taxonID	type	description	language	source
03CA87824E42FFE5CBEEB5B2DACEF936.taxon	materials_examined	Material examined: One female, Tea Research Institute-Lahijan, 37 ° 12 ’ 17.11 ” N, 50 ° 01 ’ 23.19 ” E, 9 m, collected from soil, October 2018; five females, Rasht, Bazar Jomeh, 37 ° 24 ’ 21 ” N, 49 ° 7 ’ 320 ” E, 3 m, soil, August 2018; 4 males, Rasht, Keshel Varzal Village, 37 ° 10 ’ 59 ” N 49 ° 38 ’ 42 ” E, 34 m, bell pepper, Capsicum annuum L. (Solanaceae) leaves, February 2019; 4 females, Rasht, Pasikhan Village, 37 ° 17 ’ 28 ” N 49 ° 31 ’ 46 ” E, - 13 m, litter, April 2019. Habitat and distribution in Iran: Mazandaran Province, soil and litter; Guilan Province, soil and litter (Kazemi et al., 2014; Saberi et al., 2016; Nemati et al., 2018; Hajizadeh et al., 2020; Ghasemi & Hajizadeh, 2021). World Distribution: Japan, Korea, Iran. Holotype for P. akimotoi: Japan, Ehime Prefecture, Omogo Valley, litter in cedar forest. For G. incises: Southern China, Linjiang, pine forest litter (Quintero-Gutiérrez & Halliday, 2021).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E42FFE6CBEEB623D8A2FC59.taxon	description	Idiosoma length 815 (800 – 830), width 580 (560 – 600) (Figures 1 – 2). Measurements of setae: j 1 73 (71 – 75), j 2 74 (73 – 75), j 3 69 (68 – 70), j 4 62 (61 – 63), j 5 64 (63 – 65), j 6 64 (63 – 65), z 1 78 (76 – 80), z 2 81 (80 – 82), z 3 69 (68 – 70), z 5 69 (68 – 70), z 6 69 (68 – 70), s 2 64 (66 – 64), s 3 17 (16 – 18), s 4 19 (18 – 20), s 5 77 (78 – 76), s 6 70 (69 – 71), J 1 74 (73 – 75), J 2 69 (68 – 70), J 3 66 (65 – 67), J 4 74 (73 – 75), J 5 74 (73 – 75), Z 1 72 (71 – 73), Z 2 71 (70 – 72), Z 4 74 (73 – 75), Z 5 77 (75 – 79), S 1 64 (63 – 65), S 2 71 (70 – 72), S 3 74 (73 – 75). Tritosternum 170 long, base 33 long, laciniae 138 long (Figure 5). A pair of narrow presternal shields present near the tritosternal base. Sternal shield 173 long and 140 wide, with three pairs of sternal setae, st 1 60, st 2 68, st 3 70. Metasternal setae st 4 55 long. Epigynal setae st 5 68 long; perigenital rim present. Ventrianal shield 300 long and 380 wide, with four pairs of pre-anal setae Jv 1 65, Jv 2 75, Zv 1 58, Jv 3 78, and para-anal 35, post-anal 25 setae. Stigmata situated near the antero-lateral angles of coxae IV. Peritremes extending beyond coxae I. Gnathosoma (Figure 3). Hypostome with one pair of capitular setae, pc 38, and three pairs of hypostomal setae, h 1 75, h 2 35, h 3 45 (Figure 3). Legs (Figures 7 – 11); tarsus I without pretarsus, 175 long, almost twice as long as tibia I 95 long (Figure 7), trochanter I with one median papilliform tubercle (Figure 8); femur II & III and trochanter IV each with a ventral tubercle (Figures 9 – 11).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E42FFE6CBEEB623D8A2FC59.taxon	materials_examined	Material examined: Two females, Rudbar, Rostamabad, Naqlabar village, 36 ° 96.23 ” N, 46 ° 53.61 ” E, collected from soil, June, 2011. World Distribution: Taiwan, Korea, Japan and India. Holotype from Georgia, Batumi, soil and litter under trees (Quintero-Gutiérrez & Halliday, 2021).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E41FFE6CBEEB2CED9B4FAB1.taxon	materials_examined	Material examined: Four females, Rasht, Pasikhan Village, 37 ° 17 ’ 28 ” N 49 ° 31 ’ 46 ” E, - 13 m, litter, April 2019; 6 females, Rasht, Pir Bazar Village, 37 ° 19 ’ 57 ” N 49 ° 28 ’ 59 ” E, - 18 m, soil, December 2018. Habitat and distribution in Iran: Tehran Province, soil, ornamental plants; Guilan Province, soil (Hamidi et al., 2013; Maleki et al., 2016; Nemati et al., 2018; Ghasemi & Hajizadeh, 2021). World Distribution: Argentina, U. S. A., China and Spain. Holotype for G. gamasoides from Italy (Florence, Genoa), decaying vegetables (Quintero-Gutiérrez & Halliday, 2021).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E41FFE7CBEEB7D9D985FEC5.taxon	description	World Distribution: U. S. A. (Illinois, Columbia, Maryland). Holotype for M. muscorum: U. S. A., Parker, Illinois, in moss. For H. pulchella: U. S. A., Columbia and Lake City, Florida, in humus. For N. hurlbutti: U. S. A., Maryland, Bowie, Fagus litter (Quintero-Gutiérrez & Halliday, 2021).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E40FFE7CBEEB072DE8FFD3F.taxon	description	World Distribution: Iran, Tehran, collected from soil. Type depository not stated (Quintero-Gutiérrez & Halliday, 2021). Note: Quintero-Gutiérrez & Halliday (2021) pointed out that Holaspina persicum is a nomen nudum because the authors did not say where the types are deposited (ICZN Article 16.4.2). It is therefore excluded from the key to Parholaspididae species of Iran. It may be a synonym of Parholaspulus alstoni Evans, 1956.	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E40FFE7CBEEB224D9D5FC6B.taxon	materials_examined	Type species Holostapis (Holaspulus) tenuipes Berlese, 1904, by original designation	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E40FFE7CBEEB519DF41F9FE.taxon	materials_examined	Material examined: Four females, Rasht, Jirdeh Village, 37 ° 12 ’ 43 ” N 49 ° 32 ’ 51 ” E, 22 m, litter, April 2019; 3 females, Rasht, 37 ° 11 ’ 47 ” N 49 ° 39 ’ 9 ” E, 26 m, wood sorrel, Oxalis acetosella L. (Oxalidaceae) leaves, November 2018; five females, Bijar Boneh Village, 37 ° 18 ’ 10 ” N 49 ° 39 ’ 10 ” E, - 6 m, soil, April 2019. Habitat and distribution in Iran: Isfahan Province, soil and litter; Mazandaran Province, soil, litter and humus (Ramezani & Nemati, 2010; Kazemi et al., 2012; Kazemi & Rajaei, 2013; Kazemi et al., 2014; Saberi et al., 2016; Nemati et al., 2018; Ghasemi & Hajizadeh, 2021). World Distribution: Switzerland, U. S. A. (Hawaii), U. K. (London), Mexico, Japan, Philippines. Holotype for H. tenuipes: Italy, Florence, debris in greenhouse. For M. tridentifer: U. S. A., Minnesota, greenhouse. For P. pachylaelapsoides: Switzerland, Berne Botanic Gardens, plant nursery (Quintero-Gutiérrez & Halliday, 2021).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E40FFE7CBEEB764DE0DF92B.taxon	materials_examined	Type species Krantzolaspina rebatti Datta & Bhattacharjee, 1989. by original designation	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E40FFE0CBEEB626D8BEFE75.taxon	biology_ecology	Habitat and distribution in Iran: Kerman Province, Jiroft, soil and litter at the base of Phoenix dactylifera L. and Citrus sinensis (L.); Esfahan Province, soil (Kadkhodaei et al., 2013; Hajizadeh et al., 2017; Nemati et al., 2018).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E40FFE0CBEEB626D8BEFE75.taxon	distribution	World Distribution: Philippine and Iran. Holotype for P. angustatus: Philippines, Palawan Island, Olanguan Falls, litter in tropical rainforest. For I. jiroftensis: Iran, Jiroft City, Kerman Province, soil and litter at the base of Phoenix dactylifera (Quintero-Gutiérrez & Halliday, 2021).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E40FFE0CBEEB626D8BEFE75.taxon	discussion	Note: This species was described in the genus Indutolaelaps Karg, 1997 (Leptolaelapidae) (Hajizadeh et al., 2017), but Quintero-Gutiérrez et al. (2020) synonymised Indutolaelaps jiroftensis Hajizadeh et al., 2017 with Krantzolaspina angustatus (Ishikawa, 1987).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E47FFE0CBEEB3EFD90BFAE1.taxon	materials_examined	Material examined: One female, Fuman, 37 ° 12 ’ 51.30 ” N, 49 ° 15 ’ 37.57 ” E, 70 m, collected from soil, October 2018; 1 female 3 males, Otaghvar, Langarud, 37 ° 5 ’ 46.82 ” N, 50 ° 6 ’ 45.46 ” E, 175 m, collected from soil, December 2019; one female, Vajargah, Kelachay, 37 ° 2 ’ 37.92 ” N, 50 ° 23 ’ 59.60 ” E, 4 m, collected from soil, November 2019; 1 female, Otaghvar-Langarud, 37 ° 6 ’ 1.31 ” N, 50 ° 6 ’ 22.32 ” E, 70 m, collected from soil, December 2019; 1 female, Rahimabad, Rudsar, 37 ° 0 ’ 27.61 ” N, 50 ° 16 ’ 56.13 ” E, 392 m, collected from soil, November 2019; 1 female, Amlash, 37 ° 5 ’ 32.64 ” N, 50 ° 8 ’ 57.80 ” E, 123 m, collected from soil, December 2019; 4 females, Rasht, Keshel Varzal Village, 37 ° 10 ’ 59 ” N 49 ° 38 ’ 42 ” E, 34 m, litter, February 2019; 5 females, Rasht, Bijar Boneh Village, 37 ° 18 ’ 10 ” N 49 ° 39 ’ 10 ” E, - 6 m, litter, May 2019. Habitat and distribution in Iran: Mazandaran Province, soil and litter (Kazemi et al., 2014; Saberi et al., 2016; Nemati et al., 2018; Hajizadeh et al., 2020; Ghasemi & Hajizadeh, 2021). World Distribution: Korea and Russia. Holotype Russia, Sakhalin Island, Kuril Islands; Kunashir, Shikotan Island, forest litter (Quintero-Gutiérrez & Halliday, 2021).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E47FFE0CBEEB0E8DEC7FD74.taxon	materials_examined	Type species Neparholaspis spathulatus Evans, 1956, by original designation.	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E47FFE0CBEEB0E8DEC7FD74.taxon	materials_examined	Type species Tricholaspis marginalis Evans, 1956, by original designation. Synonymy by Krantz, 1960: 405.	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E47FFE0CBEEB496D95CF983.taxon	materials_examined	Type species Parholaspulus alstoni Evans, 1956, by original designation.	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E47FFE0CBEEB496D95CF983.taxon	materials_examined	Type species: Holaspina excentricus Petrova, 1967, by original designation.	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
03CA87824E47FFE1CBEEB7BEDACEFE51.taxon	materials_examined	Material examined: Two females, two males, Tea Research Institute-Lahijan, 37 ° 12 ’ 17.11 ” N, 50 ° 01 ’ 23.19 ” E, 9 m, collected from soil, October 2018; three females, two males, Rahimabad-Rudsar, 37 ° 01 ’ 40.58 ” N, 50 ° 21 ’ 01.05 ” E, 52 m, collected from soil and leaf litter, October 2018; five females, two males, Vajargah-Kelachay, 37 ° 2 ’ 25.88 ” N, 50 ° 24 ’ 12.17 ” E, 7 m, collected from soil, November 2019. Habitat and distribution in Iran: Tehran Province, soil; Kerman Province, soil, manure and litter; Zanjan Province, soil; Mazandaran Province, soil and litter; Guilan Province, soil and litter (Kazemi & Rajaei, 2013; Hamidi et al., 2013; Bigdeli et al., 2014; Kazemi et al., 2014; Masnavipour et al., 2011; Maleki et al., 2016; Saberi et al., 2016; Nemati et al., 2018; Hajizadeh et al., 2020). World Distribution: Azerbaijan, Russia, Belgium, Germany, Ukraine, Georgia, Hungary, Korea, Vietnam, U. S. A., Iran. Holotype from U. K., London, Royal Botanic Gardens, from soil (Quintero-Gutiérrez & Halliday, 2021).	en	Hajizadeh, Jalil (2022): Review of the Iranian Parholaspidid mites (Acari: Mesostigmata) with a key to the species. Zootaxa 5178 (3): 229-240, DOI: 10.11646/zootaxa.5178.3.2
