taxonID	type	description	language	source
03CCDB58E627FF9437CD79B7FF50F826.taxon	materials_examined	Type material. Holotype: 3: " S. Afr., E. Transvaal / Berlin Forest St. / 25.33 S – 30.44 E " [white, printed], " 22.9.1986; E-Y: 2284 / litt. rock crevice / leg. Endrödy-Younga " [white, printed], " STENOMASTIGUS / berlinafricanus m. / HOLOTYPUS " [red, printed] (TMSA). Paratype: 3: " RSA: Mpumalanga / Sabie env., 1200 m / S 25 ° 13,1 '; E 30 ° 37 ' / 10 – 11. II. 04, Hlaváč lgt. " [white, printed], " under litter / on wet wood " [white, handwritten], " STENOMASTIGUS / berlinafricanus m. / PARATY- PUS " [yellow, printed] (coll. PH).	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E627FF9437CD79B7FF50F826.taxon	diagnosis	Diagnosis. Male: pronotum and elytra uniformly dark brown, head slightly darker; protrochanters with projecting distal margin forming subtriangular tooth, projection clearly shorter than remaining part of trochanter; profemora in lateral view with very shallow ventral subapical emargination and straight dorsal margin; aedeagus in abparameral view with very strong constriction between basal capsule of median lobe and base of copulatory piece, in left lateral view long paramere with very slender proximal part gradually and only slightly narrowing towards apex, evenly curved and with rounded, fin-like parameral tooth, strongly standing out from the silhouette of paramere. Females and their diagnostic characters unknown.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E627FF9437CD79B7FF50F826.taxon	description	Description. Body of male (Fig. 22) long and slender, with extremely long appendages, BL 3.43 – 3.53 mm; coloration moderately dark brown, head, pedicel, scapus and femora slightly darkened; vestiture silverish. Head broadest at eyes, HL 0.58 – 0.60 mm, HW 0.63 – 0.64 mm; occipital constriction distinctly broader than 0.5 x HW; vertex more than twice as broad as long, distinctly concave in middle, with posterior margin bearing broad subtriangular median emargination extended anterad by median longitudinal groove as long as half of vertex, lateral parts of vertex convex; frons much longer than broad, impressed between convex supraantennal tubercles, its anterior part located between antennal insertions convex and as narrow as width of supraantennal tubercle; vertex and frons uniformly covered with fine microsculpture composed of a maze-like net of grooves that gives a matt but slightly iridescent appearance; vestiture extremely short but dense, composed of recumbent setae; clypeus nearly four times as broad as long, with slightly concave anterior margin and glossy surface with only traces of microsculpture and transverse row of several very long, thick and erect macrosetae directed anterad; labrum 2.5 x as broad as long, with distinct median emargination and glossy surface bearing row of several very long and thick macrosetae directed anteriorly and curved ventrally. Eyes large and coarsely faceted, each in lateral view oval and elongate, slightly shorter than temple. Antennae very slender, distinctly longer than body, AeL 4.05 mm, lengths of antennomeres (from I to XI): 0.83 mm, 0.40 mm, 0.33 mm, 0.30 mm, 0.39 mm, 0.30 mm, 0.35 mm, 0.31 mm, 0.30 mm, 0.27 mm, 0.27 mm; scapus and pedicel strongly thickened, all flagellomeres very thin and hardly noticeably thickening towards antennal apex; scapus about 6 x as long as broad, distinctly broadened between middle and distal third, with two longitudinal ventral setal rows each composed of 6 very long, thick and strongly erect bristles and with surface covered uniformly with moderately sparse, short suberect setae; pedicel 5 x as long as broad, with two longitudinal ventral setal rows each composed of 3 long bristles, and additional single ventral apical bristle, surface of pedicel covered with moderately sparse, short suberect setae; all flagellomeres strongly elongate, much narrower than pedicel and lighter in colour, each covered with short and nearly recumbent setae denser than those on scapus and pedicel, without bristles. Pronotum pear-shaped in dorsal view, strongly elongate and broadest between middle and anterior third, PL 0.98 mm, PW 0.75 mm; anterior margin weakly arcuate; lateral margins strongly rounded in anterior half, broadly constricted just posterior to middle and weakly rounded in posterior fourth; posterior margin strongly arcuate; pronotal disc with very indistinctly marked median longitudinal carina and pair of shallow oval impressions between middle and anterior third, separated by distance slightly longer than width of impression. Surface covered with similar matt microsculpture and dense vestiture as those on vertex and frons. Elytra oval and more convex than pronotum, broadest distinctly anterior to middle, EL 1.88 – 1.95 mm, EW 1.23 – 1.25 mm, EI 1.53 – 1.56; sub-basal part of each elytron slightly impressed, so that narrow basal part appears slightly raised; posterior adsutural angle of each elytron blunt, elytra with barely noticeable shallow and diffused striae; surface covered with similar microsculpture and vestiture as pronotum. Legs very long and slender; protrochanters modified as in Fig. 12; protibiae as in Fig. 17. Aedeagus (Figs. 25, 26, 35 – 38) very slender, AeL 1.63 mm, in abparameral view with very narrow " stalk " between basal capsule and base of copulatory piece, short paramere obliterated, long paramere in lateral views evenly curved; parameral tooth fin-like, in both lateral aspects strongly standing out from the silhouette of paramere; apex of copulatory piece as in Figs. 62, 67. Female. Unknown.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E627FF9437CD79B7FF50F826.taxon	distribution	Distribution. Republic of South Africa, East Transvaal.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E627FF9437CD79B7FF50F826.taxon	etymology	Etymology. Locotypical, after the Berlin Forest, where the holotype specimen was collected.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E627FF9437CD79B7FF50F826.taxon	discussion	Remarks. Among all species of Stenomastigus with protrochanters projecting distally, S. berlinafricanus has the longest aedeagus, with the most evenly curved long paramere in left lateral view. The relatively dark body pigmentation found in this species is also characteristic of S. kosianus sp. n. (all remaining species belonging to this group are yellowish-testaceous). The latter species can easily be distinguished by its very long, rod-like protrochanters.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E624FF9B37CD7B8EFD66F945.taxon	materials_examined	Type material. Holotype: 3: " S. Afr.; KwaZulu-Natal / Kosi Bay mouth / 26.58 S – 32.50 E " [white, printed], " 13.11.2002; E-Y: 3531 / general collecting / Burger, Harrison, Müller " [white, printed], " STENOMASTIGUS / kosianus m. / HOLOTYPUS " [red, printed] (TMSA). Paratypes: 2 ƤƤ: same data as for holotype and additional label " STENOMASTIGUS / kosianus m. / PARATY- PUS " [yellow, printed] (TMSA).	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E624FF9B37CD7B8EFD66F945.taxon	diagnosis	Diagnosis. Male: head and pronotum dark brown, elytra slightly lighter; protrochanters with projecting distal margin forming slender rod, the projection as long as remaining part of trochanter; profemora in lateral view with deep ventral subapical emargination and minimally bent dorsal margin; aedeagus in abparameral view with strong constriction between basal capsule of median lobe and base of copulatory piece, in left lateral view long paramere with broad proximal part rapidly narrowing towards apex, slightly bent in two places and with rounded, fin-like parameral tooth weakly standing out from the silhouette of paramere. Female: body pigmented as in male; elytra with large but relatively shallow sub-basal impression separated from pronotal base by narrow basal ridge.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E624FF9B37CD7B8EFD66F945.taxon	description	Description. Body of male (Fig. 23) long and slender, with extremely long appendages, BL 3.33 mm; coloration moderately dark brown, with head and pronotum slightly darker than remaining body parts; vestiture silverish. Head broadest at eyes, HL 0.58 mm, HW 0.60 mm; occipital constriction distinctly broader than 0.5 x HW; vertex more than twice as broad as long, distinctly concave in middle, with posterior margin bearing broad subtriangular median emargination extended anteriorly by median longitudinal groove reaching to frons, lateral parts of vertex convex; frons much longer than broad, impressed between convex supraantennal tubercles, its anterior part located between antennal insertions convex and slightly broader than supraantennal tubercle; vertex and frons uniformly covered with very fine but very dense punctures adjacent to one another and extremely short but dense vestiture composed of recumbent setae; clypeus nearly four times as broad as long, with slightly concave anterior margin and glossy surface with only traces of microsculpture and transverse row of several very long, thick and erect macrosetae directed anteriorly; labrum 2.5 x as broad as long, with shallow median emargination and glossy surface bearing row of several very long and thick macrosetae directed anteriorly and curved ventrally. Eyes large and coarsely faceted, each in lateral view oval and elongate, as long as temple. Antennae very slender, distinctly longer than body, AeL 3.76 mm, lengths of antennomeres (from I to XI): 0.70 mm, 0.38 mm, 0.33 mm, 0.31 mm, 0.31 mm, 0.30 mm, 0.35 mm, 0.30 mm, 0.28 mm, 0.24 mm, 0.28 mm; scapus and pedicel strongly thickened, all flagellomeres very thin and hardly noticeably thickening towards antennal apex; scapus about 6 x as long as broad, slightly broadening between middle and distal third, with two longitudinal ventral setal rows each composed of 6 very long, thick and strongly erect bristles and with surface covered uniformly with moderately sparse, short suberect setae; pedicel 5 x as long as broad, with two longitudinal ventral setal rows each composed of 3 long bristles, and additional single ventral apical bristle, surface of pedicel covered with moderately sparse, short suberect setae; all flagellomeres strongly elongate, much narrower than pedicel and slightly lighter in colour, each covered with short and nearly recumbent setae denser than those on scapus and pedicel, without bristles. Pronotum pear-shaped in dorsal view, strongly elongate and broadest between middle and anterior third, PL 0.95 mm, PW 0.68 mm; anterior margin nearly straight; lateral margins strongly rounded in anterior half, broadly constricted just posterior to middle and weakly rounded in posterior fourth; posterior margin strongly arcuate; pronotal disc with indistinctly marked median longitudinal carina and pair of shallow oval impressions near middle, separated by distance much longer than width of impression. Surface covered with microsculpture composed of very dense microgranules and with matt appearance; vestiture very short, dense, recumbent. Elytra oval and more convex than pronotum, broadest distinctly anterior to middle, EL 1.80 mm, EW 1.25 mm, EI 1.44; sub-basal part of each elytron barely noticeably impressed, so that narrow basal part appears minimally raised; posterior adsutural angle of each elytron blunt, elytra with barely noticeable shallow and diffused striae visible only near suture; surface covered with microsculpture similar to that on pronotum, but microgranules are slightly larger and flatter; vestiture extremely short, dense and recumbent. Legs very long and slender; protrochanters modified as in Fig. 18; profemora as in Fig. 18. Aedeagus (Figs. 27, 28, 39 – 42) very slender, AeL 1.48 mm, in abparameral view with narrow " stalk " between basal capsule and base of copulatory piece, short paramere obliterated, long paramere in lateral views slightly bent near middle and in subapical region; parameral tooth fin-like, in both lateral views weakly standing out from the silhouette of paramere. Apex of copulatory piece as in Figs. 63, 68. FIGURES 25 – 34. Aedeagus in abparameral (25, 27, 29, 31, 33) and left lateral (26, 28, 30, 32, 34) views. 25 – 26, Stenomastigus berlinafricanus sp. n. 27 – 28, S. kosianus sp. n. 29 – 30, S. allaeri Leleup. 31 – 32, S. basilewskyi Leleup. 33 – 34, S. kochi Leleup. Abbreviations: c, constriction; cp, copulatory piece; lp, long paramere; m, median lobe; pt, parameral tooth. Scale bars: 0.5 mm. Female (Fig. 24). Similar to male in body pigmentation, microsculpture and vestiture, differs clearly in larger length, different proportions of body parts and, most strikingly, in shape of elytra. BL 3.65 – 4.15 mm, HL 0.67 – 0.68 mm, HW 0.60 – 0.73 mm, AnL 4.04 mm, lengths of antennomeres (from I to XI): 0.83 mm, 0.45 mm, 0.35 mm, 0.31 mm, 0.33 mm, 0.30 mm, 0.35 mm, 0.30 mm, 0.28 mm, 0.27 mm, 0.27 mm; PL 0.93 – 1.05 mm, PW 0.70 – 0.83 mm. Elytra very broad, broadest between middle and anterior third, EL 2.13 – 2.43 mm, EW 1.48 – 1.70 mm, EI 1.43 – 1.44; in dorsal view each elytron bears very deep and elongate sub-basal impression delimited anteriorly by narrow transverse basal ridge, posteriorly and laterally the impression gradually becoming shallower and nearly confluent with surrounding areas; additionally, adsutural parts of the impression deepened and forming indistinct longitudinal grooves. Apex of each elytron forming blunt and obtuse angle, best visible in lateral view; in dorsal view apices of elytra broadly separated from each other and space between them forms subtriangular notch. In lateral view the highest point of elytra located in anterior third, basal ridges and sub-basal impression clearly visible.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E624FF9B37CD7B8EFD66F945.taxon	distribution	Distribution. Republic of South Africa, Natal.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E624FF9B37CD7B8EFD66F945.taxon	etymology	Etymology. Locotypical, after Kosi Bay, near where the type series was collected.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E624FF9B37CD7B8EFD66F945.taxon	discussion	Remarks. See remarks for S. berlinafricanus.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E629FF9D37CD7DE4FDD0F900.taxon	materials_examined	Material studied. Paratypes: 3 33, 3 ƤƤ, Republic of South Africa, Natal, Estcourt District, Champagne Castle (RMCA). Revised diagnosis. Male: Body (Fig. 43) yellowish-testaceous, with slightly darker head and pronotum; protrochanters (Fig. 14) with projecting distal margin forming slender rod slightly broadening apically, projection as long as remaining part of trochanter; profemora (Fig. 19) in lateral view with shallow ventral subapical emargination and nearly straight dorsal margin; aedeagus (Figs. 29, 30, 45 – 49) in abparameral view with weak constriction between basal capsule of median lobe and base of copulatory piece, in left lateral view long paramere with slender proximal part gradually narrowing towards apex, nearly evenly curved and with rounded, very short and broad parameral tooth weakly standing out from the silhouette of paramere. Female (Fig. 44): body pigmented as in male; elytra with deep basal impression adjacent to pronotal base; in lateral view apex of elytra located clearly above middle of elytral height (indistinguishable from female of S. kochi, see Remarks).	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E629FF9D37CD7DE4FDD0F900.taxon	distribution	Distribution. Republic of South Africa, Natal.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E629FF9D37CD7DE4FDD0F900.taxon	discussion	Remarks. Females of this species cannot be distinguished from S. kochi and species identification is possible only when females have been collected together with males in the same locality. The key provided by Leleup (1968) is incorrect and the illustration showing the elytra of S. kochi in lateral view (Fig. 126 in Leleup, 1968) is misleading. In fact the female of S. kochi has the elytra in lateral view identical to that of S. allaeri and very similar to that illustrated by Leleup in his Fig. 125.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E62FFF9F37CD7C5CFD78FDE6.taxon	materials_examined	Material studied. Paratypes: 9 33, 6 ƤƤ, Republic of South Africa, Natal, Bergville District, Indumeni Forest (RMCA). Revised diagnosis. Male: Body (Fig. 54) yellowish-testaceous, with slightly darker head and pronotum; protrochanters (Fig. 15) with projecting distal margin forming broad rod slightly broadening apically, projection longer than remaining part of trochanter; profemora (Fig. 20) in dorsal view with deep lateral subapical emargination and broad, blunt lateral tooth at base of emargination; aedeagus (Figs. 31, 32, 50 – 53) in abparameral view with weak constriction between basal capsule of median lobe and base of copulatory piece, in left lateral view long paramere with long, subtriangular parameral tooth strongly standing out from the silhouette of paramere; apical part of paramere above tooth distinctly broadened. Female (Fig. 55): body pigmented as in male; elytra with deep basal impression adjacent to pronotal base; in lateral view apex of elytra located nearly in middle of elytral height.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E62FFF9F37CD7C5CFD78FDE6.taxon	distribution	Distribution. Republic of South Africa, Natal.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E62DFF8137CD7972FD78F829.taxon	materials_examined	Material studied. Holotype 3 and paratype Ƥ, Republic of South Africa, Natal, Bergville District, Oqalweni Forest (TMSA). Revised diagnosis. Male: Body (Fig. 60) yellowish-testaceous, with slightly darker head and pronotum; protrochanters (Fig. 14) with projecting distal margin forming moderately slender rod slightly broadening apically, projection as long as remaining part of trochanter; profemora (Fig. 19) in lateral view with very deep subapical ventral emargination, curved dorsal margin and pointed, subtriangular ventral tooth located at base of emargination; aedeagus (Figs. 33, 34, 56 – 45) in abparameral view with weak constriction between basal capsule of median lobe and base of copulatory piece, in left lateral view long paramere with very short, rounded and broad parameral tooth weakly standing out from the silhouette of paramere, apical part of paramere above tooth distinctly broadened. Female (Fig. 61): body pigmented as in male; elytra with deep basal impression adjacent to pronotal base; in lateral view apex of elytra clearly located above middle of elytral height (indistinguishable from female of S. allaeri, see Remarks for the latter species).	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
03CCDB58E62DFF8137CD7972FD78F829.taxon	distribution	Distribution. Republic of South Africa, Natal.	en	Jałoszyński, Paweł (2012): Stenomastigus Leleup (Staphylinidae, Scydmaeninae): status of subgenus Acanthostigus Leleup and revision of species with elongated male protrochanters. Zootaxa 3153: 39-56, DOI: 10.5281/zenodo.279590
