFB55FD6F1B46.taxon	etymology	Etymology. From “ beatus ”, the name of related species, and suffix “ oides ”, because new species is similar to S. beatus Chao.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC2FFD3838EFB55FD6F1B46.taxon	distribution	Distribution. China (Fujian, Guangxi).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC1FFD1838EFF45FD4B1DB4.taxon	materials_examined	Material examined. China (ZJUH). 1 female, Guangdong Prov., Yunan Tongledashan, 12 – 13. VIII. 2003, Xu Zaifu, No. 20054294.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC1FFD1838EFF45FD4B1DB4.taxon	distribution	Distribution. China (Fujian, Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC1FFD1838EFE07FC2C1E89.taxon	materials_examined	Material examined. China (ZJUH). Hainan Prov.: 1 female, Bawangling, 9 – 10. VI. 2007, Liu Jingxian, No. 200703565; 1 female, Yinggeling, 24 – 25. V. 2007, Liu Jingxian, No. 200702696.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC1FFD1838EFE07FC2C1E89.taxon	distribution	Distribution. China (Hainan) (new record); Indonesia, Philippines.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC1FFD1838EFCC0FA671873.taxon	materials_examined	Material examined. China (ZJUH). Shandong Prov.: 1 female, Wendeng, IX – X. 1988, Hou Shaojin, No. 888962. Zhejiang Prov.: 1 female, Xitianmushan Dahenglu, 9. X. 1998, Zhao Mingshui, No. 999860. Yunnan Prov.: 1 female, Xiaguan, 31. VII. 1981, Li Qiang, No. 200012433; 1 female, Lancang, 20. IV. 1981, He Junhua, No. 812250. Shaanxi Prov.: 1 female, Qinling Tiantaishan, 3. IX. 1999, He Junhua, No. 990338.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC1FFD1838EFCC0FA671873.taxon	distribution	Distribution. China (Shandong, Zhejiang, Yunnan, Shaanxi) (new record); Japan, Korea, Russia (Far East).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC1FFEC838EFB45FD4C1CE3.taxon	description	(Fig. 8) Description. Female. Body length 4.3 mm; fore wing length 2.9 mm. Head. Antennae filiform, 30 - segmented. Scape 1.6 × as long as its maximum width. First flagellar segment 5.3 × as long as its apical width, 1.2 × as long as second segment. Penultimate segment 3.0 × as long as wide, 0.9 × as long as pointed apical segment. Head width (dorsal view) 1.5 × its median length, 1.2 × as wide as mesoscutum. Head behind eyes (dorsal view) evenly and weakly roundly narrowed; transverse diameter of eye 1.5 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.15 × its sides; POL: Od: OOL = 4.0: 3.0: 10.0. Vertex entirely distinctly densely transversely striate with dense additional granulation between striae; frons entirely, coarsely, densely and weakly curvedly transversely striate. Eye with short and sparse setae, 1.1 × as high as broad. Malar space 0.5 × eye height and almost equal to basal width of mandible. Temple mainly finely or very finely reticulate-coriaceous, with fine vertical striation in posterior third. Face densely, coarsely and weakly curvedly transversely striate with additional dense granulation. Face width 1.25 × eye height and 1.1 × as long as height of face and clypeus combined. Hypoclypeal depression medium-sized and round, its width 0.7 × distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, ventrally joined with hypostomal carina upper base of mandible. Mesosoma. Length of mesosoma 2.0 × its height. Pronotal keel distinct, its posterior branch coarse, distinctly separated from posterior margin of pronotum; anterior branch of carina rather fine and situated submedially. Mesoscutum highly and roundly elevated above pronotum, its median lobe weakly convex anteriorly and without anterolateral corners. Mesoscutum distinctly and densely granulate, its median lobe anteriorly without additional striation, with long and dense rugae situated along notauli and laterally, with two distinct and weakly convergent posteriorly straight carinae in medioposterior half; mostly glabrous, with sparse, long and semi-erect or erect pale setae arranged in wide stripe along notauli and in single line laterally. Notauli distinct, wide, coarsely densely crenulate, rather deep anteriorly and shallow posteriorly. Prescutellar depression rather deep, long, with three carinae, finely rugulose, 0.4 × as long as scutellum. Scutellum entirely densely and distinctly granulate, without rugae, weakly convex, with distinct lateral carinae. Mesopleuron distinctly densely granulate with additional striation marginally. Subalar depression shallow, wide, densely and coarsely striate with granulation. Precoxal sulcus deep and oblique in anterior 0.6, shallow and curved in posterior 0.4, densely and distinctly crenulategranulate, running along entire lower part of mesopleuron. Propodeum with distinct and wide lateral tubercles, with basolateral areas distinctly delineated by carinae and densely granulate with short rugosity along carinae, rest part of propodeum densely granulate with sparse striation; areola rather wide and long; basal carina present in basal 0.2, 0.6 × as long as anterior fork of areola. Wings. Fore wing 3.6 × as long as wide. Vein r arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 21: 4: 38: 18. Second submarginal cell not narrowed distally, its length 2.9 × maximum width, 1.4 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior quarter of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.7 × as long as vein 1 - M. Vein m-cu antefurcal, strongly oblique towards base of wing, weakly infuscate. Legs. Hind coxa dorsally densely transversely striate with fine granulation, distinctly densely granulate laterally, with distinct basoventral corner and tooth. Hind femur densely granulate in upper half and coriaceous to almost smooth in lower half, 3.4 × as long as wide. Hind tibia apically with two slender spines on outer side. Dorsal surface of hind tibia with long, dense and semi-erect setae, their length 1.0 – 1.4 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 0.9 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly curved ventrally, dorsally distinctly arched in basal 0.5 and almost straight in apical 0.5, thickened in basal 0.3; widened in apical fourth (dorsal view), with distinct spiracular tubercles in widened basal third. Length of petiole 2.7 × its apical width, 1.5 × length of propodeum. Second tergite without separate laterotergites. Median length of second and third tergites combined 1.6 × basal width of second tergite, 0.8 × as long as their maximum width. Second suture absent. Petiole distinctly and sparsely longitudinally striate, with additional dense reticulation between striae. Remaining tergites entirely smooth. Ovipositor straight. Ovipositor sheath 1.6 × as long as petiole, 0.6 × as long as metasoma, 0.9 × as long as mesosoma, 0.5 × as long as fore wing. Colour. Head yellowish brown. Mesosoma and metasoma reddish brown, apical parts of metasomal tergites yellowish. Antennae yellow to brownish yellow in basal half, brown medially, 11 subapical segments yellow, last apical segment brown. Palpi whitish yellow. Legs yellow to brownish yellow, hind femur brown in apical half, hind tibia subbasally whitish yellow in wide area. Ovipositor sheath yellow to brownish yellow in basal 0.4 and dark brown in apical 0.6. Fore wing distinctly and more or less evenly infuscate. Pterostigma brown, pale yellow in basal third and apically. Variation. Body length 3.7 – 4.4 mm; fore wing length 2.7 – 3.2 mm. First flagellar segment 4.8 – 5.5 × as long as its apical width. Transverse diameter of eye 1.5 – 1.8 × length of temple. Basal carina of propodeum 0.6 – 1.0 × as long as anterior fork of areola. Hind femur 3.3 – 3.4 × as long as wide. Length of petiole 2.5 – 2.7 × its apical width, 1.5 – 1.6 × length of propodeum. Ovipositor sheath 1.4 – 2.0 × as long as petiole, 0.6 – 0.9 × as long as metasoma, 0.9 – 1.2 × as long as mesosoma, 0.4 – 0.6 × as long as mesosoma. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC1FFEC838EFB45FD4C1CE3.taxon	materials_examined	Material examined. Holotype. female, China, Guangdong Prov., Shixing Chebaling, 21. VIII. 2003, Xu Zaifu, No. 20052618 (ZJUH). Paratypes: 1 female, China, Guangdong Prov., Fuyuan Nanling, 8. V. 2004, Xu Zaifu, No. 20049601 (ZJUH); 1 female, Guangdong Prov., Dinghushan, 18 – 19. VI. 1983, Zhang Yalin, No. 200011402 (ZJUH); 1 female, Fujian Prov., Wuyishan Xianfengling, 13. IX. 1989, No. 20005838 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius cephalus sp. nov. is similar to S. montivagans Chao, but differs in having the vertex densely striate-granulate (only evenly striate in S. montivagans), mesopleuron medially densely granulate with rugosity (smooth in S. montivagans), and body paler (darker in S. montivagans). This new species also similar to S. lynceus Nixon and S. gades Nixon, but differs from it in having the vertex densely striate-granulate (only striate in S. lynceus and S. gades), head weakly narrowed behind eyes (dorsal view) and temple longer (strongly narrowed and temple shorter in S. lynceus and S. gades), scutellum distinctly and densely granulate (finely granulate to smooth in S. lynceus and S. gades), and mesopleuron medially densely granulate with rugosity (smooth in S. lynceus and S. gades).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFC1FFEC838EFB45FD4C1CE3.taxon	etymology	Etymology. From “ cephalos ” (Greece for “ head ”), because of head enlarged and coarsely sculptured. Distribution. China (Fujian, Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFCFFEC838EFEF5FC991E48.taxon	materials_examined	Material examined. Female (paratype, HNHM), “ Shaowu, Fukien, China, 3. IV. 1944, H. F. Chao Coll. ”, “ Spathius chunliuae Chao, sp. nov., Paratype ”. China (ZJUH). Guangdong Prov.: 1 female, Yingde Shimentai, 29. III. 2003, Wang Yiping, No. 20032531; 2 females, Fogang Guanyinshan, 15 – 16. IX. 2007, Xu Zaifu, Nos. 200711436 and 200711690. Guizhou Prov.: 2 females, Mayanghe Wanjia, 27 – 30. IX. 2007, Zhu Lanlan, Nos. 200708118, 200708089. Yunnan Prov.: 1 female, Xishuangbanna, 31. VII. 2003, Xu Zaifu, No. 20053595.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFCFFEC838EFEF5FC991E48.taxon	distribution	Distribution. China (Fujian, Guangdong, Guizhou, Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFCFFEA838EFD03FDF41963.taxon	description	(Fig. 9) Description. Female. Body length 5.3 mm; fore wing length 4.0 mm. Head. Antennae filiform, more than 26 - segmented (apical segments missing). Scape 1.6 × as long as its maximum width. First flagellar segment 6.7 × as long as its apical width, 1.2 × as long as second segment. Head width (dorsal view) 1.45 × its median length, 1.2 × as wide as mesoscutum. Head behind eyes (dorsal view) weakly and roundly narrowed; transverse diameter of eye 1.6 × length of temple. Ocelli medium-sized, arranged in almost equilateral triangle; POL: Od: OOL = 3.0: 3.0: 11.0. Vertex almost entirely smooth; frons entirely, coarsely, rather sparsely and straight transverse striate. Eye glabrous, 1.2 × as high as broad. Malar space 0.4 × eye height and 0.7 × basal width of mandible. Temple mainly smooth. Face sparsely, distinctly and curvedly transversely striate with additional fine reticulation, only finely granulate medially. Face width 1.1 × eye height and 1.2 × as long as height of face and clypeus combined. Hypoclypeal depression medium-sized and round, its width almost equal to distance from edge of depression to eye, 0.45 × width of face. Occipital carina complete dorsally, ventrally not joined with hypostomal carina being obliterated upper base of mandible. Mesosoma. Length of mesosoma 1.9 × its height. Pronotal keel distinct, its posterior branch coarse, connected on wide territory with posterior margin of pronotum; anterior branch of carina distinct and situated submedially. Mesoscutum highly and roundly elevated above pronotum, its median lobe weakly convex anteriorly and without anterolateral corners. Mesoscutum distinctly and densely granulate, its median lobe anteriorly with granules partly situated in fine transverse lines, with long and dense rugae along notauli and laterally, with two distinct and weakly convergent posteriorly undulate carinae in medioposterior half; mostly glabrous, with rather dense, long and semi-erect pale setae arranged in wide stripe along notauli and in narrow stripe laterally. Notauli distinct, wide, coarsely densely crenulate, deep anteriorly and more shallow posteriorly. Prescutellar depression rather deep, long, with four carinae, finely rugulose-granulate, 0.3 × as long as scutellum. Scutellum entirely densely and distinctly granulate, without rugae, weakly convex, with lateral carinae. Mesopleuron entirely densely granulate with fine striation partly. Subalar depression shallow, rather narrow, coarsely rugose-striate and with granulation. Precoxal sulcus deep, wide, sinuate, shallow and oblique in anterior 0.6, sparsely crenulate with striation in posterior 0.4, running along entire lower part of mesopleuron. Propodeum with distinct and rather wide lateral tubercles, with basolateral areas distinctly delineated by carinae and densely granulate with distinct rugosity along carinae, rest part of propodeum sparsely striate and finely coriaceous; areola rather wide and medium-sized; basal carina present in basal 0.2, 0.5 × as long as anterior fork of areola. Wings. Fore wing 4.0 × as long as wide. Vein r arising from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 24: 7: 37: 21. Second submarginal cell not narrowed distally, its length 3.2 × maximum width, 1.5 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior third of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.7 × as long as vein 1 - M. Vein m-cu antefurcal, strongly oblique towards base of wing, faintly infuscate. Legs. Hind coxa dorsally transversely striate with dense distinct granulation, distinctly densely granulate laterally, but finely granulate below, with distinct basoventral corner and tooth. Hind femur subclavate, densely and very finely granulate with fine striation dorsally, smooth on rest part, 3.7 × as long as wide. Hind tibia apically with three slender spines on outer side. Dorsal surface of hind tibia with long, dense and semi-erect setae, their length 0.9 – 1.3 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.55 × as long as second-fifth segments combined. Second segment of hind tarsus 0.6 × as long as basitarsus, 1.2 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) very weakly curved ventrally, dorsally distinctly arched in basal 0.5 and almost straight in apical 0.5, thickened in basal 0.4; weakly widened in apical fifth (dorsal view), with spiracular tubercles in weakly widened basal third. Length of petiole 2.7 × its apical width, 1.7 × length of propodeum. Second tergite with laterotergites separated in basal half only. Median length of second and third tergites combined 1.6 × basal width of second tergite, 0.8 × as long as their maximum width. Second suture absent. Petiole distinctly and rather densely longitudinally striate, with additional dense and fine reticulation between striae. Second tergite basally finely aciculate. Remaining tergites smooth. Ovipositor straight. Ovipositor sheath 2.6 × as long as petiole, 1.2 × as long as metasoma, 1.8 × as long as mesosoma, 0.8 × as long as fore wing. Colour. Head and mesonotum yellow to brownish yellow, face reddish brown. Remaining mesosoma and all metasoma reddish brown to dark reddish brown, tergites behind petiole brownish yellow apically. Antennae yellow to brownish brown, brown in apical third, possibly without pale segments (apical segments missing). Palpi whitish. Legs yellow to whitish yellow, all femora in apical thirds brown, hind coxa entirely yellow, hind tibia yellow, brown on short median area, brownish basally. Ovipositor sheath yellow to brownish yellow in basal 0.5 and brown to dark brown in apical 0.5. Fore wing very faintly infuscate, with more distinct brownish spots around vein 1 - SR, 1 - M and below pterostigma. Pterostigma brown, pale yellow in basal quarter. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFCFFEA838EFD03FDF41963.taxon	materials_examined	Material examined. Holotype. female, China, Hainan Prov., Jianfengling Mingfenggu, 25. X. 2007, Liu Jingxian, No. 200709690 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius clavator sp. nov. is similar to S. angustifemur Belokobylskij et Maeto, but differs in having the petiole and second tergite wide (slender in S. angustifemur), ovipositor sheath shorter (longer in S. angustifemur), mesopleuron medially distinctly and widely rugosity-granulate (smooth in S. angustifemur), temple short (long in S. angustifemur), scutellum distinctly densely granulate (mainly smooth in S. angustifemur), head, mesoscutum and hind coxa yellow to brownish yellow (reddish brown to dark reddish brown in S. angustifemur). The differences between S. clavator sp. nov. and S. hikoensis Belokobylskij are shown in the key.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFCFFEA838EFD03FDF41963.taxon	etymology	Etymology. From “ clavator ” (Latin for “ club ”) because of the shape of hind femur.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFCFFEA838EFD03FDF41963.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFAFFEA838EFA75FAA61B14.taxon	materials_examined	Material examined. Female (holotype of S. colophon euro, IZCAS), Yunnan Prov., Xishuangbanna Menghun, 1200 - 1400 m, 21. V. 1958, Meng Xuwu. China (ZJUH). 1 female, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802204; 1 female, Zhejiang Prov., Lin’an Qingliangfeng, 12. VIII. 2005, Shi Min, No. 200607649; 1 female, Hainan Prov., Jianfengling Tianchi, 24. IX. 2008, No. 200806013; 1 female, Yunnan Prov., Xishuangbanna Forest Park, 31. VIII. 2003, Xu Zaifu, No. 20053528.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFAFFEA838EFA75FAA61B14.taxon	distribution	Distribution. China (Zhejiang, Hainan, Yunnan); Singapore, Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFAFFEA838EFA75FAA61B14.taxon	discussion	Remarks. Sometimes the second metasomal tergite is sculptured (striate-rugulose) in its basal 0.3 – 0.5.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFAFFEB838EF8A7FCD81CCE.taxon	materials_examined	Material examined. China (ZJUH). Hainan Prov.: 1 female, Bawangling, 26. XI. 2008, Wang Manman, No. 200805630.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFAFFEB838EF8A7FCD81CCE.taxon	distribution	Distribution. China (Hainan) (new record); Indonesia.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFBFFE9838EFEECFDCE1E56.taxon	description	(Fig. 10) Description. Female. Body length 5.2 mm; fore wing length 4.7 mm. Head. Antennae slender, weakly setiform, more than 20 - segmented (apical segments missing). Scape 1.8 × as long as its maximum width. First flagellar segment 5.0 × as long as its apical width, 1.4 × as long as second segment. Head not depressed, its width (dorsal view) 1.3 × median length. Head behind eyes roundly narrowed. Transverse diameter of eye 1.3 × length of temple. Ocelli arranged in triangle with base as long as its sides; POL: Od: OOL = 6: 6: 18. Vertex densely coarsely rugose-striate. Frons densely and transversely rugose-striate. Eye glabrous, 1.3 × as high as broad. Malar space 0.4 × eye height and 0.9 × basal width of mandible. Temple distinctly striate in upper half, smooth in lower half. Face coarsely rugose-reticulate, narrowly smooth medially; its width 1.4 × eye height and 1.1 × height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.0 × its height. Pronotal keel distinct, its posterior branch widely separated from posterior margin of pronotum. Mesoscutum highly and roundly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; densely granulate, with rather long and more or less distinct rugae near notauli and laterally; lateral lobes medially with wide granulate areas, with two convergent posteriorly carinae and narrow rugosity between them in medioposterior half. Notauli complete, wide, deep anteriorly and shallow posteriorly, sparsely crenulate. Prescutellar depression shallow, with five carinae, 0.3 × as long as scutellum. Scutellum finely coriaceous-granulate, weakly convex, with distinct lateral carinae. Mesopleuron entirely granulate and rugose. Subalar depression shallow, wide, coarsely rugose-striate and granulate. Precoxal sulcus deep, sinuate, distinctly crenulate, running along entire length of lower part of mesopleuron. Propodeum with distinct tubercles, coarsely rugose-granulate in basal half, with areas more or less distinctly delineated by carinae; basal carina 2.3 × as long as anterior fork of areola; areola narrow and pentagonal; petiolate area distinctly separated from areola. Wings. Fore wing 4.1 × as long as wide. Vein r distinctly arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 23: 6: 28: 19. Second submarginal cell not narrowed distally, its length 2.8 × maximum width, 1.4 × length of first subdiscal cell. Vein cu-a weakly postfurcal. Vein m-cu postfurcal. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, weakly curved towards base of wing. Legs. Hind coxa transversely striate and granulate dorsally, only granulate laterally and ventrally, with basoventral tooth and corner. Hind femur finely striate and granulate dorsally, finely coriaceous-granulate ventrally and laterally, 3.5 × as long as wide. Hind tibia apically with two spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 1.4 – 1.6 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) straight ventrally and distinct arched dorsally, with spiracular tubercles in basal third; sparsely striate in basal 0.7, densely striate in apical 0.3, with rugulosity between striae. Length of petiole 2.6 × its apical width (dorsal view), 1.9 × length of propodeum. Second tergite without separated laterotergite. Second suture indistinct. Median length of second and third tergites combined 1.4 × basal width of second tergite, 0.7 × as long as their maximum width. Second tergite densely finely striate, smooth laterally. Remaining tergites smooth. Ovipositor sheath 2.1 × as long as petiole, as long as metasoma, 1.5 × as long as mesosoma, 0.7 × as long as fore wing. Colour. Body dark brown; head brown; scutellum brownish yellow. Petiole dark brown; second and third tergites dark brown in basal two-thirds, brownish yellow in apical third; fourth and fifth tergites dark brown basally and apically, brownish yellow medially; sixth tergite brownish yellow. Antenna brownish yellow basally, darkened toward apex. Palpi pale yellow. Legs pale brownish yellow, fore and middle coxae, all trochanters pale yellow; hind coxa brownish yellow; hind femur darken apically. Ovipositor sheath brown, but medially widely pale brown. Fore wing distinctly infuscate. Pterostigma brown, yellow basally and apically. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFBFFE9838EFEECFDCE1E56.taxon	materials_examined	Material examined. Holotype: female, China, Yunnan Prov., Pingbian Daweishan, 18. VII. 2008, Xu Zaifu, No. 20055034 (ZJUH). Comparative diagnosis. This new species belongs to the S. moderabilis Wilkinson species group. Spathius daweiensis sp. nov. is similar to S. gutianensis sp. nov., but differs in having the temple distinctly striate in upper half (smooth in S. gutianensis), temple longer with transverse diameter of eye 1.3 × length of temple (shorter, with transverse diameter of eye 1.8 × length of temple in S. gutianensis), mesopleuron almost entirely granulate and rugose (mostly smooth in S. gutianensis), third and fourth metasomal tergites smooth (third tergite densely striate and fourth tergite finely striate in basal half in S. gutianensis), ovipositor sheath longer, as long as metasoma (shorter, 0.5 × as long as metasoma in S. gutianensis), and wing distinctly infuscate (faintly infuscate in S. gutianensis).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFBFFE9838EFEECFDCE1E56.taxon	etymology	Etymology. From the name of mountain of the type locality of species.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFFBFFE9838EFEECFDCE1E56.taxon	distribution	Distribution. China (Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF9FFE9838EFD66FB7618C9.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Fujian Prov., Jianyang Huangkeng, 18. IX. 1975, Chen Jiahua. China (ZJUH). Zhejiang Prov.: 1 female, Fengyangshan, 27. VII. 2007, Liu Jingxian, No. 200809062; 1 female, Longquan Fengyangshan Datianping, 28. VII. 2007, Liu Jingxian, No. 200802750; 1 female, Gutianshan, 3. VII. 2005, Wu Qiong, No. 200616480; 1 female, Anji Longwangshan, 18. VII. 1995, Wu Hong, No. 970092; 1 female, Xitianmushan Xianrending, 2 – 4. VI. 1990, Lou Yonggen, No. 900156. Guangdong Prov.: 1 female, Meizhou Fengxi, 29. VII. 2003, Chen Jujian, No. 20048488; 1 female, Fogang Guanyinshan, 15 – 16. IX. 2007, Xu Zaifu, No. 200711650. Hainan Prov.: 1 female, Bawangling, 26. XI. 2008, Wang Manman, No. 200805638; 1 female, Jianfengling, 7. VI. 2007, Liu Jingxian, No. 200702461; 1 female, Jianfengling, 22 – 23. IX. 2008, Wang Manman, No. 200805935; 1 female, Wuzhishan, 29. IX. 2007, Liu Jingxian, No. 200710315. Sichuan Prov.: 1 female, Pingwu Baimazhai, 25. VII. 2006, Zhang Hongying, No. 200809973. Guizhou Prov.: 1 female, Guiyang, 7 – 12. X. 1983, He Junhua, No. 833696.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF9FFE9838EFD66FB7618C9.taxon	distribution	Distribution. China (Fujian, Zhejiang, Guangdong, Hainan, Sichuan, Guizhou); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF9FFE9838EFA86FBD31AB0.taxon	materials_examined	Material examined. China (ZJUH). Hebei Prov.: 1 male, Xiaowutaishan Donglingshan, 21. VIII. 2005, Zhang Hongying, No. 200802652. Zhejiang Prov.: 1 female, Songyang, 21. VIII. 1989, He Junhua, No. 895243; 1 female, Songyang Andaihou, 15 – 17. VIII. 1989, He Junhua, No. 894467; 2 females, Longquan Fengyangshan, 27. VII. 2007, Liu Jingxian, Nos. 200801386 and 200801383.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF9FFE9838EFA86FBD31AB0.taxon	distribution	Distribution. China (Hebei, Zhejiang) (new record); Japan, Russia (Far East).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF9FFE9838EF90BFC811B88.taxon	materials_examined	Material examined. China (ZJUH). Fujian Prov.: 1 female, Qingliu, 24. VI. 1986, Qi Shicheng, No. 865254. Distribution. China (Fujian, Guangxi, Taiwan); Philippines.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF7FFE7838EFF45FD6B1D43.taxon	description	(Fig. 11)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF7FFE7838EFF45FD6B1D43.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Sichuan Prov., Emeishan, 24. VI. 1955, Leng Huaiqiong. Distribution. China (Sichuan, Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF7FFE7838EFE68FC371980.taxon	materials_examined	Material examined. China (ZJUH). Zhejiang Prov.: 5 females, Gutianshan, 3. VII. 2005, Chen Xuexin, Nos. 200601641, 200601656, 200617193, 200617196 and 200617200; 19 females, Gutianshan, 1 – 3. VII. 2005, Shi Min, Nos. 200616540, 200616548, 200616552, 200616556, 200615905, 200615932, 200615935, 200616337, 200616358, 200616361, 200616370, 200616373, 200616375, 200616387, 200616397, 200616406, 200616411, 200616412 and 200616414 (2 females in ZISP); 24 females, Gutianshan, 1 – 3. VII. 2005, Wu Qiong, Nos. 200615780, 200615784, 200615787, 200615793, 200615794, 200615941, 200615968, 200616438, 200616444, 200616450, 200616453, 200616455, 200616467, 200616468, 200616469, 200616479, 200616481, 200616484, 200616485, 200616490, 200616491, 200616495, 200616498 and 200616503; 12 females, Gutianshan, 1 – 3. VII. 2005, Zhang Hongying, Nos. 200616603, 200616609, 200616028, 200616098, 200616099, 200616103, 200616110, 200616112, 200616118, 200616122, 200616125 and 200616130; 12 females, 3 males, Gutianshan, 17 – 20. VII. 1992, Chen Xuexin, Nos. 923269, 923312, 923315, 923319, 923605, 923615, 923645, 963652, 963653, 923998, 923703, 923715, 923717, 923718 and 923727; 5 females, Gutianshan, 17 – 19. VII. 1992, Ma Yun, Nos. 923773, 923844, 923870, 923875 and 924076; 2 females, 1 male, Gutianshan, 1. VIII. 1990, Ma Yun, Nos. 906174, 906221 and 906230; 3 females, Gutianshan, VII – VIII. 1990, Xu Zhihong, Nos. 905377, 905577 and 905578; 1 female, Gutianshan, VII – VIII. 1990, Ma Yun, No. 905928; 9 females, Gutianshan, 21 – 27. VIII. 1992, Wu Hong, Nos. 948989, 948991, 949002, 949136, 949137, 949141, 949152, 949198 and 949068; 1 female, Longquan Fengyangshan, 7. VIII. 2003, Liu Jingxian, No. 20055661; 1 female, Fengyangshan, 27. VII. 2007, Liu Jingxian, No. 200809015; 1 female, Longquan Fengyangshan, 28. VII. 2007, Liu Jingxian, No. 200802753. Fujian Prov.: 1 female, Wuyishan, 15. IX. 1989, Wang Jiashe, No. 964498. Guangdong Prov.: 2 females, Fuyuan Nanling, 23. VII. 2003, Xu Zaifu, Nos. 20049009, 20049045; 1 female, Meizhou Fengxi, 29. VII. 2003, Chen Jujian, No. 20048514; 1 female, Shixing Chebaling, 27. VII. 2002, Xu Zaifu, No. 20051882. Hainan Prov.: 3 females, Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, Nos. 200802180, 200802322 and 200802394; 2 females, Jianfengling, 12 – 15. VII. 2006, Liu Jingxian, Nos. 200803605, 200803630. Yunnan Prov.: 4 females, Pingbian Daweishan, 18. VII. 2003, Xu Zaifu, Nos. 20054935, 20055139, 20055158 and 20055188.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF7FFE7838EFE68FC371980.taxon	distribution	Distribution. China (Zhejiang, Fujian, Guangdong, Hainan, Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF7FFE4838EF9DBFAD81D76.taxon	materials_examined	Material examined. China (ZJUH). Fujian Prov.: 1 female, Chongan Sangang, 27. X. 1989, Wang Jiashe, No. 20007573. Guangdong Prov.: 1 female, Fuyuan Nanling, 23. VII. 2003, Xu Zaifu, No. 20049139; 2 females, Fuyuan Nanling, 4. VIII. 2004, Xu Zaifu, Nos. 20049746, 20049825. Yunnan Prov.: 1 female, Pingbian Daweishan, 18. VII. 2003, Xu Zaifu, No. 20054966. Guizhou Prov.: 1 female, Kuankuoshui Xiangshuwan, 2. VI. 2010, Tan Jiangli, No. 201002144.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF7FFE4838EF9DBFAD81D76.taxon	distribution	Distribution. China (Heilongjiang, Fujian, Hubei, Guangdong, Yunnan, Guizhou); Japan, Korea, Mongolia, Russia, all Europe, New Zealand.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF7FFE4838EF9DBFAD81D76.taxon	discussion	Remarks. According to the description and figures in Chen & Shi (2004), S. tanycoleosus Shi et Chen is characterised by having ovipositor sheath and petiole long and substraight, fourth and fifth tergites entirely smooth, hind femur slender and vertex smooth etc., which allows us to synonymize it with S. exarator (Linnaeus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF4FFE4838EFE46FB5A1ED3.taxon	materials_examined	Material examined. China (ZJUH). Hainan Prov.: 1 female, Jianfengling, 7. VI. 2007, Liu Jingxian, No. 200702479; 1 female, Wuzhishan, 16 – 20. V. 2007, Liu Jingxian, No. 200703371; 1 female, Jianfengling Tianchi, 12 – 15. VII. 2006, Chen Tianfei, No. 200803207. Guangdong Prov.: 1 female, Fuyuan Nanling, 8. V. 2004, Xu Zaifu, No. 20049441. Guangxi Prov.: 1 female, Ningming, 16. V. 1984, Yang Ding, No. 200012472.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF4FFE4838EFE46FB5A1ED3.taxon	distribution	Distribution. China (Guangdong, Guangxi, Hainan) (new record); Japan, Korea, Russia.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF4FFE4838EFCE5FBBB18C3.taxon	materials_examined	Material examined. China (ZJUH). Guangdong Prov.: 1 female, Shixing Chebaling, 27. VII. 2002, Xu Zaifu, No. 20051840. Hainan Prov.: 1 female, Jianfengling, 6. VI. 2007, Liu Jingxian, No. 200703831; 1 female, Jianfengling, 22. XI. 2008, Tan Jiangli, No. 200805313; 1 female, Jianfengling Tianchi, 12 – 15. VII. 2006, Zhang Wenyong, No. 200803490 (ZISP); 1 female, Wuzhishan, 16 – 20. V. 2007, Liu Jingxian, No. 200703173; 1 female, Wuzhishan Shuimanxiang, 15 – 20. V. 2007, Weng Liqiong, No. 200804014; 2 females, Yinggeling, 24 – 25. V. 2007, Liu Jingxian, Nos. 200702609 and 200702662; 1 female, Yinggeling, 24 – 26. V. 2007, Liu Jingxian, No. 200704091; 1 female, Yinggeling, 23 – 25. V. 2007, Xiao Bin, No. 200707380.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF4FFE4838EFCE5FBBB18C3.taxon	distribution	Distribution. China (Guangdong, Hainan) (new record); Indonesia, Philippines.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF4FFE0838EFA95FDF41D76.taxon	description	(Fig. 12) Description. Female. Body length 4.0 mm; fore wing length 2.8 mm. Head. Antennae weakly setiform, 44 - segmented. Scape 1.7 × as long as its maximum width. First flagellar segment 4.8 × as long as its apical width, 1.2 × as long as second segment. Penultimate segment 3.6 × as long as its width. Head width (dorsal view) 1.7 × its median length, 1.3 × as wide as mesoscutum. Head behind eyes (dorsal view) distinctly roundly narrowed; transverse diameter of eye 2.0 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 4.0: 5.0: 9.0. Vertex entirely smooth; frons entirely sparsely transversely curvedly striate. Eye with sparse and very short setae, 1.25 × as high as broad. Malar space 0.3 × eye height and 0.8 × basal width of mandible. Temple mainly smooth. Face densely, distinctly and undulately transversely striate, with additional reticulation. Face width equal to eye height and 1.1 × height of face and clypeus combined. Hypoclypeal depression medium sized and subround, its width 0.7 × distance from edge of depression to eye, 0.3 × width of face. Occipital carina complete and evenly curved dorsally, ventrally obliterated on short distance before hypostomal carina upper base of mandible. Mesosoma. Length of mesosoma 1.9 × its height. Pronotal keel high, its posterior branch not fused with posterior margin of pronotum and weakly separated from it, anterior branch rather fine and situated submedially. Mesoscutum highly and roundly elevated above pronotum (lateral view), its median lobe (dorsal view) distinctly convex anteriorly and without anterolateral corners. Mesoscutum densely granulate, with long, rather coarse and dense rugae situated along notauli, with two coarse, weakly undulate and convergent posteriorly carinae in medioposterior half; glabrous on rather wide areas, with sparse, long and semi-erect or erect golden setae arranged in wide stripes along notauli and in narrow stripe laterally. Notauli wide, anteriorly deep and shallow posteriorly, coarsely crenulate. Prescutellar depression rather deep, long, with three carinae, almost smooth between carinae, 0.3 × as long as scutellum. Scutellum entirely finely granulate-coriaceous, weakly convex, with fine lateral carinae. Mesopleuron mainly smooth. Subalar depression shallow, rather wide, distinctly coarsely striate. Precoxal sulcus shallow, oblique, straight, coarsely and rather densely crenulate, but smooth posteriorly, running along anterior half of lower length of mesopleuron. Propodeum with short and wide subpointed lateral tubercles, with basolateral areas distinctly delineated by coarse carinae and finely coriaceous in basal half and distinctly longitudinally striate on rest part, rest part of propodeum mainly smooth with fine and sparse striae; areola narrow and long; basal carina present in basal 0.2, about 0.6 × as long as anterior fork of areola. Wings. Fore wing about 4.5 × as long as wide. Vein r arising from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 22: 6: 35: 13. Second submarginal cell not narrowed distally, its length about 3.5 × maximum width, 1.2 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior 0.4 of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu weakly antefurcal, weakly straight, distinctly oblique towards base of wing, infuscate. Legs. Hind coxa dorsally transversely curvedly striate, densely to finely granulate at rest part, with distinct basoventral corner and tooth. Hind femur mainly smooth, rugulose in dorso-posterior half, 3.0 × as long as wide. Hind tibia apically with single slender spines on outer side. Dorsal surface of hind tibia with long, rather dense and semi-erect setae, their length 1.1 – 1.3 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.55 × as long as second-fifth segments combined. Second segment of hind tarsus 0.6 × as long as basitarsus, 0.9 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) very weakly curved ventrally, dorsally evenly and finely curved, weakly thickened in basal third; weakly widened in basal half and distinctly widened in apical one-sixth (dorsal view), with spiracular tubercles in basal third. Length of petiole 3.4 × its apical width, 1.75 × length of propodeum. Second tergite with weakly separate laterotergite. Median length of second and third tergites combined 1.6 × basal width of second tergite, 0.8 × as long as their maximum width. Second suture absent. Petiole coarsely and sparsely longitudinally striate, with dense and rather distinct reticulation between striae. Remaining tergites smooth, but second tergite mainly very finely and interruptedly reticulate-striate. Ovipositor almost straight. Ovipositor sheath 2.8 × as long as petiole, 1.3 × as long as metasoma, 2.0 × as long as mesosoma, as long as fore wing. Colour. Body reddish brown, head paler dorsally and laterally. Antenna light reddish brown, faintly darkened medially, with 16 apical yellowish white to white segments, last segments infuscate. Palpi white. Fore and middle legs brownish yellow to pale brown, coxae and trochanters pale yellow. Hind leg light reddish brown, hind tibia faintly infuscate submedially, yellow subbasally at short distance, brownish basally. Ovipositor sheath brown, pale basally. Fore wing faintly and evenly infuscate. Pterostigma dark brown, yellow basally and apically on short spaces. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF4FFE0838EFA95FDF41D76.taxon	materials_examined	Material examined. Holotype. female, China, Hainan Prov., Yinggeling, 28. V – 3. VI. 2007, Weng Liqiong, No. 200804061 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius ferrugineus sp. nov. is similar to S. erymanthus Nixon from Philippines, but differs from it in having antennae subapically with numerous white segments (without pale subapical segments in S. erymanthus), pterostigma pale brown in short basal part (pale in almost basal half in S. erymanthus), head behind eyes (dorsal view) distinctly narrowed (feebly narrowed in S. erymanthus), face and frons entirely sculptured (broadly smooth along middle lines in S. erymanthus), propodeal areola delineated by coarse carinae (almost not delineated in S. erymanthus), second segment of hind leg longer than fifth segment without pretarsus (shorter in S. erymanthus), and petiole shorter (longer in S. erymanthus). New species is also similar to S. neleiformis sp. nov., but differs in having the hypoclypeal depression short (wide in S. neleiformis), posterior branch of pronotal keel not fused with posterior margin of pronotum (broadly fused in S. neleiformis), mesoscutum with sparse and long golden setae arranged in wide stripes along notauli (with rather dense and short pale setae arranged in narrow stripes along notauli in S. neleiformis), precoxal sulcus running along only anterior half of lower length of mesopleuron (along all length in S. neleiformis), second segment of hind tarsus shorter than fifth segment without pretarsus (distinctly longer in S. neleiformis), petiole slender (wider in S. neleiformis), and antenna apically with pale segments (with dark segments in S. neleiformis).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF4FFE0838EFA95FDF41D76.taxon	etymology	Etymology. From “ ferrugineus ” (Latin for “ ferruginous ”) because of the main body colouration.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF4FFE0838EFA95FDF41D76.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF0FFE1838EFE46FD901AAE.taxon	description	(Fig. 13) Description. Female. Body length 3.8 mm; fore wing length 2.9 mm. Head. Antennae filiform, 31 - segmented. Scape 1.6 × as long as its maximum width. First flagellar segment 5.0 × as long as its apical width, 1.2 × as long as second segment. Penultimate segment 3.8 × as long as wide, almost as long as subpointed apical segment. Head width (dorsal view) 1.6 × its median length, 1.3 × as wide as mesoscutum. Head behind eyes (dorsal view) weakly convex anteriorly and roundly narrowed posteriorly; transverse diameter of eye 1.8 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 3.5: 3.0: 8.0. Vertex very finely and interruptedly reticulate-rugulose, partly smooth; frons entirely, densely, distinctly and curvedly transversely striate. Eye glabrous, 1.2 × as high as broad. Malar space 0.5 × eye height and 0.85 × basal width of mandible. Temple entirely smooth. Face mainly with dense, coarse and weakly curved transverse striation. Face width 1.1 × eye height and almost equal to height of face and clypeus combined. Hypoclypeal depression medium size and round, its width 0.9 × distance from edge of depression to eye, 0.45 × width of face. Occipital carina complete dorsally, ventrally joined with hypostomal carina by additional ruga upper base of mandible. Mesosoma. Length of mesosoma 2.1 × its height. Pronotal keel distinct, its posterior branch coarse, weakly separated from posterior margin of pronotum, but additionally joined with it by two fine additional lateral rugae; anterior branch of carina distinct and situated submedially. Mesoscutum highly and roundly elevated above pronotum, its median lobe convex anteriorly and without anterolateral corners. Mesoscutum densely granulate, but its median lobe anteriorly with additional fine transverse striation, with short and sparse or partly indistinct rugae situated along notauli and laterally, with two distinct and convergent posteriorly undulate carinae in medioposterior half; mostly glabrous, with rather sparse, long and semi-erect yellow setae arranged in wide stripe along notauli and in single line laterally. Notauli anteriorly distinct, wide and coarsely crenulate, weakly shallow posteriorly. Prescutellar depression deep, long, with four carinae, almost smooth, 0.4 × as long as scutellum. Scutellum entirely densely and rather finely granulate, weakly convex, with distinct lateral carinae. Mesopleuron finely reticulate-coriaceous with fine striation partly, smooth in small areas. Subalar depression shallow, widely and densely striate. Precoxal sulcus deep in anterior 0.7 and shallow in posterior 0.3, curved, sparsely crenulate with fine granulation, running along entire lower part of mesopleuron. Propodeum with distinct and wide lateral tubercles, with basolateral areas distinctly delineated by carinae and densely granulate with fine striation, rest part of propodeum rugose-striate with granulation; areola narrow and long; basal carina present in basal 0.3, 1.4 × longer than anterior fork of areola. Wings. Fore wing 3.7 × as long as wide. Vein r arising from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 22: 5: 40: 19. Second submarginal cell not narrowed distally, its length 3.1 × maximum width, 1.4 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior third of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu weakly antefurcal, weakly curved towards base of wing, faintly infuscate. Legs. Hind coxa dorsally transversely striate with granulation, densely granulate at rest part and reticulation basally, with distinct basoventral corner and tooth. Hind femur reticulate-granulate in upper half and almost smooth to smooth in lower half, 3.3 × as long as wide. Hind tibia apically with four spines on outer side. Dorsal surface of hind tibia with long, rather dense and semi-erect setae, their length 1.0 – 1.6 × maximum width of tibia. Hind tarsus 0.85 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 1.1 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly curved ventrally, dorsally distinctly arched in basal 0.6 and almost straight in apical 0.4, distinctly thickened in basal 0.3; widened in apical fourth (dorsal view), with small spiracular tubercles in widened basal third. Length of petiole 2.6 × its apical width, 1.7 × length of propodeum. Second tergite without separate laterotergites. Median length of second and third tergites combined 1.5 × basal width of second tergite, 0.8 × as long as their maximum width. Second suture absent. Petiole distinctly and rather densely longitudinally striate, with additional dense and distinct reticulation in basal 0.7. Remaining tergites entirely smooth. Ovipositor straight. Ovipositor sheath 1.6 × as long as petiole, 0.7 × as long as metasoma, as long as mesosoma, 0.5 × as long as fore wing. Colour. Body yellow to brownish yellow, propodeum, petiole and spots on tergites infuscate. Antennae yellow to brownish yellow in basal half, pale brown to light reddish brown in most apical half, with four whitish apical segments. Palpi whitish yellow. Legs yellow, basally white. Ovipositor sheath brown, pale yellow in basal half. Fore wing faintly infuscate, with more dark brownish spots near veins. Pterostigma brown, pale yellow in basal quarter and apically. Variation. Body length 2.7 – 4.3 mm; fore wing length 2.1 – 3.1 mm. Antenna 25 – 31 - segmented. First flagellar segment 4.5 – 5.2 × as long as its apical width, 1.10 – 1.15 × as long as second segment. Transverse diameter of eye 1.6 – 2.0 × length of temple. Vertex sometimes almost smooth. Malar space 0.50 – 0.55 × eye height. Hind femur 3.2 – 3.6 × as long as maximum width. Length of petiole 2.5 – 3.1 × its apical width, 1.5 – 1.7 × length of propodeum. Sometimes second tergite basally finely aciculate. Ovipositor sheath 1.5 – 1.6 × as long as petiole, 0.6 – 0.7 × as long as metasoma, 0.8 – 1.0 × as long as mesosoma, 0.4 – 0.5 × as long as fore wing. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF0FFE1838EFE46FD901AAE.taxon	materials_examined	Material examined. Holotype. female, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802392 (ZJUH). Paratypes: 1 female, China, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802154 (ZISP); 2 females, Yunnan Prov., Xishuangbanna Forest Park, 31. VII. 2003, Xu Zaifu, Nos 20053556 and 20053574 (ZJUH); 1 female, Hainan Prov., Yinggeling, 28. V – 3. VI. 2007, Weng Liqiong, No. 200804331 (ZJUH); 1 female, Hainan Prov., Bawangling, 26. XI. 2008, Tan Jiangli, No. 200805700 (ZJUH); 1 female, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802259 (ZJUH); 1 female, Hainan Prov., Yinggeling, 18. X. 2007, Liu Jingxian, No. 200709777 (ZJUH); 1 female, Hainan Prov., Yinggeling, 16. XI. 2008, Tan Jiangli, No. 200805797 (ZJUH); 1 female, Hainan Prov., Diaoluoshan, 28 – 31. V. 2007, Liu Jingxian, No. 200702860 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius flavicorpus sp. nov. is similar to S. erymanthus Nixon, but differs in having the hind tibia yellowish brown in apical half (brown in S. erymanthus), pterostigma pale in basal quarter (in basal half in S. rymanthus), ocelli arranged in triangle with base larger than its sides (in equilateral triangle in S. erymanthus), mesoscutum almost perpendicularly elevated above pronotum (rather gradually in S. erymanthus), areola distinctly delineated by carinae (almost absent in S. erymanthus), hind coxa with distinct basoventral tubercle (with trace of tubercle in S. erymanthus), petiole 1.7 × as long as propodeum (almost twice in S. erymanthus), and ovipositor distinctly shorter than metasoma (distinctly longer in S. erymanthus). This new species also similar to Japanese S. longisetosus Belokobylskij et Maeto, but differs from it in having the head in dorsal view behind eyes distinctly narrowed (weakly narrowed in S. longisetosus), mesoscutum with short and rather fine rugosity along notauli and laterally (with long and distinct rugae and setae in S. longisetosus), mesopleuron medially finely reticulate granulate to smooth (entirely striate to rugose-striate in S. longisetosus), setae on dorsal surface of hind tibia shorter (longer in S. longisetosus), and body mainly pale (dark in S. longisetosus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF0FFE1838EFE46FD901AAE.taxon	etymology	Etymology. From “ flavus ” (Latin for “ yellow ”) and “ corpus ” (Latin for “ body ”), because the most part of body yellowish.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF0FFE1838EFE46FD901AAE.taxon	distribution	Distribution. China (Hainan, Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF1FFE1838EF93EFC391BE0.taxon	materials_examined	Material examined. China (ZJUH). Henan Prov.: 1 female, Xixia Huangshian, 17. VII. 1998, Chen Xuexin, No. 989229; 1 female, Neixiang Baotianman, 15. VII. 1998, Ma Yun, No. 987063.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFF1FFE1838EF93EFC391BE0.taxon	distribution	Distribution. China (Henan) (new record); Korea, Russia (Far East).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEEFFFE838EFF45FC611925.taxon	materials_examined	Material examined. Female (holotype of Spathius jilinensis Chao, IZCAS), Jilin Prov., Manjiang, 8. VIII. 1955, Li Zhiyin. Female (holotype of Spathius nungdaensis Chao, IZCAS), Beijing Prov., Beijing Agricultural University Campus (China Agricultural University), 19. V. 1973, Yang Jikun. China (ZJUH). Heilongjiang Prov.: 1 female, Jingpohu, 26. VIII. 1995, Lou Juxian, No. 962063. Henan Prov.: 1 female, Jigongshan, 11. VII. 1997, Chen Xuexin, No. 973873; 1 female, Jigongshan, 12. VII. 1997, Chen Xuexin, No. 975014; 2 females, Neixiang Baotianman, 15. VII. 1998, Chen Xuexin, Nos. 988902 and 988972; 1 female, Baotianman, 13 – 15. VII. 1998, Chen Xuexin, No. 991816; 1 female, Funiushan, 11. VII. 1996, Cai Ping, No. 973884; 1 female, Yichuan Longyuwan, 11 – 12. VII. 1996, Cai Ping, No. 972607. Jiangsu Prov.: 2 females, Yangzhou, 1981, Yang Lianmin, Nos. 820098 (2). Zhejiang Prov.: 2 females, Songyang Andaihou, 15 – 17. VII. 1989, He Junhua, Nos. 894327 and 893924; 1 female, Songyang, 18 – 31. VII. 1989, He Junhua, No. 895218; 1 female, Xitianmushan Dahenglu, 9. X. 1998, Zhao Mingshui, No. 999861; 1 female, Xitianmushan Xianrending, 14. IX. 1998, Zhao Mingshui, No. 995051; 1 female, Xitianmushan Jinshanmen, 12. XII. 1998, Zhao Mingshui, No. 20002809; 1 female, Xitianmushan Houshanmen, 3. VII. 1998, Zhao Mingshui, No. 20000146; 1 female, Xitianmushan Sanmuping, 14. VII. 1998, Zhao Mingshui, No. 999146; 1 female, Xitianmushan, 17 – 18. V. 1988, Xu Weiliang, No. 885804; 1 female, Longquan Fengyangshan, 10. VIII. 2003, Liu Jingxian, No. 20048265; 1 female, Gutianshan, 18. VII. 1992, Chen Xuexin, No. 923486. Jiangxi Prov.: 1 female, Yichun, VI. 1984, No. 841670. Fujian Prov.: 2 females, Wuyishan Tongmu, 14. VII. 1994, Chen Xuexin, Nos. 942330 and 942460; 1 female, Tongmu, X. 1979, Huang Juchang, No. 20003856; 2 females, Wuyishan, 1. X. 1989, Wang Jiashe, Nos. 964341 and 964353; 1 female, Guadang, 22. VI. 1980, Lin Naiquan, No. 20003915; 1 female, Youxi Qili, 15. VIII. 1987, Fu Chaorong, No. 20005206; 1 female, Jiangle Longqishan, 8. VII. 1991, Liu Changming, No. 969915. Guangdong Prov.: 2 females, Shixing Chebaling, 25. V. 2002, Xu Zaifu, Nos. 20051673 and 20051631; 2 females, Fuyuan Nanling, 4. VIII. 2004, Xu Zaifu, Nos. 20049720 and 20049195.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEEFFFE838EFF45FC611925.taxon	distribution	Distribution. China (Beijing, Heilongjiang, Jilin, Henan, Jiangsu, Zhejiang, Jiangxi, Fujian, Guangdong, Taiwan); Japan, Korea, Russia, India.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEEFFFE838EFF45FC611925.taxon	discussion	Remarks. Spathius brevicaudis Ratzeburg was previously erroneously recorded from Taiwan (Belokobylskij, 1996); the specimens recorded from Taiwan belong to S. generosus.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEEFFFC838EFAB5FDC01A8B.taxon	description	(Fig. 14) Description. Female. Body length 4.7 mm; fore wing length 3.2 mm. Head. Antennae slender, weakly setiform, 37 - segmented. Scape 1.6 × as long as its maximum width. First flagellar segment 6.0 × as long as its apical width, 1.2 × as long as second segment. Head not depressed, its width (dorsal view) 1.3 × median length. Head behind eyes roundly narrowed. Transverse diameter of eye 1.8 × length of temple. Ocelli arranged in triangle with base as long as its sides; POL: Od: OOL = 6: 5: 18. Vertex finely striate. Frons densely and transversely striate. Eye glabrous, 1.2 × as high as broad. Malar space 0.4 × eye height and equal to basal width of mandible. Temple smooth. Face coarsely rugose-reticulate, narrowly smooth medially; its width 1.2 × eye height and 1.1 × height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.2 × its height. Pronotal keel distinct, its posterior branch situated closely to posterior margin of pronotum, but not fused with it; anterior branch fine and situated submedially. Mesoscutum rather highly and roundly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; densely granulate, with rather long and more or less distinct rugae near notauli and laterally; lateral lobes medially with wide granulate areas, with two convergent posteriorly carinae and narrow rugosity between them in medioposterior half. Notauli complete, wide, deep anteriorly and shallow posteriorly, coarsely crenulate. Prescutellar depression distinct, deep, with three carinae, 0.3 × as long as scutellum. Scutellum densely coriaceous-granulate, weakly convex, with distinct lateral carinae. Mesopleuron mainly smooth. Subalar depression shallow, wide, coarsely rugose-striate. Precoxal sulcus deep, sinuate, distinctly crenulate, running along entire length of lower part of mesopleuron. Propodeum with distinct tubercles, finely coriaceous-granulate in basal half, with areas more or less distinctly delineated by carinae; basal carina 1.2 × as long as anterior fork of areola; areola narrow and pentagonal; petiolate area distinctly separated from areola. Wings. Fore wing 4.2 × as long as wide. Vein r distinctly arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 23: 7: 43: 22. Second submarginal cell not narrowed distally, its length 3.1 × maximum width, 1.4 × length of first subdiscal cell. Vein cu-a postfurcal. Vein m-cu postfurcal. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, weakly curved towards base of wing. Legs. Hind coxa transversely striate and with granulation dorsally, coriaceous-granulate laterally and ventrally, with basoventral tooth and corner. Hind femur finely striate and granulate dorsally, finely coriaceous-granulate laterally and ventrally; 3.5 × as long as wide. Hind tibia apically with two spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 1.1 – 1.4 × maximum width of tibia. Hind tarsus 0.8 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) almost straight ventrally and distinct arched dorsally, with spiracular tubercles in basal third; sparsely striate, with rugulosity between striae. Length of petiole 2.3 × its apical width (dorsal view), 1.7 × length of propodeum. Second tergite without separated laterotergite. Second suture indistinct. Median length of second and third tergites combined 1.6 × basal width of second tergite, 0.7 × as long as their maximum width. Second and third tergites densely longitudinally striate, with additional reticulation between striae, smooth apically; fourth tergite finely striate in basal half, smooth in apical half, without separated laterotergites. Remaining tergites smooth. Ovipositor sheath 1.6 × as long as petiole, 0.5 × as long as metasoma, 0.9 × as long as mesosoma, 0.5 × as long as fore wing. Colour. Body reddish brown; mesoscutum and scutellum dark brown. Petiole reddish brown; second tergite dark brown; remaining tergites dark brown basally, reddish brown apically. Antenna brownish yellow basally, yellowish brown to brown in median 0.3, twelve subapical segments pale yellow, last apical segment brown. Palpi pale yellow. Legs yellowish brown, fore and middle coxae and all trochanters pale yellow. Ovipositor sheath brown, but medially widely pale brown. Fore wing faintly infuscate, more or less maculate. Pterostigma brown, yellow in narrow areas basally and apically. Variation. Body length 5.0 mm; fore wing length 3.4 mm. Fore wing 4.0 × as long as maximum width. Length of second submarginal cell 3.2 × its maximum width. Hind femur 2.9 × as long as wide. Length of petiole 2.2 × its apical width (dorsal view), 1.8 × length of propodeum. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEEFFFC838EFAB5FDC01A8B.taxon	materials_examined	Material examined. Holotype: female, China, Zhejiang Prov., Gutianshan, 3. VII. 2005, Shi Min, No. 200616383 (ZJUH). Paratype: 1 female, China, Zhejiang Prov., Gutianshan, 2. VII. 2005, Wu Qiong, No. 200615915 (ZJUH). Comparative diagnosis. This new species belongs to the S. moderabilis Wilkinson species group. Spathius gutianensis sp. nov. is similar to S. albithorax sp. nov., but differs in having the vertex entirely striate (mostly smooth, only finely striate medio-laterally in S. albithorax), mesopleuron almost smooth (weakly coriaceous-granulate in S. albithorax), fourth tergite only longitudinally striate (very dense semi-circularly anastomose striate and with very dense rugulosity between striae in S. albithorax), body dark (pale yellow in S. albithorax), and dorsally setae of hind tibia 1.1 – 1.4 × maximum width of tibia (1.9 – 2.2 × in S. albithorax).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEEFFFC838EFAB5FDC01A8B.taxon	etymology	Etymology. From the name of type locality of species.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEEFFFC838EFAB5FDC01A8B.taxon	distribution	Distribution. China (Zhejiang).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFECFFFC838EF8D0FCF21BDD.taxon	materials_examined	Material examined. China (ZJUH). Zhejiang Prov.: 1 female, Tianmushan Sanliting, 13. VI. 1998, Chen Xuexin, No. 980834.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFECFFFC838EF8D0FCF21BDD.taxon	distribution	Distribution. China (Zhejiang) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEAFFFA838EFF45FD421D90.taxon	description	(Fig. 15)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEAFFFA838EFF45FD421D90.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Hainan Prov., Shuiman, 26. V. 1960, Li Suofu. China (ZJUH). 1 female, Guangdong Prov., Longmen Nankunshan, 7. VIII. 2004, Xu Zaifu, No. 20053276.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEAFFFA838EFF45FD421D90.taxon	distribution	Distribution. China (Hainan, Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEAFFFB838EFE2BFDF3186E.taxon	description	(Fig. 16) Description. Female. Body length 8.8 mm; fore wing length 5.5 mm. Head. Antennae almost filiform, more than 19 - segmented (apical segments missing). Scape 1.8 × as long as its maximum width. First flagellar segment 5.5 × as long as its apical width, as long as second segment. Head width (dorsal view) 1.4 × median length. Head behind eyes (dorsal view) roundly narrowed; transverse diameter of eye 1.7 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 5: 5: 12. Vertex smooth. Frons entirely densely and regularly striate. Eye glabrous, 1.3 × as high as broad. Malar space 0.4 × eye height and 0.8 × basal width of mandible. Temple smooth. Face almost entirely transversely undulately striate, with fine rugulosity, clypeus rugose-striate; its width 1.2 × eye height and 1.2 × height of face and clypeus combined. Clypeal suture distinct, but shallow, without additional carina. Clypeus flat, with narrow ventral flange. Hypoclypeal depression rather small and round, its width 0.7 × distance from edge of depression to eye, 0.35 × width of face. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.5 × its height. Pronotal keel rather high, situated before middle of pronotum, its posterior branch distinctly separated from posterior margin of pronotum. Mesoscutum not highly and roundly elevated above pronotum, its median lobe distinctly protruding forwards, convex-rounded anteriorly (dorsal view), without anterolateral corners. Mesoscutum entirely covered by dense, semi-erect, mostly short and partly mixed with long setae; coarsely and long transversely striate with narrow granulate median longitudinal areas. Notauli deep anteriorly and shallow posteriorly, wide, complete, coarsely crenulate. Prescutellar depression shallow, long, with three carinae, finely rugulose, 0.4 × as long as scutellum. Scutellum almost smooth and flat, with lateral carinae. Mesopleuron almost entirely smooth. Subalar depression shallow, wide, coarsely and sparsely rugose-striate. Precoxal sulcus deep and wide in anterior 0.6, shallow and more or less narrow in posterior 0.4, weakly curved, coarsely crenulate, running along almost entire lower part of mesopleuron. Propodeum without tubercles, mostly coarsely rugulose-reticulate, with areas finely delineated by carinae, basolateral areas more or less granulate-punctate in basal half and coarsely rugose in posterior half; basal carina long, 2.3 × as long as anterior fork of areola; areola wide and rather short. Wings. Fore wing 4.3 × as long as wide. Vein r arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 15: 5: 37: 21. Second submarginal cell not narrowed distally, its length 3.4 × maximum width, 0.9 × length of first subdiscal cell. Vein m-cu postfurcal. Vein CU 1 a postfurcal. Hind wing vein M + CU 0.4 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, strongly oblique towards base of wing. Legs. Hind coxa dorsally transversely and rather finely semi-circularly striate, almost entirely smooth in lower half, without basoventral tooth and corner. Hind femur almost smooth, elongate-oval, 3.2 × as long as wide. Hind tibia apically without spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 1.5 – 1.6 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.6 × as long as basitarsus, 1.9 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) almost straight, more or less regularly thick, weakly widened in apical 0.2 (dorsal view), with weak or almost indistinct spiracular tubercles; with distinct rugosity and striation at least laterally and very densely almost entirely granulate-reticulate between striae. Length of petiole 7.3 × its apical width (dorsal view), 2.8 × length of propodeum. Second and third tergites with separate laterotergites. Second suture absent. Median length of second and third tergites combined 3.4 × basal width of second tergite, 1.7 × their maximum width. Second tergite densely longitudinally aciculate. Third tergite densely semi-circularly transversely aciculate. Fourth and sixth tergites entirely fine or very fine transversely aciculate. Ovipositor sheath 3.9 × as long as petiole, 1.9 × as long as metasoma, 4.2 × as long as mesosoma, 1.9 × as long as fore wing. Colour. Head brownish yellow. Mesosoma reddish brown, mesoscutum and scutellum brownish yellow. Petiole dark reddish brown; remaining tergites on wide area medially and apex of metasoma yellow, dark reddish brown laterally. Antenna brown to black. Palpi pale yellow. Legs yellow to pale yellow, femora widely infuscate medially, but faintly on fore and middle femora. Ovipositor sheath brown or pale brown, almost black apically. Fore wing more or less evenly faintly infuscate. Pterostigma brown, basally shortly yellow. Male. Body length 6.8 – 7.4 mm; fore wing length 5.1 – 5.3 mm. Antenna 38 - segmented. Length of petiole 9.2 – 9.6 × its apical width (dorsal view). Median length of second and third tergites combined 3.5 – 3.7 × basal width of second tergite. Legs often mostly pale. Otherwise similar to female.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEAFFFB838EFE2BFDF3186E.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Wuzhishan, 29. X. 2007, Liu Jingxian, No. 200710290 (ZJUH). Paratypes: 1 female, China, Hainan Prov., Wuzhishan Shuimanxiang, 16 – 18. V. 2007, Zeng Jie, No. 200807551 (ZJUH); 4 males, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, Nos. 200802386, 200802148, 200802345 and 200802481 (ZJUH, ZISP); 1 male, Hainan Prov., Bawangling, 26. XI. 2008, Wang Manman, No. 200805633 (ZJUH). Comparative diagnosis. This new species belongs to the S. imbecillus Enderlein species group. Spathius hainanicola sp. nov. is similar to S. vahalis Nixon, but differs in having the pronotal keel situated before middle of pronotum (behind middle of pronotum in S. vahalis), areola of propodeum present (absent in S. vahalis), vein r arising distinctly behind middle of pterostigma (from middle in S. vahalis), petiole with distinct rugosity and striation at least laterally (mostly strong shining and virtually without raised rugosity in S. vahalis), fourth and sixth tergites entirely finely or very finely transversely aciculate (smooth in S. vahalis), and ovipositor distinctly longer than body (as long as body in S. vahalis).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEAFFFB838EFE2BFDF3186E.taxon	etymology	Etymology. From Hainan island, the type locality of species, and “ incola ” (Latin for “ inhabitant ”) because this species was collected in Hainan Island.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEAFFFB838EFE2BFDF3186E.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEBFFFB838EFB7EFBD71983.taxon	materials_examined	Material examined. Female (holotype of Spathius xanthocephalus Chao, IZCAS), Fujian Prov., Jianyang Aotou, 21. VII. 1965, Zhuang Xingfa & She Rijian.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEBFFFB838EFB7EFBD71983.taxon	distribution	Distribution. China (Yunnan, Fujian, Hainan); Japan (?), Philippine, Vietnam.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEBFFFB838EF9D5FCA21AC4.taxon	materials_examined	Material examined. China (ZJUH). Yunnan Prov.: 1 female, Xishuangbanna, 31. VII. 2003, Xu Zaifu, No. 20053499.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEBFFFB838EF9D5FCA21AC4.taxon	distribution	Distribution. China (Yunnan) (new record); Philippines.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEBFFF9838EF897FCF21CC1.taxon	materials_examined	Material examined. China (ZJUH). 1 female, Zhejiang Prov., Songyang Ruoliaoxian, 17. VIII. 1994, Cai Ping, No. 944207.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFEBFFF9838EF897FCF21CC1.taxon	distribution	Distribution. China (Zhejiang) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE9FFF9838EFE9EFA6F1FF7.taxon	materials_examined	Material examined. China (ZJUH). Heilongjiang Prov.: 1 female, Yichun, 1985, Jin Liyuan, No. 850203. Jilin Prov.: 1 female, Tonghua, 1. VIII. 1994, Lou Juxian, No. 976611. Zhejiang Prov.: 1 female, Anji Longwangshan, 25. VI. 1996, Li Qiang, No. 963464; 1 female, Anji Longwangshan, 22. VII. 1996, Wu Hong, No. 971617; 2 females, Songyang, 18 – 31. VII. 1989, He Junhua, Nos. 895382 and 895396; 4 females, Songyang, 1 – 6. VIII. 1989, He Junhua, Nos. 895459, 895452, 895473 and 895454; 1 female, Songyang, 21. VIII. 1989, He Junhua, No. 895250; 1 female, Xitianmushan, 30. IX. 1998, Zhao Mingshui, No. 20002746; 1 female, Qingyuan, 1 – 2. VIII. 1985, Wu Quancong, No. 851946; 1 female, Fengyangshan Fengyangmiao, 26. VII. 2007, Liu Jingxian, No. 200809152; 1 female, Fengyangshan Huangmaojian, 30. VII. 2007, Liu Jingxian, No. 200801494. Fujian Prov.: 1 female, Wuyishan, 7. IX – 1. XII. 1989, Wang Jiashe, No. 964676; 1 female, Erliping, IX. 1979, Xu Jianfei, No. 20003845. Hunan Prov.: 16 females, Xinning, 25. IV. 1981, Tong Xinwang, Nos. 20044966 – 20044981 (1 female in ZISP); 2 females, Xinning, 28. IV. 1981, Tong Xinwang, Nos. 20044984 and 20044987. Guizhou Prov.: 1 female, Kuankuoshui, 9. VI. 2010, Tan Jiangli, No. 201004508. Sichuan Prov.: 1 female, Xichong, 24. V. 1938, Zhu Ruzuo, No. 6501615.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE9FFF9838EFE9EFA6F1FF7.taxon	distribution	Distribution. China (Heilongjiang, Jilin, Zhejiang, Fujian, Hunan, Guizhou, Sichuan) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE9FFF9838EFBC7FCEE18C8.taxon	materials_examined	Material examined. China (ZJUH). Fujian Prov.: 1 female, Longqishan, 30. IX. 1991, Liu Changming, No. 20007229.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE9FFF9838EFBC7FCEE18C8.taxon	distribution	Distribution. China (Fujian) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE9FFF7838EFA80FAE61896.taxon	materials_examined	Material examined. China (ZJUH). Zhejiang Prov.: 2 females, Gutianshan, 2 – 3. VII. 2005, Shi Min, Nos. 200615928 and 200616395; 1 female, Gutianshan, 3. VII. 2005, Wu Qiong, No. 200616456; 1 female, Suichang Jiulongshan, 18. VIII. 1994, Chen Xuexin, No. 944740; 1 female, Xitianmushan, 20. VI. 1996, Chen Xuexin, No. 972199. Fujian Prov.: 3 females, Yongtai Chibi, 17. IX. 2002, Liu Jingxian, Nos. 20023552, 20023533 and 20023637; 2 females, Yongtai Qingyunshan, 18. IX. 2002, Li Fangfang, Nos. 20023183 and 20023190; 1 female, Yongtai Daiyunshan, 13. IX. 2002, Liu Jingxian, No. 20025057; 1 female, Yongan Tianbaoyan, 19. VII. 2001, Piao Meihua, No. 200107044; 2 females, 1 male, Dehua Jiuxianshan, 12. IX. 2002, Xu Zaifu, Nos. 20024082, 20024081 and 20024775. Hunan Prov.: 1 female, Shimen Hupingshan, 13. VII. 2009, Ma Li, No. 200900931; 2 females, Shimen Hupingshan, 8 and 10. VII. 2009, Tang Pu, Nos. 200901276 and 200902021. 1 female, Liuyang, 30. IX. 1984, Tong Xinwang, No. 20044611; Guangdong Prov.: 5 females, Shixing Chebaling, 27. VII. 2002, Xu Zaifu, Nos. 20051711, 20051738, 20051875, 20051866 and 20051844; 3 females, Fuyuan Nanling, 8. V. 2004, Xu Zaifu, Nos. 20049546, 20049628, and 20049529; 4 females, Nanling, 16 – 21. VII. 2008, Xu Zaifu, Nos. 200809163, 200809166, 200809167 and 200807902; 1 female, Yunan Tongledashan, 12 – 13. VIII. 2003, Xu Zaifu, No. 20054533; 2 females, Fogang Guanyinshan, 12. V. 2004, Xu Zaifu, Nos. 20053329 and 20053330; 2 females, Fengkai Heishiding, 15. VIII. 2003, Chen Jujian, Nos. 20048912 and 20048920; 1 female, Zijin Qimuzhang, 31. VII. 2003, Song Yanxia, No. 20059034; 1 female, Meizhou Fengxi, 29. VII. 2003, Chen Jujian, No. 20048370; 2 females, Dinghushan, 6 – 7. IV. 2002, Xu Zaifu, Nos. 20026431 and 20026630; 1 female, Yangchun Baitong, 5 – 6. V. 2002, Xu Zaifu, No. 20028089; 1 female, Nankunshan, 8. VI. 2002, Xu Zaifu, No. 20028867; 3 females, Yangshan Chengjia, 25. VII. 2002, Xu Zaifu, Nos. 20029311, 20029290 and 20029310. Guangxi Prov.: 1 female, Shiwandashan, 29. XI. 2001, Ma Yun, No. 20021561; 1 female, Guangxi Botanical Garden, 30. X. 2002, Lin Naiquan, No. 20034964; 2 females, Nanling, 30. X. 2002, Lin Naiquan, Nos. 20035876 and 20035902. Hainan Prov.: 1 female, Bawangling, 26. XI. 2008, Tan Jiangli, No. 200805705; 1 female, Bawangling, 9 – 10. VI. 2007, Liu Jingxian, No. 200703581; 1 female, Yinggeling, 28. V – 3. VI. 2007, Weng Liqiong, No. 200804069; 2 females, Jianfengling Tianchi, 22 – 23. X. 2007, Liu Jingxian, Nos. 2007010729 and 2007010814; 4 females, Jianfengling, 5 – 7. VI. 2007, Weng Liqiong, Nos. 200806449, 200806454, 200806452 and 200806455; 5 females, Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, Nos. 200802334, 200802141, 200802144, 200802409 and 200802329; 1 female, Wuzhishan, 15 – 16. V. 2008, Liu Jingxian, No. 200905328. Yunnan Prov.: 4 females, Tengchong Jietouxiang, 11 – 12. VII. 2006, Zeng Jie, Nos. 200801835, 200801808, 200801643 and 200801841; 2 females, Yingjiang Tongbiguan, 20. V. 2009, Zeng Jie, Nos. 200905282 and 20095288; 1 female, Tengchong Datianpo, 22. V. 2009, Zeng Jie, No. 200904499; 4 females, Xishuangbanna, 31. VII. 2003, Xu Zaifu, Nos. 20053576, 20053536, 20053545 and 20053585. Guizhou Prov.: 1 female, Mayanghe Huangguatuo, 1. X. 2007, Liu Jingxian, No. 200709079; 1 female, Mayanghe Wanjia, 27 – 30. IX. 2007, Liu Jingxian, No. 200708540; 1 female, Leigongshan Xiaodanjiang, 5. VI. 2005, Zhang Hongying, No. 200606222; 1 female, Huishui Jiulongshan, 11. VI. 2010, Zeng Jie, No. 201003830.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE9FFF7838EFA80FAE61896.taxon	distribution	Distribution. China (Zhejiang, Fujian, Hubei, Hunan, Guangdong, Guangxi, Hainan, Yunnan, Guizhou); Japan, Korea.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE9FFF7838EFA80FAE61896.taxon	discussion	Remarks. The type material of S. fukienensis Chao was not found in the collection of Institute of Zoology, Beijing, China (IZCAS). However, according to the description and figures in Chao (1957), S. fukienensis has its subapical segments of antenna pale yellow, metasoma behind petiole compressed, petiole distinctly longer than propodeum, dorsal side of hind tibia with mixed short and long setae, mesoscutum and vertex distinctly striate. These characters clearly show that S. fukienensis Chao is a junior synonym of S. japonicus Watanabe.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE7FFF7838EFB3BFBE31A08.taxon	materials_examined	Material examined. China (ZJUH). Zhejiang Prov.: 1 female, Gutianshan, 3. VII. 2005, Wu Qiong, No. 200616432. Guangdong Prov.: 1 female, Nanling, 16 – 21. VII. 2008, Xu Zaifu, No. 200809165. Hainan Prov.: 1 female, Bawangling, 9 – 10. VI. 2007, Liu Jingxian, No. 200703501; 1 female, Jianfengling, 4 – 7. VI. 2007, Zeng Jie, No. 200711142; 1 female, Jianfengling Tianchi, 25. XI. 2008, Wang Manman, No. 200806076; 1 female, Yinggeling, 24 – 25. V. 2007, Liu Jingxian, No. 200702723.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE7FFF7838EFB3BFBE31A08.taxon	distribution	Distribution. China (Zhejiang, Guangdong, Hainan) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE7FFF5838EF958FF311CE5.taxon	materials_examined	Material examined. China (ZJUH). Yunnan Prov.: 1 female, Daweishan, 18. VII. 2003, Xu Zaifu, No. 20055114; 1 female, Pingbian Daweishan, 18. VII. 2003, 1970 – 2107 m, Li Tingjing, No. 20045292 (SIZP). Hainan Prov.: 1 female, Yinggeling, 23 – 25. V. 2007, Xiao Bin, No. 200807381. Zhejiang Prov.: 1 female, Lin’an Qingliangfeng, 10. VIII. 2005, Zhang Hongying, No. 200607443. Guangdong Prov.: 1 female, Fuyuan Nanling, 4. VIII. 2004, Xu Zaifu, No. 20049860. Henan Prov.: 1 female, Neixiang Baotianman, 14. VII. 1998, Chen Xuexin, No. 988297; 1 female, Neixiang Baotianman, 13 – 15. VII. 1998, Ma Yun, No. 987395; Fujian Prov.: 1 female, Nanjing Leshang 8. VII. 1991, Liu Changming, No. 20006248; 1 female, Dazhulan, 15. VI. 1980, Lin Naiquan, No. 20004027.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE7FFF5838EF958FF311CE5.taxon	distribution	Distribution. China (Henan, Zhejiang, Fujian, Guangdong, Hainan, Yunnan) (new record); Russia (Far East), Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE5FFF5838EFEF5FB531E27.taxon	materials_examined	Material examined. China (ZJUH). Guangdong Prov.: 1 female, Shixing Chebaling, 27. VII. 2002, Xu Zaifu, No. 20051750. Hainan Prov.: 1 female, Yinggeling, 24 – 26. V. 2007, Liu Jingxian, No. 200704046.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE5FFF5838EFEF5FB531E27.taxon	distribution	Distribution. China (Guangdong, Hainan) (new record); Russia (Far East), Korea, Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE5FFF5838EFDB7FDCE1F79.taxon	description	(Fig. 17)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE5FFF5838EFDB7FDCE1F79.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Yunnan Prov., Luxi Mangshi Santaishan, 18. V. 1935, Wu Le. Distribution. China (Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE5FFF5838EFC70FBF318E5.taxon	materials_examined	Material examined. China (ZJUH). 1 female, Guizhou Prov., Fanjingshan, 11. VII. 1993, Yao Songlin, No. 93706; Zhejiang Prov.: 2 females, Gutianshan, 3. VII. 2005, Wu Qiong, Nos. 200616465 and 200616096; 1 female, Gutianshan, 18. VIII. 2003, Yu Xiaoxia, No. 20043905. Guangdong Prov.: 1 female, Fuyuan Nanling, 4. VIII. 2004, Xu Zaifu, No. 20049715.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE5FFF5838EFC70FBF318E5.taxon	distribution	Distribution. China (Zhejiang, Guangdong, Guizhou) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE5FFF3838EFAF5FDF31D9B.taxon	description	(Fig. 18) Description. Female. Body length 5.2 mm; fore wing length 3.7 mm. Head. Antennae weakly setiform, 35 - segmented. Scape 1.6 × as long as its maximum width. First flagellar segment about 5.0 × as long as its apical width, 1.2 × as long as second segment. Penultimate segment about 4.0 × as long as wide, as long as subpointed apical segment. Head width (dorsal view) 1.6 × its median length, 1.2 × as wide as mesoscutum. Head behind eyes (dorsal view) roundly narrowed; transverse diameter of eye 1.6 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.1 × its sides; POL: Od: OOL = 3.5: 3.5: 10. Vertex smooth anteriorly and posteriorly, distinctly transverse striate medially; frons entirely and vertex in posterior half distinctly and rather densely transverse striate. Eye glabrous, 1.2 × as high as broad. Malar space 0.4 × eye height and 0.8 × basal width of mandible. Temple mainly smooth, finely vertically striate posteriorly near occipital carina. Face with rather dense, coarse and weakly curved transverse striation. Face width 1.1 × eye height and almost equal to height of face and clypeus combined. Hypoclypeal depression large and round, its width 0.85 × distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, ventrally joined with hypostomal carina by additional ruga upper base of mandible. Mesosoma. Length of mesosoma 2.0 × its height. Pronotal keel distinct, its posterior branch rather distinctly separated from posterior margin of pronotum, but joined with it by two additional fine sublateral rugae, anterior branch situated behind middle of pronotum. Mesoscutum highly and roundly elevated above pronotum, its median lobe convex anteriorly and without anterolateral corners. Mesoscutum densely granulate, with long and dense rugae along notauli and laterally, with two distinct and convergent posteriorly undulate carinae in medioposterior half; mostly glabrous, with rather dense, long and almost erect pale setae arranged narrowly along notauli and laterally. Notauli anteriorly distinct, wide, and crenulate, weakly shallow posteriorly. Prescutellar depression deep, long, with three carinae, almost smooth, 0.3 × as long as scutellum. Scutellum entirely densely granulate, weakly convex, with distinct lateral carinae. Mesopleuron mostly smooth. Subalar depression shallow, narrow and sparsely striate. Precoxal sulcus deep, straight, weakly oblique, coarsely crenulate, running along anterior 0.6 of lower part of mesopleuron. Propodeum with short and rather narrow lateral tubercles, with distinctly delineated by carinae and granulate with rugosity basolateral areas, its rest part mainly smooth and with coarse sparse striate; areola rather narrow and relatively short; basal carina present in basal 0.3, about as long as anterior fork of areola. Wings. Fore wing 4.2 × as long as wide. Vein r arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 26: 6: 44: 24. Second submarginal cell not narrowed distally, its length 3.0 × maximum width, 1.4 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior quarter of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.55 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, distinctly curved towards base of wing, faintly infuscate. Legs. Hind coxa dorsally coarsely transversely striate, densely granulate on rest part, with distinct basoventral corner and tooth. Hind femur mainly smooth, finely longitudinally striate dorsally, 3.5 × as long as wide. Hind tibia apically without spines on outer side. Dorsal surface of hind tibia with long, rather sparse and semi-erect setae, their length 1.0 – 1.5 × maximum width of tibia. Hind tarsus as long as hind tibia. Hind basitarsus 0.65 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 1.3 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly curved ventrally, dorsally distinctly arched in basal 0.3 and almost straight in apical 0.7, distinctly thickened in basal 0.3; widened in apical fifth (dorsal view), with small spiracular tubercles in widened basal third. Length of petiole 3.9 × its apical width, 2.6 × length of propodeum. Second tergite without separate laterotergites. Median length of second and third tergites combined 1.5 × basal width of second tergite, 0.7 × as long as their maximum width. Second suture absent. Petiole distinctly and rather sparsely longitudinally striate, densely striate apically, with dense and fine reticulation in medio-basal half. Remaining tergites entirely smooth. Ovipositor weakly curved down. Ovipositor sheath 2.5 × as long as petiole, 1.3 × as long as metasoma, 2.2 × as long as mesosoma, about as long as fore wing. Colour. Body reddish brown, with pale spots on head and mesosoma; metasoma behind petiole light reddish brown with narrow yellow posterior transverse stripes. Antennae pale brown in basal third, brown in median third, whitish yellow in apical third (about 10 segments). Palpi pale yellow. Fore and middle legs pale brown to yellowish brown, basally white. Hind leg frown to pale brown, trochanter, trochantellus and base of femur whitish; hind tibia almost evenly pale brown. Ovipositor sheath brownish yellow in basal third and brown to dark brown on remaining part. Fore wing faintly infuscate, with brownish spots near vein 1 - M and below pterostigma. Pterostigma brown, pale in basal third. Variation. Body length 4.5 – 5.6 mm; fore wing length 3.4 – 4.1 mm. First flagellar segment 5.0 – 5.6 × as long as its apical width, 1.1 – 1.2 × as long as second segment. Transverse diameter of eye 1.6 – 1.8 × length of temple. Vertex sometimes finely or very finely aciculate. Malar space 0.35 – 0.45 × eye height. Length of petiole 4.0 – 4.5 × its apical width, 2.2 – 2.5 × length of propodeum. Ovipositor sheath 1.3 – 1.5 × as long as metasoma, 2.1 – 2.3 × as long as mesosoma. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE5FFF3838EFAF5FDF31D9B.taxon	materials_examined	Material examined. Holotype. female, China, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802455 (ZJUH). Paratypes: 1 female, China, Hainan Prov., Yinggeling, 28. V – 3. VI. 2007, Weng Liqiong, No. 200804197 (ZJUH); 2 females, Hainan Prov., Jianfengling Tianchi, 12 – 15. VII. 2006, Weng Liqiong, Nos. 200803321 and 200803426 (ZJUH, ZISP). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius longulator sp. nov. is similar to S. colophon Nixon, but differs in having the hind coxa and femur brown (mainly bright yellow or honey yellow in S. colophon), base of ocellar triangle weakly larger than its sides (very clearly larger in S. colophon), pronotal keel distinct (rather week in S. colophon), mesoscutum with distinct granulation (with fine reticulation medially on mesoscutal lobes in S. colophon), scutellum distinctly granulate (almost smooth in S. colophon), basal carina of propodeum long (short in S. colophon), basolateral areas of propodeum densely and distinctly granulate with rugosity (mainly smooth in S. colophon), and petiole more than 2.0 × longer than propodeum (not longer than 2.0 × in S. colophon). This new species is also similar to the Japanese S. insulicola Belokobylskij et Maeto, but differs from it in having the petiole long and slender (shorter and wide in S. insulicola), first flagellar segment slender (thick in S. insulicola), vertex smooth in anterior part (entirely sculptured in S. insulicola), mesoscutum not highly and gently elevated above pronotum (highly and roundly elevated in S. insulicola). The differences between Chinese S. longulator sp. nov. and S. longus Chen et Shi are showed in the key.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE5FFF3838EFAF5FDF31D9B.taxon	etymology	Etymology. From “ longus ” (Latin for “ long ”), because of long petiole.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE5FFF3838EFAF5FDF31D9B.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE3FF8F838EFE2DFDC1186E.taxon	description	(Fig. 19) Description. Female. Body length 5.1 mm; fore wing length 3.8 mm. Head. Antennae weakly setiform, more than 29 - segmented (apical segments missing). Scape 1.7 × as long as its maximum width. First flagellar segment 5.5 × as long as its apical width, 1.25 × as long as second segment. Subapical segments about 3.0 × as long as their width. Head width (dorsal view) 1.7 × its median length, 1.3 × as wide as mesoscutum. Head behind eyes (dorsal view) convex-roundly narrowed; transverse diameter of eye 1.4 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 3.0: 4.0: 10.0. Vertex distinctly evenly coarsely transversely striate, without additional sculpture between striae, smooth laterally. Eye almost glabrous, 1.25 × as high as broad. Malar space 0.4 × eye height and 0.6 × basal width of mandible. Temple entirely smooth. Face with dense, distinct and weakly curved transverse striae and with fine additional reticulation. Face width equal to eye height and almost equal to height of face and clypeus combined. Hypoclypeal depression large and subround, its width about equal to distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, ventrally not joined with hypostomal carina obliterating upper base of mandible. Mesosoma. Length of mesosoma 2.1 × its height. Pronotal keel distinct, its posterior branch fused with posterior margin of pronotum on very short distance; anterior branch situated near middle of pronotum. Mesoscutum highly and evenly roundly elevated above pronotum, its median lobe convex anteriorly and without anterolateral corners. Mesoscutum densely granulate, with rather long and dense rugae situated along notauli and laterally, with two distinct and weakly convergent posteriorly, almost straight carinae in medioposterior third; mostly glabrous, with rather sparse, long and semi-erect golden setae arranged widely along notauli and in single line laterally. Notauli anteriorly distinct, rather wide, crenulate, deep anteriorly and shallow posteriorly. Prescutellar depression rather shallow, not long, with four carinae, reticulate-granulate, 0.3 × as long as scutellum. Scutellum entirely densely and finely coriaceous, weakly convex, with distinct lateral carinae. Mesopleuron almost entirely finely coriaceous, anteriorly almost smooth. Subalar depression shallow, wide, densely and curvedly striate with rugosity. Precoxal sulcus distinct and coarsely crenulate, deep, rather wide, distinctly sinuate, deep along anterior 0.5 and more shallow along posterior 0.5 of lower part of mesopleuron. Propodeum with short and wide lateral tubercles, with basolateral areas distinctly delineated by carinae and mainly rugose-striate, rest part of propodeum mainly and densely rugose-reticulate; areola narrow and relatively long; basal carina present in basal third, about as long as anterior fork of areola. Wings. Fore wing 3.8 × as long as wide. Vein r arising from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 29: 6: 56: 22. Second submarginal cell not narrowed distally, its length 3.0 × maximum width, 1.4 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising almost from middle of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu weakly antefurcal, oblique towards base of wing, weakly curved, very faintly infuscate. Legs. Hind coxa dorsally densely transversely striate with rather fine granulation, densely granulate below at rest part, with distinct basoventral corner and tooth. Hind femur mainly smooth, finely longitudinally striate dorsally, 3.2 × as long as wide. Hind tibia apically without spines on outer side. Dorsal surface of hind tibia with rather long, dense and semi-erect setae, their length 0.7 – 1.0 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.8 × as long as second-fifth segments combined. Second segment of hind tarsus 0.4 × as long as basitarsus, almost as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) very weakly curved ventrally, dorsally weakly and evenly arched, weakly thickened in anterior third; widened in apical fifth (dorsal view), with small spiracular tubercles in weakly widened basal third. Length of petiole 3.0 × its apical width, 2.0 × length of propodeum. Second tergite without separate laterotergites. Median length of second and third tergites combined 1.4 × basal width of second tergite, 0.85 × as long as their maximum width. Second suture absent. Petiole distinctly and rather densely longitudinally striate, with dense and fine reticulation between striae. Second tergite finely striate in basolateral third and interruptedly finely coriaceous submedially. Remaining tergites entirely smooth. Ovipositor straight. Ovipositor sheath 3.6 × as long as petiole, 1.8 × as long as metasoma, 2.5 × as long as mesosoma, as long as fore wing. Colour. Body dark reddish brown to almost black partly, head yellowish brown, dark posteriorly. Antennae pale brown basally, brown to dark brown mainly, without visible pale subapical segments. Palpi yellow. Legs yellow to brownish yellow, hind coxa light reddish brown, hind femur medially brown, yellow basally, hind tibia basally dark. Ovipositor sheath brown almost entirely. Fore wing faintly and almost evenly infuscate. Pterostigma brown, pale yellow in basal third and apically. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE3FF8F838EFE2DFDC1186E.taxon	materials_examined	Material examined. Holotype. female, China, Zhejiang Prov., Kaihua Gutianshan, 1. VIII. 1990, Ma Yun, No. 906119 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius marcrurus sp. nov. is similar to S. ogasawarus Belokobylskij et Maeto, but differs in having the vertex evenly and smooth striate (rugulose-striate in S. ogasawarus), temple longer (shorter in S. ogasawarus), rugae on mesoscutum along notauli short (distinctly long in S. ogasawarus), areola of propodeum short and basal carina long (areola very long and basal carina short in S. ogasawarus), hind coxa granulate laterally (rugose-striate in S. ogasawarus), and the second metasomal tergite partly with fine aciculation (entirely smooth in S. ogasawarus). This new species is also similar to Chinese S. carterus Chen et Shi, but differs in having the malar space short (long in S. carterus), mesoscutum roundly elevated above pronotum (almost sharply elevated in S. carterus), petiole 2.0 × as long as propodeum (1.2 × in S. carterus), and ovipositor as long as body (shorter than body in S. carterus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE3FF8F838EFE2DFDC1186E.taxon	etymology	Etymology. From “ macro ” (Greek for “ long ”) and “ urus ” (Greek for “ limb ”) because of ovipositor long.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFE3FF8F838EFE2DFDC1186E.taxon	distribution	Distribution. China (Zhejiang).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9FFF8F838EFB7EFD9C198C.taxon	description	(Fig. 20)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9FFF8F838EFB7EFD9C198C.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Yunnan Prov., 27. VI. 1957, Zang Linchao. China (ZJUH). Yunnan Prov.: 1 female, Jinghong, 31. VII. 2003, Lu Jia, No. 20045948.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9FFF8F838EFB7EFD9C198C.taxon	distribution	Distribution. China (Hubei, Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9FFF8F838EF9DFFBAD1B90.taxon	materials_examined	Material examined. China (ZJUH). Zhejiang Prov.: 8 females, Kaihua Gutianshan, VII – VIII. 1990, Xu Zhihong, Nos. 905339, 905634, 905402, 905573, 905443, 905664, 905575 and 905396; 4 females, Kaihua Gutianshan, VII – VIII. 1990, Ma Yun, Nos. 905742, 905896, 905770 and 905809; 4 females, Gutianshan, 1. VIII. 1990, Ma Yun, Nos. 906181, 906213, 906159 and 906179; 1 female, Gutianshan, 1. VII. 2005, Zhang Hongying, No. 200616605. Guangdong Prov.: 1 female, Shixing Chebaling, 21. VIII. 2003, Xu Zaifu, No. 20052647. Hainan Prov.: 1 female, Jianfengling Tianchi, 12 – 15. VII. 2006, Liu Jingxian, No. 200803618.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9FFF8F838EF9DFFBAD1B90.taxon	distribution	Distribution. China (Zhejiang, Guangdong, Hainan, Taiwan); India, Sri Lanka.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9DFF8D838EFF45FBCE1DB4.taxon	materials_examined	Material examined. China (ZJUH). Guangdong Prov.: 1 female, Xinyi Dawuling, 1. V – 6. V. 2002, Xu Zaifu, No. 20025944. Hainan Prov.: 1 female, Yinggeling, 18. X. 2007, Liu Jingxian, No. 200709887.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9DFF8D838EFF45FBCE1DB4.taxon	distribution	Distribution. China (Guangdong, Hainan) (new record); India, Philippines.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9DFF8D838EFE07FCDA1ECB.taxon	materials_examined	Material examined. China (ZJUH). Hainan Prov.: 1 female, Jianfengling, 4 – 7. VI. 2007, Zeng Jie, No. 200711346; 1 female, Jianfengling, 7. VI. 2007, Liu Jingxian, No. 200702518.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9DFF8D838EFE07FCDA1ECB.taxon	distribution	Distribution. China (Fujian, Yunnan, Hainan, Taiwan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9DFF8D838EFC9DFDFC180C.taxon	description	(Fig. 21)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9DFF8D838EFC9DFDFC180C.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Fujian Prov., Jianyang Guadun, 25. IX. 1975, Chen Jiahua. Distribution. China (Fujian).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9DFF88838EFB5FFD9D1F93.taxon	description	(Fig. 22) Description. Female. Body length 7.0 mm; fore wing length 5.0 mm. Head. Antennae weakly setiform, more than 36 - segmented (apical segments missing). Scape 2.0 × as long as its maximum width. First flagellar segment 5.8 × as long as its apical width, 1.3 × as long as second segment. Subapical segments about 4.0 × as long as their width. Head width (dorsal view) 1.5 × its median length, 1.2 × as wide as mesoscutum. Head behind eyes (dorsal view) weakly roundly narrowed; transverse diameter of eye 1.6 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 3.0: 3.5: 11.0. Vertex entirely smooth; frons entirely and rather densely transverse undulately striate. Eye with sparse and very short setae, 1.15 × as high as broad. Malar space 0.5 × eye height and 0.9 × basal width of mandible. Temple smooth. Face rather densely, distinctly and weakly undulately transversely striate, additionally with fine and dense granulation. Face width almost equal to eye height and 1.1 × height of face and clypeus combined. Hypoclypeal depression rather large and round, its width 0.8 × distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, ventrally not joined with hypostomal carina being obliterated upper base of mandible. Mesosoma. Length of mesosoma 2.3 × its height. Pronotal keel high, distinct, its posterior branch separated from posterior margin of pronotum, but additionally joined with it by two additional distinct lateral rugae, anterior branch distinct and situated submedially. Mesoscutum highly and roundly elevated above pronotum, its median lobe convex anteriorly and without anterolateral corners. Mesoscutum densely granulate, with rather long and dense rugae situated along notauli and laterally, with two distinct, subparallel and weakly undulate carinae in medioposterior half; mostly glabrous, with rather dense, long and semi-erect pale setae arranged widely along notauli and narrowly laterally. Notauli anteriorly distinct, wide, and crenulate, shallow posteriorly. Prescutellar depression rather shallow, long, with five carinae, finely rugulose to smooth between carinae, 0.35 × as long as scutellum. Scutellum entirely densely and distinctly granulate, almost flat, with distinct lateral carinae. Mesopleuron mostly finely reticulate-granulate. Subalar depression shallow, rather narrow, sparsely striate. Precoxal sulcus deep, distinctly sinuate, densely and coarsely crenulate, running along entire lower length of mesopleuron. Propodeum with short and wide lateral tubercles, with basolateral areas distinctly delineated by carinae and granulate with rugosity along carinae, rest part of propodeum mainly striate-rugose with dense granulation; areola wide and relatively short; basal carina present in basal 0.3, about as long as anterior fork of areola. Wings. Fore wing 3.8 × as long as wide. Vein r arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 36: 10: 66: 34. Second submarginal cell not narrowed distally, its length 3.3 × maximum width, 1.3 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior fifth of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu very weakly antefurcal, distinctly curved towards base of wing, distinctly infuscate. Legs. Hind coxa dorsally transversely striate with granulation, densely granulate at rest part, with distinct basoventral corner, but almost without tooth. Hind femur mainly smooth, longitudinally striate dorsally, 3.4 × as long as wide. Hind tibia apically with one-two spines on outer side. Dorsal surface of hind tibia with long, rather dense and semi-erect setae, their length 1.2 – 1.8 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.75 × as long as second-fifth segments combined. Second segment of hind tarsus 0.45 × as long as basitarsus, 1.3 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) almost straight ventrally, dorsally weakly arched in basal half and almost straight in apical half, thickened in basal half; widened in apical quarter (dorsal view), with small but distinct spiracular tubercles in widened basal third. Length of petiole 2.7 × its apical width, 1.9 × length of propodeum. Second tergite without separate laterotergites. Median length of second and third tergites combined 1.1 × basal width of second tergite, 0.65 × as long as their maximum width. Second suture absent. Petiole distinctly and densely longitudinally striate, with dense and fine reticulation between striae. Remaining tergites entirely smooth. Ovipositor straight. Ovipositor sheath 2.8 × as long as petiole, 1.2 × as long as metasoma, 1.8 × as long as mesosoma, 0.9 × as long as fore wing. Colour. Head pale yellow with brown face and large spots behind eye, malar area almost white. Mesosoma yellow, with a few brown to dark brown stripes and spots dorsally and laterally, dark brown to black below. Petiole dark reddish brown to black, remaining metasoma pale yellow, second tergite almost entirely and narrow transverse stripes on other tergites brown to dark brown. Antenna yellow in basal half, but scape and pedicel brownish yellow, segments behind its middle infuscate, several subapical segments whitish. Palpi pale yellow. Legs whitish yellow to yellow, fore and middle femora submedially and hind femur in apical half brown to dark brown, hind tibia almost entirely whitish yellow. Ovipositor sheath brownish yellow to yellow, brown apically. Fore wing faintly infuscate, darkened more intensive along veins. Pterostigma brown, pale basally and apically. Variation. Body length 5.1 – 7.3 mm; fore wing length 3.7 – 4.9 mm. Antennae 50 - segmented. First flagellar segment 6.0 – 6.3 × as long as its apical width, 1.25 – 1.40 × as long as second segment. Penultimate segments 4.5 × as long as wide, about as long as apical segment. Transverse diameter of eye 1.3 – 1.5 × length of temple. Length of mesosoma 2.2 – 2.3 × its height. Hind femur 3.1 – 3.6 × as long as wide. Length of petiole 2.8 – 3.1 × its apical width, 1.9 – 2.0 × length of propodeum. Median length of second and third tergites combined 1.25 – 1.35 × basal width of second tergite, 0.7 – 0.8 × as long as their maximum width. Ovipositor sheath 2.5 – 3.5 × as long as petiole, 1.0 – 1.5 × as long as metasoma, 1.8 – 2.4 × as long as mesosoma, 0.8 – 1.2 × as long as fore wing. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9DFF88838EFB5FFD9D1F93.taxon	materials_examined	Material examined. Holotype. female, China, Hainan Prov., Jianfengling, 6. VI. 2007, Liu Jingxian, No. 200703707 (ZJUH). Paratypes: 1 female, China, Fujian Prov., Wuyishan Xianfengling, 13. IX. 1989, No. 20005834 (ZJUH); 1 female, Hainan Prov., Jianfengling Tianchi, 12 – 15. VII. 2006. Zhang Wenyong, No. 200803529 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius moscoides sp. nov. is similar to S. moscus Nixon, but differs in having the colour of body mainly pale (mainly dark reddish brown in S. moscus), basal carina of propodeum long (short in S. moscus), recurrent vein of fore wing weakly postfurcal (strongly postfurcal in S. moscus), mesopleuron mat and almost entirely densely granulate (highly polished and smooth in S. moscus), pronotal keel fused by additional lateral carinae with posterior margin of mesoscutum (directly narrowly fused in S. moscus), setae on mesoscutum dense (very sparse in S. moscus), lateral lobes of mesoscutum with fine and short rugae along notauli (with well-marked rugae in S. moscus), scutellum distinctly granulate (entirely smooth in S. moscus), petiole almost twice longer than propodeum (distinctly shorter in S. moscus). New species is also similar to S. erymanthus Nixon, but differs in having the colour of body mainly pale (mainly fulvous red in S. erymanthus), hind tibia whitish yellow (brown but honey yellow basally in S. erymanthus), ocelli arranged in triangle with base larger its sides (ocellar triangle equilateral in S. erymanthus), pronotal keel fused by additional lateral carinae with posterior margin of mesoscutum (not fused in S. erymanthus), scutellum distinctly granulate (with weak rugosity in S. erymanthus), propodeal areola distinctly delineated by carinae (almost not defined in S. erymanthus), mesopleuron densely and finely granulate and without rugosity (mainly with distinct striation in S. erymanthus), ovipositor shorter (longer in S. erymanthus). S. moscoides sp. nov. differs from Japanese S. nagoyaensis Belokobylskij et Maeto in having the hind coxa whitish yellow (mostly dark reddish brown to black in S. nagoyaensis), subapical segments of antenna pale (dark in S. nagoyaensis), colour of body mainly pale (mainly fulvous red in S. nagoyaensis), mesosoma longer (shorter in S. nagoyaensis), recurrent vein of fore wing weakly postfurcal (distinctly postfurcal in S. nagoyaensis), hind coxa almost without basoventral tooth (with distinct tooth in S. nagoyaensis), and vertex entirely smooth (at least partly striate in S. nagoyaensis). According Chao (1977 a) and Chen & Shi (2004), this new species run to S. helle Nixon (S. xanthocephalus Chao), but differs from it in having the pronotal keel fused by additional lateral carinae with posterior margin of mesoscutum (directly widely fused in S. helle), mesoscutum with short rugae along notauli (with long rugae in S. helle), vertex entirely smooth (partly with fine interrupted striation in S. helle), and petiole longer (shorter in S. helle).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9DFF88838EFB5FFD9D1F93.taxon	etymology	Etymology. From “ moscus ”, the name of the most similar species, and suffix “ oides ” because of this species resemble S. moscus Nixon.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF9DFF88838EFB5FFD9D1F93.taxon	distribution	Distribution. China (Fujian, Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF98FF88838EFC25FDFC18D4.taxon	description	(Fig. 23)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF98FF88838EFC25FDFC18D4.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Fujian Prov., Xiqin, 25. VII. 1965. Distribution. China (Fujian).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF98FF87838EFAFDFDF31D9B.taxon	description	(Fig. 24) Description. Female. Body length 5.7 mm; fore wing length 3.9 mm. Head. Antennae weakly setiform, 40 - segmented. Scape 1.8 × as long as its maximum width. First flagellar segment 4.5 × as long as its apical width, 1.1 × as long as second segment. Penultimate segment 3.5 × as long as wide. Head width (dorsal view) 1.7 × its median length, 1.3 × as wide as mesoscutum. Head behind eyes (dorsal view) distinctly roundly narrowed; transverse diameter of eye about 2.0 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.1 × its sides; POL: Od: OOL = 3.0: 4.0: 11.0. Vertex almost smooth, very finely reticulate-aciculate medio-anteriorly; frons entirely and densely transversely curvedly striate. Eye with sparse and very short setae, 1.2 × as high as broad. Malar space 0.4 × eye height and 0.8 × basal width of mandible. Temple smooth. Face densely, distinctly and weakly undulately transversely striate and without additional granulation. Face width 1.1 × eye height and 1.15 × height of face and clypeus combined. Hypoclypeal depression rather large and rounded, its width 0.8 × distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, ventrally joined by additional ruga with hypostomal carina upper base of mandible. Mesosoma. Length of mesosoma 2.2 × its height. Pronotal keel high, its posterior branch distinctly separated from posterior margin of pronotum, but joined with it by two additional distinct lateral rugae, anterior branch fine and situated submedially. Mesoscutum highly and roundly elevated above pronotum, its median lobe convex anteriorly and without anterolateral corners. Mesoscutum densely granulate, with rather long and dense rugae situated along notauli and laterally, with two distinct and convergent posteriorly carinae in medioposterior half; mostly glabrous, with sparse, long and semi-erect pale setae arranged narrowly along notauli and in single line laterally. Notauli anteriorly deep, wide, coarsely crenulate, shallow and narrow posteriorly. Prescutellar depression rather deep, long, with three carinae, densely granulate-rugulose between carinae, 0.4 × as long as scutellum. Scutellum entirely densely and distinctly granulate, weakly convex, with distinct lateral carinae. Mesopleuron mainly smooth. Subalar depression rather shallow, wide, sparsely striate, but almost smooth in lower half .. Precoxal sulcus deep and oblique in anterior 0.7, shallow and curved in posterior 0.3, coarsely and usually densely crenulate, running along entire lower length of mesopleuron. Propodeum with short and wide lateral tubercles, with basolateral areas distinctly delineated by carinae and weakly granulate-rugulose to rugose posteriorly, rest part of propodeum mainly and densely striate-rugose without granulation; areola narrow and short; basal carina present in basal 0.4, about 2.0 × as long as anterior fork of areola. Wings. Fore wing 3.9 × as long as wide. Vein r arising from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 28: 5: 43: 22. Second submarginal cell not narrowed distally, its length 3.5 × maximum width, 1.4 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior third of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu strongly antefurcal, straight, distinctly oblique towards base of wing, infuscate. Legs. Hind coxa dorsally transversely curvedly striate with dense granulation, densely granulate with reticulation at rest part, with weak round basoventral corner, without tooth. Hind femur longitudinally rugulose-striate dorsally, rugulose-reticulate laterally, 3.0 × as long as wide. Hind tibia apically without spines on outer side. Dorsal surface of hind tibia with long, rather sparse and semi-erect setae, their length 1.0 – 1.4 × maximum width of tibia. Hind tarsus 0.85 × as long as hind tibia. Hind basitarsus 0.8 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 1.4 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) very weakly curved ventrally, dorsally weakly arched in basal half and straight in apical half, thickened in basal third; widened in apical fifth (dorsal view), with small spiracular tubercles in widened basal third. Length of petiole 3.4 × its apical width, 2.0 × length of propodeum. Second tergite in basal half with separate laterotergites. Median length of second and third tergites combined 1.5 × basal width of second tergite, 0.7 × as long as their maximum width. Second suture absent. Petiole distinctly and sparsely (but densely in apical fifth) longitudinally striate, with dense and fine reticulation between striae. Remaining tergites smooth, but second tergite very finely reticulate basally. Ovipositor almost straight. Ovipositor sheath 2.8 × as long as petiole, 1.35 × as long as metasoma, almost 2.0 × as long as mesosoma, almost as long as fore wing. Colour. Body light reddish brown, head yellowish brown. Antenna yellowish brown in basal half and brown to dark brown in apical half, without pale subapical segments. Palpi pale whitish. Legs yellow to yellowish brown or pale brown, fore and middle coxae and all trochanters and trochantelli white to whitish yellow, hind femur reddish brown to dark reddish brown, hind tibia basally brownish, pale yellow in subbasal half. Ovipositor sheath brownish yellow to light brown, brown apically. Fore wing distinctly infuscate, more dark along veins and below pterostigma. Pterostigma brown, pale yellow in basal third and pale brown anteriorly. Variation. Body length 3.7 – 5.0 mm; fore wing length 2.6 – 3.5 mm. Antennae 35 - segmented. First flagellar segment 5.0 × as long as its apical width. Penultimate segments 4.0 × as long as wide, about as long as apical segment. Transverse diameter of eye 1.8 – 2.0 × length of temple. Length of mesosoma 2.3 – 2.4 × its height. Hind femur 3.0 – 3.1 × as long as wide. Length of petiole 3.8 – 4.0 × its apical width, 2.0 – 2.1 × length of propodeum. Median length of second and third tergites combined 1.5 – 1.7 × basal width of second tergite, 0.7 – 0.9 × as long as their maximum width. Ovipositor sheath 2.4 – 2.6 × as long as petiole, 1.1 – 1.3 × as long as metasoma, 1.9 × as long as mesosoma, 0.9 × as long as fore wing. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF98FF87838EFAFDFDF31D9B.taxon	materials_examined	Material examined. Holotype. female, China, Hainan Prov., Diaoluoshan, 28. V – 01. VI. 2007, Zeng Jie, No. 200806716 (ZJUH). Paratypes: 3 females, China, Hainan Prov., Jianfengling Tianchi, 25. XI. 2008, Wang Manman, Nos. 200806093, 200806088 and 200806104 (ZJUH, ZISP). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius nehebrus sp. nov. is similar to S. hebrus Nixon, but differs in having the colour of body mainly light reddish brown (mainly dark reddish brown to black in S. hebrus), hind tibia whitish in basal half and brownish yellow in apical half (honey yellow and very faintly darkened medially in S. hebrus), fore wing distinctly infuscate (faintly infuscate in S. hebrus), ocelli arranged in triangle with base weakly larger its sides (considerably larger in S. hebrus), mesopleuron mainly smooth (mainly sculptured in S. hebrus), setae on dorsal surface of hind tibia shorter (longer in S. hebrus), second segment of hind tarsus longer (shorter in S. hebrus), petiole distinctly shorter and distinctly widened apically (long and weakly widened apically in S. hebrus), and ovipositor shorter (longer in S. hebrus). New species differs from S. longipetiolus Belokobylskij et Maeto in having the posterior branch of pronotal keel distinctly separated from posterior margin of pronotum (weakly separated in S. longipetiolus), hind coxa without basoventral tubercle (with distinct tubercle in S. longipetiolus), hind femur wide (more narrow in S. longipetiolus), and mesoscutum with distinct rugae (short or indistinct rugae in S. longipetiolus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF98FF87838EFAFDFDF31D9B.taxon	etymology	Etymology. From “ ne ” (Russia for “ not ”), and “ hebrus ”, the name of the most similar species.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF98FF87838EFAFDFDF31D9B.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF97FF85838EFE2DFDCE19D3.taxon	description	(Fig. 25) Description. Female. Body length 4.8 mm; fore wing length 3.1 mm. Head. Antennae weakly setiform, more than 37 - segmented (apical segments missing). Scape 1.6 × as long as its maximum width. First flagellar segment 5.0 × as long as its apical width, 1.25 × as long as second segment. Subapical segments 4.0 × as long as their width. Head width (dorsal view) 1.6 × its median length, 1.25 × as wide as mesoscutum. Head behind eyes (dorsal view) distinctly roundly narrowed; transverse diameter of eye 1.8 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.1 × its sides; POL: Od: OOL = 3.5: 4.0: 9.0. Vertex entirely smooth; frons densely transversely curvedly striate, smooth laterally. Eye with sparse and very short setae, 1.25 × as high as broad. Malar space 0.35 × eye height and 0.9 × basal width of mandible. Temple mainly smooth. Face densely, distinctly and linearly transversely striate, with additional reticulation. Face width 1.1 × eye height and 1.2 × height of face and clypeus combined. Hypoclypeal depression medium sized and suboval, its width equal to distance from edge of depression to eye, 0.45 × width of face. Occipital carina complete dorsally, ventrally obliterated on short distance, but almost joined by additional ruga with hypostomal carina upper base of mandible. Mesosoma. Length of mesosoma about 2.0 × its height. Pronotal keel rather high, its posterior branch fused with posterior margin of pronotum on wide distance, anterior branch high and situated submedially. Mesoscutum highly and roundly elevated above pronotum, its median lobe distinctly convex anteriorly and without anterolateral corners. Mesoscutum densely granulate, with short or very short and dense rugae situated along notauli, with two distinct and convergent posteriorly carinae in medioposterior 0.4; mostly glabrous, with rather dense, short and semi-erect pale setae arranged in narrow stripes along notauli and in single line laterally. Notauli anteriorly deep, wide, coarsely crenulate, shallow posteriorly. Prescutellar depression rather deep, short, with four carinae, almost smooth between carinae, 0.3 × as long as scutellum. Scutellum mainly smooth, partly with very fine reticulation, weakly convex, with fine lateral carinae. Mesopleuron mainly smooth. Subalar depression shallow, narrow, distinctly striate with rugosity. Precoxal sulcus deep, oblique in anterior 0.7, shallow and upcurved in posterior 0.3, coarsely and rather densely crenulate, running along entire lower length of mesopleuron. Propodeum with short and wide lateral tubercles, with basolateral areas distinctly delineated by carinae and finely reticulate medially and striate along carinae, rest part of propodeum mainly and rather densely reticulate-rugose, without granulation; areola narrow and long; basal carina present in basal 0.2, about 0.5 × as long as anterior fork of areola. Wings. Fore wing 4.4 × as long as wide. Vein r arising from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 24: 4: 42: 17. Second submarginal cell not narrowed distally, its length 3.2 × maximum width, 1.3 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior third of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu weakly antefurcal, straight, distinctly oblique towards base of wing, weakly infuscate. Legs. Hind coxa dorsally transversely curvedly striate, smooth or almost smooth at rest part, with distinct basoventral corner and tooth. Hind femur entirely smooth, 3.2 × as long as wide. Hind tibia apically with four spines on outer side. Dorsal surface of hind tibia with long, rather sparse and semi-erect setae, their length 0.5 – 1.1 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 1.15 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly curved ventrally, dorsally evenly and rather distinctly arched, thickened in basal third; widened in apical one-sixth (dorsal view), with spiracular tubercles in weakly widened basal third. Length of petiole 2.5 × its apical width, 1.7 × length of propodeum. Second tergite with separate laterotergite. Median length of second and third tergites combined 1.7 × basal width of second tergite, almost as long as their maximum width. Second suture absent. Petiole coarsely and sparsely longitudinally striate, with dense and distinct reticulation between striae. Remaining tergites smooth, but second tergite finely reticulate-striate basally. Ovipositor almost straight. Ovipositor sheath 3.5 × as long as petiole, 1.3 × as long as metasoma, 2.1 × as long as mesosoma, 1.1 × as long as fore wing. Colour. Head reddish brown to dark reddish brown, paler around eyes, yellowish dorso-laterally. Mesosoma and petiole black, metasoma behind petiole dark reddish brown. Antenna reddish brown to dark brown, yellow to brownish yellow basally, apically without pale segments. Palpi whitish yellow. Legs yellowish brown to pale brown, all trochanters whitish yellow, hind femur light reddish brown, hind tibia basally brownish, pale yellow in subbasal third, then light brown; hind tarsus dark reddish brown. Ovipositor sheath brown, pale basally. Fore wing faintly and evenly infuscate. Pterostigma brown, yellow in basal quarter. Variation. Body length 4.7 – 5.0 mm; fore wing length 3.3 mm. First flagellar segment 5.0 – 5.3 × as long as its apical width. Transverse diameter of eye 1.8 – 2.0 × length of temple. Mesosoma with reddish spots; length of mesosoma 1.85 – 1.95 × its height. Basal carina of propodeum 0.35 – 0.40 × as long as anterior fork of areola. Hind femur 3.0 – 3.3 × as long as wide. Length of petiole 2.2 – 2.5 × its apical width, 1.70 – 1.75 × length of propodeum. Median length of second and third tergites combined 1.5 × basal width of second tergite, 0.9 × as long as their maximum width. Ovipositor sheath 3.2 – 3.4 × as long as petiole, 1.4 – 1.6 × as long as metasoma, 1.9 – 2.1 × as long as mesosoma, 0.9 – 1.0 × as long as fore wing. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF97FF85838EFE2DFDCE19D3.taxon	materials_examined	Material examined. Holotype. female, China, Yunnan Prov., Hekou, 20 – 21. VII. 2003, Xu Zaifu, No. 20055254 (ZJUH). Paratypes: 1 female, China, Yunnan Prov., Hekou, 20 – 21. VII. 2003, Xu Zaifu, No. 20055256 (ZISP); 1 female, Yunnan Prov., Hekou, 20. VII. 2003, 107 m, Jiang Qian, No. 20045469 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius neleiformis sp. nov. is very similar to S. neleus Nixon, but differs in having the head behind eyes (dorsal view) distinctly narrowed and distinctly transverse (weakly narrowed and subcubical in S. neleus), setae on mesoscutum rather short, semi-erect and sparse (long, almost erect and rather dense in S. neleus), median mesoscutal lobe without transverse striation and with very short rugae along notauli (with vague transverse striation and with long rugae along notauli in S. neleus), and hind femur elongate-oval (subclavate in S. neleus). New species differs from S. fasciatus Walker in having the head behind eyes (dorsal view) short and distinctly narrowed (long and weakly narrowed in S. fasciatus), first flagellar segment slender (thick in S. fasciatus), second tergites with separate laterotergites and aciculate on wide basal area (without separate laterotergites and smooth in S. fasciatus), and basal carina of propodeum short (longer in S. fasciatus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF97FF85838EFE2DFDCE19D3.taxon	etymology	Etymology. From “ neleus ”, the name of the most similar species, and “ formis ” (Latin for “ form ”) because they are morphologically similar species.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF97FF85838EFE2DFDCE19D3.taxon	distribution	Distribution. China (Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF95FF85838EF9E5FD981B70.taxon	description	(Fig. 26)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF95FF85838EF9E5FD981B70.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Fujian Prov., Jianyang Huangkenggongshe Dazhulan 19. VII. 1967, Chen Jiahua.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF95FF85838EF9E5FD981B70.taxon	distribution	Distribution. China (Fujian, Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF95FF83838EF84BFED41EB3.taxon	distribution	Distribution. China (Zhejiang, Fujian, Guangdong, Guangxi, Yunnan) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF95FF83838EF84BFED41EB3.taxon	discussion	Remark. The doubtful specimens of S. opis ssp. rufovariegatus from Phu Lang Thuong, (Nixon, 1943 a: 289 – 290) is probably the same described species, but the location (Phu Lang Thuong) is actually not from China but from Vietnam.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF80838EFF45FAC71DDE.taxon	materials_examined	Material examined. China (ZJUH). Henan Prov.: 1 female, Jigongshan, 12. VII. 1997, Chen Xuexin, No. 973907. Zhejiang Prov.: 8 females, 4 males, Ningbo Xiangshan, 15. X. 2004, Wei Shujun, Nos. 20059149 – 20059160; 3 females, Ningbo Xiangshan, 13. IX. 2004, Wei Shujun, Nos. 20053605 – 20053607. Yunnan Prov.: 1 female, Anning Hotspring, 18 – 20. VII. 1988, Chen Xuexin, No. 881721.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF80838EFF45FAC71DDE.taxon	distribution	Distribution. China (Henan, Zhejiang, Fujian, Yunnan); Indonesia, Malaysia, Philippines, Vietnam.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF80838EFDEEFDCA1F7C.taxon	description	(Fig. 27)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF80838EFDEEFDCA1F7C.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Sichuan Prov., Emeishan Xianxiangchi, 17. VIII. 1957, Zhu Fuxing.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF80838EFDEEFDCA1F7C.taxon	distribution	Distribution. China (Sichuan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF80838EFC4FFC811900.taxon	description	(Fig. 28)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF80838EFC4FFC811900.taxon	distribution	Distribution. China (Jilin); Japan, Russia (Far East).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF80838EFC4FFC811900.taxon	discussion	Remark. Re-examination of the type material of S. exarator oriens Belokobylskij (Belokobylskij, 1998) described from the Russian Far East and Japan shows that previously used diagnostic features are actually characteristic at the species level of this taxon (stat. nov.). Comparison of this species with description of S. changbaishanensis Chen et Shi (type material of this taxon is not availiable for study) distinctly shows that S. changbaishanensis is the junior synonym of S. oriens (syn. nov.)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF80838EFA5BFCFE1A45.taxon	materials_examined	Material examined. China: 1 female, Guangdong Prov., Yunan Tongledashan, 12 – 13. VIII. 2003, Xu Zaifu, No. 20054410 (ZJUH).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF80838EFA5BFCFE1A45.taxon	distribution	Distribution. China (Guangdong, Yunnan, Taiwan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF9C838EF914FCF11EC6.taxon	description	(Fig. 29) Description. Female. Body length 4.9 mm; fore wing length 3.5 mm. Head. Antennae slender, weakly setiform, more than 37 - segmented (apical segments missing). Scape 1.5 × as long as its maximum width. First flagellar segment 4.4 × as long as its apical width, 1.1 × as long as second segment. Head not depressed, its width (dorsal view) 1.4 × median length. Head behind eyes roundly narrowed. Transverse diameter of eye 2.1 × length of temple. Ocelli arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 2: 2: 5. Vertex smooth. Frons densely and transversely striate. Eye glabrous, 1.2 × as high as broad. Malar space 0.4 × eye height and 0.8 × basal width of mandible. Temple smooth. Face entirely coarsely rugose-reticulate, clypeus rugose-striate; its width 1.1 × eye height and 1.2 × height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.4 × its height. Pronotal keel distinct, its posterior branch slightly fused with posterior margin of pronotum; anterior branch fine and situated submedially. Pronotal lateral depression wide, partly delineated by carinae, densely and coarsely crenulate. Mesoscutum rather highly and perpendicularly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; distinctly densely granulate, with short carinae along notauli and laterally, with wide and only granulate median areas on lateral and median lobes; with two convergent posteriorly carinae and narrow rugosity between them in medioposterior half. Notauli complete, wide, deep anteriorly and shallow posteriorly, coarsely crenulate. Prescutellar depression distinct, deep, with three carinae, 0.3 × as long as scutellum. Scutellum coriaceous-granulate, weakly convex, with distinct lateral carinae. Mesopleuron mostly smooth. Subalar depression shallow, rather wide, densely and coarsely rugose-striate. Precoxal sulcus shallow, wide, weakly oblique, crenulate, running along anterior 0.6 of lower part of mesopleuron. Propodeum without tubercles, densely and coarsely rugose-reticulate, densely granulate basally or in basal half, with areas more or less distinctly delineated by carinae; basal carina 1.5 × as long as anterior fork of areola; areola large and pentagonal; petiolate area distinctly separated from areola. Wings. Fore wing 3.8 × as long as wide. Vein r arising from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 14: 4: 28: 13. Second submarginal cell not narrowed distally, its length 3.2 × maximum width, 1.2 × length of first subdiscal cell. Vein cu-a almost interstitial. Vein m-cu postfurcal. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu interstitial, strongly oblique towards base of wing. Legs. Hind coxa finely rugose and granulate, with basoventral tooth and corner. Hind femur almost smooth, 3.2 × as long as wide. Hind tibia apically with four spines on outer margin. Dorsal surface of hind tibia with rather long, sparse and semi-erect setae, their length 1.7 – 2.0 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.8 × as long as second-fifth segments combined. Second segment of hind tarsus 0.4 × as long as basitarsus, 1.1 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) ventrally straight and distinctly arched dorsally, with small spiracular tubercles in basal third; distinctly and entirely striate, with dense rugulosity between striae. Length of petiole 2.0 × its apical width (dorsal view), 1.6 × length of propodeum. Second-fourth tergites with separated laterotergites. Second suture indistinct. Median length of second and third tergites combined 1.5 × basal width of second tergite, 1.1 × as long as their maximum width. Second and third tergites densely and anastomose striate, with very dense rugulose microsculpture between striae, reticulate-punctate subapically, apically narrowly smooth; fourth tergite very dense reticulate-areolate, with short striation basally, finely reticulate to smooth apically. Fifth tergite very finely and sparsely coriaceous-reticulate in basal half. Remaining tergites smooth. Ovipositor sheath 3.3 × as long as petiole, 1.1 × as long as metasoma, 2.0 × as long as mesosoma, 0.8 × as long as fore wing. Colour. Body yellowish brown. Mesosoma reddish brown; pronotum and mesopleuron dark brown. Metasoma almost black, with yellowish spots apically on each tergites. Antenna yellowish brown, faintly darkened toward apex. Palpi pale yellow. Legs yellowish brown, fore and middle coxae and all trochanters pale yellow; hind coxa dorsally and hind femur medially with reddish brown spots; all tibiae basally and submedially reddish brown; all tarsi pale reddish brown. Ovipositor sheath dark brown. Fore wing faintly infuscate. Pterostigma brown. Variation. Body length 4.6 – 5.6 mm; fore wing length 3.4 – 3.9 mm. First flagellar segment 4.3 – 4.6 × as long as its apical width, as long as second segment. Transverse diameter of eye 2.0 – 2.2 × length of temple. Eye 1.3 × as high as broad. Basal carina of propodeum 1.3 – 1.6 × as long as anterior fork of areola. Fore wing 3.6 – 3.9 × as long as maximum width. Length of second submarginal cell 3.0 × its maximum width. Hind femur 2.9 × as long as wide. Length of petiole 2.2 × its apical width (dorsal view), 1.8 × length of propodeum. Ovipositor sheath 3.2 × as long as petiole, 1.0 – 1.2 × as long as metasoma. Male. Body length 3.4 – 4.8 mm; fore wing length 2.8 – 3.3 mm. Hind femur 2.8 – 3.0 × as long as wide. Length of petiole 2.6 – 2.8 × its apical width (dorsal view), 1.8 × length of propodeum. Median length of second and third tergites combined 1.9 – 2.1 × basal width of second tergite, 1.3 – 1.4 × their maximum width. Black areas on metasomal tergites sometimes wide. Sometimes mesosoma or all body pale. Otherwise similar to female.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF9C838EF914FCF11EC6.taxon	materials_examined	Material examined. Holotype: female, China, Zhejiang Prov., Gutianshan, 1. VIII. 1990, Ma Yun, No. 906224 (ZJUH). Paratypes: 1 male, China, Zhejiang Prov., Tianmushan, 2 – 4. VI. 1990, He Junhua, No. 905038 (ZJUH); 1 female, 1 male, Guangdong Prov., Shixing Chebaling, 21. VIII. 2003, Xu Zaifu, Nos. 20052313 and 20052345 (ZJUH); 1 female, Guangdong Prov., Yunnan Tongledashan, 12 – 13. VIII. 2003, Xu Zaifu, No. 20054591 (ZJUH); 1 male, Guangdong Prov., Shixing Chebaling, 21. VIII. 2003, Xu Zaifu, No. 20051980 (ZISP); 1 female, Hainan Prov., Bawangling, 9 – 10. VI. 2007, Liu Jingxian, No. 200703451 (ZJUH); 1 female, Hainan Prov., Diaoluoshan, 16 – 17. VI. 2006, Liu Jingxian, No. 200802142 (ZISP); 1 female, Hainan Prov., Jianfengling, 7. VI. 2007, Liu Jingxian, No. 200702482 (ZJUH). Comparative diagnosis. This new species belongs to the S. leucippus Nixon species group. Spathius parallelus sp. nov. differs from all members of S. leucippus species group in the short temple (long in all other species), ovipositor much shorter than body (almost as long as body length in all other species), petiole strongly widened towards apex (slightly widened towards apex in all other species), legs without dark spots (with more or less dark spots in all other species), mesoscutum elevated almost perpendicularly above pronotum (roundly, not perpendicularly above pronotum in all other species), and acrosternite of first segment short (long in all other species).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF9C838EF914FCF11EC6.taxon	etymology	Etymology. From “ parallelus ” (Greek for “ parallel ”), because of its metasoma parallel-sided in posterior half.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF90FF9C838EF914FCF11EC6.taxon	distribution	Distribution. China (Zhejiang, Guangdong, Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8CFF9C838EFC96FBE3191F.taxon	materials_examined	Material examined. China (ZJUH). Zhejiang Prov.: 2 females, Kaihua Gutianshan, VII – VIII. 1990, Xu Zhihong, Nos. 905539 and 905540; 2 females, Gutianshan, 20. VII. 1992, Chen Xuexin, Nos. 923710 and 923719; 1 female, Gutianshan, 1. VIII. 1990, Ma Yun, No. 906116; 1 female, Qingyuan Baishanzu, 18. VII. 1994, Wu Hong, No. 946766; 1 female, Qingyuan Baishanzu, 20. VII. 1994, Wu Hong, No. 946942; 1 female, Tianmushan Laodian, 28. VII. 1998, Zhao Mingshui, No. 20001041. Guangdong Prov.: 2 females, Shixing Chebaling, 27. VII. 2002, Xu Zaifu, Nos. 20051868 and 20051732; 1 female, Shixing Chebaling, 21. VIII. 2003, Xu Zaifu, No. 20052294; 2 females, Meizhou Fengxi, 29. VII. 2003, Chen Jujian, Nos. 20048562 and 20048601. Hainan Prov.: 1 female, Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802326 (ZJUH); 1 female, Diaoluoshan, 16 – 17. VII. 2006, Weng Liqing, No. 200802475 (ZJUH).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8CFF9C838EFC96FBE3191F.taxon	distribution	Distribution. China (Zhejiang, Guangdong, Hainan) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8CFF9B838EFAA5FCF11D76.taxon	description	(Fig. 30) Description. Female. Body length 5.0 mm; fore wing length 3.3 mm. Head. Antennae slender, filiform, 41 - segmented. Scape 1.5 × as long as its maximum width. First flagellar segment 6.3 × as long as its apical width, 1.1 × as long as second segment. Head width (dorsal view) 1.4 × median length. Head behind eyes (dorsal view) roundly narrowed; transverse diameter of eye 1.6 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.3 × its sides; POL: Od: OOL = 6: 5: 14. Vertex smooth. Frons finely striate. Eye glabrous, 1.3 × as high as broad. Malar space 0.4 × eye height and 0.8 × basal width of mandible. Temple smooth. Face strongly transversely striate and with fine rugosity; its width 1.1 × eye height and 1.2 × height of face and clypeus combined. Clypeal suture distinct, but shallow, without additional carina. Clypeus flat, with narrow ventral flange. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.6 × its height. Pronotal keel posterior branch distinctly separated from posterior margin of pronotum. Lateral areas of pronotum distinctly striate. Mesoscutum not highly and roundly elevated above pronotum, its median lobe protruding forwards, convex-rounded anteriorly (dorsal view), without anterolateral corners. Mesoscutum with sparse long and almost erect setae situated widely along notauli and narrowly marginally; densely granulate, with additional short rugosity on lateral lobes. Notauli deep anteriorly and shallow posteriorly, wide, complete, coarsely crenulate. Prescutellar depression shallow, with three carinae, finely rugulose, 0.4 × as long as scutellum. Scutellum finely granulate-coriaceous, almost flat, with lateral carinae. Mesopleuron smooth. Subalar depression shallow, wide, almost smooth. Precoxal sulcus weakly curved, coarsely crenulate, running along almost entire lower part of mesopleuron. Propodeum without tubercles, mainly irregularly rugose-reticulate with a few striation; basal carina long, 2.4 × as long as anterior fork of areola; areola long and narrow. Wings. Fore wing 4.6 × as long as wide. Vein r arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 14: 3: 23: 12. Second submarginal cell not narrowed distally, its length 3.4 × maximum width, 0.9 × length of first subdiscal cell. Vein m-cu postfurcal. Vein CU 1 a postfurcal. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, strongly oblique towards base of wing. Legs. Hind coxa dorsally transversely and semi-circularly striate, almost entirely finely granulate in lower half, without basoventral tooth and corner. Hind femur almost smooth, elongate-oval, 3.2 × as long as wide. Hind tibia apically with two spines on outer margin. Dorsal surface of hind tibia with rather long, sparse and semi-erect setae, their length 1.8 – 2.3 × maximum width of tibia. Hind tarsus as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Second segment of hind tarsus 0.6 × as long as basitarsus, 1.6 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) almost straight, more or less regularly thick, weakly widened in apical 0.2 (dorsal view), with indistinct spiracular tubercles in basal third; sparsely and finely striate and finely coriaceous and granulate between striae in basal and apical third, only finely granulate-coriaceous on median third. Length of petiole 8.6 × its apical width (dorsal view), 3.0 × length of propodeum. Second tergite with separate laterotergite. Second suture indistinct. Median length of second and third tergites combined 3.1 × basal width of second tergite, 1.3 × their maximum width. Second tergite; third tergite finely semi-circularly transversely aciculate. Rest tergites smooth. Ovipositor sheath 3.3 × as long as petiole, 1.6 × as long as metasoma, 3.3 × as long as mesosoma, 1.5 × as long as fore wing. Colour. Head yellowish brown. Mesosoma and petiole almost reddish brown. Remaining tergites darker than petiole. Antenna brownish yellow, faintly infuscate towards apex. Palpi pale yellow. Legs brownish yellow, fore and middle coxae and trochanters pale yellow, hind coxa and femur reddish brown. Ovipositor sheath brown or pale brown, almost black apically. Fore wing more or less faintly infuscate. Pterostigma dark brown, basally shortly yellow. Variation. Body length 4.2 – 7.3 mm; fore wing length 2.9 – 5.0 mm. Antennae 49 - segmented. First flagellar segment 5.0 × as long as its apical width, as long as second segment. Face width equal to eye height and 1.1 × height of face and clypeus combined. Mesoscutum densely granulate, with short rugosity on lateral lobes. Fore wing 4.3 × as long as maximum width. Length of second submarginal cell 3.9 × its maximum width, 1.1 × length of first subdiscal cell. Hind femur 3.5 × as long as wide. Length of petiole 7.8 × its apical width (dorsal view), 2.8 × length of propodeum. Median length of second and third tergites combined 2.9 – 3.2 × basal width of second tergite, 1.4 × their maximum width. Ovipositor sheath 3.9 × as long as petiole, 1.7 × as long as metasoma, 3.7 × as long as mesosoma, 1.7 × as long as fore wing. Head blackish brown or black. Mesoscutum, scutellum and mesopleuron dark reddish brown or black. Propodeum black. Petiole reddish brown. Male. Unknown	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8CFF9B838EFAA5FCF11D76.taxon	materials_examined	Material examined. Holotype: female, China, Guangdong Prov., Fogang Guanyinshan, 15 – 16. IX. 2007, Xu Zaifu, No. 200711666 (ZJUH). Paratypes: 1 female, China, Guangdong Prov., Fogang Guanyinshan, 29. VII. 2003, Chen Jujian, No. 20048680 (ZISP); 1 female, Guangdong Prov., Nanling, 16 – 21. IX. 2008, Xu Zaifu, No. 200809164 (ZJUH); 1 female, Zhejiang Prov., Xitianmushan Jinshanmen, 4. VIII. 1998, Zhao Mingshui, No. 20002616 (ZJUH); 1 female, Hainan Prov., Yinggeling, 24 – 26. V. 2007, Liu Jingxian, No. 200704131 (ZJUH); 1 female, Hainan Prov., Yinggeling, 28. V – 3. VI. 2007, Weng Liqiong, No. 200804060 (ZJUH); 1 female, Hainan Prov., Yinggeling, 16. XI. 2008, Tan Jiangli, No. 200805780 (ZISP); 1 female, Hainan Prov., Wuzhaishan, 29. X. 2007, Liu Jingxian, No. 200710027 (ZJUH). Comparative diagnosis. This new species belongs to the S. imbecillus Enderlein species group. Spathius parimbecillus sp. nov. is similar to S. imbecillus Enderlein, but differs in having the face strongly transversely striate with fine rugosity (coarsely reticulate-rugose with fine transverse striation in S. imbecillus), lateral areas of pronotum distinctly striate (complete smooth in S. imbecillus), propodeum mainly irregularly rugose-reticulate with a few striae (strongly and characteristically transversely carinate in S. imbecillus), metapleuron entirely areolate-reticulate with rugosity (mostly smooth in S. imbecillus), and scutellum coriaceous or granulate-coriaceous with striation laterally (shining and virtually smooth in S. imbecillus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8CFF9B838EFAA5FCF11D76.taxon	etymology	Etymology. From “ par ” (Greek for “ near ”), and the specific name “ imbecillus ”, because of its similar to S. imbecillus.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8CFF9B838EFAA5FCF11D76.taxon	distribution	Distribution. China (Guangdong, Zhejiang, Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8BFF9B838EFE46FCE51E6F.taxon	materials_examined	Material examined. 1 female, China, Hainan Prov., Yinggeling, 16. XI. 2008, Tan Jiangli, No. 200805215 (ZJUH). Distribution. China (Hainan) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8BFF9B838EFD7FFA9418B1.taxon	materials_examined	Material examined. China (ZJUH). Zhejiang Prov.: 1 female, Tianmushan Laodian, 14. VII. 1998, Zhao Mingshui, No. 20000852. Fujian Prov.: 1 female, Tongmu, X. 1979, Huang Juchang, No. 20003851. Yunnan Prov.: 1 female, Xishuangbanna, 31. VII. 2003, Xu Zaifu, No. 20053526.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8BFF9B838EFD7FFA9418B1.taxon	distribution	Distribution. China (Zhejiang, Fujian, Yunnan); former Czechoslovakia, France, Hungary, Italy, Japan, Korea, Lithuania, Mongolia, Russia, Spain, Ukraine.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8BFF9B838EFD7FFA9418B1.taxon	discussion	Remarks. According to the description and figures in Chen & Shi (2004), S. applanatus Chen et Shi has its head and mesosoma distinctly depressed, ovipositor sheath not longer than metasoma and hind coxa with basoventral tooth and corner. These and other characters allow synonymising S. applanatus with S. phymatodis.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8BFF9B838EFB0EFE671A3B.taxon	materials_examined	Material examined. China (ZJUH). Yunnan Prov.: 1 female, Xishuangbanna, 31. VII. 2003, Xu Zaifu, No. 20053554.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8BFF9B838EFB0EFE671A3B.taxon	distribution	Distribution. China (Yunnan) (new record); Indonesia.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8BFF9B838EFB0EFE671A3B.taxon	discussion	Remarks. The ovipositor of this specimen from Yunnan province is slightly longer than the holotype of this species from BMNH.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8BFF9B838EF9B3FB471B7D.taxon	materials_examined	Material examined. China (ZJUH). Inner Mongolia Prov.: 1 female, Xiziwangqi, 18. VIII. 2000, Chen Xuexin, No. 20032020. Zhejiang Prov.: 1 female, Xitianmushan, 1. VIII. 1998, Zhao Mingshui, No. 20002957.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF8BFF9B838EF9B3FB471B7D.taxon	distribution	Distribution. China (Inner Mongolia, Zhejiang) (new record); Japan, Russia (Far East).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF88FF99838EFF45FDC11F0E.taxon	description	(Fig. 31) Description. Female. Body length 2.9 mm; fore wing length 2.3 mm. Head. Antennae filiform, 27 - segmented. Scape 1.6 × as long as its maximum width. First flagellar segment 4.6 × as long as its apical width, almost as long as second segment. Penultimate segment 3.5 × as long as wide, almost as long as subpointed apical segment. Head width (dorsal view) 1.5 × its median length, 1.3 × as wide as mesoscutum. Head behind eyes (dorsal view) evenly convex-roundly narrowed; transverse diameter of eye 1.6 × length of temple. Ocelli medium-sized, arranged in equilateral triangle; POL: Od: OOL = 1.5: 2.5: 9.0. Vertex entirely smooth; frons densely and linearly aciculate, smooth laterally and posteriorly. Eye glabrous, 1.15 × as high as broad. Malar space 0.4 × eye height and 0.85 × basal width of mandible. Temple entirely smooth. Face mainly dense, rather fine and straight transverse striate, smooth medially. Face width 1.3 × eye height and 1.3 × height of face and clypeus combined. Hypoclypeal depression medium size and round, its width 0.9 × distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, ventrally not joined with hypostomal carina upper base of mandible. Mesosoma. Length of mesosoma 1.9 × its height. Pronotal keel distinct, its posterior branch distinct, almost fused medially with posterior margin of pronotum; anterior branch of carina distinct and situated submedially. Mesoscutum highly and roundly elevated above pronotum, its median lobe convex anteriorly and without anterolateral corners. Mesoscutum densely and distinctly granulate, without rugae along notauli and laterally, with two distinct and strongly convergent posteriorly and almost straight carinae in medioposterior third; mostly glabrous, with sparse, long and semi-erect or erect yellow setae arranged in stripe along notauli and in single line laterally. Notauli rather deep, wide, coarsely and densely crenulate, weakly shallow posteriorly. Prescutellar depression rather deep, long, with four carinae, finely coriaceous, 0.35 × as long as scutellum. Scutellum entirely densely and finely granulate-coriaceous, weakly convex, with lateral carinae. Mesopleuron reticulate-granulate and partly with striation, smooth in small anterior area. Subalar depression shallow, widely and densely striate with granulation partly. Precoxal sulcus (sternaulus) deep in anterior half and shallow in posterior half, weakly curved, sparsely crenulate and partly with fine granulation, running along entire lower part of mesopleuron. Propodeum with very short lateral tubercles, with basolateral areas distinctly delineated by carinae and densely granulate with fine striation laterally, rest part of propodeum granulate and partly rugose-striate; areola rather wide and short; basal carina present in basal 0.3, 1.4 × longer than anterior fork of areola. Wings. Fore wing 3.4 × as long as wide. Vein r arising weakly behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 17: 3: 33: 16. Second submarginal cell weakly narrowed distally, its length 3.1 × maximum width, 1.5 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior 0.4 of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu weakly antefurcal, strongly curved towards base of wing, faintly infuscate. Legs. Hind coxa dorsally transversely striate with dense granulation, laterally densely granulate to finely granulate-reticulate, with distinct basoventral corner and fine tooth. Hind femur finely striate-reticulate in upper third and smooth on remainder part, 3.3 × as long as wide. Hind tibia apically with single spines on outer side. Dorsal surface of hind tibia with long, rather dense and almost erect setae, their length 1.0 – 1.5 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Metasoma. Petiole (lateral view) weakly curved ventrally, dorsally distinctly arched in basal half and almost straight in apical half, distinctly thickened in basal 0.5; widened in apical fourth (dorsal view), with very small spiracular tubercles in widened basal third. Length of petiole 2.2 × its apical width, 1.4 × length of propodeum. Second tergite without separate laterotergites. Median length of second and third tergites combined 1.2 × basal width of second tergite, 0.7 × as long as their maximum width. Second suture absent. Petiole distinctly and rather densely longitudinally striate, with additional dense and distinct reticulation. Remaining tergites entirely smooth. Ovipositor straight. Ovipositor sheath 1.7 × as long as petiole, 0.6 × as long as metasoma, 0.9 × as long as mesosoma, 0.4 × as long as fore wing. Colour. Head yellow. Mesosoma and metasoma light reddish brown, propodeum and petiole reddish brown. Antennae yellow to brownish yellow in basal half, brown to dark brown apical half, without pale subapical segments. Palpi whitish yellow. Legs yellow to brownish yellow, hind tibia whitish subbasally. Ovipositor sheath pale brown in basal half and brown in apical half. Fore wing faintly and almost evenly infuscate. Pterostigma brown, pale yellow in basal quarter. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF88FF99838EFF45FDC11F0E.taxon	materials_examined	Material examined. Holotype. female, China, Zhejiang Prov., Deqing Fatou, 27. V. 1995, He Junhua, No. 959008 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius proximoscus sp. nov. is very similar to S. moscus Nixon, but differs in having the head more transverse (less transverse in S. moscus), scutellum entirely and densely but not strongly granulate-coriaceous (mainly smooth in S. moscus), mesopleuron almost entirely striate-reticulate (smooth in S. moscus), ovipositor sheath distinctly shorter than metasoma (weakly longer in S. moscus), and vein 1 m-cu separated from vein 2 RS on 0.3 (on 0.7 in S. moscus). This new species also similar to S. leschii Belokobylskij, but differs from it in having the subapical segments of antenna dark (distinctly pale in S. leschii), mesopleuron entirely striate-rugulose (mainly smooth in S. leschii), first flagellar segment almost as long as second segment (distinctly longer in S. leschii), mesoscutum with long and almost erect setae (with shorter and semi-erect setae in S. leschii), body mainly yellow to light reddish brown (mainly reddish brown to black in S. leschii), and fore wing faintly infuscate (distinctly infuscate in S. leschii). S. proximoscus sp. nov. differs from S. nanpingensis Chao by completely smooth vertex (finely sculptured in S. nanpingensis), antennae 27 - segmented (21 - segmented in S. nanpingensis), setae on dorsal margin of hind tibia very long (short in S. nanpingensis), and mesopleuron mainly sculptured (mainly smooth in S. nanpingensis).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF88FF99838EFF45FDC11F0E.taxon	etymology	Etymology. From “ proxime ” (Latin for “ similar ”) and species name “ moscus ”, because new species is similar to S. moscus.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF88FF99838EFF45FDC11F0E.taxon	distribution	Distribution. China (Zhejiang).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF89FF94838EFC53FDA21A1B.taxon	description	(Fig. 32) Description. Female. Body length 5.0 mm; fore wing length 3.5 mm. Head. Antennae filiform, 42 - segmented. Scape 1.8 × as long as its maximum width. First flagellar segment 5.5 × as long as its apical width, 1.3 × as long as second segment. Penultimate segment 3.2 × as long as wide, 0.9 × as long as apical segment. Head width (dorsal view) 1.5 × its median length, 1.25 × as wide as mesoscutum. Head behind eyes (dorsal view) weakly roundly narrowed; transverse diameter of eye 1.6 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.15 × its sides; POL: Od: OOL = 3.5: 4.0: 1.1. Vertex coarsely and weakly curvedly striate, with distinct oblique striae laterally, with fine to very fine reticulation between striae. Frons entirely, distinctly, densely and weakly curvedly transversely striate. Eye almost glabrous, 1.2 × as high as broad. Malar space 0.45 × eye height and equal to basal width of mandible. Temple smooth in anterior half and distinctly subvertically striate in posterior half. Face densely and distinctly transversely striate, almost without additional reticulation, smooth in subround medial lower area. Face width 1.15 × eye height and almost as long as height of face and clypeus combined. Hypoclypeal depression medium-sized and round, its width 0.9 × distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, ventrally joined with hypostomal carina upper base of mandible by additional rugae. Mesosoma. Length of mesosoma 1.9 × its height. Pronotal keel distinct, its posterior branch coarse, fused with posterior margin of pronotum on rather wide distance; anterior branch of carina distinct and situated behind middle. Mesoscutum highly and roundly elevated above pronotum, its median lobe convex anteriorly and without anterolateral corners. Mesoscutum distinctly and densely granulate on narrow median areas of lobes, its median lobe anteriorly without granules situated in transverse lines, with long or very long and dense rugae situated along notauli and laterally, with two distinct and weakly convergent posteriorly carinae in medioposterior half; mostly glabrous, with rather dense, long and semi-erect pale setae arranged in wide stripe along notauli and in almost single line laterally. Notauli wide, coarsely and densely crenulate, deep anteriorly and more shallow posteriorly. Prescutellar depression shallow, long, with three carinae, additionally finely rugulose, 0.25 × as long as scutellum. Scutellum finely to very finely granulate in anterior half and densely distinctly granulate in posterior half, almost without rugae, weakly convex, with fine lateral carinae. Mesopleuron wide and densely curvedly striate with fine granulation partly, with two small smooth areas. Subalar depression rather deep, wide, coarsely rugose-striate. Precoxal sulcus rather wide, straight, oblique, deep, running in anterior 0.6, with additional very shallow, curved and densely rugulose depression in posterior 0.4. Propodeum with distinct and wide lateral tubercles, with basolateral areas distinctly delineated by carinae and densely finely granulate with rugosity along carinae, rest part of propodeum sparsely striate and with dense and finely reticulation; areola rather narrow and long; basal carina present in basal 0.15, 0.4 × as long as anterior fork of areola. Wings. Fore wing 3.4 × as long as wide. Vein r arising weakly behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 16: 6: 39: 18. Second submarginal cell not narrowed distally, its length 2.5 × maximum width, 1.2 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising almost from anterior third of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.7 × as long as vein 1 - M. Vein m-cu weakly postfurcal, oblique towards base of wing, faintly infuscate. Legs. Hind coxa dorsally densely undulately transversely striate with additional reticulation, finely and densely granulate-reticulate laterally, almost smooth below, with distinct basoventral corner and tooth. Hind femur elongate-oval, finely and densely striate dorsally, 3.4 × as long as wide. Hind tibia apically without spines on outer side. Dorsal surface of hind tibia with medium length, rather dense and semi-erect setae, their length 0.8 – 1.2 × maximum width of tibia. Hind tarsus almost as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 1.2 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly curved ventrally, dorsally distinctly arched in basal 0.5 and almost straight in apical 0.5, thickened in basal 0.3; distinctly widened in apical fifth (dorsal view), with weak spiracular tubercles in weakly widened basal third. Length of petiole 2.5 × its apical width, 1.4 × length of propodeum. Second tergite without laterotergites. Median length of second and third tergites combined 1.6 × basal width of second tergite, 0.75 × as long as their maximum width. Second suture absent. Petiole distinctly and rather sparsely longitudinally striate, with dense and distinct reticulation between striae. Remaining tergites entirely smooth. Ovipositor straight. Ovipositor sheath 2.8 × as long as petiole, almost as long as metasoma, 1.55 × as long as mesosoma, 0.8 × as long as fore wing. Colour. Head yellow, face brownish. Mesosoma and metasoma light reddish brown to yellowish brown, mesoscutum brownish yellow, petiole reddish brown. Antennae yellow to brownish brown in basal half, brown submedially, with 15 pale subapical segments and dark single apical segment. Palpi whitish. Legs yellow to brownish yellow, white to whitish yellow basally, hind femur in apical two-thirds brown, hind tibia almost entirely pale yellow, faintly darkened basally. Ovipositor sheath brown to pale brown, yellowish brown basally. Fore wing faintly and evenly infuscate. Pterostigma brown, pale yellow in basal quarter and apically. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF89FF94838EFC53FDA21A1B.taxon	materials_examined	Material examined. Holotype. female, China, Guangdong Prov., Yunan Tongledashan, 12 – 13. VIII. 2003, Xu Zaifu, No. 20054574 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius pseudaphareus sp. nov. is similar to S. aphareus Nixon, but differs in having the head and hind coxa yellow or whitish yellow (rust red or reddish brown in S. aphareus), head behind eyes in dorsal view weakly narrowed and temple rather long (head strongly narrowed and temple short in S. aphareus), malar space longer (shorter in S. aphareus), posterior branch of pronotal keel distinctly fused with posterior margin of pronotum (not fused in S. aphareus), basal carina of propodeum very short (long in S. aphareus), and petiole short (long in S. aphareus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF89FF94838EFC53FDA21A1B.taxon	etymology	Etymology. From “ pseudo ” (Greece for “ false ”) and the name of related species, because new species is similar to S. aphaereus Nixon.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF89FF94838EFC53FDA21A1B.taxon	distribution	Distribution. China (Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF84FF94838EF9ADFC331B53.taxon	materials_examined	Material examined. China (ZJUH). Jiangsu Prov.: 1 female, Wuxi, 18. VIII. 1987, Li Qiang, No. 200012142. Distribution. China (Jiangsu) (new record); Japan, Russia (Far East).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF85FF92838EFF45FDCE1D76.taxon	description	(Fig. 33) Description. Female. Body length 4.0 mm; fore wing length 3.3 mm. Head. Antennae slender, weakly setiform, 40 - segmented. Scape 1.4 × as long as its maximum width. First flagellar segment 6.3 × as long as its apical width, 1.3 × as long as second segment. Head not depressed, its width (dorsal view) 1.4 × median length. Head behind eyes rather strongly and roundly narrowed. Transverse diameter of eye 1.8 × length of temple. Ocelli arranged in triangle with base 1.2 × as long as its sides; POL: Od: OOL = 5: 5: 12. Vertex and frons densely and coarsely striate; Eye glabrous, 1.3 × as high as broad. Malar space 0.4 × eye height and 0.8 × basal width of mandible. Temple smooth. Face densely coarsely rugose-striate; its width 1.2 × eye height and 1.1 × height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, joined below with hypostomal carina upper base of mandible. Mesosoma. Length of mesosoma 1.8 × its height. Pronotal keel of posterior branch widely fused with posterior margin of pronotum. Mesoscutum highly and perpendicularly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; densely granulate and coarsely transversely rugose striate, rugosity absent only on narrow median areas of lateral lobes; mesoscutum medioposteriorly with two distinct and almost parallel striae and sparse rugosity near them. Notauli complete, wide, deep, coarsely crenulate. Prescutellar depression deep, with three carinae, 0.4 × as long as scutellum. Scutellum more or less densely granulate, weakly convex, with distinct lateral carinae. Mesopleuron medially smooth or with very fine reticulation. Subalar depression shallow, narrow, striate-rugose. Precoxal sulcus deep, more or less wide, sinuate, more or less distinctly shallow in posterior 0.3, coarsely and sparsely crenulate, running along almost entirely length of lower part of mesopleuron. Propodeum small and wide lateral tubercles, coriaceous or granulate; areas distinctly delineated by carinae; areola rather large, narrow, pentagonal, sparsely striate; petiolate area long and narrow, more or less widened submedially; basal carina as long as anterior fork of areola. Wings. Fore wing 3.7 × as long as wide. Vein r arising almost from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 21: 6: 40: 23. Second submarginal cell not narrowed distally, its length 3.2 × maximum width, 1.4 × length of first subdiscal cell. Vein cu-a postfurcal. Vein m-cu postfurcal. Vein CU 1 a not interstitial, arising from anterior 0.3 of distal vein of subdiscal cell. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu weakly antefurcal, oblique to base of wing. Legs. Hind coxa dorsally densely concentrically striate, smooth laterally, with basoventral tooth and corner. Hind femur smooth, 4.0 × as long as wide. Hind tibia apically with two spines on outer margin. Dorsal surface of hind tibia with almost erect, long and short setae, length of setae 0.5 – 1.3 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.6 × as long as basitarsus, 1.4 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly curved ventrally and distinct arched in basal half dorsally, with spiracular tubercles in basal third; striate, with rugulosity between striae in basal 0.7, smooth apically. Length of petiole 3.8 × its apical width (dorsal view), 2.0 × length of propodeum. Second tergite without separate laterotergites. Second suture absent. Median length of second and third tergites combined 2.4 × basal width of second tergite, as long as their maximum width. Remaining tergites smooth. Ovipositor sheath 4.9 × as long as petiole, 2.4 × as long as metasoma, 3.4 × as long as mesosoma, 1.6 × as long as fore wing. Colour. Body black; head reddish brown. Antenna reddish brown basally, darken toward apex. Palpi brown. Legs dark brown; middle and hind femora blackish brown. Ovipositor sheath brown. Fore wing faintly infuscate. Pterostigma dark brown. Variation. Body length 4.4 mm; fore wing length 3.4 mm. Scape 1.5 × as long as its maximum width. First flagellar segment 6.5 × as long as its apical width. Fore wing 3.8 × as long as maximum width. Hind femur 3.9 × as long as wide. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF85FF92838EFF45FDCE1D76.taxon	materials_examined	Material examined. Holotype: female, China, Yunnan Prov., Anning hotspring, 18 – 20. VII. 1988, Chen Xuexin, No. 881719 (ZJUH). Paratype: 1 female, China, Yunnan Prov., Yunlong Tianchi, 21. VIII. 2003, Jiang Qian, No. 20046309 (ZJUH). Comparative diagnosis. This new species belongs to the S. antennalis Szépligeti species group. Spathius pseudido sp. nov. is similar to S. dido Nixon, but differs in having the width of head 1.4 × median length (1.7 × in S. dido), mesoscutal sculpture weak (coarse in S. dido), propodeum with weak granulation (almost entirely smooth in S. dido), pronotum and mesopleuron black (reddish brown in S. dido).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF85FF92838EFF45FDCE1D76.taxon	etymology	Etymology. From “ pseudo ” (Greek for “ pseudo ”), and the specific name “ dido ”, because of its similarity to S. dido.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF85FF92838EFF45FDCE1D76.taxon	distribution	Distribution. China (Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF82FFAE838EFE46FDF31F93.taxon	description	(Fig. 34) Description. Female. Body length 4.4 mm; fore wing length 3.6 mm. Head. Antennae almost filiform, more than 19 - segmented (apical segments missing). Scape 1.8 × as long as its maximum width. First flagellar segment 5.5 × as long as its apical width, as long as second segment. Head width (dorsal view) 1.5 × median length. Head behind eyes (dorsal view) roundly narrowed; transverse diameter of eye 2.0 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.3 × its sides; POL: Od: OOL = 6: 5: 13. Vertex entirely and at least finely striate or aciculate. Frons smooth. Eye glabrous, 1.3 × as high as broad. Malar space 0.4 × eye height and equal to basal width of mandible. Temple smooth. Face with very dense, fine and absolutely even transverse aciculation (like surface of gramophone record or CD); its width 1.1 × eye height and equal to height of face and clypeus combined. Clypeal suture absent. Hypoclypeal depression rather small and round, its width 0.6 × distance from edge of depression to eye, 0.3 × width of face. Occipital carina complete dorsally, not joined with hypostomal carina being obliterated upper base of mandible. Mesosoma. Length of mesosoma 2.0 × its height. Pronotal keel coarse, its posterior branch fused medially with posterior margin of pronotum, anterior branch situated submedially on pronotum. Mesoscutum highly and roundly elevated above pronotum, its median lobe convex anteriorly and without anterolateral corners. Mesoscutum mostly glabrous, with rather sparse, more or less long and almost erect pale setae arranged narrow along notauli and laterally, entirely densely granulate; with two distinct and convergent posteriorly carinae medioposteriorly. Notauli anteriorly distinct, wide, and crenulate, very shallow in posterior half. Prescutellar depression deep, rather short, with six carinae, finely rugulose-reticulate, 0.3 × as long as scutellum. Scutellum entirely densely granulate, distinctly convex, with distinct lateral carinae. Mesopleuron densely granulate-coriaceous. Subalar depression very shallow, wide, and finely reticulate-granulate, with sparse striae. Precoxal sulcus distinct, deep, straight, crenulate, running along anterior half of lower part of mesopleuron. Propodeum with very short and wide lateral tubercles, with distinctly delineated by carinae and granulate basolateral areas, its rest part coarsely and transversely striate with rugosity; areola narrow and long; basal carina present in basal third, 2.0 × as long as anterior fork of areola. Wings. Fore wing 3.9 × as long as wide. Vein r arising behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 28: 5: 31: 24. Second submarginal cell not narrowed distally, its length 3.0 × maximum width, 1.6 × length of first subdiscal cell. Vein CU 1 a arising from anterior third of distal margin of first subdiscal cell. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, curved and strongly oblique towards base of wing. Legs. Hind coxa dorsally distinctly transversely striate, widely and densely granulate at rest part, with basoventral tooth and corner. Hind femur distinctly and densely granulate-coriaceous, 3.0 × as long as wide. Hind tibia apically without spines on outer side. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 0.9 – 1.0 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.8 × as long as second-fifth segments combined. Second segment of hind tarsus 0.45 × as long as basitarsus, 1.1 × as longer as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) almost straight ventrally, dorsally rather strongly arched in basal third and straight in apical two-thirds, thickened in basal third; widened in apical quarter (dorsal view), with spiracular tubercles in basal third; distinctly rugulose-striate with dense rugulosity between striae in basal 0.4 and finely striate apically. Length of petiole 2.3 × its apical width (dorsal view), 1.9 × length of propodeum. Second tergite with separate laterotergites. Median length of second and third tergites combined 1.6 × basal width of second tergite, 0.7 × as long as their maximum width. Second suture indistinct. Second and third tergites densely longitudinally striate with dense rugulosity between striae, almost smooth narrowly apically. Fourth tergite densely longitudinally striate in basal two-thirds with dense rugulosity between striae, smooth in apical third. Fifth tergite almost smooth, with finely sculpture medially. Remaining tergites smooth. Ovipositor straight. Ovipositor sheath 1.4 × as long as petiole, 0.7 × as long as metasoma, 0.8 × as long as mesosoma, 0.4 × as long as fore wing. Colour. Body light yellowish brown to reddish brown. Antennae yellowish brown. Palpi pale yellow. Legs pale yellowish brown, sometimes with brown tint. Ovipositor sheath brownish yellow in basal 0.6 and black in apical 0.4. Fore wing faintly infuscate, with brown spots near vein 1 - SR, 1 - M and pterostigma. Pterostima brown, pale in basal third. Variation. Body length 3.4 – 3.8 mm; fore wing length 2.9 – 3.2 mm. Mesosoma 1.9 × as long as its height. Basal carina of propodeum 1.8 × as long as anterior fork of areola. Fore wing 3.7 × as long as maximum width. Length of second submarginal cell 3.2 × its maximum width. Hind femur 2.9 × as long as wide. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF82FFAE838EFE46FDF31F93.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Wuzhishan Shuimanxiang, 16 – 18. V. 2007, Zeng Jie, No. 200807548 (ZJUH). Paratypes: 1 female, China, Hainan Prov., Wuzhishan, 29. X. 2007, Liu Jingxian, No. 200710309 (ZJUH); 1 female, Hainan Prov., Diaoluoshan, 1 – 2. VI. 2007, Liu Jingxian, No. 200703955 (ZJUH). Comparative diagnosis. This new species belongs to the S. vulnificus Wilkinson species group. Spathius pseudocritolaus sp. nov. is similar to S. critolaus Nixon, but differs in having the vertex entirely and at least finely striate or aciculate (almost smooth and sometimes maximum with traces of transverse aciculation in S. critolaus), pronotal keel coarse (very weak in S. critolaus), propodeum with areas distinctly delineated by carinae (without such areas in S. critolaus), setae on dorsal surface of hind tibia long (short, much shorter than its middle width in S. critolaus), and fourth tergite densely striate (smooth in S. critolaus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF82FFAE838EFE46FDF31F93.taxon	etymology	Etymology. From “ pseudo ” (Greek for “ false ”), and the specific name “ critolaus ”, because of its similar to S. critolaus.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF82FFAE838EFE46FDF31F93.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBEFFAE838EFC25FC4F18D7.taxon	materials_examined	Material examined. China (ZJUH). Guangdong Prov.: 1 female, Shixing Chebaling, 19. IV. 2002, Xu Zaifu, No. 20050566. Yunnan Prov.: 1 female, Xishuangbanna, 31. VII. 2003, Xu Zaifu, No. 20053438.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBEFFAE838EFC25FC4F18D7.taxon	distribution	Distribution. China (Guangdong, Yunnan) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBEFFAC838EFAE7FDF41D53.taxon	description	(Fig. 35) Description. Female. Body length 3.9 mm; fore wing length 2.9 mm. Head. Antennae weakly setiform, more than 27 - segmented (apical segments missing). Scape 1.8 × as long as its maximum width. First flagellar segment 5.5 × as long as its apical width, as long as second segment. Subapical segments about 4.5 × as long as their width. Head width (dorsal view) 1.5 × its median length, 1.2 × as wide as mesoscutum. Head behind eyes (dorsal view) roundly narrowed; transverse diameter of eye 1.9 × length of temple. Ocelli medium-sized, arranged in almost equilateral triangle; POL: Od: OOL = 3.0: 3.0: 7.0. Vertex entirely smooth; frons distinctly, densely and almost entirely transversely striate. Eye glabrous, 1.25 × as high as broad. Malar space 0.4 × eye height and 0.7 × basal width of mandible. Temple entirely smooth. Face with dense, distinct and weakly curved transverse striae with fine reticulation, smooth on wide vertical median stripe. Face width equal to eye height and 1.25 × height of face and clypeus combined. Hypoclypeal depression large and round, its width 0.8 × distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, ventrally not joined with hypostomal carina obliterating upper base of mandible. Mesosoma. Length of mesosoma 2.1 × its height. Pronotal keel distinct, its posterior branch distinctly fused with posterior margin of pronotum on short distance; anterior branch situated near middle of pronotum. Mesoscutum highly and roundly elevated above pronotum, its median lobe convex anteriorly and without anterolateral corners. Mesoscutum densely granulate, granules on median lobe situated in curved lines; with rather long and dense rugae situated along notauli and laterally, with two distinct and weakly convergent posteriorly almost straight carinae in medioposterior half; mostly glabrous, with rather sparse, long and semi-erect pale setae arranged rather wide along notauli and in single line laterally. Notauli anteriorly distinct, wide, crenulate, deep anteriorly and shallow posteriorly. Prescutellar depression rather shallow, not long, with three carinae, almost smooth, 0.3 × as long as scutellum. Scutellum entirely densely granulate, weakly convex, with distinct lateral carinae. Mesopleuron almost entirely granulate, partly with curved striation. Subalar depression rather shallow, wide, sparsely and curvedly striate-rugose. Precoxal sulcus distinct and coarsely crenulate-rugose, deep, wide, straight, oblique along anterior 0.6 and shallow, wide and weakly curved along posterior 0.4 of lower part of mesopleuron. Propodeum with short and rather wide lateral tubercles, with basolateral areas distinctly delineated by carinae and granulate with rugosity, rest part of propodeum mainly smooth to finely reticulate and with sparse striae; areola rather narrow and relatively short; basal carina present in basal quarter, about as long as anterior fork of areola. Wings. Fore wing 3.5 × as long as wide. Vein r arising almost from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 27: 5: 37: 20. Second submarginal cell not narrowed distally, its length 3.1 × maximum width, 1.5 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior third of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, distinctly oblique towards base of wing and almost straight, faintly infuscate. Legs. Hind coxa dorsally densely granulate with fine transverse striation, densely granulate to reticulate-coriaceous below at rest part, with distinct basoventral corner and tooth. Hind femur mainly smooth, very finely longitudinally striate dorsally, 3.4 × as long as wide. Hind tibia apically with two spines on outer side. Dorsal surface of hind tibia with not long, rather sparse and semi-erect setae, their length 0.8 – 1.2 × maximum width of tibia. Hind tarsus almost as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Second segment of hind tarsus 0.45 × as long as basitarsus, 1.3 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly curved ventrally, dorsally weakly and evenly arched, weakly thickened submedially; widened in apical fifth (dorsal view), with small spiracular tubercles in weakly widened basal third. Length of petiole 3.1 × its apical width, 1.8 × length of propodeum. Second tergite without separate laterotergites. Median length of second and third tergites combined 1.6 × basal width of second tergite, 0.8 × as long as their maximum width. Second suture absent. Petiole distinctly and rather densely longitudinally striate, with dense and fine reticulation between striae. Remaining tergites entirely smooth. Ovipositor straight. Ovipositor sheath 3.1 × as long as petiole, 1.2 × as long as metasoma, 1.9 × as long as mesosoma, 0.9 × as long as fore wing. Colour. Body dark reddish brown to reddish brown, with pale spots on head upper eyes and on mesoscutum along notauli. Antennae yellowish brown basally, brown to dark brown on rest part, without pale subapical segments. Palpi pale yellow. Legs pale yellow to whitish yellow, middle and hind femora in apical halves, hind tibia on short submedian sport and hind tarsus brown. Ovipositor sheath brownish yellow in basal half and brown to dark brown on apical half. Fore wing faintly infuscate, with brownish spots near 1 - M vein and below pterostigma. Pterostigma brown, pale in basal third and apically. Variation. Body length 3.0 – 3.9 mm; fore wing length 2.4 – 3.0 mm. First flagellar segment 5.0 – 5.5 × as long as its apical width. Transverse diameter of eye 1.7 – 1.8 × length of temple. Length of mesosoma 2.0 – 2.1 × its height. Basal carina of propodeum 0.6 × as long as anterior fork of areola. Hind femur 3.3 – 3.6 × as long as wide. Length of petiole 2.9 – 3.2 × its apical width, 1.80 – 1.85 × length of propodeum. Sometimes second tergite striate at short basolateral areas. Ovipositor sheath 2.5 – 3.0 × as long as petiole, 1.2 – 1.3 × as long as metasoma, 1.8 – 1.9 × as long as mesosoma, 0.8 – 0.9 × as long as fore wing. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBEFFAC838EFAE7FDF41D53.taxon	materials_examined	Material examined. Holotype. female, China, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802260 (ZJUH). Paratypes: 1 female, China, Hainan Prov., Jianfengling Tianchi, 22 – 23. X. 2007, Liu Jingxian, No. 200710731 (ZJUH); 1 female, Hainan Prov., Jianfengling, 5 – 7. VI. 2007, Weng Liqiong, No. 200806464 (ZISP); 1 female, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802279 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius quasiasander sp. nov. is similar to S. asander Nixon, but differs in having the frons with distinct striation (very finely striate in S. asander), mesoscutum almost perpendicularly elevated above pronotum (very gradually in S. asander), scutellum densely and distinctly granulate (with trace of sculpture in S. asander), mesopleuron distinctly and entirely sculptured (mainly smooth in S. asander), and setae on dorsal surface of hind tibia shorter (long in S. asander). This new species also similar to Chinese S. beatus Chao, but differs from it in having the malar space dark (pale in S. beatus), apical segments of antenna without pale segments (with pale segments in S. beatus), posterior branch of pronotal carina fused with posterior margin (separated in S. beatus), scutellum coarsely granulate (finely coriaceous to smooth in S. beatus), and ovipositor long (short in S. beatus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBEFFAC838EFAE7FDF41D53.taxon	etymology	Etymology. From “ quasi ” (Latin for “ near ”) and the name of the most related species.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBEFFAC838EFAE7FDF41D53.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBCFFAA838EFE78FD421803.taxon	description	(Fig. 36) Description. Female. Body length 6.0 mm; fore wing length 4.5 mm. Head. Antennae slender, weakly setiform, more than 47 - segmented (apical segments missing). Scape 1.5 × as long as its maximum width. First flagellar segment 3.8 × as long as its apical width, 1.2 × as long as second segment. Head not depressed, its width (dorsal view) 1.4 × median length. Head behind eyes weakly convex in anterior half, roundly narrowed in posterior half. Transverse diameter of eye 2.1 × length of temple. Ocelli arranged in triangle with base 1.3 × as long as its sides; POL: Od: OOL = 7: 7: 23. Vertex very coarsely rugose-striate. Frons densely coarsely rugose-striate. Eye glabrous, 1.3 × as high as broad. Malar space 0.4 × eye height and 0.8 × basal width of mandible. Temple coarsely vertically striate, widely medially smooth or almost smooth. Face densely coarsely rugose-striate; its width 1.1 × eye height and 1.2 × height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.0 × its height. Pronotal keel distinct, its posterior branch widely fused with posterior margin of pronotum. Mesoscutum highly and perpendicularly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; densely irregularly rugose-striate and granulate, with two convergent posteriorly carinae and narrow rugosity between them in medioposterior half; median and lateral lobes of mesoscutum with wide areas covered only granulate sculpture. Notauli complete, wide, shallow, densely crenulate. Prescutellar depression shallow, with five carinae, 0.3 × as long as scutellum. Scutellum densely granulate, flat, with distinct lateral carinae. Mesopleuron distinctly rugose and granulate; coriaceous-granulate below precoxal sulcus. Subalar depression shallow, wide, transversely rugose-striate and granulate. Precoxal sulcus deep, narrow, straight, finely coriaceous-granulate, running along anterior 0.6 of lower part of mesopleuron. Propodeum without distinct tubercles, with areas distinctly delineated by carinae; basal carina as long as anterior fork of areola; basolateral areas entirely coriaceous-granulate, rugose in apical half; rest of propodeum sparsely rugose; areola large and wide, pentagonal; petiolate area short, widened distally, distinctly separated from areola. Wings. Fore wing 3.5 × as long as wide. Vein r arising weakly before middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 20: 7: 40: 18. Second submarginal cell not narrowed distally, its length 3.0 × maximum width, 1.2 × length of first subdiscal cell. Vein cu-a postfurcal. Vein m-cu postfurcal. Vein CU 1 a not interstitial, arising from anterior 0.3 of distal vein of subdiscal cell. Hind wing vein M + CU 0.7 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, weakly curved towards base of wing. Legs. Hind coxa densely granulate and transversely striate dorsally, granulate laterally, with basoventral tooth and corner. Hind femur dorsally finely striate and coriaceous-granulate, only coriaceous-granulate laterally and ventrally, 2.8 × as long as wide. Hind tibia apically with six spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 1.5 – 1.8 × maximum width of tibia. Hind tarsus 0.8 × as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Second segment of hind tarsus 0.4 × as long as basitarsus, as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) straight ventrally, distinct arched in basal half dorsally, with spiracular tubercles in basal third; entirely finely rugose, sparsely striate laterally. Length of petiole 3.1 × its apical width (dorsal view), 1.7 × length of propodeum. Second-fifth tergites with separate laterotergites. Second suture absent. Median length of second and third tergites combined 1.6 × basal width of second tergite, 0.8 × their maximum width. Second-fourth tergites densely regularly striate, with short rugulosity between striae, smooth apically. fifth tergite densely regularly striate, with short rugulosity between striae basally, finely semi-circularly transversely aciculate apically. Ovipositor sheath 3.3 × as long as petiole, 1.3 × as long as metasoma, 2.0 × as long as mesosoma, 1.1 × as long as fore wing. Colour. Body yellowish brown; mesosoma reddish brown, media lobe of mesoscutum yellowish brown. Metasoma reddish brown. Antenna brown basally, darken toward apex. Palpi pale yellow. Legs yellowish brown; fore and middle coxae and all trochanters pale yellow; hind coxa and femur reddish brown; hind tibia and tarsus pale reddish brown. Ovipositor sheath brown, almost black apically. Fore wing faintly infuscate, with distinct maculation. Pterostigma dark brown, pale in basal 0.3. Variation. Body length 5.2 – 6.4 mm; fore wing length 4.1 – 4.6 mm. Transverse diameter of eye 2.0 × length of temple. Length of second submarginal cell 3.2 × its maximum width. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Length of petiole 3.2 × its apical width (dorsal view), 1.8 × length of propodeum. Median length of second and third tergites combined 1.4 × basal width of second tergite, 0.7 × as long as their maximum width. Ovipositor sheath 3.1 × as long as petiole, 1.2 × as long as metasoma, 1.8 × as long as mesosoma, as long as fore wing. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBCFFAA838EFE78FD421803.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Jianfengling, 6. VI. 2007, Liu Jingxian, No. 200703783 (ZJUH). Paratypes: 1 female, China, Hainan Prov., Jianfengling, 5 – 7. VI. 2007, Xiao Bin, No. 200806908 (ZJUH); 1 female, Hainan Prov., Baisha Yinggeling, 18. XI. 2006, Tan Jiangli, No. 200805253 (ZJUH); 1 female, Hainan Prov., Jianfengling, 6. VI. 2007, Liu Jingxian, No. 200703845 (ZISP); 1 female, Guangdong Prov., Shixing Chebaling, 22 – 28. VII. 2008, Hong Chundan, No. 200807857 (ZJUH). Comparative diagnosis. This new species belongs to the S. striolatus Cameron species group. Spathius rectangulus sp. nov. is similar to S. hephaestus Nixon, but differs in having the scutellum densely granulate (finely or partly very finely granulate-coriaceous in S. hephaestus), petiole distinctly widen apically and entirely coarsely striate (weakly widened apically and finely sculptured in S. hephaestus), second and third tergites short with their combined length 1.6 × basal width of second tergite and 0.8 × their maximum width (long with their combined length 2.0 × basal width and 0.9 × maximum width in S. hephaestus), ovipositor distinctly shorter than body (almost as long as body in S. hephaestus), and fifth tergite strongly convex (weakly convex in S. hephaestus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBCFFAA838EFE78FD421803.taxon	etymology	Etymology. From “ rectum ” (Latin for “ straight ”) and “ angulus ” (Latin for “ angular ”), because its mesoscutum highly and perpendicularly elevated above pronotum.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBCFFAA838EFE78FD421803.taxon	distribution	Distribution. China (Hainan, Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBAFFAA838EFB55FA7F1B76.taxon	materials_examined	Material examined. China (ZJUH). Zhejiang Prov.: 1 female, Songyang, VII. 1992, Chen Hanlin, No. 924450; 1 female, Jiulongshan, VIII. 1978, Zhou Wenbao, No. 930120; 1 female, Gutianshan, 20. VII. 1986, Xu Weiliang, No. 862805. Fujian Prov.: 1 female, Wuyishan Dazhulan, 15. VII. 1994, Chen Xuexin, No. 942007; 1 female, Jiangle Bailian, 10. VII. 1997, Liu Changming, No. 20006278; 1 female, Longqishan, 10. X. 1991, Chen Xuexin, No. 920227; 1 female, Shanghang Buyun, 22 – 24. VII. 1988, Ma Yun, No. 885219. Guangdong Prov.: 1 female, Fengkai Qiancengfeng, 14. VIII. 2003, Xu Zaifu, No. 20054837; 1 female, Meizhou Fengxi, 29. VII. 2003, Chen Jujian, No. 20048705; 1 female, Ruyuan Nanling, 8. V. 2004, Xu Zaifu, No. 20049501; 1 female, Shixing Chebaling, 19. IV. 2002, Xu Zaifu, No. 20050818; 1 female, Shixing Chebaling, 15. VII. 2003, Xu Zaifu, No. 20058758 (ZISP); 1 female, Yunan Tongledashan, 12 – 13. VIII. 2003, Xu Zaifu, No. 20054743. Hainan Prov.: 1 female, Jianfengling, 4 – 7. VI. 2007, Zeng Jie, No. 200711238; 2 females, Diaoluoshan, 29. V – 2. VI. 2007, Xiao Bin, Nos. 200804570 and 200804571; 1 female, Yinggeling, 9 – 10. VI. 2007, Zeng Jie, No. 200804539; 1 female, Yinggeling, 18. X. 2007, Liu Jingxian, No. 200709783; 1 female, Diaoluoshan, 28. V – 1. VI. 2007, Zeng Jie, No. 200806722. Guizhou Prov.: 1 female, Kuankuoshui Baishao, 7. VI. 2010, Tang Pu, No. 201004353; 1 female, Kuankuoshui Qinggangtang, 9. VI. 2010, Zeng Jie, No. 201003746.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBAFFAA838EFB55FA7F1B76.taxon	distribution	Distribution. China (Jilin, Zhejiang, Fujian, Guangdong, Hainan, Guizhou); Japan, Korea, Russia (Far East).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBAFFAB838EF846FF001E34.taxon	materials_examined	Material examined. China (ZJUH). Heilongjiang Prov.: 22 females, 4 males, Yichun, 1985, Jin Liyuan, Nos 864360 (11), 864757 (7), 864758 (2), 864361, 864362 (4) and 850203; 1 female, Yichun, VII. 1985, Jin Liyuan, No. 851843; 1 female, Daicen, 24. VII. 1977, He Junhua, No. 770452. Zhejiang Prov.: 1 female, Gutianshan, 22. VII. 1986, Lou Xiaoming, No. 863395.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBAFFAB838EF846FF001E34.taxon	distribution	Distribution. China (Heilongjiang, Zhejiang) (new record); Japan, Korea, Russia, Kazakhstan, Armenia, Europe.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBBFFA9838EFD84FDF41A3E.taxon	description	(Fig. 37) Description. Female. Body length 4.5 mm; fore wing length 3.1 mm. Head. Antennae filiform, 32 - segmented. Scape 1.7 × as long as its maximum width. First flagellar segment 5.4 × as long as its apical width, 1.35 × as long as second segment. Penultimate segment 4.0 × as long as wide, as long as pointed apical segment. Head width (dorsal view) 1.5 × its median length, 1.2 × as wide as mesoscutum. Head behind eyes (dorsal view) evenly roundly narrowed; transverse diameter of eye 1.6 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 4.0: 4.0: 11.0. Vertex coarsely reticulate-rugulose, with undulate subtransverse striation in posterior half; frons entirely, distinctly and curvedly transversely striate. Eye with short and sparse setae, 1.2 × as high as broad. Malar space 0.35 × eye height and 0.7 × basal width of mandible. Temple mainly smooth, finely vertically striate in posterior third. Face mainly with rather dense, coarse and almost linearly transversely striation. Face width 1.1 × eye height and 1.2 × as long as height of face and clypeus combined. Hypoclypeal depression weakly enlarged and oval, its width 1.4 × distance from edge of depression to eye, 0.55 × width of face. Occipital carina complete dorsally, ventrally joined with hypostomal carina upper base of mandible. Mesosoma. Length of mesosoma 1.9 × its height. Pronotal keel distinct, its posterior branch coarse, very weakly separated from posterior margin of pronotum; anterior branch of carina distinct and situated submedially. Mesoscutum highly and roundly elevated above pronotum, its median lobe weakly convex anteriorly and without anterolateral corners. Mesoscutum finely and densely granulate, its median lobe anteriorly with additional fine transverse striation, with long and rather dense rugae situated along notauli and laterally, with two distinct and convergent posteriorly undulate carinae in medioposterior half; mostly glabrous, with sparse, rather long and semi-erect yellow setae arranged in wide stripe along notauli and in single line laterally. Notauli distinct, very wide and coarsely sparsely crenulate, evenly deep. Prescutellar depression deep, long, with three carinae, finely rugulose, 0.4 × as long as scutellum. Scutellum entirely densely and distinctly granulate, without rugae, almost flat, with distinct lateral carinae. Mesopleuron smooth medially, with fine striation or rugulosity around. Subalar depression shallow, wide, sparsely and coarsely striate with rugosity. Precoxal sulcus deep and oblique in anterior 0.6 and very shallow and curved in posterior 0.4, sparsely and coarsely crenulate, running along entire lower part of mesopleuron. Propodeum with distinct and wide lateral tubercles, with basolateral areas distinctly delineated by carinae and densely rugose-reticulate, rest part of propodeum rugose-striate, without granulation; areola rather narrow and long; basal carina present in basal 0.15, 0.4 × as long as anterior fork of areola. Wings. Fore wing 3.8 × as long as wide. Vein r arising weakly behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 20: 6: 40: 23. Second submarginal cell not narrowed distally, its length 2.7 × maximum width, 1.4 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising from anterior quarter of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.7 × as long as vein 1 - M. Vein m-cu interstitial, weakly curved towards base of wing, infuscate. Legs. Hind coxa dorsally transversely striate with granulation, granulate to coriaceous below at rest part, with distinct basoventral corner and tooth. Hind femur finely striate with reticulation in upper half and smooth in lower half, 3.6 × as long as wide. Hind tibia apically with two spines on outer side. Dorsal surface of hind tibia with long, dense and semi-erect setae, their length 0.8 – 1.4 × maximum width of tibia. Hind tarsus 0.85 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 1.1 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) curved ventrally, dorsally distinctly arched in basal 0.6 and almost straight in apical 0.4, thickened in basal 0.3; widened in apical fourth (dorsal view), with small spiracular tubercles in widened basal third. Length of petiole 2.8 × its apical width, 1.7 × length of propodeum. Second tergite without separate laterotergites. Median length of second and third tergites combined 1.6 × basal width of second tergite, 0.75 × as long as their maximum width. Second suture absent. Petiole distinctly and rather densely longitudinally striate, with additional dense reticulation between striae, smooth apically. Remaining tergites entirely smooth. Ovipositor straight. Ovipositor sheath 2.8 × as long as petiole, 1.2 × as long as metasoma, 1.8 × as long as mesosoma, 0.9 × as long as fore wing. Colour. Head yellow, but brownish yellow dorsally and anteriorly, dark brown posteriorly. Mesosoma reddish brown, dark ventrally, median lobe of mesoscutum yellowish brown. Metasoma dark reddish brown, tergites posteriorly yellowish. Antennae yellow in basal half, submedially and apically brown, seven subapical segments whitish yellow. Palpi whitish yellow. Legs whitish yellow, basally white, middle and hind femora with dark lateral spots in posterior half. Ovipositor sheath brown in apical 0.7 and pale brown in basal 0.3. Fore wing faintly infuscate, with more dark narrow brownish spots near veins. Pterostigma brown, pale yellow in basal quarter and apically. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBBFFA9838EFD84FDF41A3E.taxon	materials_examined	Material examined. Holotype. female, China, Hainan Prov., Wuzhishan, 16 – 20. V. 2007, Liu Jingxian, No. 200703380 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius rugosivertex sp. nov. is similar to S. leptoceras Cameron, but differs in having the head behind eyes (dorsal view) less strongly narrowed (distinctly narrowed in S. leptoceras), vertex rugose (transversely striate in S. leptoceras), posterior branch of pronotal carina weakly separated from anterior margin of mesoscutum (fused with it in S. leptoceras), scutellum densely and finely granulate (mainly smooth with punctation in S. leptoceras), basal carina of propodeum very short (long in S. leptoceras), petiole 1.8 × as long as propodeum (1.5 × in S. leptoceras), and body short (long in S. leptoceras). This new species also similar to S. ares Nixon, but differs from it in having the subapical segments of antenna pale (dark in S. ares), head transverse (subquadrate in S. ares), ocelli arranged in triangle with base larger than sides (in almost equilateral triangle in S. ares), posterior branch of pronotal carina weakly separated from anterior margin of mesoscutum (fused in S. ares), propodeum with distinct lateral tubercles (tubercles indistinct in S. ares), setae on dorsal surface of hind tibia long (short in S. ares), and petiole 1.8 × as long as propodeum (1.5 × in S. ares). Finally, S. rugosivertex sp. nov. is similar to S. gnom Belokobylskij et Maeto, but differs from it in having the subapical segments of antenna pale (dark in S. gnom), scutellum only granulate-coriaceous (densely rugose and without granulation in S. gnom), posterior branch of pronotal carina weakly separated from anterior margin of mesoscutum (strongly separated in S. gnom), and vertex rugose (transversely striate in S. gnom).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBBFFA9838EFD84FDF41A3E.taxon	etymology	Etymology. From “ rugosus ” (Latin for “ rugosus ”) and “ vertex ” (Latin for “ vertex ”), because of vertex coarsely and irregularly rugose.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFBBFFA9838EFD84FDF41A3E.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB9FFA9838EF98EFDFC1B70.taxon	description	(Fig. 38)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB9FFA9838EF98EFDFC1B70.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Fujian Prov., Dehua, 3 – 9. XI. 1974, Chen Jiahua. Distribution. China (Fujian).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB7FFA7838EFF45FC601F10.taxon	materials_examined	Material examined. China (ZJUH). Heilongjiang Prov.: 1 female, Daicen, 24. VII. 1977, He Junhua, No. 770452; 8 females, 8 males, Yichun, 1985, Jin Liyuan, Nos. 850135 (16); 1 female, 5 males, Yichun, VII. 1985, Jin Liyuan, Nos. 851838 (6). Tianjin Prov.: 2 females, 1 male, Guangang, V. 2008, Wang Xiaoyi, Nos. 200908487, 200908488 and 200908489. Zhejiang Prov.: 1 female, Longquan Fengyangshan, 10. X. 2003, Liu Jingxian, No. 20048285. Fujian Prov.: 2 females, 3 males, Lianjiang Guantou, 28. VI. 1965, Nos. 20004004 – 20004008. Hunan Prov.: 1 female, Changsha, 29. IX. 1979, Tong Xinwang, No. 20044824; 1 female, Changsha, 6. X. 1979, Tong Xinwang, No. 20044825.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB7FFA7838EFF45FC601F10.taxon	distribution	Distribution. China (Heilongjiang, Jilin, Tianjin, Zhejiang, Fujian, Shanghai, Hunan); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB7FFA7838EFF45FC601F10.taxon	discussion	Remarks. Unfortunately, the type material of S. sinicus Chao and S. bellus Chao was not found in the collection of IZCAS. According to present descriptions of these species as well as after study of additional material, S. sinicus, S. bellus and S. agrili have the same characters, i. e. palpi mostly dark, hind tibia with short pale subbasal area, mesoscutum roundly elevated above pronotum and vertex mostly smooth. These data were reasons for the synonymisation as discussed species names under S. sinicus.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB7FFA5838EFCA8FD5919D3.taxon	description	(Fig. 39) Description. Female. Body length 7.0 mm; fore wing length 4.9 mm. Head. Antennae slender, setiform, 55 - segmented. Scape 1.5 × as long as its maximum width. First flagellar segment 3.8 × as long as its apical width, 1.2 × as long as second segment. Head not depressed, its width (dorsal view) 1.4 × median length. Head behind eyes convex-roundly narrowed. Transverse diameter of eye 2.1 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.3 × its sides; POL: Od: OOL = 6: 5: 15. Vertex smooth. Frons densely and transversely striate. Eye glabrous, 1.4 × as high as broad. Malar space 0.5 × eye height and equal to basal width of mandible. Temple smooth. Face entirely coarsely rugose-reticulate, with undulately curved striae below, clypeus rugose-striate; its width 1.1 × eye height and 1.2 × height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 1.8 × its height. Pronotal keel finely developed, its posterior branch widely fused with posterior margin of pronotum; anterior branch distinct and situated submedially. Pronotal lateral depression wide, shallow, not delineated by carinae, densely and coarsely crenulate-striate. Mesoscutum highly and almost perpendicularly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; its median lobe entirely, densely and undulately transversely striate, with fine granulation partly, lateral lobes widely finely and densely granulate, with distinct transverse striation, with two subparallel longitudinal rugae and transverse striation in medioposterior 0.6. Notauli deep anteriorly and more or less shallow posteriorly, complete, coarsely crenulate. Prescutellar depression rather deep, with three strong carina, 0.4 × as long as scutellum. Scutellum smooth, weakly convex, with lateral carinae. Mesopleuron mostly smooth. Subalar depression shallow, wide, densely and coarsely rugose-striate. Precoxal sulcus deep, narrow, almost straight, crenulate, running along anterior 0.7 of lower part of mesopleuron. Propodeum without tubercles, but with areas distinctly delineated by carinae, basolateral areas granulate in rather narrow basomedian part and widely coarsely rugose along carina; areola transversely striate; petiolate area rather long and more or less distinctly separated from areola; basal carina 0.6 × as long as fork of areola. Wings. Fore wing 4.6 × as long as wide. Vein r arising from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 20: 6: 40: 19. Second submarginal cell not narrowed distally, its length 3.1 × maximum width, equal to length of first subdiscal cell. Vein cu-a distinctly postfurcal. Vein m-cu postfurcal. Hind wing vein M + CU 0.9 × as long as vein 1 - M. Vein m-cu interstitial, strongly oblique towards base of wing. Legs. Hind coxa dorsally transversely and semi-circularly striate, finely granulate laterally and ventrally, with basoventral tooth and corner. Hind femur almost smooth, 2.9 × as long as wide. Hind tibia apically with eight spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 1.2 – 1.5 × maximum width of tibia. Hind tarsus 0.8 × as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Second segment of hind tarsus 0.7 × as long as basitarsus, 1.1 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly concave ventrally, dorsally slightly and regularly arched in basal third, with spiracular tubercles in basal fifth; coarsely rugose-striate. Length of petiole 3.3 × its apical width (dorsal view), 2.3 × length of propodeum. Second and third tergites with separated laterotergites. Second suture absent. Median length of second and third tergites combined 2.0 × basal width of second tergite, 0.8 × their maximum width. Second and third tergites densely striate, with dense rugulosity between striae, smooth in apical 0.2; fourth tergite finely punctate, with very fine striation medially; fifth-seventh tergites finely punctate, smooth apically. Ovipositor sheath 3.8 × as long as petiole, 1.8 × as long as metasoma, 2.8 × as long as mesosoma, 1.4 × as long as fore wing. Colour. Head black. Mesosoma and petiole reddish brown. Second and third tergites reddish brown in basal fifth, remaining metasoma black. Antenna brown, faintly darkened toward apex, without pale subapical segments. Palpi pale yellow. Legs reddish brown, all femora and tibiae basally darken. Ovipositor sheath light brown, almost black apically. Fore wing faintly infuscate, with a dark spot below pterostigma. Pterostigma dark brown, yellow in basal 0.3. Variation. Body length 5.2 – 5.6 mm. Fore wing length 3.8 – 4.0 mm. Width of head (dorsal view) 1.5 × median length. Eye 1.3 × as high as broad. Malar space 0.4 × eye height and 0.9 × basal width of mandible. Face width equal to eye height and 1.1 × height of face and clypeus combined. Length of mesosoma 1.9 × its height. Precoxal sulcus almost smooth, finely crenulate. Length of second submarginal cell 3.0 – 3.2 × its maximum width. Hind wing vein M + CU 0.8 × as long as vein 1 - M. Hind femur 3.0 × as long as wide. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB7FFA5838EFCA8FD5919D3.taxon	materials_examined	Material examined. Holotype: female, China, Guangdong Prov., Longmen Nankunshan, 7. VIII. 2004, Xu Zaifu, No. 20053241 (ZJUH). Paratypes: 1 female, China, Guangdong Prov., Shixing Chebaling, 22 – 28. VII. 2008, Hong Chundan, No. 200807853 (ZJUH); 1 female, Sichuan Prov., Yaan Zhangjianshan, 14. VII. 2006, Zhang Hongying, No. 200613575 (ZJUH). Comparative diagnosis. This new species belongs to the S. philoctetes Nixon species group. Spathius spinosus sp. nov. is similar to S. ishigakus Belokobylskij, but differs in having the second and third tergites densely striate and with dense rugulosity between striae (entirely very small and dense reticulate-areolate and more finely sculptured posteriorly in S. ishigakus), mesopleuron mostly smooth (widely striate-rugose in S. ishigakus), hind coxa with basoventral tooth (without tooth in S. ishigakus), head, femora and tibiae dark brown (yellowish brown or light reddish brown in S. ishigakus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB7FFA5838EFCA8FD5919D3.taxon	etymology	Etymology. From “ spinosus ” (Latin for “ spinous ”), because of its hind tibia apically with numerous spines on outer margin.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB7FFA5838EFCA8FD5919D3.taxon	distribution	Distribution. China (Guangdong, Sichuan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB5FFA3838EF9F8FDF31BA3.taxon	description	(Fig. 40) Description. Female. Body length 5.6 mm; fore wing length 4.1 mm. Head. Antennae slender, weakly setiform, 52 - segmented. Scape 1.4 × as long as its maximum width. First flagellar segment 4.0 × as long as its apical width, 1.3 × as long as second segment. Head not depressed, its width (dorsal view) 1.4 × median length. Head behind eyes weakly convex in anterior half, roundly narrowed in posterior half. Transverse diameter of eye 1.6 × length of temple. Ocelli arranged in triangle with base 1.3 × its sides; POL: Od: OOL = 8: 7: 21. Vertex mostly smooth, rugose-striate anteriorly. Frons densely and coarsely rugose-striate. Eye glabrous, 1.4 × as high as broad. Malar space 0.4 × eye height and 0.9 × basal width of mandible. Temple smooth. Face densely coarsely rugose-striate; its width 1.2 × eye height and 1.2 × height of face and clypeus combined. Clypeal suture entirely distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 1.9 × its height. Pronotal keel distinct, its posterior branch widely fused with posterior margin of pronotum. Mesoscutum highly and perpendicularly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; densely and irregularly rugose-striate with granulation, with two convergent posteriorly carinae and narrow rugosity between them in medioposterior half. Notauli complete, wide, deep anteriorly, shallow posteriorly, sparsely crenulate. Prescutellar depression shallow, with four carinae, 0.3 × as long as scutellum. Scutellum almost smooth, flat, with distinct lateral carinae. Mesopleuron finely striate-rugose, coriaceous-granulate below precoxal sulcus. Subalar depression shallow, wide, transversely rugose-striate. Precoxal sulcus deep, narrow, straight, finely crenulate, running along anterior 0.6 of lower part of mesopleuron. Propodeum without distinct tubercles, with areas distinctly delineated by carinae; basal carina 1.5 × as long as anterior fork of areola; basolateral areas granulate basally, rugose in apical half; rest of propodeum sparsely rugose; areola large and wide, pentagonal; petiolate area short, widened distally, indistinctly separated from areola. Wings. Fore wing 3.5 × as long as wide. Vein r arising weakly before middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 20: 6: 33: 18. Second submarginal cell not narrowed distally, its length 2.9 × maximum width, equal to length of first subdiscal cell. Vein cu-a postfurcal. Vein m-cu postfurcal. Vein CU 1 a not interstitial, arising from anterior 0.3 of distal vein of subdiscal cell. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu interstitial, weakly curved towards base of wing. Legs. Hind coxa densely granulate and transversely striate dorsally, granulate laterally, with basoventral tooth and corner. Hind femur finely striate and coriaceous-granulate dorsally, almost smooth laterally and ventrally, 2.6 × as long as wide. Hind tibia apically with five spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 1.0 – 1.2 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.7 × as long as second-fifth segments combined. Second segment of hind tarsus 0.4 × as long as basitarsus, as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly curved ventrally and distinct arched in basal half dorsally, with spiracular tubercles in basal third; entirely striate, with rugulosity between striae. Length of petiole 3.9 × its apical width (dorsal view), 2.0 × length of propodeum. Second-fifth tergites with separate laterotergites. Second suture absent. Median length of second and third tergites combined 1.6 × basal width of second tergite, 0.9 × as long as their maximum width. Second and third tergites densely regularly striate, with short rugulosity between striae, smooth apically. fourth tergite coriaceous-rugose, smooth in apical fifth; fifth tergite finely punctate in basal two-thirds, smooth in apical third. Ovipositor sheath 3.9 × as long as petiole, 1.4 × as long as metasoma, 2.1 × as long as mesosoma, 0.7 × as long as fore wing. Colour. Body reddish brown; vertex and frons yellowish brown. Antenna pale reddish brown basally, darken towards apex. Palpi pale yellow. Legs reddish brown; all coxae posteriorly and femora basally faintly dark brown. Ovipositor sheath brown, almost black apically. Fore wing faintly infuscate, with indistinct maculation. Pterostigma dark brown, pale in basal 0.3. Male. Body length 4.8 mm; fore wing length 3.8 mm. Scape 1.5 × as long as its maximum width. First flagellar segment 4.8 × as long as its apical width. Length of mesosoma 1.8 × its height. Length of petiole 4.0 × its apical width (dorsal view). Body entirely pale. Otherwise similar to female.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB5FFA3838EF9F8FDF31BA3.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Wuzhishan, 15 – 16. V. 2008, Liu Jingxian, No. 200905336 (ZJUH). Paratype: 1 male, China, Hainan Prov., Diaoluoshan, 28 – 31. V. 2007, Liu Jingxian, No. 200702855 (ZJUH). Comparative diagnosis. This new species belongs to the S. striolatus Cameron species group. Spathius striolatiformis sp. nov. is similar to S. striolatus Cameron, but differs in having the vertex anteriorly sculptured (complete smooth except for a few punctures in S. striolatus), mesopleuron entirely sculptured (with smooth area in S. striolatus), precoxal sulcus finely crenulate (smooth in S. striolatus), CU 1 a arising from anterior 0.3 of distal part of subdiscal cell (from middle in S. striolatus), and ovipositor as long as body (distinctly shorter in S. striolatus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB5FFA3838EF9F8FDF31BA3.taxon	etymology	Etymology. From “ striolatus ”, the name of the most similar species, and “ formis ” (Latin for “ form ”) because they are morphologically similar species.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB5FFA3838EF9F8FDF31BA3.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB0FFA1838EFF45FD5018DE.taxon	description	(Fig. 41) Description. Female. Body length 6.0 mm; fore wing length 4.5 mm. Head. Antennae slender, weakly setiform, 50 - segmented. Scape 1.6 × as long as its maximum width. First flagellar segment 3.8 × as long as its apical width, 1.2 × as long as second segment. Head not depressed, its width (dorsal view) 1.4 × median length. Head behind eyes roundly narrowed. Transverse diameter of eye 1.7 × length of temple. Ocelli arranged in triangle with base 1.3 × its sides; POL: Od: OOL = 6: 5: 12. Vertex smooth. Frons densely and transversely striate. Eye glabrous, 1.3 × as high as broad. Malar space 0.5 × eye height and 0.8 × basal width of mandible. Temple smooth. Face entirely coarsely rugose-reticulate, clypeus rugose-striate; its width equal to eye height and equal to height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.2 × its height. Pronotal keel distinct, its posterior branch fused with posterior margin of pronotum at short distance; anterior branch fine and situated submedially. Pronotal lateral depression wide, partly delineated by carinae, densely and coarsely crenulate. Mesoscutum rather highly and roundly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; distinctly densely granulate, with short carinae along notauli and laterally, with wide and only granulate median areas on lateral and median lobes; with two convergent posteriorly carinae and narrow rugosity between them in medioposterior half. Notauli complete, wide, deep anteriorly and shallow posteriorly, coarsely crenulate. Prescutellar depression distinct, deep, with six carinae, 0.3 × as long as scutellum. Scutellum coriaceous-granulate, weakly convex, with distinct lateral carinae. Mesopleuron weakly sculptured or almost smooth medially. Subalar depression shallow, rather wide, coarsely rugose-striate. Precoxal sulcus shallow, weakly curved, finely crenulate, running along entire length of lower part of mesopleuron. Propodeum without tubercles, densely and coarsely rugose-reticulate, densely granulate basally or in basal half, with areas more or less distinctly delineated by carinae; basal carina 1.3 × as long as anterior fork of areola; areola large and pentagonal; petiolate area distinctly separated from areola. Wings. Fore wing 3.8 × as long as wide. Vein r arising before middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 14: 4: 23: 10. Second submarginal cell not narrowed distally, its length 3.0 × maximum width, 1.2 × length of first subdiscal cell. Vein cu-a slightly postfurcal. Vein m-cu postfurcal. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu weakly antefurcal, strongly oblique towards base of wing. Legs. Hind coxa transversely striate dorsally, densely granulate laterally and ventrally, with basoventral tooth and corner. Hind femur almost smooth, finely striate dorsally, 2.8 × as long as wide. Hind tibia apically with four spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 1.2 – 1.6 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.8 × as long as second-fifth segments combined. Second segment of hind tarsus 0.4 × as long as basitarsus, 1.1 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) straight ventrally and distinct arched dorsally, with small spiracular tubercles in basal third; distinctly and entirely striate, with dense rugulosity between striae. Length of petiole 2.1 × its apical width (dorsal view), 1.8 × length of propodeum. Second-fourth tergites with separated laterotergites. Second suture indistinct. Median length of second and third tergites combined 1.3 × basal width of second tergite, as long as their maximum width. Second and third tergites densely and anastomose striate, with very dense rugulosity between striae, reticulate-punctate subapically, apically narrowly smooth; fourth tergite very dense semi-circularly anastomose striate, with very dense rugulose microsculpture between striae. Fifth tergite finely and sparsely coriaceous-reticulate basally, finely semi-circularly striate apically. Remaining tergites smooth. Ovipositor sheath 4.2 × as long as petiole, 1.6 × as long as metasoma, 2.6 × as long as mesosoma, 1.2 × as long as fore wing. Colour. Head pale yellow, with brown or brownish frons at least medially, vertex with brown patch behind ocelli and on temples medially. Mesosoma dark reddish brown to almost black, with large yellow or pale yellow: areas along notauli and sometimes on anterior part of median lobe of mesoscutum, pronotum in lateral upper corner, propleuron posteriorly, most part of meso- and metapleuron, and propodeum laterally in anterior 0.7 – 0.8. Petiole black; second and third tergites mostly black with yellow narrow lateral (partly) and wide posterior parts, second tergite without yellow spots; fourth tergite black in basal 0.4 – 0.5, then medially brown and widely yellow laterally; following tergite brown, often laterally more or less widely yellow, apex of metasoma yellow. Scape and pedicel almost entirely brownish; basal half of flagellum brownish yellow to pale brown, apical half almost black. Palpi pale yellow. Fore and middle coxae and trochanters pale yellow; all femora yellow basally and apically and more or less widely brown medially; all tibiae more or less widely pale yellow, basally on short area and rather widely medially infuscate to almost black or dark brown; fore tarsus with almost black two basal segments and light reddish brown remaining segments; middle and hind tarsi mostly yellowish brown or pale brown, with first and often second segments distinctly infuscate in basal half. Ovipositor sheath brown or dark brown, but medially widely pale brown. Fore wing faintly infuscate, with more or less distinct maculation. Pterostigma dark brown, yellow in basal 0.25 – 0.30 and apically. Variation. Body length 5.2 – 5.8 mm; fore wing length 4.0 – 4.4 mm. Antennae 48 – 54 - segmented. Transverse diameter of eye 1.8 × length of temple. Mesosoma 2.1 × as long as its height. Basal carina of propodeum 1.5 × as long as anterior fork of areola. Fore wing 3.6 × as long as maximum width. Length of second submarginal cell 3.2 × its maximum width. Hind femur 2.9 × as long as wide. Length of petiole 2.2 × its apical width (dorsal view). Median length of second and third tergites combined 1.4 × basal width of second tergite. Ovipositor sheath 4.5 × as long as petiole, 1.7 × as long as metasoma, 2.8 × as long as mesosoma, 1.3 × as long as fore wing. Male. Unknown	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB0FFA1838EFF45FD5018DE.taxon	materials_examined	Material examined. Holotype: female, China, Zhejiang Prov., Gutianshan, 2. VII. 2005, Wu Qiong, No. 200615961 (ZJUH). Paratypes: 1 female, China, Zhejiang Prov., Gutianshan, 2. VII. 2005, Wu Qiong, No. 200615963 (ZJUH); 1 female, Zhejiang Prov., Gutianshan, 3. VII. 2005, Wu Qiong, No. 200616494 (ZISP); 1 female, Guangdong Prov., Shixing Chebaling, 22 – 28. VII. 2008, Wen Min, No. 200807885 (ZJUH); 1 female, Guangdong Prov., Shixing Chebaling, 22 – 28. VII. 2008, Hong Chundan, No. 200807854 (ZJUH). Comparative diagnosis. This new species belongs to the S. leucippus Nixon species group. Spathius subcyparissus sp. nov. is similar to S. cyparissus Nixon, but differs in having the petiole wide and 2.1 × as long as apical width (narrow and 2.6 × as long as apical width in S. cyparissus), second tergite almost entirely dark (with two rather large medioposterior yellow spots in S. cyparissus), pronotal carina fused medially on narrow area with posterior margin of pronotum (rather distinctly separated in S. cyparissus), lateral lobes of mesoscutum with short lateral rugae and with wide only granulate median areas (with long rugae and narrow granulate areas in S. cyparissus), vertex smooth (transverse striate in anterior 0.6 in S. cyparissus), hind femur wide, 2.8 × as long as maximum width (narrow, 3.4 × as long as maximum width in S. cyparissus), and mesopleuron weakly sculptured or almost smooth medially (entirely coarsely rugose-striate in S. cyparissus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB0FFA1838EFF45FD5018DE.taxon	etymology	Etymology. From “ sub ” (Latin for “ under ”) because it is similar to to S. cyparissus Nixon.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB0FFA1838EFF45FD5018DE.taxon	distribution	Distribution. China (Zhejiang, Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB1FFBC838EFAE3FDA11BA3.taxon	description	(Fig. 42) Description. Female. Body length 4.5 mm; fore wing length 3.3 mm. Head. Antennae filiform, 35 - segmented. Scape 1.7 × as long as its maximum width. First flagellar segment 6.0 × as long as its apical width, 1.2 × as long as second segment. Penultimate segment 4.2 × as long as wide, 1.1 × as long as apical segment. Head width (dorsal view) 1.45 × its median length, 1.25 × as wide as mesoscutum. Head behind eyes (dorsal view) weakly convex and roundly narrowed; transverse diameter of eye 1.6 × length of temple. Ocelli small, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 3.0: 3.0: 9.0. Vertex smooth, medially with fine to very fine transverse aciculation; frons entirely, distinctly, densely and weakly curvedly transversely striate. Eye glabrous, 1.1 × as high as broad. Malar space 0.55 × eye height and basal width of mandible. Temple smooth. Face in sublateral stripes densely and distinctly transversely striate with additional fine reticulation, smooth medially and finely rugulose to smooth laterally. Face width 1.25 × eye height and 1.25 × as long as height of face and clypeus combined. Hypoclypeal depression medium-sized and round, its width 0.9 × distance from edge of depression to eye, 0.45 × width of face. Occipital carina complete dorsally, ventrally not joined with hypostomal carina being obliterated upper base of mandible. Mesosoma. Length of mesosoma 2.1 × its height. Pronotal keel distinct, its posterior branch rather coarse, weakly separated from posterior margin of pronotum; anterior branch of carina distinct and situated submedially. Mesoscutum not highly and gently-roundly elevated above pronotum, its median lobe weakly convex anteriorly and without anterolateral corners. Mesoscutum distinctly and densely granulate, its median lobe anteriorly without granules situated in fine transverse lines, with long and dense rugae along notauli and laterally, with three distinct and convergent posteriorly carinae in medioposterior half; mostly glabrous, with rather dense, long and semi-erect or erect golden setae arranged in wide stripe along notauli and in single line laterally. Notauli wide, coarsely densely crenulate, deep anteriorly and more shallow posteriorly. Prescutellar depression rather deep, long, with five curved carinae, finely and densely rugulose, 0.4 × as long as scutellum. Scutellum entirely densely and finely to very finely granulate-coriaceous, without rugae, weakly convex, with lateral carinae. Mesopleuron entirely densely granulate-coriaceous with distinct striation below and posteriorly. Subalar depression shallow, rather narrow, coarsely rugose-striate and with granulation. Precoxal sulcus rather wide, sinuate, deep, straight and oblique in anterior 0.6, shallow and curved in posterior 0.4, densely crenulate, running along entire lower part of mesopleuron. Propodeum with low and wide lateral tubercles, with basolateral areas distinctly delineated by carinae and densely granulate with rugosity along carinae, rest part of propodeum sparsely striate and with dense and finely reticulation; areola rather wide and medium-sized; basal carina present in basal quarter, 0.7 × as long as anterior fork of areola. Wings. Fore wing 4.0 × as long as wide. Vein r arising weakly behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 25: 7: 43: 18. Second submarginal cell not narrowed distally, its length about 4.0 × maximum width, 1.3 × length of first subdiscal cell. Vein CU 1 a not interstitial, arising almost from middle of posterior margin of first subdiscal cell. Hind wing vein M + CU 0.55 × as long as vein 1 - M. Vein m-cu interstitial, oblique towards base of wing, faintly infuscate. Legs. Hind coxa dorsally weakly rugulose-striate with fine additional granulation, finely and densely granulate laterally, with distinct basoventral corner and tooth. Hind femur elongate-oval, almost entirely smooth, 3.5 × as long as wide. Hind tibia apically with three slender spines on outer side. Dorsal surface of hind tibia with very long, rather sparse and semi-erect or erect setae, their length 1.1 – 1.6 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.55 × as long as basitarsus, 1.6 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) almost straight ventrally, dorsally very weakly arched in basal 0.5 and straight in apical 0.5, thickened in basal 0.3; distinctly widened in apical fifth (dorsal view), with weak spiracular tubercles in weakly widened basal third. Length of petiole 2.7 × its apical width, 1.9 × length of propodeum. Second tergite without laterotergites. Median length of second and third tergites combined 1.4 × basal width of second tergite, 0.55 × as long as their maximum width. Second suture absent. Petiole distinctly and sparsely longitudinally striate, with dense and distinct reticulation between striae. Remaining tergites entirely smooth. Ovipositor straight. Ovipositor sheath 3.2 × as long as petiole, 1.3 × as long as metasoma, 2.2 × as long as mesosoma, as long as fore wing. Colour. Head yellow with brownish yellow spots on face and behind eyes. Mesosoma yellowish brown, laterally with darkened stripes. Petiole dark reddish brown to black, remaining metasoma yellowish brown with brownish spots. Antennae yellow to brownish brown in basal third, brown in apical two-thirds, without pale subapical segments. Palpi whitish. Legs yellow to brownish yellow, all femora (especially hind one) in apical thirds or half pale brown to brown, hind coxa entirely brownish yellow, hind tibia mainly yellow, brown on short basal and submedian area. Ovipositor sheath yellow in basal half and brown in apical half. Fore wing faintly and evenly infuscate. Pterostigma brown, pale yellow in basal third. Male Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB1FFBC838EFAE3FDA11BA3.taxon	materials_examined	Material examined. Holotype. female, China, Guangdong Prov., Shixing Chebaling, 21. VIII. 2003, Xu Zaifu, No. 20052613 (ZJUH). Comparative diagnosis. This new species belongs to the S. fasciatus Walker species group. Spathius suberymanthus sp. nov. is similar to S. erymanthus Nixon, but differs in having the body paler (darker in S. erymanthus), vertex partly finely aciculate (entirely smooth in S. erymanthus), antennae arranged not in equilateral triangle (arranged in equilateral triangle in S. erymanthus), mesoscutum with dense and long rugae (with short and fine rugae in S. erymanthus), propodeal areola distinctly delineated (not defined in S. erymanthus), and recurrent vein weakly postfurcal (distinctly postfurcal in S. erymanthus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB1FFBC838EFAE3FDA11BA3.taxon	etymology	Etymology. From “ sub ” (Latin for “ under ”) because of it is similar to S. erymanthus.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFB1FFBC838EFAE3FDA11BA3.taxon	distribution	Distribution. China (Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFAAFFBA838EFF45FD3A1DFC.taxon	description	(Fig. 43)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFAAFFBA838EFF45FD3A1DFC.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Fujian Prov., Jianyang Guadun, 25. IX. 1965, Chen Jiahua. China (ZJUH). 1 female, Yunnan Prov., Yingjiang Tongbiguan, 17. V. 2009, Zeng Jie, No. 200904269; 1 female, Hainan Prov., Jianfengling Tianchi, 12 – 15. VII. 2006, Weng Liqiong, No. 200803320.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFAAFFBA838EFF45FD3A1DFC.taxon	distribution	Distribution. China (Fujian, Yunnan, Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFAAFFBA838EFDCFFD011811.taxon	materials_examined	Material examined. China (ZJUH). Zhejiang Prov.: 1 female, Gutianshan, 3. VII. 2005, Zhang Hongying, No. 200616085. Hainan Prov.: 2 females, Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, Nos. 200802170 and 200802312; 1 female, Jianfengling, 6. VI. 2007, Liu Jingxian, No. 200703717; 3 females, Jianfengling, 7. VI. 2007, Liu Jingxian, Nos. 200702359, 200702483 and 200702521 (1 female in ZISP); 2 females, Jianfengling, 5 – 7. VI. 2007, Weng Liqiong, Nos. 200806458 and 200806487; 2 females, 1 male, Jianfengling, 4 – 7. VI. 2007, Zeng Jie, Nos. 200710867, 200711335 and 200711343; 1 male, Jianfengling Tianchi, 12 – 15. VII. 2006, Chen Tianfei, No. 200803175; 1 female, 1 male, Jianfengling Tianchi, 12 – 15. VII. 2006, Liu Jieming, Nos. 200803039 and 200803056; 1 female, Jianfengling Tianchi, 22 – 23. X. 2007, Liu Jingxian, No. 200710768; 1 female, Jianfengling Tianchi, 25. XI. 2008, Tan Jiangli, No. 200805088; 1 female, 1 male, Jianfengling Tianchi, 12 – 15. VII. 2006, Weng Liqiong, Nos. 200803331 and 200803344.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFAAFFBA838EFDCFFD011811.taxon	distribution	Distribution. China (Zhejiang, Hainan, Taiwan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFAAFFB8838EFBA8FDF31906.taxon	description	(Fig. 44) Description. Female. Body length 3.8 mm; fore wing length 2.6 mm. Head. Antennae slender, weakly setiform, more than 30 - segmented (apical segments missing). Scape 1.2 × as long as its maximum width. First flagellar segment 5.6 × as long as its apical width, 1.3 × as long as second segment. Head slightly depressed, its width (dorsal view) 1.2 × median length. Head behind eyes convex in anterior half, roundly narrowed in posterior half; transverse diameter of eye 1.1 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.4 × its sides; POL: Od: OOL = 6: 3: 9. Vertex with dense transverse striation, without additional ground sculpture between striae. Frons very densely and finely striate. Eye glabrous, 1.3 × as high as broad. Malar space 0.4 × eye height and 0.9 × basal width of mandible. Temple semi-circularly densely striate, almost smooth below. Face transversely striate, with dense microsculpture between striae; its width 1.1 × eye height and 1.3 × height of face and clypeus combined. Clypeal suture distinct. Clypeus flat, with narrow ventral flange. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.7 × its height. Pronotal keel with single, very high and far separated from posterior part of pronotum branch. Pronotal lateral depression rather narrow, deep, marginate, smooth, with several sparse striae. Mesoscutum weakly and roundly elevated above pronotum. Mesoscutum with sparse and semi-erect setae along notauli and laterally; entirely densely granulate. Notauli wide, deep, complete, coarsely crenulate. Prescutellar depression long, rather deep, with 3 strong carinae, 0.3 × as long as scutellum. Scutellum granulate, flat, with lateral carinae. Mesopleuron coarsely striate in upper half, granulate in lower half. Subalar depression shallow, wide, densely and coarsely striate. Precoxal sulcus deep, almost straight, crenulate and granulate, running along anterior 0.7 of lower part of mesopleuron. Propodeum without tubercles, coarsely striate-rugose and granulate; with areas indistinctly delineated by carinae. Wings. Fore wing 4.2 × as long as wide. Vein r arising before middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 14: 3: 18: 11. Second submarginal cell not narrowed distally, its length 3.7 × maximum width, 1.2 × length of first subdiscal cell. Vein cu-a slightly postfurcal. Vein m-cu postfurcal. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, strongly oblique towards base of wing. Legs. Hind coxa densely and distinctly granulate, without basoventral tooth and corner. Hind femur coriaceous, 2.9 × as long as wide. Hind tibia apically with five spines on outer margin. Dorsal surface of hind tibia with long, sparse and semi-erect setae, their length 1.4 – 1.6 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.4 × as long as second-fifth segments combined. Second segment of hind tarsus 0.6 × as long as basitarsus, 1.5 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) almost straight ventrally, arched dorsally, weakly thickened in basal fifth; distinctly widened in apical quarter (dorsal view), with small spiracular tubercles in basal third; densely rugulose, with striation laterally and apically. Length of petiole 2.9 × its apical width (dorsal view), 1.7 × length of propodeum. Second tergite with separate laterotergite. Second suture indistinct. Median length of second and third tergites combined 1.3 × basal width of second tergite, 0.7 × their maximum width. Second and third tergites entirely semi-circularly and finely transversely aciculate; fourth-sixth tergites in apical halves finely transversely aciculate, almost smooth in basal half. Ovipositor sheath 2.0 × as long as petiole, 0.8 × as long as metasoma, 1.1 × as long as mesosoma, 0.6 × as long as fore wing. Colour. Mesosoma and petiole black, head and remaining metasomal tergites reddish brown. Antennae yellowish brown, darkened toward apex. Palpi reddish brown, apical segment of maxillary palpi yellow. Tegula black. Legs reddish brown, fore and middle coxae pale yellow; all tarsi (except dark fifth segment) yellow; hind tibia entirely (including base) reddish brown. Ovipositor sheath white yellow in basal half, brown to dark brown in apical half. Fore wing distinctly infuscate, with hyaline transverse stripes apically, near base of pterostigma and subbasally. Pterostigma brown, white in basal quarter. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFAAFFB8838EFBA8FDF31906.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Bawangling, 26. XI. 2008, Tan Jiangli, No. 200805701 (ZJUH). Comparative diagnosis. This new species belongs to the S. ruficeps (Smith) species group. Spathius tanae sp. nov. is similar to S. manni Brues, but differs in having the precoxal sulcus running along anterior 0.7 of lower part of mesopleuron (running along entire lower length of mesopleuron in S. manni), propodeum with areas indistinctly delineated by carinae (with areas distinctly delineated by carinae in S. manni), head slightly depressed (not depressed in S. manni), and middle coxa pale yellow (reddish brown in S. manni).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFAAFFB8838EFBA8FDF31906.taxon	etymology	Etymology. This species is named in honour of Dr Tan Jiangli, the collector of the type specimen.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFAAFFB8838EFBA8FDF31906.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA8FFB8838EFAA4FBC31A8B.taxon	materials_examined	Material examined. China (ZJUH). Hainan Prov.: 1 female, Yinggeling, 24 – 25. V. 2007, Liu Jingxian, No. 200702738.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA8FFB8838EFAA4FBC31A8B.taxon	distribution	Distribution. China (Hainan) (new record); Indonesia; Malaysia, Singapore.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA8FFB6838EF8DDFDAC1EC6.taxon	description	(Fig. 45)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA8FFB6838EF8DDFDAC1EC6.taxon	distribution	Distribution. China (Jilin, Shanxi, Henan, Zhejiang, Fujian, Hubei, Hainan, Yunnan, Ningxia); Japan, Korea, Russia (Far East).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA8FFB6838EF8DDFDAC1EC6.taxon	discussion	Remarks. We examined the holotypes of S. verustus Chao and S. lunganjiding Chao from China. They all have the same characters, i. e short ovipositor sheath, palpi light reddish brown or yellow, hind tibia with long pale subbasal area, mesopleuron always smooth in lower half and fore wing with darkenings along veins, which allows it to be synonymised. Comparison of the material from the East Palaearctic determined as S. radzayanus Ratzeburg (Belokobylskij, 1996, 1998) with type material of S. radzayanus from the Deutsches Entomologisches Institut (Müncheberg, Germany) clearly showed that it is a different species. So, the Eastern Palaearctic material is treated here as belonging to S. verustus Chao.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA6FFB4838EFC96FDF3194E.taxon	description	(Fig. 46) Description. Female. Body length 4.0 mm; fore wing length 3.0 mm. Head. Antennae slender, setiform, 33 - segmented. Scape 1.4 × as long as its maximum width. First flagellar segment 4.0 × as long as its apical width, as long as second segment. Head not depressed, its width (dorsal view) 1.5 × median length. Head behind eyes roundly narrowed. Transverse diameter of eye 2.5 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 5: 5: 8. Vertex mostly coriaceous-granulate. Frons densely and transversely striate. Eye glabrous, 1.3 × as high as broad. Malar space 0.4 × eye height and 1.1 × basal width of mandible. Temple finely coriaceous-granulate. Face finely rugose-granulate, clypeus rugose-striate; its width equal to eye height and 1.2 × height of face and clypeus combined. Clypeal suture distinct. Occipital carina complete dorsally, not joined below with hypostomal carina being obliterated on short distance upper base of mandible. Mesosoma. Length of mesosoma 2.1 × its height. Pronotal keel distinct, its posterior branch widely fused with posterior margin of pronotum; anterior branch situated in posterior 0.3 of pronotum. Mesoscutum highly and roundly elevated above pronotum, with sparse and semi-erect setae along notauli and laterally; densely and coarsely granulate, without short rugae near notauli and laterally. Notauli shallow and narrow, complete, coarsely crenulate. Prescutellar depression shallow, with five strong carina, 0.4 × as long as scutellum. Scutellum densely granulate, weakly convex, with lateral carinae. Mesopleuron entirely granulate. Subalar depression shallow, wide, sparsely striate. Precoxal sulcus deep, wide, almost straight, crenulate, running along anterior 0.6 of lower part of mesopleuron. Propodeum without tubercles, but with areas distinctly delineated by carinae, basolateral areas densely granulate and striate along carina, densely rugose posteriorly; areola transversely striate; petiolate area distinctly separated from areola; basal carina 1.4 × as long as fork of areola. Wings. Fore wing 2.8 × as long as wide. Vein r arising slightly behind middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 21: 5: 35: 17. Second submarginal cell not narrowed distally, its length 3.1 × maximum width, 1.6 × length of first subdiscal cell. Vein cu-a distinctly postfurcal. Vein m-cu postfurcal. Hind wing vein M + CU 0.6 × as long as vein 1 - M. Vein m-cu antefurcal, slightly curved towards base of wing. Legs. Hind coxa dorsally striate and with granulation, coriaceous-granulate laterally and ventrally, with basoventral tooth and corner. Hind femur coriaceous, 3.2 × as long as wide. Hind tibia apically with four spines on outer margin. Dorsal surface of hind tibia with short, sparse and semi-erect setae, their length 0.6 – 0.8 × maximum width of tibia. Hind tarsus 0.8 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, 0.9 × as long as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) slightly curved ventrally, dorsally regularly arched in basal half, with spiracular tubercles in basal third; densely and coarsely striate, with reticulation between striae. Length of petiole 1.7 × its apical width (dorsal view), 1.4 × length of propodeum. Second and third tergites with separated laterotergites. Second suture absent. Median length of second and third tergites combined 1.1 × basal width of second tergite, 0.7 × their maximum width. Second-fourth tergites densely striate, with rugulosity between striae. Fifth tergite finely coriaceous in basal half. Remaining tergites smooth. Ovipositor sheath 1.2 × as long as petiole, 0.4 × as long as metasoma, 0.6 × as long as mesosoma, 0.3 × as long as fore wing. Colour. Head and mesocutum yellowish brown; scutellum and propodeum reddish brown; mesopleuron brown; petiole dark brown; second and third tergites dark brown in basal three-quarters, yellowish brown in apical quarter; remaining tergites dark brown in basal halves, yellowish brown in apical valves. Antenna yellowish brown basally, regularly infuscate toward subapex, eight apical segments pale yellow. Palpi pale yellow. Legs yellow, all tarsi brown. Ovipositor sheath light brown, black apically. Fore wing faintly infuscate. Pterostigma brown, pale yellow basally. Variation. Body length 2.8 – 3.6 mm. Scape 1.5 × as long as its maximum width. Transverse diameter of eye 2.8 × length of temple. Malar space 0.3 × eye height and equal to basal width of mandible. Length of mesosoma 2.2 × its height. Fore wing length 3.0 mm. Length of second submarginal cell 3.0 – 3.2 × its maximum width. Hind femur 3.0 × as long as wide. Second segment of hind tarsus 0.6 × as long as basitarsus. Length of petiole 1.6 – 1.8 × its apical width (dorsal view), 1.5 × length of propodeum. Median length of second and third tergites combined 1.0 – 1.2 × basal width of second tergite, 0.8 × their maximum width. Male. Unknown	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA6FFB4838EFC96FDF3194E.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802143 (ZJUH). Paratypes: 1 female, China, Hainan Prov., Diaoluoshan, 16 – 17. VII. 2006, Liu Jingxian, No. 200802175 (ZJUH); 2 females, Hainan Prov., Jianfengling Tianchi, 12 – 15. VII. 2006, Chen Tianfei, Nos. 200803138 and 200803196 (ZJUH, ZISP); 1 female, Hainan Prov., Jianfengling Tianchi, 12 – 15. VII. 2006, Liu Jingxian, No. 200803623 (ZJUH); 1 female, Hainan Prov., Jianfengling Tianchi, 24. XI. 2008, Tan Jiangli, No. 200806056 (ZJUH); 2 females, Hainan Prov., Jianfengling Tianchi, 12 – 15. VII. 2006, Zhang Wenyong, Nos. 200803427 and 200803532 (ZJUH); 1 female, Hainan Prov., Jianfengling, 4 – 7. VI. 2007, Zeng Jie, No. 200710938 (ZJUH). Comparative diagnosis. This new species belongs to the S. piperis Wilkinson species group. Spathius virgulatus sp. nov. is similar to S. piperis Wilkinson, but differs in having the vertex mostly coriaceous-granulate (striate in S. piperis), mesoscutum without strong rugosity on coarsely granulation (with strong rugosity in S. piperis), mesopleuron only entirely granulate (granulate and with striation in S. piperis), and basolateral areas of propodeum most densely granulate (transverse rugose in S. piperis).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA6FFB4838EFC96FDF3194E.taxon	etymology	Etymology. From “ virgulatus ” (Latin for “ striped ”), because of its metasoma distinctly striped.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA6FFB4838EFC96FDF3194E.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA4FFB4838EFA1EFC641AEF.taxon	materials_examined	Material examined. China (ZJUH). 1 female, Zhejiang Prov., Anji Longwangshan, 22. IX. 2004, Ma Yun, No. 20050040; 1 female, Guizhou Prov., Kuankuoshui, 4. VI. 2010, Zeng Jie, No. 201000587; 1 female, Zhejiang Prov., Kaihua Gutianshan, 21. VII. 1992, Wu Hong, No. 948985.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA4FFB4838EFA1EFC641AEF.taxon	distribution	Distribution. China (Zhejiang, Guizhou) (new record); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA4FFB3838EF8FFFDF31D76.taxon	description	(Fig. 47) Description. Female. Body length 4.0 mm; fore wing length 2.5 mm. Head. Antennae almost filiform, more than 14 - segmented (apical segments missing). Scape 1.8 × as long as its maximum width. First flagellar segment 5.8 × as long as its apical width, 1.2 × as long as second segment. Head width (dorsal view) 1.4 × median length. Head behind eyes (dorsal view) roundly narrowed; transverse diameter of eye 1.5 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.2 × its sides; POL: Od: OOL = 6: 5: 12. Vertex entirely, coarsely, weakly undulately and curvedly striate. Frons entirely, coarsely and curvedly striate. Eye glabrous, 1.2 × as high as broad. Malar space 0.6 × eye height and 1.2 × basal width of mandible. Temple mostly finely striate-reticulate. Face with very dense, fine and absolutely even transverse aciculation (like surface of gramophone record or CD); its width 1.3 × eye height and 1.1 × height of face and clypeus combined. Clypeal suture mostly absent. Hypoclypeal depression rather small and round, its width 0.7 × distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, not joined with hypostomal carina being obliterated upper base of mandible. Mesosoma. Length of mesosoma 1.8 × its height. Pronotal keel rather fine, almost absent, situated in middle of pronotum. Mesoscutum highly and perpendicularly elevated above pronotum, its median lobe weakly protruding forwards, weakly convex anteriorly, and with short and wide obtuse anterolateral corners (dorsal view). Mesoscutum mostly glabrous, with rather sparse, more or less long and almost erect pale setae arranged narrow along notauli and laterally; entirely densely granulate; with two distinct and convergent posteriorly carinae medioposteriorly. Notauli anteriorly distinct, wide and crenulate, very shallow in posterior 0.3. Prescutellar depression deep, rather short, with three carinae, finely rugulose-reticulate, 0.3 × as long as scutellum. Scutellum entirely densely and rather finely granulate, convex, with distinct lateral carinae. Mesopleuron mostly finely reticulate-coriaceous, with small and almost smooth areas. Subalar depression very shallow, wide, finely reticulate-areolate. Precoxal sulcus distinct, narrow, curved, crenulate, running along anterior half of lower part of mesopleuron. Propodeum with very short and wide lateral tubercles, with more or less distinctly delineated by carinae and granulate basolateral areas, its rest part coarsely and transversely striate with rugosity, with areola delineated by rather fine carinae. Wings. Fore wing 3.9 × as long as wide. Vein r arising almost from middle of pterostigma. 3 - SR: r: SR 1: 2 - SR = 21: 4: 20: 14. Second submarginal cell not narrowed distally, its length 3.8 × maximum width, 1.5 × length of first subdiscal cell. 1 - SR + M vein weakly curved. Vein CU 1 a almost interstitial. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu distinctly antefurcal, weakly curved and strongly oblique towards base of wing. Legs. Hind coxa dorsally distinctly transversely striate, widely and densely granulate at rest part, without basoventral tooth but with corner. Hind femur distinctly and densely granulate-coriaceous, 3.2 × as long as wide. Hind tibia apically without spines on outer side. Dorsal surface of hind tibia with rather short, dense and semi-erect setae, their length 0.7 – 1.0 × maximum width of tibia. Hind tarsus 0.8 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.5 × as long as basitarsus, as longer as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) almost straight ventrally, dorsally rather strongly arched in basal third and straight in apical two-thirds, thickened in basal third; weakly widened in apical fifth (dorsal view), with distinct wide spiracular tubercles in basal third; densely and distinctly striate with dense rugulosity between striae in basal 0.4 and finely striate apically. Length of petiole 3.0 × its apical width (dorsal view), 1.5 × length of propodeum. Second and following tergites without separate laterotergites. Median length of second and third tergites combined 1.7 × basal width of second tergite, as long as their maximum width. Second suture absent. Second and third tergites almost entirely and very densely finely granulate, basally with fine and rather short striation, almost smooth narrowly apically. Fourth-sixth tergites very finely and very densely granulate in basal halves and almost smooth in apical halves. Ovipositor straight. Ovipositor sheath 2.1 × as long as petiole, 0.8 × as long as metasoma, 1.4 × as long as mesosoma, 0.7 × as long as fore wing. Colour. Body reddish brown. Antennae yellow to pale brown, flagellar segments narrowly infuscate apically. Palpi dark brown. Legs reddish brown, middle coxa almost entirely yellow. Ovipositor sheath brownish yellow in basal 0.6 and black in apical 0.4. Fore wing distinctly and widely maculate; subbasal cell crossed by dark fascia on its narrowest part. Pterostigma dark brown, pale yellow in basal 0.4. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA4FFB3838EF8FFFDF31D76.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Jianfengling, 22. XI. 2008, Tan Jiangli, No. 200805314 (ZJUH). Comparative diagnosis. This new species belongs to the S. rusticulus Wilkinson (S. urios Nixon) species group. Spathius wuae sp. nov. is similar to S. prusias Nixon, but differs in having the notauli finely present in posterior 0.3 (absent in S. prusias), scutellum not strongly convex (strongly convex in S. prusias), propodeum with areas rather distinctly delineated by carinae (indistinctly delineated in S. prusias), and hind coxa with basoventral corner (without corner in S. prusias).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA4FFB3838EF8FFFDF31D76.taxon	etymology	Etymology. The species is named in honour of Prof. Wu Yanwu, the well-known Chinese hymenopterist.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA4FFB3838EF8FFFDF31D76.taxon	distribution	Distribution. China (Hainan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA3FFB3838EFE46FD241E4B.taxon	materials_examined	Material examined. China (ZJUH). Hainan Prov.: 2 females, Jianfengling, 7. VI. 2007, Liu Jingxian, Nos. 200702421 and 200702601; 1 female, Jianfengling, 4 – 7. VI. 2007, Zeng Jie, No. 200711098.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA3FFB3838EFE46FD241E4B.taxon	distribution	Distribution. China (Hainan, Taiwan); Japan.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA3FF4E838EFD03FDA11826.taxon	description	(Fig. 48) Description. Female. Body length 3.8 mm; fore wing length 2.1 mm. Head. Antennae 26 - segmented. Scape 1.8 × as long as its maximum width. First flagellar segment 5.5 × as long as its apical width, 1.25 × as long as second segment. Head width (dorsal view) 1.5 × median length. Head behind eyes (dorsal view) roundly narrowed; transverse diameter of eye 1.1 × length of temple. Ocelli medium-sized, arranged in triangle with base 1.1 × its sides; POL: Od: OOL = 10: 9: 25. Vertex almost entirely coriaceous. Frons entirely coriaceous, with coarse and curved striae. Eye glabrous, 1.5 × as high as broad. Malar space 0.4 × eye height and equal to basal width of mandible. Temple almost smooth. Face with very dense, fine and absolutely even transverse aciculation (like surface of gramophone record or CD); its width 1.1 × eye height and 1.1 × height of face and clypeus combined. Clypeal suture absent. Hypoclypeal depression rather small and round, its width 0.7 × distance from edge of depression to eye, 0.4 × width of face. Occipital carina complete dorsally, not joined with hypostomal carina being obliterated upper base of mandible. Mesosoma. Length of mesosoma 2.2 × its height. Pronotal keel rather fine, its posterior branch almost fused medially with posterior margin of pronotum, anterior branch situated submedially on pronotum. Mesoscutum not highly and roundly elevated above pronotum; its median lobe convex anteriorly and without anterolateral corners. Mesoscutum densely granulate-reticulate, without striae and rugosity. Notauli anteriorly rather deep, wide, crenulate, almost completely absent in posterior half. Prescutellar depression rather shallow, short, with three carinae, rugulose-reticulate, 0.4 × as long as scutellum. Scutellum convex, with distinct lateral carinae, entirely distinctly and densely granulate. Mesopleuron rather widely reticulate-coriaceous. Subalar depression shallow, wide, and finely reticulate-striate. Precoxal sulcus distinct, narrow, straight, finely reticulate-crenulate, running along anterior 0.5 of lower part of mesopleuron. Propodeum with very small lateral tubercles. Propodeum with areas more or less distinctly delineated by rather fine carinae, rugose-reticulate with granulation and especially densely in anterior half; areola delineated by rather fine carinae, narrow and long, basal carina almost as long as anterior fork of areola. Wings. Fore wing 4.5 × as long as wide. Vein r arising almost from middle of pterostigma. 3 - SR: r: SR 1 = 12: 2: 14. 2 - SR vein almost entirely absent. Second submarginal cell not narrowed distally, its length 3.3 × maximum width, 1.5 × length of first subdiscal cell. Vein m-cu interstitial. Vein cu-a interstitial; Vein CU 1 a almost interstitial. Hind wing vein M + CU 0.5 × as long as vein 1 - M. Vein m-cu interstitial, weakly curved towards base of wing. Legs. Hind coxa densely granulate, without basoventral tooth and corner. Hind femur granulate-coriaceous, almost smooth below, 3.3 × as long as wide. Hind tibia apically without spines on outer side. Dorsal surface of hind tibia with rather short, sparse and semi-erect setae, their length 0.7 – 1.0 × maximum width of tibia. Hind tarsus 0.9 × as long as hind tibia. Hind basitarsus 0.6 × as long as second-fifth segments combined. Second segment of hind tarsus 0.45 × as long as basitarsus, as longer as fifth segment (without pretarsus). Metasoma. Petiole (lateral view) weakly curved ventrally, dorsally weakly arched in basal third and straight in apical two-thirds, thickened in basal third; weakly widened in apical quarter (dorsal view), with spiracular tubercles in basal third, distinctly striate, distinctly rugulose between striae in anterior 0.7. Length of petiole 2.6 × its apical width (dorsal view), 1.6 × length of propodeum. Second tergite basally with separate laterotergite. Median length of second and third tergites combined 1.9 × basal width of second tergite, 0.9 × their maximum width. Second suture absent. Second and third tergites almost entirely with very dense and fine semi-circular aciculation. Fourth-fifth tergites with very fine coriaceous in basal halves and almost smooth in apical halves. Remaining tergites smooth. Ovipositor straight. Ovipositor sheath 1.8 × as long as petiole, 0.8 × as long as metasoma, as long as mesosoma, 0.55 × as long as fore wing. Colour. Body light reddish brown, partly only yellow, metasoma in posterior half dark reddish brown. Antennae yellow to brown, distinctly infuscate apically. Palpi brown, apical segments of maxillary palpi weakly paler. Legs brownish yellow, partly yellow. Ovipositor sheath brownish yellow in basal 0.7 and black in apical 0.3. Fore wing distinctly and widely maculate; subbasal cell crossed by dark fascia before its narrowest part. Pterostigma dark brown, pale yellow in basal 0.4. Male. Unknown.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA3FF4E838EFD03FDA11826.taxon	materials_examined	Material examined. Holotype: female, China, Guangdong Prov., Yunan Tongledashan, 12 – 13. VIII. 2003, Xu Zaifu, No. 20054357 (ZJUH). Comparative diagnosis. This new species is characterised by the absence of the vein 2 - SR and the presence of excessively fine, very dense and absolutely even transverse aciculation on face. Spathius xui sp. nov. is similar to S. buonluoicus Belokobylskij, but differs in having the vein cu-a interstitial (distinctly antefurcal in S. buonluoicus), propodeum with areola delineated by rather fine carina (without areola in S. buonluoicus), vertex without striation (with fine transverse striation in S. buonluoicus), and mesopleuron rather widely reticulate-coriaceous (smooth in S. buonluoicus).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA3FF4E838EFD03FDA11826.taxon	etymology	Etymology. This species is named in honour of Prof. Xu Zaifu, the collector of the holotype.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FFA3FF4E838EFD03FDA11826.taxon	distribution	Distribution. China (Guangdong).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF5EFF4E838EFBB6FDCE1944.taxon	description	(Fig. 49)	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF5EFF4E838EFBB6FDCE1944.taxon	materials_examined	Material examined. Female (holotype, IZCAS), Yunnan Prov., Xishuangbanna Menghun, 14. V. 1958, 1200 – 1400 m, Hong Chunpei.	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
03C0A869FF5EFF4E838EFBB6FDCE1944.taxon	distribution	Distribution. China (Yunnan).	en	Tang, Pu, Belokobylskij, Sergey, Chen, Xue-Xin (2015): Spathius Nees, 1818 (Hymenoptera: Braconidae, Doryctinae) from China with a key to species. Zootaxa 3960 (1): 1-132, DOI: 10.11646/zootaxa.3960.1.1, URL: https://doi.org/10.11646/zootaxa.3960.1.1
