taxonID	type	description	language	source
03CD3E7A7115A717FF60AA9FD1A9F76F.taxon	materials_examined	Type material. Holotype. Adult male, complete specimen with large mite attached under right wing, preserved in 14 x 11 x 2 mm piece of amber (Eocene, ~ 45 – 40 Ma, Baltic amber, Gulf of Gdańsk; MAI- 4295 a; Fig. 3 A); animal syninclusions: Acari (2 ind., MAI- 4295 a), Mycetophilidae (1 ind., MAI- 4295 b), Aphidoidea (1 ind., MAI- 4295).	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7115A717FF60AA9FD1A9F76F.taxon	etymology	Derivatio nominis. Adjective in feminine form derived from the Latin noun ‘ electrum ’, amber.	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7115A717FF60AA9FD1A9F76F.taxon	diagnosis	Diagnosis. Antennal flagellum composed of 13 segments, flagellomeres 11 – 13 fused in part. Anal point with slender distal section and strong crests. Superior volsella elongated, with round apex, bearing 2 setae. Stem of median volsella club-shaped, with wide pectinate and foliate lamellae. Inferior volsella arcuate, tapering to pointed apex armed with 2 stout setae.	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7115A717FF60AA9FD1A9F76F.taxon	description	Description. Adult male (n = 1) Total length c. 1.5 mm; wing length c. 890 µm. Head (Fig. 3 C – E). Eyes bare, small, ovoid, broadly separated. Frontal tubercles conical, c. 10 µm long. Antennal flagellum composed of 13 segments of which 11 are well discernible, flagellomeres 11 – 13 fused in part, AR 0.76 (when flagellum measured as 11 - segmented), AR 0.62 (as 12 - segmented), AR 0.50 (as 13 - segmented); plume fully developed (Fig. 3 D, E). Length of palpomeres 2 – 5 (in µm): c. 30, 64, 84, 133. Clypeals present, but cannot be counted. Thorax (Fig. 3 B). Ac at least 11, Dc at least 7 - 8 on each side, Pa 1 on each side, Scts at least 4. Wing (Fig. 3 F). Slender, with anal lobe weak, broadest at 2 / 3 length; width: 285 µm, length / width ratio: 3.12. Sc ending slightly distal of FCu, R 2 + 3 fading, poorly visible. RM slightly oblique relative to R. FCu placed well distally of RM; VR Cu 1.38. Veins ending as follows (in order from base to tip): An, Sc, Cu 1, R 1, R 4 + 5 and M 3 + 4, M 1 + 2. Wing covered with dense macrotrichia in distal half at least. Legs. Tibial apices including combs of mid and hind legs weakly visible. Spurs not observed on forelegs. Only one spur visible on mid leg (c. 20 µm long) and hind leg (c. 28 µm long). Sensilla chaetica on ta 1 of p 2 not observed. Lengths of leg segments and leg ratios in Table 1. Hypopygium (Fig. 4 A – D). Gonostylus c. 50 Μm long, shorter than gonocoxite, broadest at mid-length, tapering to blunt apex armed with strong apical setae. Anal tergite subtriangular, with several median setae and at least 7 posterolateral setae on each side of anal point (poorly visible on photographs). Anal point broad at base, distinctly narrowed distally, with slender distal section and strong crests tapering towards anal point tip. Superior volsella elongated, with broadened round apex, bearing 2 setae on median margin. Digitus absent. Stem of median volsella club-shaped, slightly broadened apically, c. 20 µm long, with several setiform and 4 – 5 wide pectinate and foliate lamellae (Fig. 4 A, B, D). Inferior volsella reaching 2 / 3 length of gonostylus at most, somewhat arcuate, tapering to pointed posteromedially directed apex, with several strong setae including 2 stout setae on apex (Fig. 4 A – C).	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7115A717FF60AA9FD1A9F76F.taxon	discussion	Remarks. Stempellinella electra is the third fossil species of the genus found in Eocene amber, along with S. bicorna Seredszus et Wichard, 2007 and S. ivanovae Giłka et Zakrzewska, 2014 (Seredszus & Wichard 2007, Zakrzewska & Giłka 2014). The new species fits well the emended generic diagnosis for Stempellinella Brundin, 1947 (Ekrem 2007) — the adult male has bare ovoid and broadly separated eyes (Fig. 3 C), the wing vein R 4 + 5 ending opposite to M 3 + 4 (Fig. 3 F), the broadened superior volsella, and the gonostylus shorter than the gonocoxite (Fig. 4). Significant differences in the hypopygium structure have been observed between the three fossil Stempellinella species. S. electra is distinct in having a long anal point, broad at base and narrowed distally (in contrast to the short anal point in S. bicorna) bearing strong anal point crests (Fig. 4 B, C) — not observed in the two fossil relatives, but resembling those known from several extant species (cf. Ekrem 2007). The best diagnostic characters for S. electra is the superior volsella elongated and broadened apically, the inferior volsella arcuate and pointed, and the median volsella bearing several setiform and 4 – 5 wide pectinate and foliate lamellae (Fig. 4 A, B, D) — the combination of shapes not recorded neither in fossil or extant Stempellinella (the pectinate lamellae are known only from S. reissi Casas et Vilchez-Quero, 1991). It is worth noting that we observed a tendency to formation of a fully-segmented antennal flagellum in Stempellinella electra, similar to that known from S. ivanovae. However, when comparing this character in these two species, the higher number of incompletely fused flagellomeres may indicate more advanced state in S. electra (flagellomeres 11 – 13 fused in part, Fig. 3 D, E) than that plesiomorphic state known from S. ivanovae (only flagellomeres 12 and 13 fused in part; Zakrzewska & Giłka 2014: fig. 3 C).	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7112A717FF60A889D6E7F5B7.taxon	materials_examined	Material examined. Adult male (tarsi of left legs separated or not visible) preserved in 50 x 20 x 12 mm piece of amber (Eocene, ~ 45 – 40 Ma, Baltic amber, Gulf of Gdańsk; MAI- 1140); animal syninclusions: Diptera (numerous ind.): Chironomidae, Ceratopogonidae, Psychodidae, Empididae + Thysanoptera (1 ind.).	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7112A717FF60A889D6E7F5B7.taxon	discussion	Remarks. The structure of the hypopygial anal point, the presence of the short nipple-like process on the gonostylus, the wing venation pattern (except the presence of a thin and semitransparent vein R 2 + 3) are the characters found in the presently examined individual, which indicate the recently described species — Stempellinella ivanovae. Unfortunately, the remaining diagnostic structures are too weakly observable to complement description of the species (see Zakrzewska & Giłka 2014). The total number of all known Stempellinella increases now to 22 (see the list below), including the three fossil and 18 extant species listed by Ekrem (2007) + 1 species presently proposed to be transferred from the genus Stempellina Thienemann et Bause, 1913. S. sofiae, known from adult male (Fusari & Lamas 2014), fits the generic diagnosis of Stempellinella, including several crucial characters discussed above (e. g. gonostylus distinctly shorter than gonocoxite; superior volsella rounded, extensive), thus is listed in a new combination.	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7111A714FF60AFA9D120F71A.taxon	materials_examined	Material examined. Holotype (MAI- 4356) (see Giłka 2011 a).	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7111A71BFF60A89CD0CCF0D5.taxon	materials_examined	Type material. Holotype. Adult male (tarsi of left fore and hind leg missing, tarsi of mid legs broken and separated) preserved in 21 x 13 x 1 mm piece of amber (Eocene, ~ 45 – 40 Ma, Baltic amber, Gulf of Gdańsk; MAI- 415 a; Fig. 5 A); animal syninclusions: Ceratopogonidae, Formicidae, Mordellidae, Thysanoptera, Psocoptera (all as single ind., MAI- 415).	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7111A71BFF60A89CD0CCF0D5.taxon	etymology	Derivatio nominis. Adjective derived from the Latin noun ‘ glaesum ’, amber.	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7111A71BFF60A89CD0CCF0D5.taxon	diagnosis	Diagnosis. Frontal tubercles stout, cylindrical. Antenna with flagellomeres 10 – 11 fused in part and 12 – 13 completely fused. Wing vein FCu placed far distally of RM (VR Cu 1.52). Anal point wide at base, distinctly narrowed subapically, tapering to pointed apex. Superior volsella with 2 setae on apex. Digitus long, extending beyond superior volsella, curved, with blunt apex. Stem of median volsella bulbous, with slender foliate lamellae.	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7111A71BFF60A89CD0CCF0D5.taxon	description	Description. Adult male (n = 1) Total length c. 1.2 mm; wing length c. 820 Μm. Head (Fig. 5 C – E). Eyes bare, reniform, with dorsomedian extensions (Fig. 5 C). Frontal tubercles stout, cylindrical with blunt apices, c. 15 - 20 Μm long (Fig. 5 D). Antennal flagellum composed of 11 segments of which 10 are well discernible, flagellomeres 10 – 11 fused in part (borders between remaining flagellomeres not visible, but distribution of setal tubercles indicates 13 segments), AR 0.82 (when flagellum measured as 10 - segmented), AR 0.65 (as 11 - segmented); plume fully developed (setae separated, but setal tubercles well developed) (Fig. 5 E). Length of palpomeres 2 – 5 (Μm): c. 30, 60, 76, 129. At least 15 clypeals. Thorax. Ac and Dc not visible (thorax damaged in dorsal part), Pa 3 on each side, Scts 6 at least. Wing (Fig. 5 F). Slender, with anal lobe weak, broadest at 2 / 3 length, width: 260 Μm, length / width ratio 3.15. FCu placed far distally of RM; VR Cu 1.52. Veins ending as in most extant Tanytarsus (from base to tip): An, Sc, Cu 1, R 1, R 2 + 3, M 3 + 4, R 4 + 5, M 1 + 2; distances between ends of R 1 – R 2 + 3 and R 2 + 3 – R 4 + 5 unequal (VR C c. 2.15). Almost whole wing (except base) covered with dense macrotrichia. Legs. Tibia of fore leg without spur. Tibial combs of mid and hind legs separated, fan-shaped, teeth up to 16 Μm long; each comb with slender spur, 26 Μm long (mid leg) and 20 – 32 Μm long (hind leg). Sensilla chaetica on ta 1 of p 2 not observed. Lengths of leg segments and leg ratios in Table 2. Hypopygium (Fig. 6 A – E). Gonostylus stout, slightly arcuate, c. 50 Μm (as long as gonocoxite), with sparse setae placed on median margin in distal half. At least 2 - 3 tubercles bearing posterolateral setae on each side of anal point. Anal point wide at base, distinctly narrowed subapically, tapering to pointed apex; spinulae or crests unobservable from ventral side. Superior volsella (visible only in distal part) with 2 setae on apex; digitus long, extending beyond superior volsella, curved, with blunt apex (Fig. 6 D). Stem of median volsella bulbous, c. 15 Μm long, with 3 slender foliate lamellae (Fig. 6 E). Inferior volsella reaching half length of gonostylus, stout, with slightly swollen apical half posteromedially directed, armed with several curved setae on apex and strong setiform microtrichia on posterolateral margin (Fig. 6 A – C).	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A7111A71BFF60A89CD0CCF0D5.taxon	discussion	Remarks. A set of characters given in the diagnosis and description (except the number of antennal flagellomeres, see remarks below), indicate that the new species belongs to the genus Tanytarsus. Several significant characters, i. e. the shape of the hypopygial superior volsella (with 2 setae on apex), the digitus (long, extending far beyond superior volsella), and the median volsella (with stout bulbous stem and slender foliate lamellae) are known from species of different systematic groups, e. g. the eminulus- mendax - or chinyensis group (cf. Reiss & Fittkau 1971, Ekrem 2003, Giłka & Paasivirta 2009). However, these characters form a combination unknown from any extant or fossil Tanytarsus, thus a group membership of Tanytarsus glaesarius remains open. Tanytarsus glaesarius is the first Eocene species of this genus known from the adult male having the antenna with the reduced number of flagellomeres (Fig. 5 E). This character is similar to that discussed above (see remarks on Stempellinella electra), but we propose to weigh it differently on a background of all known Tanytarsus, including extant species of this genus (males with 13 flagellomeres). Since we have not observed other characters that might suggest non-typical structure (e. g. deformations, asymmetry, brachyptery or other associated features known from non-flying chironomids; cf. Giłka 2011 b, Giłka et al. 2013 a), we assume that the reduced number of flagellomeres observed in T. glaesarius should be treated as a species-specific feature rather than a character which might be considered in the context of phylogenetic trends.	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A711EA71EFF60AFD0D3E9F655.taxon	materials_examined	Type material. Holotype. Adult male (tarsi of hind legs broken and separated) preserved in 15 x 13 x 5 mm piece of amber (Eocene, ~ 45 – 40 Ma, Baltic amber, Gulf of Gdańsk; MAI- 4325 a; Fig. 7 A); animal syninclusion: Sciaridae (1 ind., MAI- 4325). Paratypes. Two adult males (tarsi of all legs broken and separated and / or missing, wings poorly preserved), as syninclusions, slightly compressed, preserved in 16 x 18 x 4 mm piece of clarified amber (MAI- 4352, same data as holotype; Fig. 7 B).	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A711EA71EFF60AFD0D3E9F655.taxon	etymology	Derivatio nominis. Combination of two words: ‘ protoplastus ’ and ‘ gregarius ’. The species is the oldest known representative of the Tanytarsus gregarius species group.	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A711EA71EFF60AFD0D3E9F655.taxon	diagnosis	Diagnosis. Femur of mid leg longer than femur of hind leg. Gonostylus with subapical tooth-like process. Anal point slender, tapering to blunt apex, bearing spinulae arranged in row. Superior volsella posteriorly directed, with nipple-like extension on apex and 4 strong medially directed setae on median margin. Digitus not observed. Stem of median volsella long, straight, with 5 – 6 slender foliate lamellae.	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A711EA71EFF60AFD0D3E9F655.taxon	description	Description. Adult male (n = 1 – 3) Total length 2.6 – 3.1 mm; wing length 1475 – 1575 Μm. Head (Fig. 7 D, E). Eyes bare, with well developed dorsomedian extensions. Frontal tubercles not observed. Antenna with 13 well discernible flagellomeres, AR 0.92 – 0.97, plume fully developed (Fig. 7 E). Length of palpomeres 2 – 5 (Μm): 40 – 48, 113 – 137, 115 – 125, 149 – 185. Clypeals present but poorly visible (at least 4 – 6 in paratype specimens). Thorax. Ac at least 12, Dc 13 on each side, Pa 3 on each side, Scts 12 – 13. Wing (Fig. 7 F, G). Ellipse-shaped, with anal lobe weak, broadest at 2 / 3 length, width: 385 – 395 Μm, length / width ratio 3.83 – 3.98. FCu placed well distally of RM; VR Cu 1.33. Veins ending as in most extant Tanytarsus (from base to tip): An, Sc, Cu 1, R 1, R 2 + 3, M 3 + 4, R 4 + 5, M 1 + 2; distances between ends of R 1 – R 2 + 3 and R 2 + 3 – R 4 + 5 unequal (VR C 1.35). Almost whole wing (except base) covered with dense macrotrichia. Legs. Tibia of fore leg with single, 28 – 30 Μm long spur (a vestigial comb composed of 2 – 3 teeth, observed on one leg of paratype specimen is recognized as artefact). Tibial combs of mid and hind legs separated, fan-shaped, teeth up to 20 Μm long (mid leg) and c. 20 – 24 Μm long (hind leg); each comb with slender spur, up to 30 Μm long (mid leg) and 36 – 48 Μm long (hind leg). Sensilla chaetica on ta 1 of p 2 not observed. Lengths of leg segments and leg ratios in Table 3. Hypopygium (Fig. 8 A – F). Gonostylus stout, slightly curved at mid length, c. 95 – 115 Μm, longer than gonocoxite, bearing several setae placed at median margin of distal half, with subapical anteromedially directed tooth-like process. Median setae on anal tergite not observed, at least 3 posterolateral setae on each side of anal point. Anal point slender, tapering to blunt apex, bearing at least 3 spinulae arranged in row, crests tapering towards anal point apex. Superior volsella posteriorly directed, oval, slightly elongated, with nipple-like extension on apex and 4 strong medially directed setae placed on distinct protuberances on median margin (Fig. 8 D). Digitus not observed. Stem of median volsella c. 25 Μm long, straight, bearing 5 – 6 slender foliate lamellae (Fig. 8 E, F). Inferior volsella reaching 1 / 3 length of gonostylus at most, club-shaped, with posteromedially turned head-like apical part, armed with several stout curved setae.	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A711EA71EFF60AFD0D3E9F655.taxon	discussion	Remarks. Tanytarsus protogregarius is here proposed to be included in the gregarius species group due to a distinct similarity to the extant species — Tanytarsus gregarius Kieffer, 1909. Both the species are similar in the shape of the anal point and the superior volsella that is posteriorly directed and armed with nipple-like apical extension (cf. Fig. 8 and Reiss & Fittkau 1971, fig. 26). The best characters separating the new species from other members of the group is the subapical tooth-like process of the gonostylus and the presence of four strong medially directed setae on median margin of the superior volsella (three strong setae are known from extant species). Moreover, in the adult male of T. protogregarius we observed interesting proportions of leg segments. In all examined individuals femora of mid legs are longer than those of hind legs (Table 3). Similar proportions we found in other fossil species — Tanytarsus congregabilis Giłka et Zakrzewska, 2013 of the lugens group. It may indicate close relations between the two groups, which were previously proposed also to be treated as one — the concept based on adult male morphological characters of extant species (Giłka 2000). The latter character discussed, however, is not known from any extant European species of these two groups [T. gregarius, T. aberrans Lindeberg, 1970, T. inaequalis Goetghebuer, 1921, and T. lugens (Kieffer, 1916), T. bathophilus Kieffer, 1911, T. latiforceps Edwards, 1941, T. trux Giłka et Paasivirta, 2007; all examined], in which femora of mid legs are typically shorter than those of hind legs.	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
03CD3E7A711BA71EFF60A957D3E3F5F1.taxon	materials_examined	Material examined. Holotype (MAI- 5157 a), paratype (MAI- 5157 b) (see Giłka 2010). Additional specimens: 1 male (tarsus of right hind leg missing) preserved in 35 x 28 x 18 mm piece of amber (Eocene, ~ 45 – 40 Ma, Baltic amber, Gulf of Gdańsk; MAI- 275); animal syninclusion: Aphidoidea (1 ind.); 1 male preserved in 22 x 13 x 4 mm piece of amber (Eocene, ~ 45 – 40 Ma, Baltic amber, Gulf of Gdańsk; MAI- 5184), animal syninclusions: Formicidae (1 ind.), Acari (1 ind.).	en	Zakrzewska, Marta, Giłka, Wojciech (2015): The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk. Zootaxa 3946 (3): 347-360, DOI: 10.11646/zootaxa.3946.3.3
