identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03CF87EEFF88D647B38CFDC4FD89B8D1.text	03CF87EEFF88D647B38CFDC4FD89B8D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cymonomus suluensis Takeda & Ohtsuchi & Komatsu 2021	<div><p>Cymonomus suluensis sp. nov.</p> <p>(Figs. 2–3)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 08 (Sulu <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.03&amp;materialsCitation.latitude=2.03" title="Search Plazi for locations around (long -2.03/lat 2.03)">Sea</a>; 08°44.6′N, 119°05.4′E – 08°44.8′N, 119°06.2′E; 2,030 – 2,030 m deep); 3 m beam trawl; 25 May, 1972; ˂ holotype, NSMT-Cr 28965.</p> <p>Measurements. CB 5.1 mm; CL 4.4 mm excluding rostrum; length of rostrum, 2.5 mm; length of eyestalk, 1.5 mm including basal segment; length of ambulatory leg, 14.5 mm.</p> <p>Diagnosis. Small species. Carapace subquadrate in outline, slightly wider than long, with weakly convex branchial regions; carapace dorsal surface and lateral margins covered and fringed with many spinules and granules. Rostrum long, about one-third as long as carapace, directed forward, weakly tapering, armed with three spinules at each side along distal half. Inner half of supraorbital margin with some tubercles; external orbital tooth spiniform, about half as long as rostrum, directed obliquely outward. Eyestalk fixed, slender, directed obliquely outward along whole length, obliquely downward for its basal half, horizontal for its distal half; in dorsal view, distal end of eyestalk just at level of basal one third of rostrum; eyestalk armed with 2 distant spinules on upper surface; cornea distinct at tip of eyestalk, without pigments. Chelipeds armed with spinules along margins of merus, carpus and palm; distal part of carpus and proximal part of palm deeply excavated to form a deep cavity. Ambulatory leg slender.</p> <p>Description of holotype (Female). Carapace (Figs. 2A, 3A) subquadrate in outline, only slightly wider than long; carapace lateral margins weakly convex outward along hepatic and branchial parts, shallowly concave between both parts; carapace dorsal surface (Fig. 2C–D) not convex as a whole, roughened with small, sharp dispersed granules mainly on branchial regions; gastric, branchial and cardiac regions weakly demarcated with shallow depressions or indistinct furrows; anterior extension of mesogastric region narrow, reaching nearly to basal part of rostrum; a suberect spinule at anterior part of protogastric region; protogastric and hepatic regions indistinctly confluent, separated from branchial region by an oblique furrow from shallow depression between hepatic and branchial outer margins to a transverse shallow, short furrow separating cardiac and intestinal regions; gastro-branchial oblique furrow turned obliquely outward to lateral end of carapace posterior margin along intestinal region.</p> <p>Rostrum (Figs. 2A, 3A–C) long, about one-third as long as carapace, directed forward, weakly tapering, with 3 spinules on each margin along distal half; proximal spinules of both sides alternate in position, diminishing in length distally. Supraorbital margin (Figs. 2A, 3A–C) nearly concave along inner half, with some marginal spinules weakly directed outward; outer half of supraorbital margin unarmed, weakly curving downward toward external orbital tooth. External orbital tooth (Fig. 3A–C) spiniform, slender, with tip bifid in left side, broken off in right side, extending beyond tip of eyestalk. Subhepatic tooth (Fig. 3A–B) shorter than external orbital and hepatic spiniform teeth, but sharp, directed obliquely outward. Hepatic margin (Figs. 2A, 3A) weakly convex, with a strong spine at anterior one third, a shorter spine on posterior slope and some subsidiary spinules. Carapace branchial margin (Figs. 2A, C, 3A) twice as long as hepatic margin, armed with a series of 10–15 spinules of approximately similar size. Carapace posterior margin (Figs. 2A, C–D, 3A) as long as front-orbital margin, rather strongly concave at median part, with a narrow but deep marginal furrow along whole length of posterior margin.</p> <p>Eyestalk (Fig. 3A–C) slender, directed obliquely outward for whole length along external orbital tooth, obliquely downward for its basal half, horizontally for its distal half, distal end of eyestalk attaining to tip of inner infraorbital tooth; in dorsal view, distal end of eyestalk slightly exceeding one-third of, or slightly less than half of rostrum; cornea distinct at distal end of eyestalk, without pigment; upper surface of eyestalk armed with 2 outward-directed spinules with some distance.</p> <p>Third maxilliped (Fig. 2B) pediform; ischium elongated, rectangular, curved outward distally; merus half as long as ischium, weakly curved inward as a whole, with rounded disto-lateral angle; merus with spine at median part of basal one-third of outer surface; outer margin armed with 4 strong spines, basal 2 directed laterally, distal 2 obliquely forward, inner margin with a series of spines, 2 sub-erect spines at subdistal part much stronger than others; outer margin of exopod armed with 3 erect equidistant spines at side of ischium, each side of merus with an obliquely-forward directed spinule.</p> <p>Pleon with 6 pleomeres and telson (Fig. 2B, D); pleomere 1 small, quadrate, lateral margins weakly concave for whole length; pleomere 2 prominent, widening posteriorly, with 2 spines along median line and a cluster of some spines at each lateral part; pleomere 3 similar to pleomere 2 in shape and armature, with much smaller spines and spinules; pleomere 4 as wide as pleomere 3, with vestigial tubercles in median line; pleomeres 5–6 and telson weakly tapering distally, with thin plate-like appearance, unarmed, each pleomere distinctly articulated, separated laterally from other pleomeres by V-shaped notch; telson obtuse at tip, forming rounded triangle as a whole.</p> <p>Both chelipeds (Figs. 2E–F, 3D) equal in size and shape, not inflated. Merus weakly curved, with about equidistant 10 spinules of alternate sizes on posterior margin of merus. Carpus armed with some longer spinules and some smaller spinules on outer and upper surfaces, respectively; inner angle produced to be a tubercle directed forward. Palm and fingers together weakly curved inward; inner proximal part of palm (Figs. 2E–F, 3D) deeply excavated together with distal inner part of carpus to form a deep cavity. Palm and fingers (Fig. 3E) subequal in length, with untoothed cutting edges of fingers. A detached ambulatory leg of left side long, slender (Fig. 3F).</p> <p>Remarks. Ng et al. (2008) enumerated 24 species in the genus Cymonomus A. Milne-Edwards, 1880 and five species in the genus Cymonomoides Tavares, 1993. Both genera are distinguished only by the fusion of the telson and the sixth pleomere being indistinguishably fused as a pleotelson in Cymonomus, but separated by a visible suture in Cymonomoides. Ahyong and Ng (2009), however, doubted the validity of Cymonomoides on the basis that in Cymonomus diogenes newly described by them from the Philippines, the sixth pleomere and the telson are immovably fused with a clear suture in males, but completely fused without a visible suture in females. Recently, Ahyong (2019) decidedly returned Cymonomoides delli (Griffin and Brown, 1976), C. cubensis (Chace, 1940) and C. valdiviae (Lankester, 1903) to the original position in the genus Cymonomus. As rightly mentioned by Ahyong and Ng (2017), the genus Cymonomoides may be restricted to the type species, C. guinotae (Tavares, 1991) from Brazil, and C. fitoi Lemaitre and Bermudez, 2000 from the Caribbean coast of Colombia.</p> <p>Since the enumeration of the species by Ng et al. (2008), altogether 18 species were additionally described as follows: 1) one species from New Zealand (Ahyong, 2008); 2) four species from the Philippines (Ahyong and Ng, 2009); 3) one species from the Philippines (Ahyong and Ng, 2011); 4) one species from off Madagascar (Ahyong, 2014); 5) two species from Taiwan and Japan (Ahyong and Ng, 2017); 6) eight species from Australia and New Zealand (Ahyong, 2019); 7) one species from Indonesia (Ahyong et al., 2020).</p> <p>Ahyong and Ng (2017) reduced Cymonomus sagamiensis Sakai, 1983 from Japan to a synonym of C. umitakae Takeda, 1981 from Japan. As a result, the genus Cymonomus is composed of 31 Indo-West Pacific, 11 West Atlantic, and 2 Northeast Atlantic and Mediterranean species, and in this paper, C. suluensis is described as the 45 th Cymonomus species and 32 nd Indo-West Pacific species.</p> <p>Ahyong (2019) revised the New Zealand and Australian cymonomid crabs and distinguished six species groups in the genus Cymonomus: 1) C. bathamae group, 2) C. curvirostris group, 3) C. delli group, 4) C. granulatus group, 5) C. karenae group, and 6) C. soela group.</p> <p>The new species described in this paper, C. suluensis, is quite distinctive in having the long rostrum and the long eyestalks directed obliquely outward as a whole, and downward along the basal half and horizontally along the distal half, with the distinct transparent cornea, and readily distinguished from all the known species. As far as the long rostrum concerned, the new species corresponds to the C. granulatus group which is composed of six species with the combination of a long rostrum that overreaches the eyestalks and well-developed outer orbital tooth. The six species referred to this group by Ahyong (2019) are C. aequilonius Dell, 1971 from New Zealand, C. alius Ahyong, 2019 from New Zealand, C. granulatus (Norman, in Wyville Thomson, 1873) from the Northeast Atlantic and the Mediterranean Sea, C. indicus Ihle, 1916 from Indonesia, C. japonicus Balss, 1922 from Japan, and C. magnirostris Tavares, 1991 from Brazil. Of them, C. granulatus and C. magnirostris from the Atlantic Ocean are closer to the new species rather than four species from the Pacific Ocean.</p> <p>The new species, C. suluensis, however, is quite characteristic in the carapace margin fringed with many spinules, and the rostrum armed with some spinules of good length at each side, and each eyestalk directed obliquely outward as a whole and downward for its basal half and horizontally for its distal half. These characters make this species distinct from all the species including the C. granulatus group and the sole representative of the seventh group in the genus Cymonomus, herein called the C. suluensis group.</p> <p>Etymology. Named after the type locality, the Sulu Sea.</p> <p>Distribution. Known only from the type locality. The bathymetric record, 2,030 m, is the deepest for any cymonomid species.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF88D647B38CFDC4FD89B8D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF8CD646B3A4FAA1FD80BB41.text	03CF87EEFF8CD646B3A4FAA1FD80BB41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicranodromia foersteri Guinot 1993	<div><p>Dicranodromia foersteri Guinot, 1993</p> <p>(Figs. 4–5)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 20 (Sibutu Passage; 05°40.9′N, 119°46.3′E – 05°43.1′N, 119°47.0′E; 460–514 m deep); otter trawl; 10 June, 1972; 1˂ (ovig.), NSMT-Cr 28966.</p> <p>Measurements. CB 33.3 mm; CL in median line, 33.7 mm; length of rostrum, 3.3 mm; length of first ambulatory leg, 62.2 mm; diameter of egg, 2.2 mm.</p> <p>Remarks. In the ovigerous female at hand, the carapace was cracked into two, with somewhat damaged posterior part (Fig. 4A), but there is no problem to depict the species characteristics. The key prepared by Guinot (1995) was used to identify the present specimen, with combination of the characters that 1) the carapace dorsal surface is smooth and not disguised by the setae of variable lengths, 2) the palm is granulated for its whole outer surface, 3) the last leg is rather long and slender. As a result, it was keyed out to D. martini Guinot, 1995 from the Philippines, or D. foersteri Guinot, 1993 from the Chesterfield Islands, New Caledonia and Vanuatu. Some differences between the two species were mentioned by Guinot (1995), viz. the antero-external tooth of the basal antennal segment is strong and long in D. martini (stout and short in D. foersteri), the infra-orbital tooth is large and bifid in D. martini (pointed and simple in D. foersteri), the epistome is armed with some sharp granules in D. martini (unarmed in D. foersteri), the anterior margin of the buccal frame is fringed with spinules in D. martini (only indistinctly dentated in D. foersteri), and the dactyli of the first two ambulatory legs are relatively long and subequal to the carpi in D. martini (comparatively short and slightly shorter than the carpi in D. foresteri). These differences are appli- cable to the ovigerous female examined, and the different length and stoutness of the ambulatory legs are the important criteria among the differences. It is recorded at present that the ambulatory dactyli are half as long as the propodi in D. foersteri, rather than two-thirds in D. martini, although Ng and Naruse (2007) found some variations in the length of the ambulatory carpi in C. martini.</p> <p>Ng and McLay (2005) described a new species, D. danielae from Balicasag Island in the Bohol Sea, 200–300 m deep, on the comparison with D. doederleini Ortmann, 1892 and D. martini Guinot, 1995. In D. danielae, however, the outer surface of the chela is smooth, the external orbital tooth is armed with spinules, the posterior margin of the epistome is spinulate, and the meri and propodi of the subdorsal legs are proportionately shorter. In D. danielae, the outer margin of the frontal lobe is straight, differing from the weakly concave margin of D. doederleini and D. martini.</p> <p>Distribution. This species has been previously recorded from Vanuatu, the Chesterfield Islands and New Caledonia, 495–660 m deep (Guinot, 1993, 1995), and the close congener, C. martini, from the Sulu and Bohol Seas, 437–930 m deep (Guinot, 1995; Ng and Naruse, 2007).</p> </div>	https://treatment.plazi.org/id/03CF87EEFF8CD646B3A4FAA1FD80BB41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF8DD64DB070F98FFD0FBD11.text	03CF87EEFF8DD64DB070F98FFD0FBD11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homolodromia hakuhoae Takeda & Ohtsuchi & Komatsu 2021	<div><p>Homolodromia hakuhoae sp. nov.</p> <p>(Figs. 6–8)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 08 (Sulu <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.03&amp;materialsCitation.latitude=2.03" title="Search Plazi for locations around (long -2.03/lat 2.03)">Sea</a>; 08°44.6′N, 119°05.4′E – 08°44.8′N, 119°06.2′E; 2,030 – 2,030 m deep); 3 m beam trawl; 25 May, 1972; ˂ (ovig.) holotype, NSMT-Cr 28967.</p> <p>Measurements. CB 12.8 mm; CL in median line, 17.3 mm; length of rostrum, 2.8 mm; length of first ambulatory leg, 67.0 mm; diameter of egg, 2.0– 2.1 mm.</p> <p>Diagnosis. Carapace, chelipeds and ambulatory legs thickly covered with short stiff setae; carapace narrow, convex dorsally, weakly widening posteriorly, with surface smooth. Front with deep U-shaped excavation, with a pair of strong, spiniform teeth directed forwards. External orbital tooth as long as, but narrower than frontal tooth, directed obliquely outward. Chelipeds slender, with weakly widening palm; fingers deeply excavated along grasping edges, fixed finger with bifid tip. Ambulatory legs long, cylindrical. Subchelae at tips of last 2 legs armed with 2 strong spines at distal part of the upper margin and 4 longer spines at truncated part of distal extension of palm.</p> <p>Description of holotype (Ovigerous female). Carapace, chelipeds and ambulatory legs uniformly and thickly covered with short, stiff setae, without long setae (Fig. 6A–B). Carapace (Fig. 6C) narrowly oblong, strongly convex dorsally, weakly widening posteriorly, widest at posterior parts of branchial regions of both sides; carapace dorsal areolation obscured by setae, but surface (after denudation) smooth, without granules or spinules, roughly divided into three parts: 1) anterior part including protogastric and hepatic regions in front of a pair of median small pits side by side at boundary between meso- and metagastric regions, 2) median part including metagastric, cardiac and mesobranchial regions, and 3) posterior part including intestinal and metabranchial regions; in dorsal view, a flattened hepatic swelling lateral to the buccal flame. Frontal teeth strong, tuberculate, tapering to sharp tip (Fig. 6C) separated by deep U-shaped sinus. Each supraorbital margin (Fig. 6C) provided with 2 granules at medially, followed by obliquely-outward directed spiniform tooth at external angle. Eyestalk (Fig. 7D) short, cornea distinct, mostly obscured by setae. Antennal flagellum (Fig. 6A) slightly longer than carapace length, each segment provided with some longish setae; basal segment armed with strong spine at distal part of outer margin.</p> <p>Third maxilliped (Fig. 7A) long, rather pediform, with quadrate ischium and merus; outer margin of merus armed with some spinules; exopod tapers, not reaching merus distal margin.</p> <p>Pleon (Fig. 7E) seven-segmented, pleomeres 1–6 and telson strongly developed; surfaces of all pleomeres and telson smooth, covered with setae, longitudinally convex along median line; telson subequal in length to pleomeres 1–6 combined.</p> <p>Chelipeds mostly subcylindrical, palm (Fig. 7C) only slightly widening distally, but not inflated at all; upper inner margin of movable finger strongly crested as a longitudinal edge, both of lower margins entire, resembling bird beak; immovable finger deeply excavated to receive movable finger, bifid at tip and toothed along inner margin (Fig. 7A–B).</p> <p>Ambulatory legs (Fig. 6A) cylindrical, remarkably long, first leg ca 3.8 times as long as carapace. Last 2 pairs subdorsal as usual, about half as long as anterior 2 pairs; subchela of each pair similar in shape and armature, with 2 strong spines at distal part of propodus upper margin, 4 longer spines at truncated part of distal extension of propodus (Fig. 8).</p> <p>Etymology. Named after the RV Hakuhō Maru of the Ocean Research Institute, University of Tokyo.</p> <p>Remarks. The genus Homolodromia was extensively studied by Guinot (1995) who distinguished five species in the genus (H. paradoxa A. Milne-Edwards, 1880 and H. monstrosa Martin, Christiansen and Trautwein, 2001 from the West Atlantic; H. robertsi Garth, 1973 from the East Pacific; H. kai Guinot, 1993 from the West and South Pacific; H. bouvieri Doflein, 1904 from the western Indian Ocean). Recently, Padate et al. (2020) described the sixth species, H. rajeevani from the eastern Arabian Sea and the southwestern Bay of Bengal. All of the six species are deep-water inhabitants, with bathymetric range, 375–914 m (H. paradoxa), 631– 814 m (H. monstrosa), 560–880 m (H. robertsi), 680–850 m (H. kai), 492–960 m (H. bouvieri), and 645–957 m (H. rajeevani).</p> <p>Padate et al. (2020) prepared the key to the six species based on Guinot (1995) and Tavares and Lemaitre (2014). Following the key, the present new species, H. hakuhoae, is keyed out as follows: 1) lPropodal thumbs of P4–P5 pseudochelae terminating in more than 2 curved distal spines. Pollex of cheliped without occlusal notch; dactylus without proximal elevation,z differing from H. rajeevani and H. bouvieri. 2) lCarapace and pereopods smooth, carpus of cheliped bearing short disto-lateral spine; anterolateral teeth oriented obliquely,z differing from H. robertsi. 3) lCarapace and appendages covered with short, stiff setae; pseudo-rostral horns separated by V-shaped gap,z differing from H. kai. 5) The remaining two species, H. paradoxa and H. monstrosa are differentiated in the characters that lMales having distinct supraorbital spine; antennal spine bifurcated. Mature females with relatively short P5, P5 meri not reaching the level of gastric pits on carapace in longitudinal position in H. paradoxa, and that lMales lacking supraorbital spine; antennal spine undivided. Mature females with relatively short P5, P5 meri the overreaching the level of gastric pits in folded position in H. monstrosa.z</p> <p>Apart from the key quoted above, the new species, H. hakuhoae, which is represented only by the female holotype, is seemingly close to H. kai in having the narrow carapace without a constriction between the hepatic and branchial margins, although as mentioned above, the carapace and appendages are covered with short stiff setae different from long soft hairs in H. kai.</p> <p>Distribution. The holotype, a female, was obtained at 2,030 m deep in the Sulu Sea, as the deepest record among species of Homolodromia.</p> </div>	https://treatment.plazi.org/id/03CF87EEFF8DD64DB070F98FFD0FBD11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF86D64DB3BAFE4FFE2BB989.text	03CF87EEFF86D64DB3BAFE4FFE2BB989.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paromolopsis boasi Wood-Mason	<div><p>Paromolopsis boasi Wood-Mason, in Wood-Mason and Alcock, 1891</p> <p>(Fig. 11D)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 20 (Sibutu Passage; 05°40.9′N, 119°46.3′E – 05°43.1′N, 119°47.0′E; 460–514 m deep); otter trawl; 10 June, 1972; 1˂ (CB 32.0 mm; CL 35.3 mm including rostrum), NSMT-Cr 28968.</p> <p>Remarks. This monotypic species of the genus Paromolopsis is rather well known as one of the representative deep-water crabs, and finely represented by Alcock (1901: Pl. 5 fig. 23). Then, this species was illustrated in color by Sakai (1976: Pl. 15 fig. 2), and photographed by Serène and Lohavanijaya (1973: Pl. 3 fig. D), Guinot and Richer de Forges (1981: Pl. 6 fig. 3), Miyake (1983: Pl. 5 fig. 4 in color), Guinot (1984: Pl. 3 fig. A), Davie and Short (1989: Fig. 1C), Ng et al. (2001: Fig. 1b in color), Ahyong et al. (2009: Fig. 82 in color), Ng and Richer de Forges (2017: Fig. 24H in color), Guinot and Richer de Forges (1995: Fig. 18f, g), Padate et al. (2020: Fig. 3k), and Richer de Forges and Ng (2020: Fig. 1E).</p> <p>Distribution. Widely distributed in the Indo-West Pacific waters from Japan to Madagascar through the western Pacific and eastern Indian Oceans; 280–1,380 m deep.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF86D64DB3BAFE4FFE2BB989	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF86D64EB273F9F9FE1EBF7B.text	03CF87EEFF86D64EB273F9F9FE1EBF7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lysirude goekei Takeda & Ohtsuchi & Komatsu 2021	<div><p>Lysirude goekei sp. nov.</p> <p>(Figs. 9–10)</p> <p>Lyreidus channeri: Griffin, 1970, p. 107, figs. 6d, i, r, 7f, 8e, pl. 2 fig. B. [Not Lyreidus channeri Wood-Mason, 1885]</p> <p>Lysirude channeri: Goeke, 1985 [1986], p. 215, fig. 6: Feldman, 1992, fig. 10: Tucker, 1998, fig. 1 (3–4); Ng et al., 2008, p. 42 (in list), fig. 17: Guinot et al., 2013, p. 289, fig. 40C–D. [Not Lysirude channeri (Wood-Mason, 1885)]</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 13 (Sulu <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.31333&amp;materialsCitation.latitude=8.343333" title="Search Plazi for locations around (long 118.31333/lat 8.343333)">Sea</a>, 08°20.8′N, 118°19.8′E – 08°20.6′N, 118°18.8′E; 730–738 m deep); 3 m beam trawl; 27 May, 1972; ˁholotype, NSMT-Cr 28969.</p> <p>Measurements. CB 16.6 mm excluding carapace anterolateral spines; CL 26.4 mm excluding rostrum; length of rostrum, 4.1 mm; length of eyestalk, 2.3 mm.</p> <p>Diagnosis. Carapace pyriform, smooth, convex dorsally sideways; rostrum, external orbital spines and anterolateral spines of carapace elongate. Pleomeres 3–4 each with medial spine. Cheliped merus with proximal spine; carpus with distal spine; palm without tooth on upper margin, with 3 triangular teeth on lower margin, proximal tooth rudimental. Carpus of first ambulatory leg with subdistal tooth on upper margin. Propodus of third ambulatory leg strongly lobate on inner margin; dactylus paddle-shaped. Fourth ambulatory leg reduced. G1 with trilobate apex.</p> <p>Description of holotype (Male). Carapace (Fig. 9A) narrowly pyriform, 1.59 times as long as broad, dorsally convex transversely, widest at anterolateral corner; anterolateral margin divergent toward anterolateral spine, with median inconspicuous hump, covered with minute granules; lateral margins of both sides subparallel in anterior half, convergent in posterior third, beaded with small granules in posterior half of lateral margin and posterior margin; posterior margin wider than frontal neck; dorsal surface seemingly smooth, but entirely covered with microscopic pits, with pair of shallow subparallel furrows along center of carapace. Rostrum (Figs. 9, 10A) elongate, triangular, ca. 1.7 times as long as broad, ca. 1.8 times as long as eyestalk. Supraorbital margin (Figs. 9–10A) U-shaped, provided with a very short, obliquely-outward directed furrow at base of external orbital spine; external orbital spine elongated, slightly longer than rostrum, ca. 2.0 times as long as eyestalk, slightly directed outward. Eyestalk (Fig. 10A) short, lanceolate, movable, with small, pyriform, pigmented cornea on lateral side of apex.</p> <p>Pleon (Fig. 9A–B) with free 6 slender pleomeres and small triangular telson; first 3 pleomeres visible in dorsal view; surfaces of all pleomeres and telson smooth, longitudinally convex along median line, furnished with short setae along lateral margins; pleomeres 3–4 each with porximally recurved, medial spine.</p> <p>Third maxilliped (Fig. 9B) elongate, narrow, with oblongly quadrate ischium and merus; ischium with transverse suture on mesial half of proximal 0.2; exopod reaching beyond ischium.</p> <p>Cheliped segments (Fig. 9) subcylindrical to compressed; merus with short spine on proximal 0.3 of dorsal surface; carpus with strong spine on distal part of dorsal surface; palm (Fig. 10B) compressed, crested on upper margin, with 3 triangular teeth on lower margin, teeth becoming larger distally, proximal tooth rudimentry; upper margin of movable finger crested, cutting edge weak, without teeth; fixed finger broad, compressed, cutting edge irregularly crenulate.</p> <p>Ambulatory legs (Fig. 9) different in shape from each other. First leg rather compressed; merus with a median line of scattered granules and very short setae on lower surface; carpus crested on upper margin with distal denticle; propodus broadened, crested on both margins; dactylus lanceolate. Second leg longest, subcylindrical, weakly crested on both margins of propodus, with lanceolate dactylus. Third leg short, rather compressed; ischium with a small distal tooth; propodus (Fig. 10C) short, crested on outer margin, with strongly expanded, ovate lobe on inner margin, lobe entirely fringed with long setae; dactylus foliaceous, paddle-shaped, entirely fringed with long setae along expanded inner margin. Fourth leg reduced, fringed with short setae on propodus and dactylus.</p> <p>G1 (Fig. 10D–E) stout, trilobate at apex; distolateral lobe of apex rounded, with aperture; mesial lobe tongue-shaped; medial lobe small, triangular. G2 (Fig. 10F) stout, slightly shorter than G1, acuminate at tip,</p> <p>Etymology. Named after Dr. Gary D. Goeke for his great contributions to the taxonomy on raninid crabs.</p> <p>Remarks. Currently, the genus Lysirude comprises three recent species, viz., L. nitidus (A. Milne-Edwards, 1880) from the western Atlantic, L. channeri (Wood-Mason, 1885) (with L. gracilis Wood-Mason, 1888 as a synonym) from the Indo-West Pacific and L. griffini Goeke, 1985 from the Philippines. However, the recent redescription of L. channeri on the basis of the holotype and recently collected specimens from the Bay of Bengal by Rozario et al. (2017) and the present study reveal that the records of L. channeri from the West Pacific are due to mis- identification, representing a new species distinct from the Indian Ocean form. In this paper we described L. goekei sp. nov. based on the Sulu Sea specimen, with photographs and line drawings. The records of L. channeri listed in the above synonymy list (p. 77) are referrable to the new species.</p> <p>Lysirude goekei is readily distinguished from L. channeri by the following features: 1) the rostrum and external orbital spine are elongate and ca. 1.7 and 2.0 times as long as the eyestalk, respectively (vs. slightly longer than the eyestalk in L. channeri); 2) the marginal spine between the external orbital spine and the anterolateral spine of the carapace is represented by a low swelling (vs. strong in L. channeri); 3) the upper margin of the cheliped palm has no tooth (vs. with a subterminal tooth in L. channeri); 4) proximal-most tooth of the lower margin of the cheliped palm is rudimentary, tubercle-like (vs. distinct and triangular in L. channeri). The pigmentation of cornea may also differentiate both species (unpigmented in L. channeri).</p> <p>Griffin (1970) considered that the lack of the intercalated spine of the carapace in the specimen from the South China Sea may be due to breakage during life, but the lack of this spine is constant as observed among the specimens from the Philippines (Goeke, 1985 [1986]; Feldman, 1992; Tucker, 1998; Ng et al., 2008; Guinot et al., 2013; this study). Goeke (1985 [1986]: fig. 6B) showed a variation in the presence of the intercalated spine on the carapace in the Philippine juvenile specimen, and also mentioned that the presence of the intercalated spine is also rarely seen among juveniles in the Atlantic species, L. nitidus.</p> <p>Distribution. South China Sea and Philippines, 366–820 m deep.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF86D64EB273F9F9FE1EBF7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF85D64EB3AEFC0AFC4EBE61.text	03CF87EEFF85D64EB3AEFC0AFC4EBE61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ethusa sexdentata (Stimpson 1858)	<div><p>Ethusa sexdentata (Stimpson, 1858)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 12 (Sulu <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.14167&amp;materialsCitation.latitude=8.3116665" title="Search Plazi for locations around (long 118.14167/lat 8.3116665)">Sea</a>; 08°19.0′N, 118°09.1′E – 08°18.7′N, 118°08.5′E; 495–500 m deep); 3 m beam trawl; 26 May, 1972; 1 ˁ (CB 5.5 mm; CL 6.0 mm), NSMT-Cr 28970.</p> <p>Remarks. The male specimen at hand agrees well with the males recorded by Stimpson (1858, 1907) in having the sharp external orbital tooth directed obliquely outward, and differs from the females illustrated by Sakai (1937, 1965, 1976), in which the external orbital tooth is stout and not tuberculated. In the present male, four frontal teeth are spiniform, weakly directed obliquely outward, and similar to, but shorter than the external orbital tooth; the tip of the frontal inner tooth slightly exceeds the tip of the frontal outer tooth; the tip of the external orbital tooth does not reach the basal part of the frontal teeth. The lateral margin of the carapace is directed obliquely outward, with the laterally convex branchial margin. The distal part of the whip-like G2 is sticking out from the subterminal part of the G1.</p> <p>In the revision of the subfamily Ethusinae, Castro (2005) extensively studied all the Indo-West Pacific species of Ethusa including E. sexdentata. According to the monograph, E. sexdentata reported by Chen (1986) was misidentified and represents a new species, named by Castro (2005) as E. abbreviata.</p> <p>Distribution. Known from Japan, Taiwan, the East and South China Seas, and the Philippines; 30–ca. 500 m deep.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF85D64EB3AEFC0AFC4EBE61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF85D650B069FD11FEB6BEC9.text	03CF87EEFF85D650B069FD11FEB6BEC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Urashima pustuloides (Sakai 1961)	<div><p>Urashima pustuloides (Sakai, 1961)</p> <p>(Fig. 11A–C)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 11 (Sulu <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.975&amp;materialsCitation.latitude=8.196667" title="Search Plazi for locations around (long 117.975/lat 8.196667)">Sea</a>; 08°12.7′N, 117°59.6′E – 08°11.8′N, 117°58.5′E; 285–306 m deep); 3 m beam trawl; 26 May, 1972; 1 ˁ (CB 42.4 mm including lateral teeth; CL 35.7 mm in median line), NSMT-Cr 28971.</p> <p>Remarks. The male at hand is without doubt identified as Urashima pustuloies (Sakai, 1961) which was described in detail by Sakai (1961 and 1976, as Randallia), diagnosed briefly by Chen (1989) and Tan (1996) as Randallia, and studied deeply and designated as the type species of the new genus Urashima by Galil (2003). This species is comparatively large in the family Leucosiidae, having a nearly diamond-shaped carapace. The remarkable characters are noted and shown in Fig. 11A–C. The dorsal surface is separated into regions by the linear furrows and covered with the dispersed obtuse tubercles of variable sizes. The epibranchial tubercle at the anterolateral and posterolateral junction protrudes laterally and is bifid at the tip. The intestinal region is thick, more or less tuberculate as a whole, and tipped with a small tubercle. The male pleon narrows strongly toward the telson, the tip of which is distinctly bifid; the fused pleomere is armed with a sharp erect spine at the median part close to the telson. The color in life of an immature female was given by Richer de Forges and Ng (2020: Fig. 5H).</p> <p>Another congener is U. lamellidentata (Wood-Mason, 1892) known from the Andaman and Maldive archipelagoes and finely illustrated by Kumar et al. (2013), being characteristic in having the lamellate crest on the carapace anterolateral margin and also lamellate, rounded teeth on the carapace posterior margin.</p> <p>Distribution. From Japan to Australia through East and Southeast Asian waters (Galil and Ng, 2015); 85–839 m deep. For records from the Philippines, see Chen (1989), Tan (1996), Galil (2003), Komatsu et al. (2005), and Galil and Ng (2007).</p></div> 	https://treatment.plazi.org/id/03CF87EEFF85D650B069FD11FEB6BEC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF9BD650B3BEFCB9FCF2BFC4.text	03CF87EEFF9BD650B3BEFCB9FCF2BFC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyrtomaia horrida Rathbun 1916	<div><p>Cyrtomaia horrida Rathbun 1916</p> <p>(Figs. 12A–C)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 12 (Sulu <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.14167&amp;materialsCitation.latitude=8.3116665" title="Search Plazi for locations around (long 118.14167/lat 8.3116665)">Sea</a>; 08°19.0′N, 118°09.1′E – 08°18.7′N, 118°08.5′E; 495–500 m deep); 3 m beam trawl; 26 May, 1972; 1 ˁ(Fig. 12A) (CB 15.2 mm excluding branchial spines; CL 15.0 mm excluding pseudorostral spines), NSMT-Cr 28972; 1 ˁ (Fig. 12B–C) (CB 9.0 mm; CL 8.3 mm), NSMT-Cr 28973.</p> <p>Remarks. Among the known species of the genus Cyrtomaia, the taxonomic validity and status of C. horrida pilosa Ihle and Ihle-Landenberg, 1931 known only by the holotype from the Kei Islands, has been long discussed. It was considered as synonymous with C. horrida by Griffin (1976), but Guinot and Richer de Forges (1982) examined the holotype and tentatively elevated this taxon to the full species. Later, Guinot and Richer de Forges (1986) and Griffin and Tranter (1986) also considered it as a synonym of C. horrida. Recently, however, Richer de Forges and Ng (2007), who examined a long series of the specimens from the Bohol Sea, confirmed that the females of C. horrida are somewhat more pilose than the males especially in the small specimens, and decidedly synonymized C. horrida pilosa with C. horrida. At present, the genus Cyrtomaia is comprised of 29 species from the Indo-Pacific waters (Richer de Forges and Ng, 2007, 2009a; Ng et al., 2008).</p> <p>In the specimens at hand, all the chelipeds and ambulatory legs are missing, but the spinulation of the carapace surface agrees well with the description and figures by Guinot and Richer de Forges (1982: Fig. 24, as? Cyrtomaia pilosa), though the carapace (Fig. 12A–B) is not pilose as in the holotype. They also agree essentially with the description by Guinot and Richer de Forges (1982), though the G1 may be not fully developed and rather similar to that of C. suhmi Miers, 1886 in general structure. According to Griffin and Tranter (1986), C. suhmi, C. smithii Rathbun, 1893 and C. owstoni Terazaki, 1903 have a slen- der simple type of the G1.</p> <p>Distribution. West Pacific from Japan to the Solomon Islands through the Philippines, the South China Sea and Indonesian waters; 24–787 m deep.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF9BD650B3BEFCB9FCF2BFC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF9BD653B1A5FC0BFE17BE0C.text	03CF87EEFF9BD653B1A5FC0BFE17BE0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyrtomaia largoi Richer de Forges and Ng 2007	<div><p>Cyrtomaia largoi Richer de Forges and Ng, 2007</p> <p>(Fig. 12D–E)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 20 (Sibutu Passage; 05°40.9′N, 119°46.3′E – 05°43.1′N, 119°47.0′E; 460–514 m deep); otter trawl; 10 June, 1972; 1 ˁ (Fig. 12E) (CB 7.7 mm excluding branchial spines; CL 8.3 mm excluding pseudorostral spines), NSMT-Cr 28991; 1 ˂ (Fig. 12D) (CB 22.7 mm; CL 20.9 mm), NSMT-Cr 28974.</p> <p>Remarks. The specimens at hand agree well with the original description. This species is readily distinguished from congeners by having subparallel pseudorostral spines. In the smaller male specimen (Fig. 12E), the pseudorostral spines are proportionately shorter and not subparallel, and also the protogastric spines are slightly divergent as noted by Richer de Forges and Ng (2007).</p> <p>According to Richer de Forges and Ng (2007), C. suhmi Miers, 1886, C. curviceros Bouvier, 1915 (at present known as a junior subjective synonym of C. suhmi), C. maccullochi Rathbun, 1918 and this species form a group commonly lacking the intermediate ocular spines and preocular spines. However, our specimens, as well as the holotype and also the holotype of C. suhmi, have very small but distinct intermediate ocular spine (Fig. 12D in the present study; Guinot and Richer de Forges 1982, fig. 11A; Richer de Forges and Ng, 2007, fig. 4B–D, 5A).</p> <p>Distribution. Bohol Sea (type locality) and Sulu Sea, central Philippines; Taiwan (Richer de Forges et al., 2009), South China Sea (Lee et al., 2017); 437–443 m deep.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF9BD653B1A5FC0BFE17BE0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF98D653B3BAFD11FB8DBDA4.text	03CF87EEFF98D653B3BAFD11FB8DBDA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Platymaia bartschi Rathbun 1916	<div><p>Platymaia bartschi Rathbun, 1916</p> <p>(Fig. 13)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 20 (Sibutu Passage; 05°40.9′N, 119°46.3′E – 05°43.1′N, 119°47.0′E; 460–514 m deep); otter trawl; 10 June, 1972; 1 young ˁ (CB 36.1 mm; CL 34.9 mm excluding rostral spine, soft shell), NSMT-Cr 28975.</p> <p>Remarks. In addition to the soft shell male recorded above, the right half of the carapace and left cheliped are in the same vial. Considering the size of the half carapace with 73.7 mm in CL and the shape of the slender cheliped, the specimen is fully developed female. Although the rostral tooth is unfortunately broken off, the specimen is safely identified as Platymaia bartschi. In the soft shell male (Fig. 13), the rostral tooth is warped for its distal half, but it is clear that the main rostrum is slender and much longer than the lateral rostral teeth.</p> <p>Griffin (1976) distinguished Platymaia bartschi from P. wyvillethomsoni Miers, 1886 based mainly on the Philippine specimens; in P. bartschi the inter-antennular spine is almost three times as long as the rostral spines (about twice as long as in P. wyvillethomsoni). Griffin (1976) and Griffin and Tranter (1986) also reported develop- mental variation, namely that in juveniles the carapace bears numerous spines including six lateral branchial spines, two on each protogastric, mesogastric, cardiac, epibranchial and mesobranchial regions. Guinot and Richer de Forges (1986) finely illustrated the front-orbital details and had little doubt about the occurrence of this species in Japan.</p> <p>Distribution. Philippines (Rathbun, 1916; Griffin, 1976; Guinot and Richer de Forges, 1986), South China Sea (Griffin and Tranter, 1986), Taiwan (Griffin, 1976), and Japan (Sakai, 1965b, 1976), 185–592 m deep.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF98D653B3BAFD11FB8DBDA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF98D653B06EFDAAFBFABB41.text	03CF87EEFF98D653B06EFDAAFBFABB41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Platymaia remifera Rathbun 1916	<div><p>Platymaia remifera Rathbun, 1916</p> <p>(Fig. 12F)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 11 (Sulu <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.975&amp;materialsCitation.latitude=8.196667" title="Search Plazi for locations around (long 117.975/lat 8.196667)">Sea</a>; 08°12.7′N, 117°59.6′E – 08°11.8′N, 117°58.5′E; 285–306 m deep); 3 m beam trawl; 26 May, 1972; 1 ˁ (CB 29.4 mm; CL 27.4 mm including rostral spine), 1˂ (ovig.) (CB 37.1 mm; CL 35.6 mm), NSMTCr 28976; 1 ˂ (Fig. 10F) (CB 31.6 mm; CL 28.7 mm), NSMT-Cr 28977.</p> <p>RV Hakuhō Maru KH-72-1 cruise, sta. 12 (Sulu <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.14167&amp;materialsCitation.latitude=8.3116665" title="Search Plazi for locations around (long 118.14167/lat 8.3116665)">Sea</a>; 08°19.0′N, 118°09.1′E – 08°18.7′N, 118°08.5′E; 495–500 m deep); 3 m beam trawl; 26 May, 1972; 6 ˁˁ (CB 9.2 mm; CL 8.4 mm – CB 21.8 mm; CL 20.1 mm), 6˂˂ (CB 9.9 mm; CL 9.3 mm – CB 21.5 mm; CL 19.3 mm), NSMTCr 28992.</p> <p>Remarks. The genus Platymaia is comprised of 10 species from Indo-West Pacific (Griffin and Tranter, 1986; Ng et al., 2008). We have many specimens of various sizes (CB 9.2–37.1 mm), which agree well with the description by Guinot and Richer de Forges (1986).</p> <p>Lee et al. (2017) recently recorded P. aff. remifera from the South China Sea, noting that the interior border of the branchial region is armed with three strong spines in the South China Sea specimens and five weak spines in the Taiwanese specimens. Based on our specimens, however, this character seems variable. There is one strong spine followed by one to four spines of variable distinctness.</p> <p>Distribution. Philippines (Guinot and Richer de Forges, 1986; this study), Taiwan (Ng and Huang, 1997), and South China Sea (Serène and Lohavanijaya, 1973); 100–700 m deep.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF98D653B06EFDAAFBFABB41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF9ED654B3BBFBDFFE3FBDDB.text	03CF87EEFF9ED654B3BBFBDFFE3FBDDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleistacantha oryx Ortmann 1893	<div><p>Pleistacantha oryx Ortmann, 1893</p> <p>(Fig. 14A–B)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 11 (Sulu <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.975&amp;materialsCitation.latitude=8.196667" title="Search Plazi for locations around (long 117.975/lat 8.196667)">Sea</a>; 08°12.7′N, 117°59.6′E – 08°11.8′N, 117°58.5′E; 285–306 m deep); 3 m beam trawl; 26 May, 1972; 1 ˁ (CB 13.7 mm excluding branchial spinules; CL 18.3 mm excluding pseudorostral spines), NSMT-Cr 28978.</p> <p>RV Hakuhō Maru KH-72-1 cruise, sta. 20 (Sibutu Passage; 05°40.9′N, 119°46.3′E – 05°43.1′N, 119°47.0′E; 460–514 m deep); otter trawl; 10 June, 1972; 2 ˂˂ (CB 9.3 mm; CL 6.4 mm; broken, soft shell), NSMT-Cr 28993.</p> <p>Remarks. Ng et al. (2008) listed 14 species in the genus Pleistacantha. Recently, two synonymous genera of Pleistacantha, Pleisticanthoides Yokoya, 1933 and Parapleisticantha Yokoya, 1933, were reappraised as valid (Ng and Richer de Forges, 2012; Richer de Forges et al., 2013). In addition, Pleistacantha kannu Ng, Ravinesh and Ravichandran, 2017 was described from the Indian Ocean, and Ahyong et al. (2019) recognized Echinoplax rubida Alcock, 1895 as a valid species of Pleistacantha. Consequently, the genus Pleistacantha at present consists of 12 species from the Indo-West Pacific region.</p> <p>Pleistacantha oryx was confused with P. maxima Ahyong and Lee, 2006, but distinguished by the pseudorostral spines which are broadly convex in lateral view, the less spinulate carapace surface, and the different G1 structure (Ahyong and Lee, 2006). Otherwise, the smaller size at maturity can be a help to distinguish P. oryx from P. maxima.</p> <p>Distribution. Most of the previous records of Pleistacantha oryx from Japan, the East China Sea, the Philippines and Australia were referred to those of P. maxima by Ahyong and Lee (2006), and therefore, the exact distributional range of P. oryx is uncertain at present. Although the distributional ranges of both species may widely overlap each other, the reliable records of P. oryx are only those by Ortmann (1893: Sagami Bay, 90–180 m deep) and Ahyong and Lee (2006: Sagami Sea, 540 m deep, and off Kaohsiung County, southern Taiwan).</p> </div>	https://treatment.plazi.org/id/03CF87EEFF9ED654B3BBFBDFFE3FBDDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF9FD657B3CEFE4FFCBCBAD9.text	03CF87EEFF9FD657B3CEFE4FFCBCBAD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleisticanthoides simplex (Rathbun 1932)	<div><p>Pleisticanthoides simplex (Rathbun, 1932)</p> <p>(Fig. 14C)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 20 (Sibutu Passage; 05°40.9′N, 119°46.3′E – 05°43.1′N, 119°47.0′E; 460–514 m deep); otter trawl; 10 June, 1972; 1 ˁ (CB 5.9 mm; CL 8.2 mm excluding pseudorostral spines), NSMT-Cr 28979.</p> <p>Description of a male from the Sibutu Passage. Carapace pyriform, dorsal surface covered with long hooked setae, posterior surface sparsely covered with granules, gastric, hepatic, anterior cardiac regions smooth; regions well-defined: gastric region slightly elevated; branchial region inflated, separated from cardiac, gastric regions by deep groove. Front with 2 sharp, short, diverging pseudorostral spines; pseudorostral spine basally with 4 sharp accessory spines (1 on ventolateral surface, 3 directed anteriorly); supraocular eave fringed with 5 spines (anterior 3 long, posterior 2 short); postocular spine strong, with small accessory spines anteriorly, dorsally, posteriorly; interocular spine strong, directed dorsally, proximally with small spine; subhepatic region weakly inflated, with long spine bearing some accessory spines, followed by 4 long spines bearing stiff setae (shorter than supraocular tooth). Eyestalk relatively long, slender, subdistally with small spine ventrally; cornea rounded, diameter greater than that of peduncle. Inter-antennular spine (=true rostrum) pointed downwards, medially divided by U-shaped concavity to form 2 spines. Antennae almost twice as long as pseudorostral spines; basal antennal article slender, fused with carapace, with long, sharp spine on distolateral angle; urinary article with large green gland opening, surrounded by raised rounded margin, outer border with strong flattened triangular tooth. Epistome smooth; buccal frame quadrangular. Third maxilliped with segments flattened, covered with long setae; outer border of carpus crenulated.</p> <p>Chelipedal merus moderately inflated, dor- somesial margin armed with long, sharp spines. Carpus globular. Chela inflated, with 6 spines on dorsal surface, row of 7 strong spines on outer surface, row of 4 spines on inner surface, immovable finger, dactylus smooth on surface.</p> <p>Ambulatory legs conspicuously long, length decreasing posteriorly. Meri and carpi with row of long hooked setae and/or long stiff setae. Propodi and dactyli with row of very long, relatively rigid setae on flexor surface.</p> <p>Male pleon with 6 free pleomeres and telson, covered with subacute granules and long stiff setae; pleomeres 4–5 with large, elongated median tubercle on distal margin; pleomere 6 with distolateral margins not expanded; telson with rounded distal margin. G1 curved in entire length, distal third dorsolaterally flattened.</p> <p>Remarks. The genus Pleisticanthoides was established by Yokoya (1933) for this species by monotypy. This genus had once synonymized with Pleistacantha Miers, 1879 by Sakai (1938), but recently, Ng and Richer de Forges (2012) recognized Pleisticanthoides as a distinct genus from Pleistacantha and added two species from the Philippines, Papua New Guinea, and Vanuatu to this genus. The genus Pleistacanhoides is distinguished from Pleistacantha by 1) the carapace dorsal surface with few spinules or setae (versus with numerous spines in Pleistacantha), 2) the margins of the meri and propodi of the second to fourth ambulatory legs lined with only stiff setae (versus lined with hard spines, spinules and stiff setae in Pleistacantha), and 3) the G1 evenly curved, with the distal third dorsoventrally flattened, and without trace of a subdistal process (versus the relatively slender and straight, distal third is evenly cylindrical and there is a recurved process just before the tip in Pleistacantha).</p> <p>The present specimens generally agree with the description and illustrations given by Yokoya (1933) and Ng and Richer de Forges (2012), and keyed out correctly to l Pleistacantha simplex z following Ahyong et al. (2005). The distinguishing characters of Pleisticanthoides simplex from the other two congeners were discussed by Ng and Richer de Forges (2012), and recently, Takeda and Komatsu (2020) recorded a pair of specimens from Amami-Oshima Island in the northern Ryukyu Islands, ca. 200 m deep.</p> <p>Distribution. This species is known from Japan (from Sagami Bay to Amami-Oshima Island) (Sakai, 1976; Ng and Richer de Forges, 2012; Takeda and Komatsu, 2020), and the Sulu Sea and Indonesian waters (Griffin and Tranter, 1986). The known bathymetric range is from 60 to 540 m.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF9FD657B3CEFE4FFCBCBAD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF9DD656B3AFFEFBFC6AB9CC.text	03CF87EEFF9DD656B3AFFEFBFC6AB9CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laubierinia nodosa (Rathbun 1916)	<div><p>Laubierinia nodosa (Rathbun, 1916)</p> <p>(Fig. 15E)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 20 (Sibutu Passage; 05°40.9′N, 119°46.3′E – 05°43.1′N, 119°47.0′E; 460–514 m deep); otter trawl; 10 June, 1972; 1 ˁ (CB 11.0 mm including branchial tubercles; CL 16.5 mm excluding pseudorostral spines), 1˂ (ovig.) (CB 15.0 mm; CL 21.4 mm), NSMT-Cr 28980.</p> <p>Remarks. This species was originally described in Sphenocarcinus by Rathbun (1916), but Garth (1958) regarded the genera Sphenocarcinus A. Milne-Edwards, 1878 and Oxypleurodon Miers, 1886 as synonymous with the genus Rochinia A. Milne-Edwards, 1875. However, Richer de Forges (1995) transferred this species, Rochinia nodosa, to Oxypleurodon reappraised by Tavares (1991b). Richer de Forges and Ng (2009b) further transferred this species, Oxypleurodon nodosus, to their new genus Laubierinia. The genus Laubierinia, the type species of which is Rochinia carinata Griffin and Tranter, 1986, was defined as distinct from the closest genus Rochinia, in having the rounded carapace (versus pyriform in Rochinia), the flattened pseudorostral spines (versus long slender spines in Rochinia), and the large tubercles on the hepatic and branchial regions (versus large spines in Rochinia).</p> <p>As a result through the current studies, this species was assigned to the genus Laubierinia as L. nodosa (Rathbun, 1916), together with the type species, L. carinata (Griffin and Tranter, 1986). Recently, Pugettia globulifera Wood-Mason, in Wood-Mason and Alcock, 1891, which has been recorded by Alcock (1895) and Alcock and Anderson (1895) as Scyramathia globulifera, and by Griffin and Tranter (1986) as Rochinia globulifera, was transferred to Laubierinia by Lee et al. (2021). Laubierinia nodosa is characteristic among the three congeners in having the broad blunt tubercles instead of well-developed islets in the midline of the mesogastric, cardiac and intestinal regions, the absence of a small central mesogastric tubercle, the large nodular projection instead of strong epibranchial spine, and the non-carinate extensor surfaces of the chelipeds and ambulatory legs.</p> <p>The present specimens show good matches with Rathbun (1916, as Sphenocarcinus), Griffin (1976, as Sphenocarcinus), and Richer de Forges and Ng (2009b).</p> <p>Scyra tuberculata Yokoya, 1933, which is known only by the original description based on the specimens from the south of Satsuma, Kagoshima-ken (= -Prefecture), 133 m deep, and near the Koshiki Islands, 300 m deep, was remarked to be synonymous with Sphenocarcinus nodosa Rathbun, 1916 (Sakai, 1976, as Rochinia; Miyake, 1983, as Rochinia), though it is not referenced in recent studies (Ng et al., 2008; Richer de Forges and Ng, 2009b). The present specimens also agree well with the original description (Yokoya, 1933: 156, fig. 55), but could not be compared with the type specimens. With kind help of Dr. F. Takeshita, the type specimens of S. tuberculata were determined to be not located in the Kitakyushu Museum of Natural History and Human History (KMNH), where Yokoya`s extant specimens are deposited.</p> <p>Distribution. Japan (from Mikawa Bay at the Pacific coast of central Honshu to Shimo-Koshiki Island in the west of Kyushu), Philippines, Indonesia, northwestern Australia, Papua New Guinea, Solomon Island, Vanuatu, and New Caledonia; 135–905 m deep.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF9DD656B3AFFEFBFC6AB9CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF9DD658B075FA73FC34B8D1.text	03CF87EEFF9DD658B075FA73FC34B8D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oxypleurodon Miers 1886	<div><p>Genus Oxypleurodon Miers, 1886</p> <p>Oxypleurodon sphenocarcinoides (Rathbun, 1916)</p> <p>(Fig. 15F)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 11 (Sulu <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.975&amp;materialsCitation.latitude=8.196667" title="Search Plazi for locations around (long 117.975/lat 8.196667)">Sea</a>; 08°12.7′N, 117°59.6′E – 08°11.8′N, 117°58.5′E; 285–306 m deep); 3 m beam trawl; 26 May, 1972; 1 ˁ (CB 7.4 mm excluding branchial tubercles; CL 11.8 mm excluding pseudorostral spines), NSMTCr 28981.</p> <p>Remarks. This species was originally described from the Philippines as Chorilia sphenocarcinoides, but transferred to the genus Sphaerocarcinus by Griffin (1976), to the genus Rochinia by Griffin and Tranter (1986), and then, to the present genus, Oxypleurodon, by Tavares (1991b). The specimen examined agrees well with the original description (Rathbun, 1916), and the later accounts (Griffin, 1976; Richer de Forges and Ng, 2009b).</p> <p>Distribution. Only known from the Philippines at the depths of 200– 300 m.</p> <p>Oxypleurodon wilsoni Richer de Forges and Poore, 2008 (Figs. 15A–D, 16)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 20 (Sibutu Passage; 05°40.9′N, 119°46.3′E – 05°43.1′N, 119°47.0′E; 460–514 m deep); otter trawl; 10 June, 1972; 1 ˁ(Fig. 15A) (CB 10.6 mm excluding branchial tubercles; CL 16.9 mm excluding pseudorostral spines), NSMT-Cr 28982; 1 ˂ (Figs. 15B, D) (CB 6.0 mm; CL 10.1 mm), NSMT-Cr 28983; 1 ˂ (Fig. 15C) (CB 8.2 mm; CL 14.0 mm), NSMT-Cr 28984; 1 ˂ (ovig.) (CB 9.6 mm; CL 15.5 mm), NSMT-Cr 28985.</p> <p>Remarks. Our specimens generally agree with the original description, but the supraorbital plates are more strongly thickened as shown in Fig. 15A, and in our smallest specimen (Fig. 15B, D), the dorsal plates are not fully developed, and look more sparsely distributed.</p> <p>The present species closely resembles O. luzonicum Rathbun, 1916 distributed in the West Pacific from the Kii Peninsula in the Pacific coast of central Japan to northwestern Australia (Richer de Forges and Poore, 2008; Marumura and Takeda, 2012). According to Richer de Forges and Poore (2008), O. wilsoni is distinguished from O. luzonicum by 1) the anteriorly sharpened supraocular plate, 2) the lozengeshaped mesogastric plate, 3) the small, round and medially elevated cardiac plate, 4) the oblong, externally pointing epibranchial plates, and 5) the presence of a small tubercle between the mesogastric and hepatic spines. In addition, the following characters can be also helpful to differentiate the two species: the longer pseudorostral spines, and absence of the subhepatic plate and a tubercle among the closely gathered hepatic, subbranchial and anterior epibranchial plates. The G1 (Fig. 16) also differs from the G1 drawings of O. luzonicum represented by Guinot and Richer de Forges (1985: Fig. 21C–D).</p> <p>Distribution. Western Australia and the Sulu Sea; 329–514 m deep.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF9DD658B075FA73FC34B8D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
03CF87EEFF93D65BB199FAA1FD87BEB1.text	03CF87EEFF93D65BB199FAA1FD87BEB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoplax surugensis (Rathbun 1932)	<div><p>Pycnoplax surugensis (Rathbun, 1932)</p> <p>Material examined. RV Hakuhō Maru KH-72-1 cruise, sta. 20 (Sibutu Passage; 05°40.9′N, 119°46.3′E – 05°43.1′N, 119°47.0′E; 460–514 m deep); otter trawl; 10 June, 1972; 1˂ (CB 19.2 mm including second anterolateral teeth; CL 17.3 mm), NSMT-Cr 28986.</p> <p>Remarks. This species has been placed in Carcinoplax since the original description in 1932, and wholly figured in some monographic works such as Guinot (1989), Sakai (1976), Chen (1984a, b), and Dai and Yang (1991) together with several taxonomic records. In the monographic work of the family Goneplacidae, however, Castro (2007) designated this species as the type species of the new genus Pycnoplax, and referred C. bispinosa Rathbun, 1914, C. meridionalis Rathbun, 1923, C. vioctoriensis Rathbun, 1923 and P. latifolia Castro, 2007 to the new genus. Some characters important to distinguish P. surugensis from the congeners are that the external orbital angle is triangular with an obtuse tip, the carapace first anterolateral tooth is narrow, sharp and directed forward, and the second (last) anterolateral tooth is markedly strong, sharply tuberculate and directed obliquely forward.</p> <p>Distribution. West to South Pacific (Japan, East China Sea, Taiwan, Philippines, Indonesia, and New Caledonia); 65–496 m deep.</p></div> 	https://treatment.plazi.org/id/03CF87EEFF93D65BB199FAA1FD87BEB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Masatsune;Ohtsuchi, Naoya;Komatsu, Hironori	Takeda, Masatsune, Ohtsuchi, Naoya, Komatsu, Hironori (2021): Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2): 65-97, DOI: 10.50826/bnmnszool.47.2-65
