taxonID	type	description	language	source
03CF3761FFFD762FFF605282FAA0FACD.taxon	diagnosis	Diagnosis. This genus is distinguished from the other elaterid genera by the deep antennal grooves between pronotum and hypomeron and the deeply grooved pronotosternal sutures receiving pro-tarsi. Included species. Seven species: S. bakeri Fleutiaux, 1916 (Philippines: Luzon), S. guttatus Fleutiaux, 1919 (Singapore), S. hisamatsui Nakane, 1987 (Japan: Ogasawara Islands), S. japonicus Nakane and Hisamatsu, 1991 (Japan: Honshu), S. lambirensis Arimoto, sp. nov. (Malaysia: Borneo), S. miriensis Arimoto, sp. nov. (Malaysia: Borneo), S. williamsi Van Zwaluwenberg, 1941 (New Caledonia).	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFFD762FFF605282FAA0FACD.taxon	discussion	Remarks. Nakane (1987 b) reported a specimen from Honshu, Japan as Subprotelater sp. Nakane (1991) described the species as new, S. japonicus, with the authority as “ Nakane et Hisamatsu ”. Lawrence et al. (2000 onwards) showed a species from Australia without assigning it a species name. Subprotelater hisamatsui was proposed as a junior synonym of S. bakeri based on a comparison of a dorsalview photograph of the holotype of S. bakeri (Suzuki, 2003). However, a detailed direct comparison between these species has not been conducted. Suzuki (2022, 2024) adhered to this suggestion and treated species from the Ogasawara Islands, Japan, as S. bakeri. Makihara and Ôhira (2006) examined a specimen from Babeldaob Island, Palau, but did not assign it to a specific species. Suzuki (2024) subsequently treated this specimen as S. bakeri, although it has not been re-examined.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFFD762FFF605282FAA0FACD.taxon	biology_ecology	Ecology. The two new species, Subprotelater lambirensis and Subprotelater miriensis, were collected in primary lowland dipterocarp forest using FITs at heights of 30 – 50 m and ultraviolet light traps at 17 m and 35 m, respectively (this study). Subprotelater hisamatsui was collected using Malaise traps (Suzuki, 2003), black light traps (Makihara and Ôhira, 2006), and sticky traps placed on branches of trees, particularly Schima mertensiana (Suzuki, 2024). Subprotelater japonicus has been collected in various settings, for example, on rotting wood with bracket fungi likely belonging to the family Polyporaceae (Nakane, 1987 b) and on the decaying wood of Quercus gilva (Arimoto, 2019), and using different methods such as FITs (Watanabe, 2008) and vegetation beating (Takahashi, 2010; Noto, 2024). This species was found from the lowland forest to natural forest consisting mainly of beech and mizunara oak around 800 m (Watanabe, 2008; Takahashi, 2010). The northernmost distribution of this species, and of the genus, is approximately 39.6 ° N, 141.1 ° E (Takahashi, 2010). Subprotelater williamsi was collected while being on the bark of the weeping paperbark tree (Melaleuca leucodendron) in lowland areas (Van Zwaluwenberg, 1941).	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFFC762AFF6052CAFE3BFD6C.taxon	description	(Figures 1 – 3)	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFFC762AFF6052CAFE3BFD6C.taxon	materials_examined	Type material. Holotype. Male (AA 2737), Malaysia, Sarawak, Miri, Lambir Hills National Park, 22 IX 2010, Keiko Kishimoto-Yamada and Takao Itioka leg., by FIT in the canopy.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFFC762AFF6052CAFE3BFD6C.taxon	diagnosis	Male. Diagnosis. This species is characterized by the following features: punctures small; interspaces between punctures generally larger than puncture diameter; body black; elytron with four sub-circular yellow spots; antennomeres I and II dark red; antennomeres III – IX black; legs dark red but coxae and femurs black tinged with red; setae yellow brown; pronotum 1.2 × longer than wide; prosternal process 1.8 × longer than procoxal cavity length in ventral view; mesal posterior angle of hypomeron strongly protruding; posterior part of scutellar shield not visible in lateral view; elytron 4.8 × longer than wide, 2.0 × longer than pronotal length; metepisternum narrower than width of elytral epipleuron; basal struts 0.15 × total length of median lobe; apices of parameres roundly truncate.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFFC762AFF6052CAFE3BFD6C.taxon	description	Measurements. BL: 4.52, BW: 1.25, MAE: 0.78, MBE: 0.53, OI: 146, PL: 1.48, PML: 1.31, PW: 1.23, PAW: 0.79, PLI: 120, PWI: 156, EL: 2.92, EW: 0.61, EI: 479, BI: 198. Description. Body (Fig. 1) elongate; surface generally smooth, shining, but rough on depression of hypomeron for reception of pro-femur and tibia; punctures small; depressions of hypomeron for reception of proleg and of mesepimeron and metasternum for reception of midleg without punctures; interspaces between punctures generally larger than puncture diameter. Color. Body black. Elytron with four sub-circular yellow spots, of which one spot medially on humerus, two spots side-by-side on anterior 1 / 3, and one spot medially on posterior 1 / 3. Pronotal posterior lateral apices, posterior external edge of antennal groove of hypomeron, apical part of prosternal process, distal edge of elytron, posterior margins of metasternum and metacoxal plates and posterior part of abdomen tinged with red. Mouthparts yellowish with labrum and mandibles dark red; antennomeres I and II dark red; III – IX black; legs dark red but coxae and femurs black tinged with red. Body covered with yellow brown setae (Fig. 1 A, B). Head. Frontal margin broadly rounded in dorsal view (Fig. 2 A); frons distinctly depressed mesally and longitudinally (Fig. 2 A, B); supra-antennal carina absent (Fig. 2 B, C); frontoclypeal region most protruding between antennal sockets (Fig. 2 D), grooved between eyes and mandibles (Fig. 2 C); the grooves receive basal two antennomeres, opened posteriorly. Eyes 0.15 × longer than interocular distance in dorsal view. Mouthparts directed ventrally (Fig. 2 D). Labrum semioval (Fig. 2 E), 0.5 × longer than wide. Antennae extended beyond half-length of pronotum and not reaching pronotal posterior lateral apices by antennomere XI (Fig. 1 A); relative antennomere lengths: II <III <IV – X <XI <I (Fig. 2 F); antennomere I elongate and curved cylindrical; II short cylindrical, 0.9 × longer than wide; III trapezoidal, 1.2 × longer than wide, 1.9 × longer than II; IV – X parallelogram, weakly rounded distally, 1.1 – 1.2 × longer than wide; IV 1.2 × longer than III, 0.8 × longer than II – III combined; V 0.95 × longer than IV; XI bullet shaped, 1.8 × longer than wide, 1.45 × longer than X; each antennomeres with a longitudinal carina distally; the carinae on VIII – X not reaching their anterior edge, and carina on XI short. Mandibles unidentate (Fig. 2 G); penicillus short, formed by brush of short setae; dorsal sinuous carina developed; with setae of various lengths on outside of the carina; outside concave basally (Fig. 2 C). Maxilla (Fig. 2 H): cardo constricted posteriorly; basistipe triangular; mediostipe bullet-shaped; galea short, brush-like; lacinia short, brush-like; palpomeres short and broad; palpomeres II – III shorter than wide; palpomere IV fusiform, 1.4 × longer than wide. Labium (Fig. 2 I): mentum wide trapezoidal, with long setae; prementum with cluster of short setae medially; apical palpomere cylindrical. Prothorax widened posteriad but almost parallel-sided on posterior half, widest on posterior 1 / 5, and then weakly narrowed ahead of posterior angles (Fig. 1 A). Antennal grooves between pronotum and hypomeron deep and opened anteriorly, not reaching posterior lateral apices of prothorax, visible in lateral view (Fig. 2 D); sides of antennal grooves carinate. Pronotum sub-hexagonal, 1.2 × longer than wide, roundly convex, tallest posterior to the center (Fig. 2 D), with median longitudinal depression posteriorly; each lateral carina extending from posterior angles along antennal grooves (Fig. 2 D: black arrow), not reaching anterior angles; anterior angles obtuse (Fig. 2 A); anterior edge straight in dorsal view (Fig. 2 A); posterior angles extending posteriad, moderately broad, unicarinate; posterior edge without sublateral incision near each hind angle, lobated and protruding medially. Hypomeron depressed for reception of pro-femur and tibia, carinate behind the depression; anterior angle sharply pointed (Fig. 2 J: white arrow); posterior edge broadly rounded mesally and weakly emarginate near distal posterior angle; mesal posterior angle strongly and sharply protruding (Fig. 2 J: black arrow). Pronotosternal sutures (Fig. 1 B) deeply grooved to receive pro-tarsi; tarsal groove opened posteriorly and connected to hypomeron depression for reception of pro-femur and tibia (Fig. 2 J), becoming shallower anteriad, not reaching anterior edges of hypomeron and prosternum. Prosternum: anterior edge carinate, roundly projecting medially, depressed posterior to prosternal lobe; prosternal lobe short, not reaching the level of anterior edge of prothorax (Fig. 2 D). Prosternal process 1.8 × longer than procoxal cavity length, depressed between procoxae, strongly curved dorsad behind procoxae (Fig. 2 D); dorsal margin broadly rounded but with shallow concave medially in lateral view (Fig. 2 D); ventral lobe almost parallel sided and gradually narrowed near apex in ventral view, with median longitudinal carina on posterior 1 / 3 (Fig. 1 C); the carina forming pointed subapical tooth (Fig. 2 D: white arrow); ventral margin rounded inward anteriorly and straight posteriorly in lateral view (Fig. 2 D); ventral apex reaching dorsal apex; apex rounded in ventral and lateral views (Figs. 1 C, 2 D). Scutellar shield strongly inclined anterior-downwards (Fig. 1 A), tongue-shaped in anterodorsal view (Fig. 2 K), narrowed posteriad but almost parallel-sided around its half-length, 1.1 × longer than wide, convex (Fig. 2 L); anterior part protruding anteriad (Figs. 1 A, 2 L); posterior part not visible in lateral view (Fig. 2 D); anterior angles rounded; anterior edge straight; apex rounded. Mesosternum (Fig. 3 A): anterior edge straight but sinuate laterally, lobate on each side; mesosternal cavity floor without median band of setae; mesosternal cavity almost parallel-sided in ventral view; borders of mesosternal cavity in lateral view rounded anteriorly, straight medially and then curved ventrad obtusely (Fig. 2 D); mesosternal process between mesocoxae lower than mesocoxae, not visible in lateral view (Fig. 2 D); posterior edge 0.2 × wider than total width of mesosternum, almost straight (Fig. 3 A); mesosternum and metasternum connate medially (Fig. 2 J). Mesepisternum excavate anteriorly (Fig. 2 J). Mesepimeron depressed for reception of mid-femur (Fig. 2 J). Mesocoxal cavity closed to mesepisternum by mesosternum and mesepimeron (Fig. 2 J). Elytron 4.8 × longer than wide, 2.0 × longer than pronotal length, parallel-sided but narrowed from posterior 1 / 2, rounded apically, broadly convex but plane medially (Fig. 1 C), with striae. Elytral apices not meeting at midline (Fig. 1 A). Hind wings (Fig. 3 B) fully developed, without veins CuA 1 and CuA 3 + 4; RA 3 + 4 vestigial (Fig. 3 B: white arrow); r 4 translucent; mediomedial-cross vein between MP 1 + 2 and MP 3 + 4 translucent (Fig. 3 B: black arrow); mediocubital cross-vein between MP 3 + 4 and CuA 2 absent (Fig. 3 B: grey arrow); AA 3 vestigial; radial cell 4.0 × longer than wide. Metepisternum narrower than width of elytral epipleuron (Fig. 2 J). Metasternum depressed for reception of mid-tibia and tarsi, carinate ahead of depression for reception of mid-tibia and ventrad of depression for reception of mid-tarsi (Fig. 2 J: grey arrow), sulcate medially and ahead of metacoxal cavities (Fig. 1 B). Metacoxal plate narrowed medially; lateral edge distinctly longer than width of metepisternum; lateral anterior angle rounded; lateral posterior angle broadly pointed (Fig. 3 C: arrow). Tibiae with a pair of simple spurs (Fig. 3 D: arrow); relative tarsomere lengths: IV <III <II <I <V (Fig. 3 D); tarsi and claws simple. Abdomen. Ventrite V semicircular, broadly rounded apically (Fig. 1 B), 0.55 × longer than wide. Tergites and sternites VIII ‒ X yellow. Tergite VIII (Fig. 3 E) as long as wide, almost parallel-sided basally but weakly sinuate, broadly rounded apically. Sternite VIII (Fig. 3 F) triangular, abruptly narrowed at its half-length, shorter than wide; apex rounded. Tergite IX (Fig. 3 G) trapezoidal; anterior edge widely emarginate; median translucent area reaching tergite X. Tergite X (Fig. 3 G) curved triangular, widely emarginate anteriorly, rounded apically, 1.2 × longer than wide, 0.45 × length of tergite IX. Sternite IX (Fig. 3 H) elongate oval, widest at its half-length, broadly rounded apically. Aedeagus (Fig. 3 I, J) yellow, elongate. Phallobase 0.3 × total length of aedeagus, 1.1 × longer than wide. Median lobe gradually tapering to apex, exceeding apices of parameres by its apex; basal struts 0.15 × total length of median lobe. Parameres separated ventrally; apical parts without lateral subapical barbs, roundly truncate apically, without setae (probably due to damage). Female. Unknown.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFFC762AFF6052CAFE3BFD6C.taxon	etymology	Etymology. The specific epithet derives from Lambir Hills National Park, the type locality.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFFC762AFF6052CAFE3BFD6C.taxon	distribution	Distribution. Malaysia (Borneo).	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFFC762AFF6052CAFE3BFD6C.taxon	biology_ecology	Ecology. The life history and ecology of this species remain unknown, except that the holotype was collected via FIT in the canopy.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFF87626FF6050A7FA9DF9B0.taxon	description	(Figures 4 – 6)	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFF87626FF6050A7FA9DF9B0.taxon	materials_examined	Type material. Malaysia, Sarawak, Miri, Lambir Hills National Park, by ultraviolet light traps. Holotype. Male (AA 2739), 17 m, 23 – 24 VIII 1998. Paratypes. 1 male (AA 2738), 17 m, 14 – 15 XII 1996; 1 male (AA 2740), 35 m, 18 – 19 XII 1998; 1 male (AA 2741), 17 m, 17 – 18 V 1999; 1 male (AA 2742), 35 m, 15 – 16 II 2002.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFF87626FF6050A7FA9DF9B0.taxon	diagnosis	Male. Diagnosis. This species is characterized by the following features: punctures large and umbilicate; interspaces between punctures generally shorter than puncture diameter; body brown; antennae and legs orange; elytron with four sub-circular yellow spots; setae yellow; pronotum 1.3 × longer than wide; prosternal process 2.0 – 2.3 × longer than procoxal cavity length in ventral view; mesal posterior angle of hypomeron obtuse, not protruding; posterior part of scutellar shield visible in lateral view; elytron 5.2 – 5.5 × longer than wide, 2.0 – 2.2 × longer than pronotal length; metepisternum narrower than width of elytral epipleuron; basal struts 0.3 × total length of median lobe; apices of parameres tapering to apex.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFF87626FF6050A7FA9DF9B0.taxon	description	Measurements (5 specimens; holotype in parentheses). BL: 4.61 – 6.13 (4.61), BW: 1.11 – 1.54 (1.11), MAE: 0.90 – 1.11 (0.90), MBE: 0.54 – 0.66 (0.54), OI: 160 – 171 (169), PL: 1.41 – 1.99 (1.41), PML: 1.28 – 1.75 (1.28), PW: 1.10 – 1.56 (1.10), PAW: 0.87 – 1.12 (0.87), PLI: 128 – 129 (129), PWI: 126 – 146 (126), EL: 3.01 – 4.02 (3.01), EW: 0.55 – 0.75 (0.55), EI: 521 – 545 (545), BI: 197 – 216 (213). Description. Body (Fig. 4) elongate; surface generally smooth, shining but rough on depression of hypomeron for reception of pro-femur and tibia and on depressions of mesepimeron and metasternum for reception of midleg; punctures large and umbilicate, but on elytra small; depressions of hypomeron for reception of proleg and of mesepimeron and metasternum for reception of midleg without punctures; interspaces between punctures generally shorter than puncture diameter, but on metasternum, elytra and abdominal ventrites larger than puncture diameter and partly shorter than puncture diameter. Color. Body brown. Elytron with four sub-circular yellow spots, of which one spot medially on humerus, two spots side-by-side on anterior 1 / 3, and one spot medially on posterior 1 / 3. External edge of mandibles, mesal edge of antennal groove of hypomeron, sides of tarsal groove of pronotosternal sutures, external edge of scutellar shield, metasternal carina ventrad of depression for reception of mid-tarsi, striae and distal edge of elytron, and posterior edge of each abdominal ventrite blackish. Mouthparts yellowish with labrum and mandibles red-brown; antennae and legs orange with yellow tarsi. Body covered with yellow setae (Fig. 4). Head. Frontal margin broadly rounded in dorsal view (Fig. 5 A); frons weakly depressed mesally (Fig. 5 A, B); supra-antennal carina absent (Fig. 5 B, C); frontoclypeal region most protruding between antennal sockets (Fig. 5 D), grooved between eyes and mandibles (Fig. 5 C); the grooves receive basal two antennomeres, opened posteriorly. Eyes 0.25 – 0.35 × longer than interocular distance in dorsal view. Mouthparts directed ventrally (Fig. 5 D). Labrum elliptical, anterior edge more tapered (Fig. 5 E), 0.5 × longer than wide. Antennae extended beyond half-length of pronotum and not reaching pronotal posterior lateral apices by antennomere XI (Fig. 4 A, B); relative antennomere lengths: II <III <IV – X <XI <I (Fig. 5 F); antennomere I elongate and curved cylindrical; II short cylindrical, 0.8 – 0.85 × longer than wide; III trapezoidal, 1.1 – 1.4 × longer than wide, 2.0 – 2.4 × longer than II; IV – X parallelogram, weakly rounded distally, 1.1 – 1.3 × longer than wide; IV 1.1 – 1.2 × longer than III, 0.75 – 0.9 × longer than II – III combined; V 1.0 – 1.05 × longer than IV; XI bullet shaped, 1.7 – 2.1 × longer than wide, 1.3 – 1.5 × longer than X; each I – X with longitudinal carina distally; carinae on VI – X not reaching to their anterior edge. Mandibles unidentate (Fig. 5 G); penicillus short, formed by brush of short setae; dorsal sinuous carina developed; with setae of various lengths on outside of the carina; outside concave basally (Fig. 5 C). Maxilla (Fig. 5 H): cardo constricted posteriorly; basistipe triangular; mediostipe rectangular; galea short, brush-like; lacinia short, brush-like; palpomeres short and broad; palpomeres II – III shorter than wide; palpomere IV fusiform, 1.2 – 1.3 × longer than wide. Labium (Fig. 5 I): mentum wide trapezoidal, with long setae; prementum with cluster of short setae medially; apical palpomere subcylindrical. Prothorax widened posteriad on anterior half, slightly narrowed on posterior half ahead of posterior angles (Fig. 4 A), widest on its half-length or across posterior lateral apices, broadly rounded laterally on posterior 2 / 3. Antennal grooves between pronotum and hypomeron deep and opened anteriorly, not reaching posterior lateral apices of prothorax, visible in lateral view (Fig. 5 D); sides of antennal grooves carinate. Pronotum sub-hexagonal, 1.3 × longer than wide, roundly convex but weakly constricted posterior to anterior edge (Fig. 4 A, B), tallest around the center (Fig. 5 D), with median longitudinal depression posteriorly; each lateral carina extending from posterior angles along antennal grooves (Fig. 5 D: black arrow), not reaching anterior angles; anterior angles acute (Fig. 5 A); anterior edge straight in dorsal view (Fig. 5 A); posterior angles extending posteriad, acute, with median longitudinal carina vestigial; posterior edge without sublateral incision near each hind angle, lobated and protruding medially. Hypomeron depressed for reception of pro-femur and tibia, carinate behind the depression; anterior angle sharply pointed (Fig. 5 J: white arrow); posterior edge broadly rounded mesally and weakly emarginate near distal posterior angle; mesal posterior angle obtuse, not protruding (Fig. 5 J: black arrow). Pronotosternal sutures (Fig. 4 C) deeply grooved to receive pro-tarsi; tarsal groove opened posteriorly and connected to hypomeron depression for reception of pro-femur and tibia (Fig. 5 J), becoming shallower anteriad, not reaching anterior edges of hypomeron and prosternum. Prosternum: anterior edge carinate, roundly projecting medially, depressed posterior to prosternal lobe; prosternal lobe short, not reaching the level of anterior edge of prothorax (Fig. 5 D). Prosternal process 2.0 – 2.3 × longer than procoxal cavity length, depressed between procoxae, strongly curved dorsad behind procoxae (Fig. 5 D); dorsal margin broadly rounded but with shallow concave medially in lateral view (Fig. 5 D); ventral lobe almost parallel sided and gradually narrowed near apex in ventral view, with median longitudinal carina on posterior 1 / 3 (Fig. 4 C); the carina forming pointed subapical tooth (Fig. 5 D: white arrow); ventral margin rounded inward anteriorly and straight posteriorly in lateral view (Fig. 5 D); ventral apex reaching dorsal apex; apex rounded in ventral and lateral views (Figs. 4 C, 5 D). Scutellar shield distinctly inclined anterior-downwards but posterior part parallel to elytral surface (Fig. 4 A, B), tongue-shaped in anterodorsal view (Fig. 5 K), narrowed posteriad but almost parallel-sided posteriorly, 1.1 – 1.3 × longer than wide, convex (Fig. 5 L); anterior part strongly protruding anteriad (Figs. 4 A, 5 L); posterior part visible in lateral view (Fig. 5 D: gray arrow); anterior angles rounded; anterior edge weakly and widely emarginated from anterior view; apex rounded or narrowly emarginate apically. Mesosternum (Fig. 6 A): anterior edge straight but anterior edge of mesosternal cavity weakly protruding, lobate on each side; mesosternal cavity floor without median band of setae; mesosternal cavity almost parallel-sided anteriorly in ventral view; borders of mesosternal cavity in lateral view rounded anteriorly, straight medially and then curved ventrad obtusely (Fig. 5 D); mesosternal process between mesocoxae lower than mesocoxae, not visible in lateral view (Fig. 5 D); posterior edge 0.15 – 0.2 × wider than total width of mesosternum, almost straight (Fig. 6 A); mesosternum and metasternum connate medially (Fig. 6 A). Mesepisternum excavate anteriorly (Fig. 6 A). Mesepimeron depressed for reception of mid-femur (Fig. 5 J). Mesocoxal cavity closed to mesepisternum by mesosternum and mesepimeron (Fig. 5 J). Elytron 5.2 – 5.45 × longer than wide, 2.0 – 2.2 × longer than pronotal length, parallel-sided but narrowed from posterior 1 / 3, rounded apically, broadly convex but plane medially (Fig. 4 A), with striae. Elytral apices not meeting at midline (Fig. 4 A, B). Hind wings (Fig. 6 B) fully developed, without veins CuA 1 and CuA 3 + 4; RA 3 + 4 vestigial (Fig. 6 B: white arrow); r 4 translucent; mediomedial-cross vein between MP 1 + 2 and MP 3 + 4 vestigial (Fig. 6 B: black arrow); mediocubital cross-vein between MP 3 + 4 and CuA 2 anteriad to contact point between MP 3 and MP 4, not complete (Fig. 6 B: grey arrow); AA 3 located at contact point between CuA 2 and CAS; radial cell 3.7 – 4.2 × longer than wide. Metepisternum narrower than width of elytral epipleuron (Fig. 5 J). Metasternum depressed for reception of mid-tibia and tarsi, carinate ahead of depression for reception of mid-tibia and ventrad of depression for reception of mid-tarsi (Fig. 5 J: grey arrow), sulcate medially and ahead of metacoxal cavities (Fig. 4 C). Metacoxal plate narrowed medially; lateral edge distinctly longer than width of metepisternum; lateral anterior angle rounded; lateral posterior angle broadly pointed (Fig. 6 C: arrow). Tibiae with a pair of simple spurs (Fig. 6 D: arrow); relative tarsomere lengths: IV <III <II <V <I (Fig. 6 D); tarsi and claws simple. Abdomen. Ventrite V curved triangular, rounded apically (Fig. 4 C), 0.55 – 0.6 × longer than wide. Tergites and sternites VIII ‒ X yellow. Tergite VIII (Fig. 6 E) longer than wide, almost paradell-sided basally but weakly sinuate, broadly rounded apically. Sternite VIII (Fig. 6 F) triangular, abruptly narrowed at its half-length, shorter than wide; apex rounded. Tergite IX (Fig. 6 G) trapezoidal; anterior edge widely emarginate; posterior median notch moderate (0.1 – 0.15 × total length of tergite IX). Tergite X (Fig. 6 G) curved triangular but lateral edges sinuate in holotype, widely emarginate anteriorly, rounded apically, 0.9 – 1.2 × longer than wide, 0.4 – 0.45 × length of tergite IX. Sternite IX (Fig. 6 H) parallel-sided on anterior 2 / 5, expanded just behind anterior half, and then roundly converging posteriad. Aedeagus (Fig. 6 I, J) yellow, elongate. Phallobase 0.25 – 0.3 × total length of aedeagus, 0.9 – 0.95 × longer than wide. Median lobe gradually tapering to apex, exceeding apices of parameres by its apex; basal struts 0.3 × total length of median lobe. Parameres separated ventrally; apical parts without lateral subapical barbs, tapering to apex, each with less than two long setae dorsally and less than two long setae ventrally. Female. Unknown.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFF87626FF6050A7FA9DF9B0.taxon	etymology	Etymology. The specific epithet derives from Miri, the type locality.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFF87626FF6050A7FA9DF9B0.taxon	distribution	Distribution. Malaysia (Borneo).	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
03CF3761FFF87626FF6050A7FA9DF9B0.taxon	biology_ecology	Ecology. This species may be observed year-round in lowland rainforests, with specimens collected in February, May, August, and December. All specimens were captured using light traps set on trees during the night.	en	Arimoto, Kôichi, Kaliang, Clement Het, Mizuno, Takafumi, Meleng, Paulus, Kishimoto-Yamada, Keiko, Itioka, Takao (2025): First record and two new species of the genus Subprotelater Fleutiaux (Coleoptera, Elateridae, Subprotelaterinae) from Borneo. Zootaxa 5673 (4): 557-570, DOI: 10.11646/zootaxa.5673.4.4, URL: https://doi.org/10.11646/zootaxa.5673.4.4
