taxonID	type	description	language	source
073B253007DD921C2022FD0DE18F01C7.taxon	materials_examined	Material examined. Type material. Holotype ♂ (Fig. 13); CHINA, Guangdong: Chebaling National Nature Reserve, Shixing County, 24.72 N, 114.26 E, alt. 496 m, 28. V. 2017, leg. Chen Kai. Paratypes: Hunan: 1 ♂, Mt. Huilongshan, Zixing, 26.08 N, 113.39 E, alt. 886 m, 8. VI. 2016, leg. Chen Kai and Duan Yongjiang; 1 ♂, Jinyinpu, Bamianshan Reserve, Guidong County, 25.97 N, 113.71 E, alt. 973 m, 16. VI. 2015, leg. Chen Kai; 2 ♀, Gaowangjie National Nature Reserve, Guzhang County, 28.66 N, 110.08 E, alt. 890 m, 18. VI. 2017, leg. Zhang Dandan, genitalia slides no. SYSU 0994 (molecular voucher no. SYSU-LEP 0191), 0957; Guangdong: 1 ♂, same data as holotype; 2 ♂, idem except leg. Duan Yongjiang, genitalia slide no. SYSU 0993, molecular voucher no. SYSU-LEP 0190; 2 ♂, idem except leg. Kou Zongqing, genitalia slide no. SYSU 0933.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
073B253007DD921C2022FD0DE18F01C7.taxon	diagnosis	Diagnosis. In appearance, E. crassihamata resembles E. cruoralis in the reddish brown subterminal band, but can still be recognized by the predominantly yellow basal 2 / 3 of the forewing sprinkled with reddish brown scales and the presence of a faint antemedial line on the forewing and postmedial lines on both wings. In male genitalia it differs from E. diaphana and E. xanthocrossa by the distally rounded, moderately setose uncus, the pointed and recurved dorsal process of the sacculus, and the long and slender phallus, which is longer than the length of the valva; from E. cruoralis it differs by the wider distal uncus, the small triangular, strongly sclerotized process near the distal sacculus as well as the hook-like ventral sella. In female genitalia, the sclerotized antrum is approximately 1.5 x as long as the anterior apophysis whereas in E. cruoralis the sclerotized antrum is as long as the anterior apophysis.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
073B253007DD921C2022FD0DE18F01C7.taxon	description	Description (Figs 3, 13). Head. Frons and vertex yellowish brown, frons with cream white stripe laterally. Antenna yellowish brown, cilia in male as long as width of corresponding flagellomeres. Labial palpus brown with white scales at base. Maxillary palpus brown. Thorax. Saffron dorsally, pale yellow ventrally. Foreleg: femur brown; tibia brown and white alternately; tarsi white except distal three brown. Midleg: femur pale brown; tibia yellow dorsally, white ventrally, outer spur half as long as inner one; tarsi white ventrally, pale yellow dorsally. Hindleg: femur pale brown; tibia yellow, basal inner spur in male about three times as long as basal outer spur, apical inner spur about twice as long as apical outer spur; tarsi pale yellow. Wingspan 17.5 - 18.5 mm. Forewing yellow edged by reddish brown subterminal band, sprinkled with reddish brown scales from base to postmedial line, slightly darkened along costal margin, veins covered with reddish brown scales terminally, terminal band narrow, saffron; antemedial line reddish brown, curved outwards from basal fourth of costa to about basal third of dorsum; orbicular stigma faint, dark brown; reniform stigma straight, strip-like, dark brown; postmedial line reddish brown, weakly sinuate from 3 / 4 of costa to base of M 2, bent inwards to base of CuA 2, then curved outwards to about middle of dorsum; inner margin of subterminal band nearly parallel to postmedial line; underside with ground colour as on upperside but paler; fringe saffron mixed with pale yellow scales, mostly reddish brown at tornus. Hindwing with costal margin translucent white to 2 / 3 of costa, basal half medially pale reddish brown, followed by pale yellow band, outer margin sinuate, edged by reddish brown subterminal band, terminal band narrow, saffron, veins with reddish brown scales terminally; postmedial line indistinct; fringe as in forewing; underside paler than upperside especially in basal half. Abdomen. Brown dorsally, whitish ventrally, apical margins of segments tinged with white.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
073B253007DD921C2022FD0DE18F01C7.taxon	etymology	Etymology. The specific name is derived from the Latin crassi- = thick and hamata = hook-like, referring to the thick, hook-like ventral sella.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
073B253007DD921C2022FD0DE18F01C7.taxon	distribution	Distribution (Fig. 24). China (Hunan, Guangdong).	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
B71FE7DAA72FEA9130093BE8859DA240.taxon	materials_examined	Material examined. Type material. Lectotype (here designated) ♂: INDIA, Meghalaya: Khasis, Mar. 1894, Nat. Coll., Pyralidae NHMUK Slide no. 10935 (NHMUK).	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
B71FE7DAA72FEA9130093BE8859DA240.taxon	materials_examined	Other material examined. INDIA, Meghalaya: 7 ♂ (Fig. 14), Khasia Hills, Assam, Nissary (NHMUK); 8 ♂, Assam, Khasis, Nat. Coll. (NHMUK); 1 ♂, same data as type (NHMUK); 1 ♂, Khasis Hills, Assam (NHMUK); 1 ♂, Khasis Hills (NHMUK); CHINA, Tibet: 1 ♀, air-raid shelter, Beibeng Village, Medog County, 29.24 N, 95.17 E, alt. 750 m, 31. VII. 2018, leg. Qi Mujie, genitalia slide no. ZDD 12100 (molecular voucher no. SYSU-LEP 0377) (NKU).	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
B71FE7DAA72FEA9130093BE8859DA240.taxon	diagnosis	Diagnosis. Emphylica cruoralis resembles E. crassihamata in the reddish brown subterminal band and the saffron fringe. The differences between the two species are provided in the diagnosis of E. crassihamata. In appearance, E. cruoralis can be best recognized within the genus by the yellow postmedial band of the forewing, in male genitalia by the narrow trapezoid uncus with hair-like setae at distal third, the large, thumb-shaped, weakly sclerotized process of the ventral valva near the distal sacculus as well as the triangular ventral sella overlaid by a folded, distally blunt process. In female genitalia, it resembles E. crassihamata except for the less sinuate distolateral antrum and the relatively shorter and wider antrum anteriorly.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
B71FE7DAA72FEA9130093BE8859DA240.taxon	distribution	Distribution (Fig. 24). India (Meghalaya), China (Tibet).	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
159CE0C81CE56669A97DE9704C68B34C.taxon	materials_examined	Material examined. CHINA, Fujian: 1 ♂, Letu rain forest, Hexi, Nanjing, 24.90 N, 117.22 E, alt. 270 m, 10. VII. 2014, leg. Zhang Dandan, genitalia slide no. SYSU 1040, molecular voucher no. SYSU-LEP 0263; Guangdong: 1 ♂, Sanyue Reserve, 24.03 N, 111.57 E, alt. 272 m, 6. VII. 2013, leg. Chen Xiaohua, genitalia slide no. SYSU 1041, molecular voucher no. SYSU-LEP 0264; Hainan: 1 ♂, Mt. Limushan, 5. V. 2011, leg. Zhang Dandan and Yang Lijun; Chongqing: 1 ♀, Daheba, Mt. Jinfoshan, alt. 800 - 850 m, 15. VII. 2010, leg. Du Xicui and Song Lifang, genitalia slides no. SYSU 0969; 1 ♂, Daheba, Mt. Jinfoshan, alt. 800 - 850 m, 16. VII. 2010, leg. Du Xicui and Song Lifang, genitalia slides no. SYSU 0965.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
159CE0C81CE56669A97DE9704C68B34C.taxon	diagnosis	Diagnosis. Emphylica diaphana resembles other Emphylica species in the conical frons and the scale-like setae of the sella. It can be best distinguished from its congeners by the whitish ground colour suffused with pale brown scales on both wings, the dark brown lines at termen, in male genitalia by the triangular, distally narrowly-rounded uncus bearing only few short setae and the long, strongly sclerotized ventral sella usually projecting beyond the ventral margin of the valva. In female genitalia of E. diaphana, the antrum is strongly sclerotized, shorter than the length of the anterior apophysis, slightly wider than the ductus bursae; the maximal length of the signum is approximately 2 / 3 as long as the diameter of the corpus bursae; the two opposing angles of the signum without carinae are well-developed, almost as long as the other two. In E. xanthocrossa the antrum is broad, lightly sclerotized, no more than twice as wide as the ductus bursae; the two opposing angles of the signum without carinae are fairly short; in E. crassihamata the antrum is longer than the anterior apophysis, in E. cruoralis the antrum is as long as the anterior apophysis and the signum of both species is small (less than half of the diameter of the corpus bursae).	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
159CE0C81CE56669A97DE9704C68B34C.taxon	distribution	Distribution (Fig. 24). China (Fujian, Guangdong, Hainan, Chongqing).	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
1B86A00EE9821114361CE1BA0C7CFB74.taxon	diagnosis	Diagnosis. Species of Emphylica have a conical frons (Figs 2 - 5), similar to species of Achyra, Loxostege and Sitochroa (Figs 6 - 10), and by this differing from most genera of Pyraustinae. They can be best distinguished from the genera mentioned above in male genitalia by the narrowly triangular to trapezoid, sparsely to moderately setose uncus, the scale-liked editum, the sclerotized ventral process of the sella pointing towards the ventral margin of the valva, the nearly U-shaped juxta, the well-developed, distally rounded saccus, and the interlaced spicules in the phallus. In female genitalia, the antrum is sclerotized and the signum is rhombic.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
1B86A00EE9821114361CE1BA0C7CFB74.taxon	description	Description. Head. Frons conical. Vertex with moderately raised scales projecting between antennae. Labial palpus slightly upwardly curved, approximately twice as long as diameter of eye; first segment with white scales at base; second segment obliquely directed upward; third segment long, porrect. Maxillary palpus prominent, curved upward. Proboscis well developed, with creamy white scales at base. Antenna in male with cilia shorter than or as long as width of corresponding flagellomeres. Thorax. Dorsal side whitish brown to brown; ventral side whitish to pale yellow. Legs unmodified, hindleg with basal inner spur longer than apical inner spur, approximately three times as long as basal outer spur. Wings. Forewing elongate-triangular, costa straight to near apex, then slightly arched to apex; apex sharp; termen weakly arched, oblique to tornus; dorsum straight; upperside usually with reddish or pale brown scales; frenulum hook in male well developed, retinaculum made up a tuft of curved bristles from below base of discal cell. Hindwing broad, fan-shaped; terminal margin usually brown; frenulum simple in male, with 2 acanthae in female. Wing venation as in Fig. 11. Abdomen. Apical margin of segments tinged with yellowish white. Male genitalia. Uncus narrowly triangular to narrowly trapezoid, more or less bulging near base. Tegumen trapezoid. Vinculum U-shaped. Saccus well developed, rounded triangular, approximately as long as uncus. Valva of medium width, tongue-shaped, slightly narrowed or tapering to rounded apex, ventral margin straight to slightly curved; transtilla short, triangular, usually with sclerotized ventral process extending to distal end of juxta; costal sclerotized band broad, slightly curved; dorsal sella short, lamellar, set with thick scale-like setae forming editum, more or less curved, apically with several filaments; ventral sella strongly sclerotized, usually perpendicularly pointing towards ventral margin of valva, usually curved apically; sacculus broad, usually with pointed sclerotized dorsal process (absent in E. xanthocrossa). Juxta usually U-shaped, distal arms sclerotized. Phallus tubular, usually with interlaced cornuti, in distal end with spine-like or teeth-like area of teeth. Female genitalia. Ovipositor lobes flat, densely setose. Posterior apophysis simple, anterior apophysis usually bulging near basal third. Antrum sclerotized. Ductus seminalis originating from anterior end of colliculum. Ductus bursae long and slender, more than 1.5 x as long as diameter of corpus bursae. Corpus bursae globular, spinulose; accessory bursa present, arising from corpus bursae mediolaterally; signum narrowly rhombic to sea-star-shaped.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
1B86A00EE9821114361CE1BA0C7CFB74.taxon	biology_ecology	Biology. All of the Chinese material has been collected during the night at light. Host information is currently unavailable.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
1B86A00EE9821114361CE1BA0C7CFB74.taxon	distribution	Distribution (Fig. 24). India, China, Australia.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
5D54262FE346B215D85EB93C091BBCE1.taxon	materials_examined	Material examined. Type material. Holotype, ♀: AUSTRALIA, Northern Territory: P [ort]. Darwin, Nov. [19] 08, leg. F. P. Dodd, genitalia slide no. P 232 (ANIC).	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
5D54262FE346B215D85EB93C091BBCE1.taxon	materials_examined	Other material examined (ANIC). AUSTRALIA, Northern Territory: 1 ♂, 16.19 S, 136.05 E, 36 km SW of Borroloola, NT, 4. Nov. 1975, leg. E. D. Edwards, K. Maes Gen. Prep. nr.: 20741; genitalia slide no. ANIC 18161; 1 ♂, 16.10 S, 136.15 E, Goose Lagoon, 11 km SW by S Borroloola, NT, 31. Oct. 1975, E. D. Edwards leg.; 1 ♂, Humpty Doo, N. T., Light Trap, 10. Nov. 1959, E. B. Boerema leg.; 2 ♂, 16.40 S, 135.51 E, Bessie Spring, 8 km ESE of Cape Crawford, NT, 26. Otc. 1975, E. D. Edwards leg., genitalia slide no. P 707; 1 ♀, 16.41 S, 135.44 E, Cape Crawford road junction, NT, 29. Mar. 1995, E. D. Edwards and M. Matthews leg.; Queensland: 1 ♂, 15.45 S, 144.15 E, 2 km NNW of Jowabinna, 17. I. 1994, E. D. Edwards and P. Zborowski leg., genitalia slide no. ANIC 21185; 1 ♀, 12.42 S, 142.30 E, Moonlight creek, QLD, 13. Nov. 1993, at light, P. Zborowski and M. Horak leg., K. Maes Gen. Prep. nr.: 20742, genitalia slide no. ANIC 18162; 1 ♂, 12.40 S, 142.40 E, Batavia Downs, QLD, 22 - 23. Nov. 1992, at light, P. Zborowski and A. Calder leg.; 1 ♂, 12.40 S, 142.41 E, Batavia Downs, QLD, 11. Dec. 1992, at light, P. Zborowski and W. Dressler leg.; Western Australia: 1 ♀, 15.77 S, 128.75 E, Hidden Valley, Kununurra, III. 2016, P. M. Heath leg., genitalia slide no. ANIC 21184, molecular voucher no. SYSU-LEP 0307; 1 ♂, Kunnunurra, W. A., 9. Apr. 1962, I. F. B. Common leg.; 1 ♂, Wyndham, W. A.,?.?. [19] 30, T. G. Campbell leg.; 1 ♂, 16.10 S, 128.23 E, nr Dunham River crossing, WA, 6. Apr. 1995, E. D. Edwards and M. Matthews leg.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
5D54262FE346B215D85EB93C091BBCE1.taxon	diagnosis	Diagnosis. Emphylica xanthocrossa resembles E. crassihamata and E. cruoralis in the saffron fringe, the conical frons and the U-shaped juxta. It can be best distinguished from its congeners by the smaller wingspan (less than 15 mm), the triangular saffron spot on the forewing costa postmedially, the smoky brown subterminal margin of the hindwing, in male genitalia by the distally concave uncus, the spinulose ventral sella, the absence of a dorsal process on the sacculus, the larger juxta, the broad and slightly sinuate phallus and the ductus ejaculatorius originating from the middle of the phallus. In female genitalia, the antrum is moderately sclerotized, bottle-shaped, the two opposing angles of the signum without carinae are short, whereas in E. diaphana, E. crassihamata and E. cruoralis the antrum is strongly sclerotized and the two opposing angles of the signum without carinae are almost as long as the other two.	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
5D54262FE346B215D85EB93C091BBCE1.taxon	distribution	Distribution (Fig. 24). Australia (Northern Territory, Queensland, Western Australia).	en	Chen, Kai, Liu, Qingming, Jin, Jianhua, Zhang, Dandan (2019): Revision of the genus Emphylica Turner, 1913 based on morphology and molecular data (Lepidoptera, Crambidae, Pyraustinae). ZooKeys 836: 113-133, DOI: http://dx.doi.org/10.3897/zookeys.836.32796, URL: http://dx.doi.org/10.3897/zookeys.836.32796
