taxonID	type	description	language	source
03CF6A0FFFFC4B6BFF109654FDF4FD23.taxon	materials_examined	Type material. Holotype: CHINA (Gansu Prov.): ♂, two labels: " CHINA: S-Gansu [CH 12 - 12] / W-Qinling Shan, 128 km NW Longnan / Lazikou pass, S-side, Laolong valley / 34 ° 08 ' 14 '' N, 103 ° 51 ' 57 '' E, 2300 m, / S-slope with pine and spruce forest, litter sifted, 3. VIII. 2012, M. Schülke " [white, printed]; " EUTHEIA \ laolongiana m. \ det. P. Jałoszyński, ' 12 \ HOLOTYPUS " [red, printed] (MNHW).	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFC4B6BFF109654FDF4FD23.taxon	diagnosis	Diagnosis. Body large, broad and relatively convex; entire dorsum finely, but distinctly and densely punctate, with pronotal punctation gradually becoming finer and sparser towards anterior and lateral pronotal margins; aedeagus with complex apical structures, with a pair of long sub-apical lateral projections and large apical lateral lobes.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFC4B6BFF109654FDF4FD23.taxon	description	Description. BL 1.63 mm. Body of male (Fig. 1) moderately convex, slender, dark brown with slightly darker head, covered with light brown vestiture. Head broadest, in dorsal view, at strongly elongate and large convex eyes, HL 0.20 mm, HW 0.30 mm; vertex and frons confluent and weakly convex; supraantennal tubercles not marked. Punctures on head dorsum distinct, sharply marked and dense, of various diameters and unevenly distributed, separated by spaces 0.5 – 3 × as wide as puncture diameters; setae short, sparse and suberect. Antennae slender and gradually thickened distally, AnL 0.63 mm, antennomere I nearly 3 × as long as broad; II much shorter and slightly narrower than I, nearly twice as long as broad; III slightly transverse; IV – VII each slightly elongate; VIII slightly smaller than VII, about as long as broad; IX – X each slightly transverse; XI about twice as long as broad. Pronotum subtrapezoidal, broadest between middle and posterior third; PL 0.38 mm, PW 0.45 mm; anterior and lateral margins broadly and evenly rounded; lateral margins indistinctly convergent towards obtuse hind angles; posterior margin shallowly bisinuate; pronotal base with well-defined and nearly circular median ante-basal pit and two lateral pairs of larger, irregular and diffused pits. Punctures on pronotal disc distinct but small, sharply marked, in middle separated by spaces as wide as 1 – 2 × puncture diameters, punctures gradually reducing in diameter and depth and becoming sparser towards anterior and lateral margins of pronotum. Setae short, sparse and suberect. Elytra more convex than pronotum, oval, broadest distinctly anterior to middle; EL 0.80 mm, EW 0.63 mm, EI 1.28; humeral calli well-marked, elongate. Punctures on median part of elytra slightly smaller and distinctly shallower than those in middle of pronotum, separated by spaces subequal to puncture diameters, punctures gradually becoming finer towards lateral and posterior elytral margins; setae similar to those on pronotum. Hind wings well developed. Legs moderately long and slender, non-modified. Aedeagus (Figs. 2 – 3) moderately slender; AeL 0.41 mm; in ventral view apical part of median lobe distinctly demarcated from bulbous basal part by deep lateral constriction, sub-apical region with a pair of long lateral projections each bearing two apical setae, median apical part of median lobe (columella) with large lateral lobes strongly projecting laterally and distally, apex of columella with median notch; basal part of median lobe strongly elongate and with ventral margin of membranous area broadly and deeply emarginated; parameres short, not reaching apex of median lobe, each with three apical setae. Female. Unknown.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFC4B6BFF109654FDF4FD23.taxon	distribution	Distribution. China, Gansu Prov. (Fig. 12).	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFC4B6BFF109654FDF4FD23.taxon	etymology	Etymology. After the type locality, Laolong Valley.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFC4B6BFF109654FDF4FD23.taxon	discussion	Remarks. Species of Eutheia are extremely uniform in the external morphology and although some differences can be seen in the proportions of body parts, pigmentation, sculpture of dorsum or proportions of antennomeres, species diagnoses are primarily based on genital structures. The aedeagus in Eutheia shows far larger diversity than any external characters and often externally nearly identical species have clearly different aedeagi (e. g., Jałoszyński 2011). Eutheia laolongiana, a large and relatively stout species externally highly similar to previously known Chinese and Taiwanese congeners, has one of the most complex aedeagus within the genus, and clearly belongs to a group of East Palaearctic species with more or less bifurcate apex and variously developed lateral apical or sub-apical lobes. This group comprises E. horii Jałoszyński, 2004 (Japan), E. rufa Jałoszyński, 2004 (Japan), E. yoshidai Hoshina, 2007 (Japan), E. puetzi Jałoszyński, 2008 a (China), E. gramme Jałoszyński, 2008 b (Taiwan), E. klapperichi Franz, 1985 (Taiwan), E. latissima Jałoszyński, 2008 b (Taiwan), E. simillima Franz, 1985 (Taiwan) and E. taiwanensis Franz, 1985 (Taiwan). All Western Palaearctic and all other Eastern Palaearctic species have the apical part of median lobe simple, with rounded or distally projected apical margin, without large lateral lobes or with only slightly expanded lateral margins in the apical or sub-apical region, forming small, usually angulate lobes. An interesting exception is E. hariola Kurbatov, 1991 from the Russian Far East (Primorie), which has the apex of median lobe slightly projecting distally in middle, but its lateral parts are strongly expanded laterally and forming large, rounded lobes. Interestingly, this is also the only species other than E. laolongiana with sub-apical lateral projections bearing apical setae. Some other species of Eutheia also have the lateral sensory setae in the sub-apical region of the median lobe, e. g., E. linearis Mulsant & Rey, 1861 (Europe and W Siberia), E. exortiva Kurbatov, 1990 (Primorie) and some undescribed species from Tunisia and Texas (Jałoszyński, unpublished data), but the setae are inserted on not expanded or slightly expanded, rounded or angulate lateral margins of median lobe and not on distinct projections. The projections in E. hariola are much shorter than those in E. laolongiana.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFE4B6DFF109674FE81FBB8.taxon	materials_examined	Type material. Holotype: CHINA (Gansu Prov.): ♂, two labels: " CHINA: S-Gansu [CH 12 - 12] / W-Qinling Shan, 128 km NW Longnan / Lazikou pass, S-side, Laolong valley / 34 ° 08 ' 14 '' N, 103 ° 51 ' 57 '' E, 2300 m, / S-slope with pine and spruce forest, litter sifted, 3. VIII. 2012, M. Schülke " [white, printed]; " VERAPHIS \ gansuanus m. \ det. P. Jałoszyński, ' 12 \ HOLOTYPUS " [red, printed] (cMS). Paratype (♀): three labels; " CHINA: S-Gansu [CH 12 - 11] / W-Qinling Shan, 125 km NW / Longnan, Lazikou pass, S-side / Zhuli valley, 34 ° 07 ' 57 '' N, 103 ° 56 ' 15 '' E, 2260 m, " [white, printed], " N-slope, mixed forest with oak / and pine near creek, litter and / dead wood sifted, 3. VIII. 2012, / leg. M. Schülke " [white, printed], " VERAPHIS \ gansuanus m. \ det. P. Jałoszyński, ' 12 \ PARATYPUS " [yellow, printed] (cPJ).	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFE4B6DFF109674FE81FBB8.taxon	diagnosis	Diagnosis. Male protrochanter with subrectangular apex; aedeagus in ventral view with distinctly asymmetrical, broader than long, deeply and narrowly emarginate apical part and distinct internal armature located sub-apically; in lateral view apex of median lobe evenly rounded and curved ventrally. Females unremarkable, on the basis of morphological characters can be identified only by direct comparison to males, preferably when collected together.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFE4B6DFF109674FE81FBB8.taxon	description	Description. BL 1.58 mm. Body of male (Fig. 4) weakly convex, slender, moderately dark brown, covered with yellowish vestiture, legs and antennae only minimally lighter. Head broadest at large strongly convex eyes, HL 0.18 mm, HW 0.28 mm; vertex with distinct pair of median pits, each prolonged by shallow longitudinal groove running anterad and becoming gradually shallower and broader to disappear at posterior margins of barely marked supraantennal tubercles; frons flattened, clypeus steeply declining anterad, flattened; sides of vertex and frons confluent and convex. Punctures on head dorsum fine and indistinct; setae short, sparse and suberect. Antennae slender, with indistinctly demarcated club composed of antennomeres IX – XI, AnL 0.63 mm, antennomere I nearly 3 × as long as broad; II distinctly shorter and slightly narrower than I, 2.2 × as long as broad; III nearly as long as broad; IV – VII each elongate (V longest); VII slightly transverse and distinctly shorter than VII; IX about as long as broad; X slightly transverse; XI only about 1.3 × as long as broad. Pronotum nearly subquadrate, broadest near anterior fourth; PL 0.38 mm, PW 0.40 mm, anterior margin broadly and evenly rounded; lateral margins strongly rounded in anterior third, behind middle indistinctly constricted and in posterior fourth distinctly convergent towards obtuse and blunt hind angles; posterior margin shallowly bisinuate. Pronotal base with distinct, deep and nearly circular median ante-basal pit and lateral pair of elongate impressions running along lateral pronotal margins. Punctures on pronotal disc large and dense but shallow and diffused, in middle separated by spaces subequal to puncture diameters, punctures gradually becoming finer towards pronotal margins. Setae short and sparse, suberect. Elytra more convex than pronotum, oval, broadest slightly anterior to middle; EL 0.78 mm, EW 0.58 mm, EI 1.35; humeral calli well-marked, elongate. Punctures on median part of elytra similar to those on pronotal disc but slightly smaller and yet more diffused; setae similar to those on pronotum. Hind wings well developed. Legs moderately long and slender; protrochanter with distinctly angulate margin, with sub-rectangular apex, protibia with sub-apical fin-like projection (Fig. 6). Metaventrite with median impression nearly as long as ventrite, margins of impression diffused. Aedeagus (Figs. 8 – 9) slender; AeL 0.33 mm; in ventral view median lobe distinctly constricted in sub-median region, with asymmetrical apical part bearing deep and narrow median apical emargination; in lateral view, apex of median lobe folded ventrally; internal structures distinct, located in sub-apical region; parameres exceeding half length of median lobe. Female. Similar to male, except for non-modified fore legs. BL 1.65 mm; HL 0.18 mm, HW 0.28 mm, AnL 0.58 mm; PL 0.40 mm, PW 0.43 mm; EL 0.83 mm, EW 0.60 mm, EI 1.38.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFE4B6DFF109674FE81FBB8.taxon	distribution	Distribution. China, Gansu Prov. (Fig. 13).	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFE4B6DFF109674FE81FBB8.taxon	etymology	Etymology. After the Chinese province Gansu.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFFE4B6DFF109674FE81FBB8.taxon	discussion	Remarks. Veraphis gansuanus is most similar to its Chinese congener V. qinghaiensis and the V. japonicus species group known from Japan and China. It shares with V. qinghaiensis a similar subrectangular male protrochanter, but clearly differs in much stouter aedeagus strongly constricted in the sub-apical region and a different shape of its apical region. Veraphis gansuanus shares with the japonicus group (i. e., V. m o d e s t u s, V. japonicus (K. Sawada, 1962), V. odaigaharensis Jałoszyński & Hoshina 2005, V. tottoriensis Jałoszyński & Hoshina 2005 and V. h i s a m a t s u i Jałoszyński & Hoshina 2005) a similar shape of the aedeagus, with abruptly demarcated apical part, but differs in the shape of male protrochanter, which in nearly all species belonging to that group is rounded, not angulate. The only exception is V. h i s a m a t s u i with indistinctly angulate protrochanter, similar to that of V. gansuanus, but apical parts of aedeagi of these two species are clearly different (short, asymmetrical and with a narrow median emargination in V. gansuanus vs. long, symmetrical and with a straight apical margin in V. h i s a m a t s u i). Differences between V. gansuanus and all other Palaearctic species of Veraphis are also distinct: the new species differs from the V. irkutensis species group and from all subspecies of V. engelmarki Franz, 1971 in the apex of aedeagus without setae and the antennomere XI without the sub-apical cavity; from the V. s a w a d a i species group in the angulate male protrochanter and the aedeagus with short (and not long) and abruptly (and not gradually) demarcated apical part; and from V. s pi n os u s and V. calcarifer in the male protrochanter without pointed apical projection.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFF84B6EFF1091CAFDAFFD00.taxon	materials_examined	Type material. Holotype: CHINA (Sichuan Prov.): ♂, two labels: " CHINA [24]: N-Sichuan / pass NW Songpan, / 3600 m, 32 ° 55 ' 32 '' N, / 103 ° 25 ' 56 '' E, sifted, / 11. VIII. 2012, V. Assing " [white, printed]; " VERAPHIS \ assingi m. \ det. P. Jałoszyński, ' 12 \ HOLOTYPUS " [red, printed] (MNHW).	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFF84B6EFF1091CAFDAFFD00.taxon	diagnosis	Diagnosis. Male protrochanter with subtriangular apex; aedeagus in ventral view with asymmetrical, about as long as broad apical part slightly expanded distally in middle and distinct internal armature located sub-apically; in lateral view apical part of median lobe recurved, with apex directed dorsally.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFF84B6EFF1091CAFDAFFD00.taxon	description	Description. BL 1.45 mm. Body of male (Fig. 5) weakly convex, slender, moderately dark brown, covered with yellowish vestiture, legs and antennae only minimally lighter. Head broadest at large strongly convex eyes, HL 0.18 mm, HW 0.25 mm; vertex with distinct pair of median pits not prolonged by longitudinal grooves; frons flattened, clypeus steeply declining anterad and flattened; sides of vertex and frons confluent and convex; supraantennal tubercles weakly raised. Punctures on head dorsum fine and indistinct; setae short, sparse and suberect. Antennae slender, with distinctly demarcated club composed of antennomeres IX – XI, AnL 0.53 mm, antennomere I 2.5 × as long as broad; II distinctly shorter and slightly narrower than I, 2.5 × as long as broad; III distinctly transverse; IV – V each elongate (V longest); VI about as long as broad; VII slightly transverse; VIII distinctly shorter and slightly narrower than VII, strongly transverse; IX – X each distinctly transverse; XI barely longer than broad. Pronotum nearly subquadrate, broadest near anterior fourth; PL 0.33 mm, PW 0.35 mm, anterior margin broadly and evenly rounded; lateral margins strongly rounded in anterior third, from widest place distinctly convergent towards obtuse and blunt hind angles; posterior margin shallowly bisinuate. Pronotal base with distinct, deep and nearly circular median ante-basal pit and lateral pair of elongate impressions running along lateral pronotal margins. Punctures on pronotal disc large and dense but shallow and diffused, in middle separated by spaces equal to 1 – 1.5 puncture diameters, punctures gradually becoming finer towards pronotal margins. Setae short and sparse, suberect. Elytra more convex than pronotum, oval, broadest slightly anterior to middle; EL 0.73 mm, EW 0.53 mm, EI 1.38; humeral calli well-marked, elongate. Punctures on median part of elytra slightly denser, smaller and more diffused than those on pronotum; setae similar to those on pronotum. Hind wings well developed. Legs moderately long and slender; protrochanter with rounded margin and subtriangular apex, protibia with sub-apical fin-like projection (Fig. 7). Metaventrite with median impression nearly as long as ventrite, margins of impression diffused. Aedeagus (Figs. 10 – 11) slender; AeL 0.35 mm; in ventral view median lobe gradually broadening from subbasal to sub-apical regions, with asymmetrical apical part slightly expanded distally and strongly curved dorsally; internal structures distinct, located in sub-apical region; parameres shorter than half length of median lobe. Female. Unknown.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFF84B6EFF1091CAFDAFFD00.taxon	distribution	Distribution. China, Sichuan Prov. (Fig. 13).	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFF84B6EFF1091CAFDAFFD00.taxon	etymology	Etymology. This species is dedicated to its collector and a specialist on Staphylinidae, Volker Assing.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
03CF6A0FFFF84B6EFF1091CAFDAFFD00.taxon	discussion	Remarks. Veraphis assingi differs from all Palaearctic species of Veraphis in the strongly recurved apical part of median lobe in lateral view, with the apex directed dorsally. The new species is most similar to the V. japonicus species group, to which it can be included.	en	Jałoszyński, Paweł (2013): Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa 3609 (5): 495-503, DOI: 10.11646/zootaxa.3609.5.5
