taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962899/files/figure.png	https://doi.org/10.5281/zenodo.16962899	Figures 1–4: A. argoubi sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4. Female paratopotype in dorsal (3) and ventral (4) views. Scale bars: 10 mm (1–2, 3–4).	Figures 1–4: A. argoubi sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4. Female paratopotype in dorsal (3) and ventral (4) views. Scale bars: 10 mm (1–2, 3–4).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962901/files/figure.png	https://doi.org/10.5281/zenodo.16962901	Figures 5–12: A. argoubi sp. n. Figures 5, 8–12. Male holotype, carapace and tergites I–III (5), coxosternal area and sternites (8), right legs I– IV, retrolateral aspect (9–12 respectively). Figures 6–7. Female paratopotype, carapace and tergites I–II (6), coxosternal area and sternites (7).	Figures 5–12: A. argoubi sp. n. Figures 5, 8–12. Male holotype, carapace and tergites I–III (5), coxosternal area and sternites (8), right legs I– IV, retrolateral aspect (9–12 respectively). Figures 6–7. Female paratopotype, carapace and tergites I–II (6), coxosternal area and sternites (7).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962903/files/figure.png	https://doi.org/10.5281/zenodo.16962903	Figures 13–18: A. argoubi sp. n. Figures 13–15. Male holotype, metasoma and telson lateral (13), dorsal (14), and ventral (15) views. Figures 16–18. Female paratopotype, metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bars: 10 mm (13–15, 16–18).	Figures 13–18: A. argoubi sp. n. Figures 13–15. Male holotype, metasoma and telson lateral (13), dorsal (14), and ventral (15) views. Figures 16–18. Female paratopotype, metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bars: 10 mm (13–15, 16–18).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962905/files/figure.png	https://doi.org/10.5281/zenodo.16962905	Figures 19–40: A. argoubi sp. n., pedipalp segments. Figures 19–29. Male holotype, chela dorsal (19), external (20) and ventral (21) views. Patella dorsal (22), external (23) and ventral (24) views. Trochanter and femur internal (25), dorsal (26) and ventral (27) views. Movable (28) and fixed (29) finger dentition. Figures 30–40. Female paratopotype, chela dorsal (30), external (31) and ventral (32) views. Patella dorsal (33), external (34) and ventral (35) views. Trochanter and femur internal (36), dorsal (37) and ventral (38) views. Movable (39) and fixed (40) finger dentition. Trichobothrial pattern is indicated by white circles (19–23, 25–26).	Figures 19–40: A. argoubi sp. n., pedipalp segments. Figures 19–29. Male holotype, chela dorsal (19), external (20) and ventral (21) views. Patella dorsal (22), external (23) and ventral (24) views. Trochanter and femur internal (25), dorsal (26) and ventral (27) views. Movable (28) and fixed (29) finger dentition. Figures 30–40. Female paratopotype, chela dorsal (30), external (31) and ventral (32) views. Patella dorsal (33), external (34) and ventral (35) views. Trochanter and femur internal (36), dorsal (37) and ventral (38) views. Movable (39) and fixed (40) finger dentition. Trichobothrial pattern is indicated by white circles (19–23, 25–26).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962907/files/figure.png	https://doi.org/10.5281/zenodo.16962907	Figure 41. A. sergenti, male from Morocco, Anti Atlas mts., 62 km SE of Agadir, in vivo habitus.	Figure 41. A. sergenti, male from Morocco, Anti Atlas mts., 62 km SE of Agadir, in vivo habitus.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962909/files/figure.png	https://doi.org/10.5281/zenodo.16962909	Figures 42–45: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 42–43. Male in dorsal (42) and ventral (43) views. Figures 44–45. Female in dorsal (44) and ventral (45) views. Scale bars: 10 mm.	Figures 42–45: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 42–43. Male in dorsal (42) and ventral (43) views. Figures 44–45. Female in dorsal (44) and ventral (45) views. Scale bars: 10 mm.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962911/files/figure.png	https://doi.org/10.5281/zenodo.16962911	Figures 46–52: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 48–50. Male, coxosternal area and sternites (48), carapace and tergites I–III (49), metasoma V and telson lateral (50). Figures 46–47, 51–59. Female, carapace and tergites I–III (46), coxosternal area and sternites III–V (47), metasoma V and telson lateral (51), metasoma I–III dorsal (52).	Figures 46–52: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 48–50. Male, coxosternal area and sternites (48), carapace and tergites I–III (49), metasoma V and telson lateral (50). Figures 46–47, 51–59. Female, carapace and tergites I–III (46), coxosternal area and sternites III–V (47), metasoma V and telson lateral (51), metasoma I–III dorsal (52).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962913/files/figure.png	https://doi.org/10.5281/zenodo.16962913	Figures 53–58: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 53–55. Female, metasoma and telson lateral (53), ventral (54), and dorsal (55) views. Figures 56–58. Male, metasoma and telson lateral (56), ventral (57), and dorsal (58) views. Scale bars: 10 mm.	Figures 53–58: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 53–55. Female, metasoma and telson lateral (53), ventral (54), and dorsal (55) views. Figures 56–58. Male, metasoma and telson lateral (56), ventral (57), and dorsal (58) views. Scale bars: 10 mm.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962915/files/figure.png	https://doi.org/10.5281/zenodo.16962915	Figures 59–76: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 59–70: pedipalp segments. Figures 59–60, 69–70. Male, chela dorsal (59), external (60). Movable finger (69) and fixed (70) dentition. Figures 61–68. Female, chela dorsal (61), external (62) and ventral (63) views. Patella dorsal (64) and external (65) views. Trochanter and femur internal (66) and dorsal (67). Movable finger dentition (69). Trichobothrial pattern is indicated by white circles (61–67). Figures 71–76. Left legs I–IV, retrolateral aspect in male (71–74) and right legs III–IV, retrolateral aspect in female (75–76).	Figures 59–76: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 59–70: pedipalp segments. Figures 59–60, 69–70. Male, chela dorsal (59), external (60). Movable finger (69) and fixed (70) dentition. Figures 61–68. Female, chela dorsal (61), external (62) and ventral (63) views. Patella dorsal (64) and external (65) views. Trochanter and femur internal (66) and dorsal (67). Movable finger dentition (69). Trichobothrial pattern is indicated by white circles (61–67). Figures 71–76. Left legs I–IV, retrolateral aspect in male (71–74) and right legs III–IV, retrolateral aspect in female (75–76).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962919/files/figure.png	https://doi.org/10.5281/zenodo.16962919	Figures 77–83: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Left hemispermatophores from two males. Figures 77, 80. Whole hemispermatophore, convex views. Figures 78, 81. Capsule region, posterior views. Figures 79, 82. Capsule region, convex views. Figure 83. Capsule region, anterior view. Scale bars: 1 mm (77, 80), 500 μm (78–79, 81–83).	Figures 77–83: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Left hemispermatophores from two males. Figures 77, 80. Whole hemispermatophore, convex views. Figures 78, 81. Capsule region, posterior views. Figures 79, 82. Capsule region, convex views. Figure 83. Capsule region, anterior view. Scale bars: 1 mm (77, 80), 500 μm (78–79, 81–83).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962924/files/figure.png	https://doi.org/10.5281/zenodo.16962924	Figures 84–87: A. stockmanni sp. n., male holotype. Figures 84–86: Right hemispermatophore in convex view (84), and hemispermatophore capsule in posterior (85) and convex (86) views. Scale bars: 1 mm (84), 500 μm (85–86). Figure 87. In vivo habitus (photographed on different substrate in laboratory).	Figures 84–87: A. stockmanni sp. n., male holotype. Figures 84–86: Right hemispermatophore in convex view (84), and hemispermatophore capsule in posterior (85) and convex (86) views. Scale bars: 1 mm (84), 500 μm (85–86). Figure 87. In vivo habitus (photographed on different substrate in laboratory).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962926/files/figure.png	https://doi.org/10.5281/zenodo.16962926	Figures 88–91: A. stockmanni sp. n. Figures 88–91. Male holotype in dorsal (88) and ventral (89) views. Figures 90–91. Female paratopotype in dorsal (90) and ventral (91) views. Scale bars: 10 mm (88–89, 90–91).	Figures 88–91: A. stockmanni sp. n. Figures 88–91. Male holotype in dorsal (88) and ventral (89) views. Figures 90–91. Female paratopotype in dorsal (90) and ventral (91) views. Scale bars: 10 mm (88–89, 90–91).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962928/files/figure.png	https://doi.org/10.5281/zenodo.16962928	Figures 92–95: A. stockmanni sp. n. Figures 92, 94, 96–99. Male holotype, carapace and tergites I–III (92), coxosternal area and sternites (94), left legs I–IV, retrolateral aspect (96–99 respectively). Figures 93, 95. Female paratopotype, carapace and tergites I–IV (93), coxosternal area and sternites (95).	Figures 92–95: A. stockmanni sp. n. Figures 92, 94, 96–99. Male holotype, carapace and tergites I–III (92), coxosternal area and sternites (94), left legs I–IV, retrolateral aspect (96–99 respectively). Figures 93, 95. Female paratopotype, carapace and tergites I–IV (93), coxosternal area and sternites (95).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962930/files/figure.png	https://doi.org/10.5281/zenodo.16962930	Figures 100–107: A. stockmanni sp. n. Figures 100, 102–104. Male holotype. metasoma IV–V and telson lateral (100), metasoma and telson lateral (102), dorsal (103), and ventral (104) views. Figures 101, 105–107. Female paratopotype, metasoma IV–V and telson lateral (101), metasoma and telson lateral (105), dorsal (106), and ventral (107) views. Scale bars: 10 mm (102–104, 105–107).	Figures 100–107: A. stockmanni sp. n. Figures 100, 102–104. Male holotype. metasoma IV–V and telson lateral (100), metasoma and telson lateral (102), dorsal (103), and ventral (104) views. Figures 101, 105–107. Female paratopotype, metasoma IV–V and telson lateral (101), metasoma and telson lateral (105), dorsal (106), and ventral (107) views. Scale bars: 10 mm (102–104, 105–107).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962932/files/figure.png	https://doi.org/10.5281/zenodo.16962932	Figures 108–129: A. stockmanni sp. n., pedipalp segments. Figures 108–118. Male holotype, chela dorsal (108), external (109) and ventral (110) views. Patella dorsal (111), external (112) and ventral (113) views. Trochanter and femur internal (114), dorsal (115) and ventral (116) views. Movable (117) and fixed (118) finger dentition. Figures 119–129. Female paratopotype, chela dorsal (119), external (120) and ventral (121) views. Patella dorsal (122), external (123) and ventral (124) views. Trochanter and femur internal (125), dorsal (126) and ventral (127)	Figures 108–129: A. stockmanni sp. n., pedipalp segments. Figures 108–118. Male holotype, chela dorsal (108), external (109) and ventral (110) views. Patella dorsal (111), external (112) and ventral (113) views. Trochanter and femur internal (114), dorsal (115) and ventral (116) views. Movable (117) and fixed (118) finger dentition. Figures 119–129. Female paratopotype, chela dorsal (119), external (120) and ventral (121) views. Patella dorsal (122), external (123) and ventral (124) views. Trochanter and femur internal (125), dorsal (126) and ventral (127)	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962934/files/figure.png	https://doi.org/10.5281/zenodo.16962934	Figures 130–131. A. vaneki sp. n., male holotype in dorsal (130) and ventral (131) views. Scale bar: 10 mm.	Figures 130–131. A. vaneki sp. n., male holotype in dorsal (130) and ventral (131) views. Scale bar: 10 mm.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962936/files/figure.png	https://doi.org/10.5281/zenodo.16962936	Figures 132–137. A. vaneki sp. n., male holotype, carapace and tergites (132), coxosternal area and sternites (133), metasoma IV–V and telson lateral (134), metasoma and telson lateral (135), dorsal (136), and ventral (137) views. Scale bar: 10 mm (135–137).	Figures 132–137. A. vaneki sp. n., male holotype, carapace and tergites (132), coxosternal area and sternites (133), metasoma IV–V and telson lateral (134), metasoma and telson lateral (135), dorsal (136), and ventral (137) views. Scale bar: 10 mm (135–137).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962938/files/figure.png	https://doi.org/10.5281/zenodo.16962938	Figures 138–152: A. vaneki sp. n., male holotype. Figures 138–148. Pedipalp segments, chela dorsal (138), external (139) and ventral (140) views. Patella dorsal (141), external (142) and ventral (143) views. Trochanter and femur internal (144), dorsal (145) and ventral (146) views. Movable (147) and fixed (148) finger dentition. Figures 149–152. Right legs I–IV, retrolateral aspect.	Figures 138–152: A. vaneki sp. n., male holotype. Figures 138–148. Pedipalp segments, chela dorsal (138), external (139) and ventral (140) views. Patella dorsal (141), external (142) and ventral (143) views. Trochanter and femur internal (144), dorsal (145) and ventral (146) views. Movable (147) and fixed (148) finger dentition. Figures 149–152. Right legs I–IV, retrolateral aspect.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962942/files/figure.png	https://doi.org/10.5281/zenodo.16962942	Figures 153–164: Figures 153–161: Comparative figures of pedipalp chela dorsal and external views of males. Figure 153. A. mauritanicus (Pocock, 1902) from Morocco, Taroudant, Figure 154. A. bourdoni Vachon, 1948 from Morocco, Biougra. Figure 155. A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W. Figure 156–157. A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figure 158–159. A. argoubi sp. n., holotype. Figure 160–162. A. vaneki sp. n., holotype. Figures 162–163. Metasoma I–III in male of A. bourdoni Vachon, 1948 from Morocco, Biougra (162) and male of A. liouvillei (Pallary, 1924) from Morocco, Taroudant (163). Figure 164. Metasoma in ventral view, A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W.	Figures 153–164: Figures 153–161: Comparative figures of pedipalp chela dorsal and external views of males. Figure 153. A. mauritanicus (Pocock, 1902) from Morocco, Taroudant, Figure 154. A. bourdoni Vachon, 1948 from Morocco, Biougra. Figure 155. A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W. Figure 156–157. A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figure 158–159. A. argoubi sp. n., holotype. Figure 160–162. A. vaneki sp. n., holotype. Figures 162–163. Metasoma I–III in male of A. bourdoni Vachon, 1948 from Morocco, Biougra (162) and male of A. liouvillei (Pallary, 1924) from Morocco, Taroudant (163). Figure 164. Metasoma in ventral view, A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962944/files/figure.png	https://doi.org/10.5281/zenodo.16962944	Figures 165–172: Comparative figures of metasoma. Figures 165–168: Metasoma IV–V and telson lateral, A. argoubi sp. n., holotype male (165) and paratopotype female (166), A. stockmanni sp. n., holotype male (167), A. vaneki sp. n., holotype male (168). Figures 169–172: Metasoma ventral, A. argoubi sp. n., holotype male (169), A. sergenti male from Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (170), A. stockmanni sp. n., holotype male (171), and A. vaneki sp. n., holotype male (172).	Figures 165–172: Comparative figures of metasoma. Figures 165–168: Metasoma IV–V and telson lateral, A. argoubi sp. n., holotype male (165) and paratopotype female (166), A. stockmanni sp. n., holotype male (167), A. vaneki sp. n., holotype male (168). Figures 169–172: Metasoma ventral, A. argoubi sp. n., holotype male (169), A. sergenti male from Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (170), A. stockmanni sp. n., holotype male (171), and A. vaneki sp. n., holotype male (172).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962948/files/figure.png	https://doi.org/10.5281/zenodo.16962948	Figures 173–177: Male mitotic metaphases (173, 175, 177) and postpachytene (174, 176) of Androctonus species. Arrows show extra large chromosomes. Figures 173, 174. A. argoubi sp. n. (2n=24, 12 II). Figures 175, 176. A. stockmanni sp. n. (2n=24, 12 II). Figure 177. A. vaneki sp. n. (2n=24). Scale bar: 5 μm.	Figures 173–177: Male mitotic metaphases (173, 175, 177) and postpachytene (174, 176) of Androctonus species. Arrows show extra large chromosomes. Figures 173, 174. A. argoubi sp. n. (2n=24, 12 II). Figures 175, 176. A. stockmanni sp. n. (2n=24, 12 II). Figure 177. A. vaneki sp. n. (2n=24). Scale bar: 5 μm.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFDFF82FED5FC4AD3FAF9EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962955/files/figure.png	https://doi.org/10.5281/zenodo.16962955	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFEFF82FEC5F9C8D6A7F82F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962899/files/figure.png	https://doi.org/10.5281/zenodo.16962899	Figures 1–4: A. argoubi sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4. Female paratopotype in dorsal (3) and ventral (4) views. Scale bars: 10 mm (1–2, 3–4).	Figures 1–4: A. argoubi sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4. Female paratopotype in dorsal (3) and ventral (4) views. Scale bars: 10 mm (1–2, 3–4).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFEFF82FEC5F9C8D6A7F82F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962901/files/figure.png	https://doi.org/10.5281/zenodo.16962901	Figures 5–12: A. argoubi sp. n. Figures 5, 8–12. Male holotype, carapace and tergites I–III (5), coxosternal area and sternites (8), right legs I– IV, retrolateral aspect (9–12 respectively). Figures 6–7. Female paratopotype, carapace and tergites I–II (6), coxosternal area and sternites (7).	Figures 5–12: A. argoubi sp. n. Figures 5, 8–12. Male holotype, carapace and tergites I–III (5), coxosternal area and sternites (8), right legs I– IV, retrolateral aspect (9–12 respectively). Figures 6–7. Female paratopotype, carapace and tergites I–II (6), coxosternal area and sternites (7).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFEFF82FEC5F9C8D6A7F82F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962903/files/figure.png	https://doi.org/10.5281/zenodo.16962903	Figures 13–18: A. argoubi sp. n. Figures 13–15. Male holotype, metasoma and telson lateral (13), dorsal (14), and ventral (15) views. Figures 16–18. Female paratopotype, metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bars: 10 mm (13–15, 16–18).	Figures 13–18: A. argoubi sp. n. Figures 13–15. Male holotype, metasoma and telson lateral (13), dorsal (14), and ventral (15) views. Figures 16–18. Female paratopotype, metasoma and telson lateral (16), dorsal (17), and ventral (18) views. Scale bars: 10 mm (13–15, 16–18).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFEFF82FEC5F9C8D6A7F82F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962905/files/figure.png	https://doi.org/10.5281/zenodo.16962905	Figures 19–40: A. argoubi sp. n., pedipalp segments. Figures 19–29. Male holotype, chela dorsal (19), external (20) and ventral (21) views. Patella dorsal (22), external (23) and ventral (24) views. Trochanter and femur internal (25), dorsal (26) and ventral (27) views. Movable (28) and fixed (29) finger dentition. Figures 30–40. Female paratopotype, chela dorsal (30), external (31) and ventral (32) views. Patella dorsal (33), external (34) and ventral (35) views. Trochanter and femur internal (36), dorsal (37) and ventral (38) views. Movable (39) and fixed (40) finger dentition. Trichobothrial pattern is indicated by white circles (19–23, 25–26).	Figures 19–40: A. argoubi sp. n., pedipalp segments. Figures 19–29. Male holotype, chela dorsal (19), external (20) and ventral (21) views. Patella dorsal (22), external (23) and ventral (24) views. Trochanter and femur internal (25), dorsal (26) and ventral (27) views. Movable (28) and fixed (29) finger dentition. Figures 30–40. Female paratopotype, chela dorsal (30), external (31) and ventral (32) views. Patella dorsal (33), external (34) and ventral (35) views. Trochanter and femur internal (36), dorsal (37) and ventral (38) views. Movable (39) and fixed (40) finger dentition. Trichobothrial pattern is indicated by white circles (19–23, 25–26).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFEFF82FEC5F9C8D6A7F82F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962942/files/figure.png	https://doi.org/10.5281/zenodo.16962942	Figures 153–164: Figures 153–161: Comparative figures of pedipalp chela dorsal and external views of males. Figure 153. A. mauritanicus (Pocock, 1902) from Morocco, Taroudant, Figure 154. A. bourdoni Vachon, 1948 from Morocco, Biougra. Figure 155. A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W. Figure 156–157. A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figure 158–159. A. argoubi sp. n., holotype. Figure 160–162. A. vaneki sp. n., holotype. Figures 162–163. Metasoma I–III in male of A. bourdoni Vachon, 1948 from Morocco, Biougra (162) and male of A. liouvillei (Pallary, 1924) from Morocco, Taroudant (163). Figure 164. Metasoma in ventral view, A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W.	Figures 153–164: Figures 153–161: Comparative figures of pedipalp chela dorsal and external views of males. Figure 153. A. mauritanicus (Pocock, 1902) from Morocco, Taroudant, Figure 154. A. bourdoni Vachon, 1948 from Morocco, Biougra. Figure 155. A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W. Figure 156–157. A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figure 158–159. A. argoubi sp. n., holotype. Figure 160–162. A. vaneki sp. n., holotype. Figures 162–163. Metasoma I–III in male of A. bourdoni Vachon, 1948 from Morocco, Biougra (162) and male of A. liouvillei (Pallary, 1924) from Morocco, Taroudant (163). Figure 164. Metasoma in ventral view, A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFEFF82FEC5F9C8D6A7F82F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962944/files/figure.png	https://doi.org/10.5281/zenodo.16962944	Figures 165–172: Comparative figures of metasoma. Figures 165–168: Metasoma IV–V and telson lateral, A. argoubi sp. n., holotype male (165) and paratopotype female (166), A. stockmanni sp. n., holotype male (167), A. vaneki sp. n., holotype male (168). Figures 169–172: Metasoma ventral, A. argoubi sp. n., holotype male (169), A. sergenti male from Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (170), A. stockmanni sp. n., holotype male (171), and A. vaneki sp. n., holotype male (172).	Figures 165–172: Comparative figures of metasoma. Figures 165–168: Metasoma IV–V and telson lateral, A. argoubi sp. n., holotype male (165) and paratopotype female (166), A. stockmanni sp. n., holotype male (167), A. vaneki sp. n., holotype male (168). Figures 169–172: Metasoma ventral, A. argoubi sp. n., holotype male (169), A. sergenti male from Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (170), A. stockmanni sp. n., holotype male (171), and A. vaneki sp. n., holotype male (172).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFEFF82FEC5F9C8D6A7F82F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962948/files/figure.png	https://doi.org/10.5281/zenodo.16962948	Figures 173–177: Male mitotic metaphases (173, 175, 177) and postpachytene (174, 176) of Androctonus species. Arrows show extra large chromosomes. Figures 173, 174. A. argoubi sp. n. (2n=24, 12 II). Figures 175, 176. A. stockmanni sp. n. (2n=24, 12 II). Figure 177. A. vaneki sp. n. (2n=24). Scale bar: 5 μm.	Figures 173–177: Male mitotic metaphases (173, 175, 177) and postpachytene (174, 176) of Androctonus species. Arrows show extra large chromosomes. Figures 173, 174. A. argoubi sp. n. (2n=24, 12 II). Figures 175, 176. A. stockmanni sp. n. (2n=24, 12 II). Figure 177. A. vaneki sp. n. (2n=24). Scale bar: 5 μm.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFFEFF82FEC5F9C8D6A7F82F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962955/files/figure.png	https://doi.org/10.5281/zenodo.16962955	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8DFC4EFDD4D0BCF92E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962942/files/figure.png	https://doi.org/10.5281/zenodo.16962942	Figures 153–164: Figures 153–161: Comparative figures of pedipalp chela dorsal and external views of males. Figure 153. A. mauritanicus (Pocock, 1902) from Morocco, Taroudant, Figure 154. A. bourdoni Vachon, 1948 from Morocco, Biougra. Figure 155. A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W. Figure 156–157. A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figure 158–159. A. argoubi sp. n., holotype. Figure 160–162. A. vaneki sp. n., holotype. Figures 162–163. Metasoma I–III in male of A. bourdoni Vachon, 1948 from Morocco, Biougra (162) and male of A. liouvillei (Pallary, 1924) from Morocco, Taroudant (163). Figure 164. Metasoma in ventral view, A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W.	Figures 153–164: Figures 153–161: Comparative figures of pedipalp chela dorsal and external views of males. Figure 153. A. mauritanicus (Pocock, 1902) from Morocco, Taroudant, Figure 154. A. bourdoni Vachon, 1948 from Morocco, Biougra. Figure 155. A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W. Figure 156–157. A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figure 158–159. A. argoubi sp. n., holotype. Figure 160–162. A. vaneki sp. n., holotype. Figures 162–163. Metasoma I–III in male of A. bourdoni Vachon, 1948 from Morocco, Biougra (162) and male of A. liouvillei (Pallary, 1924) from Morocco, Taroudant (163). Figure 164. Metasoma in ventral view, A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8DFC4EFDD4D0BCF92E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962955/files/figure.png	https://doi.org/10.5281/zenodo.16962955	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8EFC71F897D6A7F93D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962907/files/figure.png	https://doi.org/10.5281/zenodo.16962907	Figure 41. A. sergenti, male from Morocco, Anti Atlas mts., 62 km SE of Agadir, in vivo habitus.	Figure 41. A. sergenti, male from Morocco, Anti Atlas mts., 62 km SE of Agadir, in vivo habitus.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8EFC71F897D6A7F93D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962909/files/figure.png	https://doi.org/10.5281/zenodo.16962909	Figures 42–45: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 42–43. Male in dorsal (42) and ventral (43) views. Figures 44–45. Female in dorsal (44) and ventral (45) views. Scale bars: 10 mm.	Figures 42–45: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 42–43. Male in dorsal (42) and ventral (43) views. Figures 44–45. Female in dorsal (44) and ventral (45) views. Scale bars: 10 mm.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8EFC71F897D6A7F93D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962911/files/figure.png	https://doi.org/10.5281/zenodo.16962911	Figures 46–52: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 48–50. Male, coxosternal area and sternites (48), carapace and tergites I–III (49), metasoma V and telson lateral (50). Figures 46–47, 51–59. Female, carapace and tergites I–III (46), coxosternal area and sternites III–V (47), metasoma V and telson lateral (51), metasoma I–III dorsal (52).	Figures 46–52: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 48–50. Male, coxosternal area and sternites (48), carapace and tergites I–III (49), metasoma V and telson lateral (50). Figures 46–47, 51–59. Female, carapace and tergites I–III (46), coxosternal area and sternites III–V (47), metasoma V and telson lateral (51), metasoma I–III dorsal (52).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8EFC71F897D6A7F93D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962913/files/figure.png	https://doi.org/10.5281/zenodo.16962913	Figures 53–58: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 53–55. Female, metasoma and telson lateral (53), ventral (54), and dorsal (55) views. Figures 56–58. Male, metasoma and telson lateral (56), ventral (57), and dorsal (58) views. Scale bars: 10 mm.	Figures 53–58: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 53–55. Female, metasoma and telson lateral (53), ventral (54), and dorsal (55) views. Figures 56–58. Male, metasoma and telson lateral (56), ventral (57), and dorsal (58) views. Scale bars: 10 mm.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8EFC71F897D6A7F93D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962915/files/figure.png	https://doi.org/10.5281/zenodo.16962915	Figures 59–76: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 59–70: pedipalp segments. Figures 59–60, 69–70. Male, chela dorsal (59), external (60). Movable finger (69) and fixed (70) dentition. Figures 61–68. Female, chela dorsal (61), external (62) and ventral (63) views. Patella dorsal (64) and external (65) views. Trochanter and femur internal (66) and dorsal (67). Movable finger dentition (69). Trichobothrial pattern is indicated by white circles (61–67). Figures 71–76. Left legs I–IV, retrolateral aspect in male (71–74) and right legs III–IV, retrolateral aspect in female (75–76).	Figures 59–76: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figures 59–70: pedipalp segments. Figures 59–60, 69–70. Male, chela dorsal (59), external (60). Movable finger (69) and fixed (70) dentition. Figures 61–68. Female, chela dorsal (61), external (62) and ventral (63) views. Patella dorsal (64) and external (65) views. Trochanter and femur internal (66) and dorsal (67). Movable finger dentition (69). Trichobothrial pattern is indicated by white circles (61–67). Figures 71–76. Left legs I–IV, retrolateral aspect in male (71–74) and right legs III–IV, retrolateral aspect in female (75–76).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8EFC71F897D6A7F93D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962919/files/figure.png	https://doi.org/10.5281/zenodo.16962919	Figures 77–83: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Left hemispermatophores from two males. Figures 77, 80. Whole hemispermatophore, convex views. Figures 78, 81. Capsule region, posterior views. Figures 79, 82. Capsule region, convex views. Figure 83. Capsule region, anterior view. Scale bars: 1 mm (77, 80), 500 μm (78–79, 81–83).	Figures 77–83: A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Left hemispermatophores from two males. Figures 77, 80. Whole hemispermatophore, convex views. Figures 78, 81. Capsule region, posterior views. Figures 79, 82. Capsule region, convex views. Figure 83. Capsule region, anterior view. Scale bars: 1 mm (77, 80), 500 μm (78–79, 81–83).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8EFC71F897D6A7F93D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962942/files/figure.png	https://doi.org/10.5281/zenodo.16962942	Figures 153–164: Figures 153–161: Comparative figures of pedipalp chela dorsal and external views of males. Figure 153. A. mauritanicus (Pocock, 1902) from Morocco, Taroudant, Figure 154. A. bourdoni Vachon, 1948 from Morocco, Biougra. Figure 155. A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W. Figure 156–157. A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figure 158–159. A. argoubi sp. n., holotype. Figure 160–162. A. vaneki sp. n., holotype. Figures 162–163. Metasoma I–III in male of A. bourdoni Vachon, 1948 from Morocco, Biougra (162) and male of A. liouvillei (Pallary, 1924) from Morocco, Taroudant (163). Figure 164. Metasoma in ventral view, A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W.	Figures 153–164: Figures 153–161: Comparative figures of pedipalp chela dorsal and external views of males. Figure 153. A. mauritanicus (Pocock, 1902) from Morocco, Taroudant, Figure 154. A. bourdoni Vachon, 1948 from Morocco, Biougra. Figure 155. A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W. Figure 156–157. A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figure 158–159. A. argoubi sp. n., holotype. Figure 160–162. A. vaneki sp. n., holotype. Figures 162–163. Metasoma I–III in male of A. bourdoni Vachon, 1948 from Morocco, Biougra (162) and male of A. liouvillei (Pallary, 1924) from Morocco, Taroudant (163). Figure 164. Metasoma in ventral view, A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8EFC71F897D6A7F93D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962944/files/figure.png	https://doi.org/10.5281/zenodo.16962944	Figures 165–172: Comparative figures of metasoma. Figures 165–168: Metasoma IV–V and telson lateral, A. argoubi sp. n., holotype male (165) and paratopotype female (166), A. stockmanni sp. n., holotype male (167), A. vaneki sp. n., holotype male (168). Figures 169–172: Metasoma ventral, A. argoubi sp. n., holotype male (169), A. sergenti male from Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (170), A. stockmanni sp. n., holotype male (171), and A. vaneki sp. n., holotype male (172).	Figures 165–172: Comparative figures of metasoma. Figures 165–168: Metasoma IV–V and telson lateral, A. argoubi sp. n., holotype male (165) and paratopotype female (166), A. stockmanni sp. n., holotype male (167), A. vaneki sp. n., holotype male (168). Figures 169–172: Metasoma ventral, A. argoubi sp. n., holotype male (169), A. sergenti male from Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (170), A. stockmanni sp. n., holotype male (171), and A. vaneki sp. n., holotype male (172).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF1FF8EFC71F897D6A7F93D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962955/files/figure.png	https://doi.org/10.5281/zenodo.16962955	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF2FF9DFC6EF885D110F81A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962924/files/figure.png	https://doi.org/10.5281/zenodo.16962924	Figures 84–87: A. stockmanni sp. n., male holotype. Figures 84–86: Right hemispermatophore in convex view (84), and hemispermatophore capsule in posterior (85) and convex (86) views. Scale bars: 1 mm (84), 500 μm (85–86). Figure 87. In vivo habitus (photographed on different substrate in laboratory).	Figures 84–87: A. stockmanni sp. n., male holotype. Figures 84–86: Right hemispermatophore in convex view (84), and hemispermatophore capsule in posterior (85) and convex (86) views. Scale bars: 1 mm (84), 500 μm (85–86). Figure 87. In vivo habitus (photographed on different substrate in laboratory).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF2FF9DFC6EF885D110F81A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962926/files/figure.png	https://doi.org/10.5281/zenodo.16962926	Figures 88–91: A. stockmanni sp. n. Figures 88–91. Male holotype in dorsal (88) and ventral (89) views. Figures 90–91. Female paratopotype in dorsal (90) and ventral (91) views. Scale bars: 10 mm (88–89, 90–91).	Figures 88–91: A. stockmanni sp. n. Figures 88–91. Male holotype in dorsal (88) and ventral (89) views. Figures 90–91. Female paratopotype in dorsal (90) and ventral (91) views. Scale bars: 10 mm (88–89, 90–91).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF2FF9DFC6EF885D110F81A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962928/files/figure.png	https://doi.org/10.5281/zenodo.16962928	Figures 92–95: A. stockmanni sp. n. Figures 92, 94, 96–99. Male holotype, carapace and tergites I–III (92), coxosternal area and sternites (94), left legs I–IV, retrolateral aspect (96–99 respectively). Figures 93, 95. Female paratopotype, carapace and tergites I–IV (93), coxosternal area and sternites (95).	Figures 92–95: A. stockmanni sp. n. Figures 92, 94, 96–99. Male holotype, carapace and tergites I–III (92), coxosternal area and sternites (94), left legs I–IV, retrolateral aspect (96–99 respectively). Figures 93, 95. Female paratopotype, carapace and tergites I–IV (93), coxosternal area and sternites (95).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF2FF9DFC6EF885D110F81A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962930/files/figure.png	https://doi.org/10.5281/zenodo.16962930	Figures 100–107: A. stockmanni sp. n. Figures 100, 102–104. Male holotype. metasoma IV–V and telson lateral (100), metasoma and telson lateral (102), dorsal (103), and ventral (104) views. Figures 101, 105–107. Female paratopotype, metasoma IV–V and telson lateral (101), metasoma and telson lateral (105), dorsal (106), and ventral (107) views. Scale bars: 10 mm (102–104, 105–107).	Figures 100–107: A. stockmanni sp. n. Figures 100, 102–104. Male holotype. metasoma IV–V and telson lateral (100), metasoma and telson lateral (102), dorsal (103), and ventral (104) views. Figures 101, 105–107. Female paratopotype, metasoma IV–V and telson lateral (101), metasoma and telson lateral (105), dorsal (106), and ventral (107) views. Scale bars: 10 mm (102–104, 105–107).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF2FF9DFC6EF885D110F81A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962932/files/figure.png	https://doi.org/10.5281/zenodo.16962932	Figures 108–129: A. stockmanni sp. n., pedipalp segments. Figures 108–118. Male holotype, chela dorsal (108), external (109) and ventral (110) views. Patella dorsal (111), external (112) and ventral (113) views. Trochanter and femur internal (114), dorsal (115) and ventral (116) views. Movable (117) and fixed (118) finger dentition. Figures 119–129. Female paratopotype, chela dorsal (119), external (120) and ventral (121) views. Patella dorsal (122), external (123) and ventral (124) views. Trochanter and femur internal (125), dorsal (126) and ventral (127)	Figures 108–129: A. stockmanni sp. n., pedipalp segments. Figures 108–118. Male holotype, chela dorsal (108), external (109) and ventral (110) views. Patella dorsal (111), external (112) and ventral (113) views. Trochanter and femur internal (114), dorsal (115) and ventral (116) views. Movable (117) and fixed (118) finger dentition. Figures 119–129. Female paratopotype, chela dorsal (119), external (120) and ventral (121) views. Patella dorsal (122), external (123) and ventral (124) views. Trochanter and femur internal (125), dorsal (126) and ventral (127)	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF2FF9DFC6EF885D110F81A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962944/files/figure.png	https://doi.org/10.5281/zenodo.16962944	Figures 165–172: Comparative figures of metasoma. Figures 165–168: Metasoma IV–V and telson lateral, A. argoubi sp. n., holotype male (165) and paratopotype female (166), A. stockmanni sp. n., holotype male (167), A. vaneki sp. n., holotype male (168). Figures 169–172: Metasoma ventral, A. argoubi sp. n., holotype male (169), A. sergenti male from Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (170), A. stockmanni sp. n., holotype male (171), and A. vaneki sp. n., holotype male (172).	Figures 165–172: Comparative figures of metasoma. Figures 165–168: Metasoma IV–V and telson lateral, A. argoubi sp. n., holotype male (165) and paratopotype female (166), A. stockmanni sp. n., holotype male (167), A. vaneki sp. n., holotype male (168). Figures 169–172: Metasoma ventral, A. argoubi sp. n., holotype male (169), A. sergenti male from Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (170), A. stockmanni sp. n., holotype male (171), and A. vaneki sp. n., holotype male (172).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF2FF9DFC6EF885D110F81A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962948/files/figure.png	https://doi.org/10.5281/zenodo.16962948	Figures 173–177: Male mitotic metaphases (173, 175, 177) and postpachytene (174, 176) of Androctonus species. Arrows show extra large chromosomes. Figures 173, 174. A. argoubi sp. n. (2n=24, 12 II). Figures 175, 176. A. stockmanni sp. n. (2n=24, 12 II). Figure 177. A. vaneki sp. n. (2n=24). Scale bar: 5 μm.	Figures 173–177: Male mitotic metaphases (173, 175, 177) and postpachytene (174, 176) of Androctonus species. Arrows show extra large chromosomes. Figures 173, 174. A. argoubi sp. n. (2n=24, 12 II). Figures 175, 176. A. stockmanni sp. n. (2n=24, 12 II). Figure 177. A. vaneki sp. n. (2n=24). Scale bar: 5 μm.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFF2FF9DFC6EF885D110F81A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962955/files/figure.png	https://doi.org/10.5281/zenodo.16962955	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFE1FF9FFC0FF9E1D635FA6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962934/files/figure.png	https://doi.org/10.5281/zenodo.16962934	Figures 130–131. A. vaneki sp. n., male holotype in dorsal (130) and ventral (131) views. Scale bar: 10 mm.	Figures 130–131. A. vaneki sp. n., male holotype in dorsal (130) and ventral (131) views. Scale bar: 10 mm.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFE1FF9FFC0FF9E1D635FA6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962936/files/figure.png	https://doi.org/10.5281/zenodo.16962936	Figures 132–137. A. vaneki sp. n., male holotype, carapace and tergites (132), coxosternal area and sternites (133), metasoma IV–V and telson lateral (134), metasoma and telson lateral (135), dorsal (136), and ventral (137) views. Scale bar: 10 mm (135–137).	Figures 132–137. A. vaneki sp. n., male holotype, carapace and tergites (132), coxosternal area and sternites (133), metasoma IV–V and telson lateral (134), metasoma and telson lateral (135), dorsal (136), and ventral (137) views. Scale bar: 10 mm (135–137).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFE1FF9FFC0FF9E1D635FA6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962938/files/figure.png	https://doi.org/10.5281/zenodo.16962938	Figures 138–152: A. vaneki sp. n., male holotype. Figures 138–148. Pedipalp segments, chela dorsal (138), external (139) and ventral (140) views. Patella dorsal (141), external (142) and ventral (143) views. Trochanter and femur internal (144), dorsal (145) and ventral (146) views. Movable (147) and fixed (148) finger dentition. Figures 149–152. Right legs I–IV, retrolateral aspect.	Figures 138–152: A. vaneki sp. n., male holotype. Figures 138–148. Pedipalp segments, chela dorsal (138), external (139) and ventral (140) views. Patella dorsal (141), external (142) and ventral (143) views. Trochanter and femur internal (144), dorsal (145) and ventral (146) views. Movable (147) and fixed (148) finger dentition. Figures 149–152. Right legs I–IV, retrolateral aspect.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFE1FF9FFC0FF9E1D635FA6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962942/files/figure.png	https://doi.org/10.5281/zenodo.16962942	Figures 153–164: Figures 153–161: Comparative figures of pedipalp chela dorsal and external views of males. Figure 153. A. mauritanicus (Pocock, 1902) from Morocco, Taroudant, Figure 154. A. bourdoni Vachon, 1948 from Morocco, Biougra. Figure 155. A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W. Figure 156–157. A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figure 158–159. A. argoubi sp. n., holotype. Figure 160–162. A. vaneki sp. n., holotype. Figures 162–163. Metasoma I–III in male of A. bourdoni Vachon, 1948 from Morocco, Biougra (162) and male of A. liouvillei (Pallary, 1924) from Morocco, Taroudant (163). Figure 164. Metasoma in ventral view, A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W.	Figures 153–164: Figures 153–161: Comparative figures of pedipalp chela dorsal and external views of males. Figure 153. A. mauritanicus (Pocock, 1902) from Morocco, Taroudant, Figure 154. A. bourdoni Vachon, 1948 from Morocco, Biougra. Figure 155. A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W. Figure 156–157. A. sergenti from Morocco, Tafengha, 2 km S Irherm, 30.073246°N 8.461766°W. Figure 158–159. A. argoubi sp. n., holotype. Figure 160–162. A. vaneki sp. n., holotype. Figures 162–163. Metasoma I–III in male of A. bourdoni Vachon, 1948 from Morocco, Biougra (162) and male of A. liouvillei (Pallary, 1924) from Morocco, Taroudant (163). Figure 164. Metasoma in ventral view, A. gonneti Vachon, 1948 from Morocco El Ouatia env., 28.53°N 11.29°W.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFE1FF9FFC0FF9E1D635FA6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962944/files/figure.png	https://doi.org/10.5281/zenodo.16962944	Figures 165–172: Comparative figures of metasoma. Figures 165–168: Metasoma IV–V and telson lateral, A. argoubi sp. n., holotype male (165) and paratopotype female (166), A. stockmanni sp. n., holotype male (167), A. vaneki sp. n., holotype male (168). Figures 169–172: Metasoma ventral, A. argoubi sp. n., holotype male (169), A. sergenti male from Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (170), A. stockmanni sp. n., holotype male (171), and A. vaneki sp. n., holotype male (172).	Figures 165–172: Comparative figures of metasoma. Figures 165–168: Metasoma IV–V and telson lateral, A. argoubi sp. n., holotype male (165) and paratopotype female (166), A. stockmanni sp. n., holotype male (167), A. vaneki sp. n., holotype male (168). Figures 169–172: Metasoma ventral, A. argoubi sp. n., holotype male (169), A. sergenti male from Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (170), A. stockmanni sp. n., holotype male (171), and A. vaneki sp. n., holotype male (172).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFE1FF9FFC0FF9E1D635FA6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962948/files/figure.png	https://doi.org/10.5281/zenodo.16962948	Figures 173–177: Male mitotic metaphases (173, 175, 177) and postpachytene (174, 176) of Androctonus species. Arrows show extra large chromosomes. Figures 173, 174. A. argoubi sp. n. (2n=24, 12 II). Figures 175, 176. A. stockmanni sp. n. (2n=24, 12 II). Figure 177. A. vaneki sp. n. (2n=24). Scale bar: 5 μm.	Figures 173–177: Male mitotic metaphases (173, 175, 177) and postpachytene (174, 176) of Androctonus species. Arrows show extra large chromosomes. Figures 173, 174. A. argoubi sp. n. (2n=24, 12 II). Figures 175, 176. A. stockmanni sp. n. (2n=24, 12 II). Figure 177. A. vaneki sp. n. (2n=24). Scale bar: 5 μm.	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
03CC2222FFE1FF9FFC0FF9E1D635FA6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16962955/files/figure.png	https://doi.org/10.5281/zenodo.16962955	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	Figures 178–179: Confirmed distribution of specimens of dark species of genus Androctonus in Morocco and Western Sahara examined for this study and deposited in the first author collection (178) and the locality of A. sergenti in Morocco, Anti Atlas mts., 62 km SE of Agadir, 30.05°N 9.07°W (179).	2025-12-31	Kovařík, František;Lowe, Graeme;Šťáhlavský, František		Zenodo	biologists	Kovařík, František;Lowe, Graeme;Šťáhlavský, František			
