taxonID	type	description	language	source
03CC0D78FFCCFF83FF751EF8FE00F876.taxon	materials_examined	Type species: Cragandhara himalaya sp. n., by present designation.	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFCCFF83FF751EF8FE00F876.taxon	diagnosis	Diagnosis. Species of the new genus (Figs 1 – 3) are externally reminiscent of the genera Gandhara (Figs 4, 5, 13, 16) and Collita Moore, 1878 (Figs 6, 7, 14, 18) but differ clearly in their genitalia structures. In the male genitalia of Cragandhara gen. n. (Figs 10, 11), the following features are characteristic of the genus. (1) The anellus bears two dense lateral bunches of spinules (vs. a couple of serrulate plates in Gandhara and a membranous anellus in Collita). (2) The ventral plate of the costa (‘ clasper’) bears a swollen longitudinal crest densely covered with minute spinules. (3) The dorsal surface of the distal saccular process is densely setose. (4) The phallus has a pair of weakly sclerotised and longitudinally rugose distal processes, which are unknown in other Asiatic Lithosiina but found in several Afrotropical genera, and are most similar to those of the genus Cragia Birket-Smith, 1965, which is fundamentally different from the new genus in its external appearance as well as the male genital capsule and female genitalia structures (Figs 8, 9, 12, 17). Unlike in Cragandhara gen. n., in the latter genus, those processes are situated laterally and the vesica is elongate, bearing granulation and lacks cornuti whereas the new genus has a single robust cornutus in the short and membranous vesica. The vesica configuration of Cragandhara gen. n. is also vaguely reminiscent of certain species of Gandhara, namely G. vietnamica Dubatolov, 2012 (Fig. 13), but distinguished by the broader main chamber and the considerably larger cornutus situated on the lateral diverticulum whereas Gandhara has a cornutus on the tip of the distal diverticulum. The female genitalia of Cragandhara gen. n. (Figs 15) are reminiscent of Gandhara (Gig. 16) but differ in the membranous posterior section of the corpus bursae bearing a heavily sclerotised pocket anterio-laterally whereas the posterior section of the corpus bursae of Gandhara is gelatinous and bears a weakly sclerotised pocket anterio-laterally.	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFCCFF83FF751EF8FE00F876.taxon	description	Description. External morphology of adults (Figs 1 – 3). Sexual dimorphism limited: female somewhat larger and with slightly longer forewing and narrower yellow costal stripe than male. Antenna ciliate in both sexes. Head ochreous yellow. Thorax greyish-brown, patagia ochreous yellow, tegula edged with ochreous yellow scales. Forewing distally dilated, its costal margin posteriorly convex. Forewing ground colour greyish-brown or brown, with ochreous yellow costal stripe tapering distally. Forewing cilia ochreous yellow. Hindwing unicolorous pale ochreous yellow. Abdomen pale ochreous yellow proximally and intense ochreous yellow distally. Male genitalia (Figs 10, 11). Uncus moderately long, slender, cylindrical, proximally straight and distally downcurved, with tiny claw-shaped tip. Tegumen short, weakly sclerotised, its arms fused in posterior ¾. Vinculum somewhat shorter than tegumen, with caliciform saccus bearing small trapezoidal tip. Valva lobular, its dorsal section apically rounded. Ventral plate of costa bears longitudinal swollen crest densely covered with spinules at base of distal saccular process. Sacculus narrow, its distal process short but robust, rostrum-shaped, gently upcurved, its dorsal surface setose. Juxta weakly sclerotised, inverted V-shaped. Anellus membranous, bearing a couple of lateral dense bunches of spinules. Phallus short and broad, somewhat medially dilated, with two distal-lateral, weakly sclerotised, longitudinally rugose, ribbon-like processes. Vesica short and broad, membranous, with several broad conical diverticula, subbasal diverticulum bearing long and robust, spike-like cornutus. Female genitalia (Fig. 15). Papilla analis trapezoidal with rounded corners, weakly setose. Apophyses long and thin, equal in length. Postvaginal area long and broad, with weakly sclerotised plate. Ostium bursae broad. Antrum short, caliciform, gelatinous. Ductus bursae tubular, narrow and short. Corpus bursae pyriform and membranous, its posterior section bearing conical heavily sclerotised lateral pocket on left side. Anterior section of corpus bursae strongly dilated and bearing elliptical signum medially. Appendix bursae shortly conical, membranous, situated postero-laterally.	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFCCFF83FF751EF8FE00F876.taxon	distribution	Distribution. Both known species of the genus are distributed in Northeast India.	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFCCFF83FF751EF8FE00F876.taxon	etymology	Etymology. The new genus name is an aggregate of the genus-group names Cragia and Gandhara. Gender is feminine.	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFC9FF86FF751CA0FF5EFDB2.taxon	description	(Figs 1, 2, 10, 15)	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFC9FF86FF751CA0FF5EFDB2.taxon	materials_examined	Type material. Holotype (Figs 1, 10): male, [NE India, north of West Bengal] “ Gopaldhara, | Darjeeling, | 3440 – 5800 '. | (H. Stevens). | 22. IX. [19] 18. ” / “ Rothschild | Bequest | B. M. 1939 - 1. ” / QR-code label with unique ID “ NHMUK 010292457 ” | “ Slide | NHMUK 0104331720 ” (NHMUK). Paratype. INDIA: female, [NE India, north of West Bengal] Gopaldhara, Mirik, Sikkim, H. Stevens / Rothschild Bequest B. M. 1939 - 1 / QR-code label with unique ID: NHMUK 010916194, gen. prep. No.: NHMUK 0104331732 (NHMUK).	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFC9FF86FF751CA0FF5EFDB2.taxon	diagnosis	Diagnosis. The forewing length is 11.0 mm in the male holotype and 12.0 mm in the female paratype. The male of C. himalaya sp. n. is externally distinguished from C. khasia sp. n. by its slightly smaller size, and the narrower forewing with a less distally convex costal margin and a narrower ochreous yellow costal stripe. The male genital capsule of C. himalaya sp. n. differs from C. khasia sp. n. in the larger spinulose bunches of the anellus, the narrower distal section of the valva, the longer setose crest of the ventral plate of the costa, and the longer distal saccular process. The phallus of the new species is somewhat proximally narrower than in C. khasia sp. n. Compared to C. khasia sp. n., the vesica of C. himalaya sp. n. has somewhat shorter and narrower diverticula, and bears a slightly longer and thinner cornutus. The female genitalia of the new species were characterised and diagnosed above under the new genus.	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFC9FF86FF751CA0FF5EFDB2.taxon	distribution	Distribution. Northeast India (north of West Bengal).	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFC9FF86FF751CA0FF5EFDB2.taxon	etymology	Etymology. The specific epithet is derived from the Himalaya mountain system and refers to the occurrence of the new species in it. The name is a noon in nominative singular in apposition to the generic name.	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFCBFF86FF751EDAFABEFB4D.taxon	description	(Figs 3, 11)	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFCBFF86FF751EDAFABEFB4D.taxon	materials_examined	Type material. Holotype (Figs 3, 11): male, [NE India, Meghalaya, Khasi Hills] “ Khasis, | Feb. 1897. | Nat. Coll. ” / “ Rothschild | Bequest | B. M. 1939 - 1. ” / QR-code label with unique ID “ NHMUK 010292439 ” | “ Slide | NHMUK 0104331715 ” (NHMUK).	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFCBFF86FF751EDAFABEFB4D.taxon	diagnosis	Diagnosis. The forewing length is 12.0 mm in the male holotype. The male of C. khasia sp. n. externally differs from C. himalaya sp. n. in its somewhat larger size, the broader forewing with a more distally convex costal margin, and the broader ochreous yellow costal stripe. The detailed comparison of the male genitalia of the two species is provided above in the diagnosis of C. himalaya sp. n. The female is unknown.	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFCBFF86FF751EDAFABEFB4D.taxon	distribution	Distribution. Northeast India (Meghalaya).	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
03CC0D78FFCBFF86FF751EDAFABEFB4D.taxon	etymology	Etymology. The specific epithet is derived from Khasi Hills, where the new species is found.	en	Volynkin, Anton V. (2023): Cragandhara, a new genus for two new species from India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina 69: 84-90, DOI: 10.37828/em.2023.69.8, URL: http://dx.doi.org/10.37828/em.2023.69.8
