identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C16916A34AFFF1FCEFF918FAADFEFE.text	03C16916A34AFFF1FCEFF918FAADFEFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mojsisovicsteuthis Jeletzky 1966	<div><p>Genus Mojsisovicsteuthis Jeletzky, 1966</p> <p>Type species: Orthoceras convergens von Hauer, 1847; p. 259, Tab. 7, figs. 1–2.</p> <p>Diagnosis: Phragmocone longi- to orthoconic, medium-sized (up to 40 cm), laterally compressed, apical angle 10–20°, chamber length-to-diameter ratio 0.16–0.30, final chamber tubular (= proostracum absent), its relative length uncertain; sutures simple or with lateral lobes, septal necks short, prochoanitic?, connecting rings swollen; siphuncle at the ventral narrow side; sheath investment-like, smooth, rostrum solidum apparently absent.</p> <p>Remarks: Despite an unusual set of characters, Jeletzky (1966) originally (and later authors such as Rieber, 1973 as well) placed the genus Mojsisovicsteuthis along with aulacoceratid belemnoids (see also Rieber, 1973). Prochoanitic septal neck as well as “conothecal growth lines” led them to this conclusion. Doyle (1990) placed Mojsisovicsteuthis to the family Xiphoteuthididae (Naef, 1922) within Aulacoceratida Stolley, 1919. By contrast, Mariotti and Pignatti (1992) and Mariotti et al. (2021) excluded Mojsisovicsteuthis from the Aulacoceratida mainly owing to the absence of a rostrum proper and untypically dense (short) chambers. Pohle &amp; Klug (2024) recently revisited the affinities of Mojsisovicsteuthis and concluded that the systematic affiliation are still puzzling. We follow this open nomenclature, because a tubular (ventrally closed) final chamber excludes this taxon from the Phragmoteuthida, whose members are typified by a ventrally opened final chamber. The overall mantle length in aulacoceratids and Mojsisovicsteuthis is difficult to estimate for the following reasons: (1) we do not know the proostracum in both taxa (if it was originally present) or its length. (2) large aulacoceratids s. l. are more than 50 cm long (Mariotti &amp; Pignatti, 1992)—but this length represents mainly the phragmocone. The largest size of the phragmocone within Mojsisovicsteuthis is estimated to be about 35 cm (see below).</p> <p>Other species previously assumed to be close to Mojsisovicsteuthis convergens (von Hauer, 1847) are M. elliptica (Mojsisovics, 1871), M. meneghinii (Salomon, 1895), M. subrotundus (Salomon, 1895), M. boeckhi (Stürzenbaum, 1875) and Mojsisovicsteuthis ? n. sp. (Jeletzky, 1966, Pl. 5, Fig. 1).</p> <p>Stratigraphic and geographic occurrences: So far known from the Anisian (Middle Triassic) through the Hettangian (Lower Jurassic).</p> <p>According to previous workers and their ambiguous samples, representatives of this rare genus are known from the Middle Triassic (Anisian to Ladinian) through the Lower Jurassic (Hettangian) of the Alpine region, Hungary, Indo-Pacific Region (Timor), with question mark also from British Columbia (Jeletzky, 1966) and also other regions (see below— M. boeckhi). In Slovakia (this report), it comes from dark-grey to black biodetritic limestones of the Ráztoka Limestone Member of Zámostie Limestone Formation, dated to lower Illyrian (upper Anisian), uppermost part of the Trinodosus Zone (Pseudohungaricum Subzone) —probably lowermost part of the Reitzi Zone (Felsoeoersensis Subzone?).</p> </div>	https://treatment.plazi.org/id/03C16916A34AFFF1FCEFF918FAADFEFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Košťák, Martin;Schlögl, Ján;Fuchs, Dirk;Havrila, Milan;Kolar-Jurkovšek, Tea;Vörös, Attila;Havelcová, Martina;Šurka, Juraj;Havrila, Jakub;Holcová, Katarína	Košťák, Martin, Schlögl, Ján, Fuchs, Dirk, Havrila, Milan, Kolar-Jurkovšek, Tea, Vörös, Attila, Havelcová, Martina, Šurka, Juraj, Havrila, Jakub, Holcová, Katarína (2024): Rare Middle Triassic coleoids from the Alpine-Carpathian system: new records from Slovakia and their significance. Swiss Journal of Palaeontology (19) 143 (1): 1-19, DOI: 10.1186/s13358-024-00316-7, URL: http://dx.doi.org/10.1186/s13358-024-00316-7
03C16916A348FFFDFCEFFED3FF77F8B4.text	03C16916A348FFFDFCEFFED3FF77F8B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mojsisovicsteuthis boeckhi (Sturzenbaum 1875)	<div><p>Mojsisovicsteuthis boeckhi (Stürzenbaum, 1875)</p> <p>(Fig. 5, A–Q; 6; 7 A–D)</p> <p>Synonymy:</p> <p>*1875 Atractites böckhi Stürzenbaum, p. 254, Tab. V, fig. 1.</p> <p>1882 Atractites boeckhi (Stürzenbaum). Mojsisovics, p. 302, pl. XCIII, figs. 12–13.</p> <p>? 1914 Atractites böckhi Stürzenbaum. Smith, p. 138, pl. XCIV, figs. 20–21.</p> <p>1915 Atractites Boeckhi Stürzenbaum, Diener, p. 18 (with additional/full synonymy).</p> <p>1920 Atractites Boeckhi Stürzenbaum; Büllow-Trummer, p. 65 (with additional synonymy).</p> <p>1973 Mojsisovicsteuthis cf. M.? boeckhi (Stürzenbaum, 1875); Rieber, p. 73–74, fig. 20, a–g.</p> <p>? 1973 Mojsisovicsteuthis meneghinii (Salomon, 1895); Rieber, p. 74, fig. 20, n–o.</p> <p>? 1975 Ausseites bilhynicus Arthaber, 1915; Özdemir, p. 137, pl. 4, figs. 1–4.</p> <p>2024 Mojsisovicsteuthis boeckhi. Pohle &amp; Klug, XXXX</p> <p>Studied material: A single specimen (Fig. 5A–D) from Harmanecká Cave—Kozelník (GPS 48°48,582 ′ N 19°02,265 ′ E, Slovakia, No. KGP-KO-001; stored at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.03775&amp;materialsCitation.latitude=48.8097" title="Search Plazi for locations around (long 19.03775/lat 48.8097)">Comenius University in Bratislava</a>). Five specimens from the type locality (Additional file 1: Table S 1) stored in the Mining and Geological Survey of Hungary in Budapest: including the holotype No. T.829 (Fig. 5J–P); specimens Nos T.3593, T.3206, T.3137 and two fragments referred to the taxon (T.81, and T.371).</p> <p>Description: The steinkern of the phragmocone fragment is 42 mm long and includes 7–8 chambers. Apical and apertural chambers are missing. The width (dimension of the flattened phragmocone) of the largest chambers measures 19 mm. The straight phragmocone appears longi- to orthoconic (apical angle 11–12°, Fig. 6). Thanks to a circular structure in the periphery of the smallest chamber determinable as the former position of the siphuncle, one can define the anatomical orientation. The phragmocone diameter is accordingly laterally compressed. Poorly defined remains of sutures, which are visible only on one side of the steinkern, are c. 4 mm apart implicating a ratio chamber length to chamber diameter of about 0.19. This suggests comparatively low chambers. Lateral lobes are perceptible. There is no evidence of preserved shell material. As mentioned above, the siphuncle is only slightly indicated in Slovakian specimen.</p> <p>The material stored in the collections of Mining and Geological Survey of Hungary in Budapest provided unambiguous evidence of the siphuncle position (Fig. 5G, H) which lies in the narrow ventral side. The diameter of the siphuncle is ~ 1.3 mm (specimen No. T.3593, Fig. 5G, H).</p> <p>In specimens Nos T.3206 (Fig. 7A) and T.81 (Fig. 7B), the tubular final chamber is partly preserved. The preserved length measures 25 and 30 mm (Additional file 1: Table S1). In these specimens, a marked constriction of the last septum prior the final chamber is seen.</p> <p>Comparing the specimen No. KGP-PO-0001 with the Hungarian material including the holotype, we observed stability in the phragmocone angle reaching 11–12°.</p> <p>With the phragmocone diameter and the apical angle of 12°, one can calculate the maximum phragmocone length by the help of trigonometric laws [diameter/2 × sin (84)/sin (12/2)]. Accordingly, specimen T.3206 with a maximum phragmocone diameter of 37 mm had a phragmocone length of approx 180 mm (length of the final chamber is not known). Assuming a final chamber that reaches a similar length (in specimens with this part preserved, Fig. 6), the total shell length of this specimen was about 350 mm, which classifies it as a medium-sized coleoid.</p> <p>Comparison: We exclude affinities to the order Phragmoteuthida, owing to the absence of a three-lobed proostracum and the presence of a longiconic (rather than brevi—or cyrtoconic) phragmocone. Despite our observation of an aulacoceratid-like final chamber, the overall phragmocone characteristics (dense chambers, apical angle, lateral lobus) also prevent a placement within the order Aulacoceratida.</p> <p>Regarding chamber distance, the specimen from Slovakia resembles the holotype of M. convergens, the type species of Mojsisovicsteuthis. However, in M. convergens, the cross section is more circular or even almost circular, as figured in von Hauer (1847, pl. VII, fig. 2), later corrected by Mojsisovics (1902, Pl. XVI, fig. 1). The apical angle of the phragmocone in the dorso-ventral view is moreover significantly higher in M. convergens (20°).</p> <p>The Slovakian specimen and the holotypes of M. boeckhi (Stürzenbaum, 1875) and M. meneghinii (Salomon, 1895, p. 195) share a laterally compressed phragmocone, but M. meneghinii differs from our specimen in a higher apical angle (16–19° according to Salomon, 1895) and more densely spaced chambers (0.13–0.17).</p> <p>In terms of apical angle, our specimen is almost identical with the holotype of M. boeckhi (Fig. 6). In addition, the Slovakian and the Hungarian specimens share a compressed phragmocone, a well developed lateral lobus, and a chamber distance ranging from 4 to 12 mm. The chamber distances just prior the final chanber are significantly constricted (in specimen T.81 ~ 3 mm).</p> <p>Recently, Vörös et al. (2022) published record of cephalopod fauna from the Middle Anisian locality Akol Hill at Barnag (Hungary) including specimen described as Mosjsisovicsteuthis sp. The specimen clearly differs from our material by having significantly more densely spaced septa.</p> <p>Specimens similar to Slovakian specimen and referred to genus Ausseites (Flower, 1944; rejected as nomen dubius by Mariotti et al., 2021) were figured by Özdemir (1975) from Ladinian/Carnian transition of Koaceli (Turkey). A. bilhynicus (Arthaber, 1915) resembles our specimen in shape, but the apical angle is higher in A. bilhynicus (~ 20–21°).</p> <p>Remarks: The relatively large size (up to 30 cm) of the phragmocone (Fig. 6) is comparable to larger specimens of aulacoceratids. The reconstruction of the length is based on the stability in the phragmocone angle in all ontogenetical stages studied.</p> <p>Stratigraphy and palaeogeography: M. boeckhi is known from the Anisian—Ladinian strata of the Alpine-Carpathian-Dinaridic region (Trinodosus—Lomelli zones; Alma, 1926). Apart of the here described material, it is reported from Wettersteinkalk in Germany (Reis, 1901), Austria (Alma, 1926 —Schreyeralmkalke, Marmolatkalke; Schnetzer, 1934 —Muschelkalk), Italy (Marmolada— Salomon, 1895; Monte Clapsavon— Canavari, 1890), Montenegro (Martelli, 1904),? Romania (Braşov, Ladinian, see on-line report: https://formatiunigeolo gice.igr.ro/formatiune/42), Hungary (Stürzenbaum, 1875; Vörös, 1987, 2018), Bosnia and Herzegovina (Kraus, 1916; von Hauer, 1888), Greece (Renz, 1909), Bulgarian/ Romanian borders (Dobrudzha; Büllow-Trummer, 1920) and? Turkey (Koaceli—Ladinian/Carnian transition; Özdemir, 1975). Smith (1914) reported M. boeckhi from the Trinodosus Zone of North America. However, these old records (predominantly attributed to “ Atractites boeckhi ” or? Ausseites in the latter case) were not studied/verified by the authors of this article.</p> <p>Crown Neocoleoidea Haas, 1997 (proostracum-bearing coleoids)</p></div> 	https://treatment.plazi.org/id/03C16916A348FFFDFCEFFED3FF77F8B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Košťák, Martin;Schlögl, Ján;Fuchs, Dirk;Havrila, Milan;Kolar-Jurkovšek, Tea;Vörös, Attila;Havelcová, Martina;Šurka, Juraj;Havrila, Jakub;Holcová, Katarína	Košťák, Martin, Schlögl, Ján, Fuchs, Dirk, Havrila, Milan, Kolar-Jurkovšek, Tea, Vörös, Attila, Havelcová, Martina, Šurka, Juraj, Havrila, Jakub, Holcová, Katarína (2024): Rare Middle Triassic coleoids from the Alpine-Carpathian system: new records from Slovakia and their significance. Swiss Journal of Palaeontology (19) 143 (1): 1-19, DOI: 10.1186/s13358-024-00316-7, URL: http://dx.doi.org/10.1186/s13358-024-00316-7
03C16916A344FFFDFF55F8CCFAF2F9B5.text	03C16916A344FFFDFF55F8CCFAF2F9B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Breviconoteuthis Rieber 1973	<div><p>Genus Breviconoteuthis Rieber, 1973</p> <p>Type species: Atractites breviconus Reis, 1907; p. 148, pl. 3. fig. 3 by monotypy.</p> <p>Diagnosis (after Fuchs &amp; Donovan, 2018): Phragmocone very small to small, brevi- to slightly cyrtoconic, circular in cross section, apical angle 38°–40°; ratio of chamber length to diameter about 0.15; pro-ostracum three-lobed, longer than phragmocone, anterior median field rounded; siphuncle marginal; rostrum unknown.</p> <p>Diagnosis (emended): Phragmocone very small to small sized, brevi- to slightly cyrtoconic, circular in cross section, apical angle 30–40°; ratio of chamber length to diameter between 0.15–0.25; pro-ostracum three-lobed, longer than phragmocone, anterior median field rounded; siphuncle marginal; rostrum unknown.</p> <p>Stratigraphic and geographic occurrences: So far known only from the upper Anisian (Wettersteinkalk) of Austria (Tyrol), Switzerland (Fuchs &amp; Donovan, 2018) and Slovakia (Anisian—lower Illyrian; this paper).</p> <p>Remarks: Breviconoteuthis and the slightly younger genus Phragmoteuthis Mojsisovics, 1882 (Carnian), the eponym of the order Phragmoteuthida, share a three-lobed proostracum morphology. Therefore, both genera can presently be distinguished only by phragmocone characters. The phragmocone of Breviconoteuthis displays a ventral curvature that is so far unknown in Phragmoteuthis.</p> <p>The genus Zugmontites Reis, 1907 (upper Anisian) is based on a single specimen whose phragmocone parameters are close to those of Breviconoteuthis. Fuchs and Donovan (2018) interpreted Zugmontites as a putative phragmoteuthid owing to the absence of a proostracum. Due to the lack of available specimens, it can be presently not excluded that Zugmontites mojsisovicsi Reis, 1907, the type spieces of Zugmontites, is identical to Breviconoteuthis breviconus. One feature delimiting the two type species might be the ventral curvature that seems to be more distinct in Breviconoteuthis than in Zugmontites.</p> </div>	https://treatment.plazi.org/id/03C16916A344FFFDFF55F8CCFAF2F9B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Košťák, Martin;Schlögl, Ján;Fuchs, Dirk;Havrila, Milan;Kolar-Jurkovšek, Tea;Vörös, Attila;Havelcová, Martina;Šurka, Juraj;Havrila, Jakub;Holcová, Katarína	Košťák, Martin, Schlögl, Ján, Fuchs, Dirk, Havrila, Milan, Kolar-Jurkovšek, Tea, Vörös, Attila, Havelcová, Martina, Šurka, Juraj, Havrila, Jakub, Holcová, Katarína (2024): Rare Middle Triassic coleoids from the Alpine-Carpathian system: new records from Slovakia and their significance. Swiss Journal of Palaeontology (19) 143 (1): 1-19, DOI: 10.1186/s13358-024-00316-7, URL: http://dx.doi.org/10.1186/s13358-024-00316-7
03C16916A344FFFAFC80F9AEFB91FBB8.text	03C16916A344FFFAFC80F9AEFB91FBB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Breviconoteuthis breviconus (Reis 1907)	<div><p>Breviconoteuthis aff. breviconus (Reis, 1907)</p> <p>(Fig. 7E–L).</p> <p>Studied material: A single specimen from the locality Podhradie (Slovakia, No. KGP-PO-001; stored at Comenius University in Bratislava). The specimen was compared to the type material—e.g. B. breviconus (No. PIMUZ M12; Fig. 7M) stored in the Paläontologisches Institut und Museum der Universität Zürich, Switzerland; and Zugmontites mojsisovicsi Reis, 1907 (No. 1901-II- 508; Fig. 7N–O) stored in the Bayerische Staatssammlung für Paläontologie und Historische Geologie München, Germany.</p> <p>Locality: Podhradie (GPS 48° 39 ′ 27.2 ″ N 18° 03 ′ 11.2 ″ E), eastern part of the Považský Inovec Mts.</p> <p>Description: The specimen represents a steinkern of a brevi- to slightly endogastric cyrtoconic phragmocone. Its apical angle measures 34° in dorsoventral and 32° in lateral views. Neither mineralized (e.g., conotheca, septa, sheath) nor nonmineralized shell material (e.g., proostracum) is preserved. Also, the ontogenetically oldest chambers are missing. The preserved part is up to 65 mm long and 45 mm wide and includes 11 chambers. Therefore, the phragmocones diameter seems to be dorsoventrally flattened; at least in the ontogentically youngest chambers. Annular constrictions on the outer surface, which correspond to former mural ridges, suggest a septal distance that gradually increases from 4 mm (apical part) to 8 mm at the anterior part. The ratio chamber length to chamber diameter varies from 0.19 to 0.23 (average 0.22). The sutures are unusually inclined towards the venter (backwards inclined). They appear to be simple without lobes or saddles. In ventral view, one can recognize a longitudinal depression that likely correlates with the former position of the siphuncle. This observation implicates that the septal necks were in contact with the inner surface of the conotheca, which in turn suggests a marginal rather than a submarginal siphuncle. There is no evidence of a longitudinal keel along the dorsum.</p> <p>Comparisons: In having a brevi- to slightly cyrtoconic phragmocone (Fig. 7M herein), the here reported Breviconoteuthis aff. breviconus is similar especially to the holotype of Breviconoteuthis breviconus Rieber, 1973. Taking into consideration that our specimen might have suffered a faint compaction, their apical angles do not differ significantly. Major differences between our Breviconoteuthis aff. breviconus and Breviconoteuthis breviconus concern the chamber distance (longer in our specimen ~ 4–8 mm vs. 2–3 mm in the holotype) and in the siphuncle characteristics (its wider cross-section in our specimen). The inclination of the septa, which is backward in our specimen, might represent the main difference, but this character is unfortunately poorly known in the type specimens.</p> <p>Mojsisovics (1871, tab 2, fig. 9) described a similar taxon Aulacoceras ellipticum. The figured specimen resembles Breviconoteuthis aff. breviconus in the septal and cameral arrangement (cf. Fig. 7E, F). However, the chamber distances are visibly larger in B. aff. breviconus. Furthermore, A. ellipticum is laterally compressed (in this respect, it more resembles genus Mosisovicsteuthis, see above), whereas B. aff. breviconus is dorsolaterally flattened. Regarding this aspect, A. ellipticum may represent another and younger Breviconoteuthis species (Austriacum Zone, Carnian; Mojsisovics, 1871), although the lateral compression also links this specimen to Zugmontites (Fig. 7N, O, herein).</p> <p>The slight phragmocone compaction may have resulted in different compressions of phragmocones in several phragmoteuthid taxa. We assume our specimen might be slightly dorso-ventrally flattened, therefore, the original cross-section should actually be more circular. However, the compression is very slight as it is seen in septal shapes, showing no significant deformation (Fig. 7 K, L).</p> <p>Stratigraphy and palaeogeography: The species is known from the Middle Triassic (upper Anisian) of Switzerland, Austria (Fuchs &amp; Donovan, 2018) and Slovakia (this report). Here, it comes from dark-grey to black biodetritic limestones of the Ráztoka Limestone Member of Zámostie Limestone Formation, dated to lower Illyrian (upper Anisian), uppermost part of the Trinodosus Zone (Pseudohungaricum Subzone) —lowermost part of the Reitzi Zone.</p> <p>Palaeobiogeographic distribution of recorded coleoids is shown on the Fig. 8.</p></div> 	https://treatment.plazi.org/id/03C16916A344FFFAFC80F9AEFB91FBB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Košťák, Martin;Schlögl, Ján;Fuchs, Dirk;Havrila, Milan;Kolar-Jurkovšek, Tea;Vörös, Attila;Havelcová, Martina;Šurka, Juraj;Havrila, Jakub;Holcová, Katarína	Košťák, Martin, Schlögl, Ján, Fuchs, Dirk, Havrila, Milan, Kolar-Jurkovšek, Tea, Vörös, Attila, Havelcová, Martina, Šurka, Juraj, Havrila, Jakub, Holcová, Katarína (2024): Rare Middle Triassic coleoids from the Alpine-Carpathian system: new records from Slovakia and their significance. Swiss Journal of Palaeontology (19) 143 (1): 1-19, DOI: 10.1186/s13358-024-00316-7, URL: http://dx.doi.org/10.1186/s13358-024-00316-7
