taxonID	type	description	language	source
03CA878ABA68DE3BFF4FF9EB73C0F86E.taxon	type_taxon	Type species. Alvania supranitida Wood, 1842 [= Aclis minor (Brown, 1827)], by monotypy. Eastern Atlantic and Mediterranean. Syntypes of A. minor were figured by Oliver et al. (2020: Fig. 13).	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA68DE3BFF4FF9EB73C0F86E.taxon	description	Description. Shell vitreous, usually elongated; protoconch smooth, globose, dome shaped, paucispiral or multispiral; teleoconch with convex whorls, commonly with one or more spiral keels, presence of microscopic spiral striae close to the sutures; aperture wide, outer lip usually opisthocline, inner lip distinctly reflected, umbilical fissure commonly present, most anterior region of aperture commonly drawn out towards the center of the shell and flattened (adapted from Bouchet & Warén 1986).	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA68DE3BFF4FF9EB73C0F86E.taxon	discussion	Remarks. A complete list of synonyms was presented by Bouchet & Warén (1986). Only Cyclonidea Laseron, 1956 (Type species: Cyclonidea carina Laseron, 1956, by original designation), which was previously synonymized with Aclis is currently considered as valid (Poppe & Tagaro 2016).	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA69DE37FF4FFF4075D6FC3A.taxon	description	Figs 2, 3	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA69DE37FF4FFF4075D6FC3A.taxon	materials_examined	Type material. Lectotype (herein designated and examined): NHMUK 1887.2.9.1576; H. M. S. Challenger coll., 10 / xi / 1873. Paralectotypes (herein examined): Brazil: Pernambuco state: type locality: NHMUK 1887.2. 9.1577 [1]; type locality: NMW 1955.158. 26255 [3]. Holotype of Aclis carolinensis (herein examined): USA: North Carolina state, Cape Hatteras, USFC sta. 2595; 35 ° 08 ′ 00 ″ N, 75 ° 05 ′ 30 ″ W, 115 m; Steamer Albatross col., 17 / x / 1885: USNM 83743 [3]. Type locality. Brazil: Pernambuco state: Challenger sta. 122; 09 ° 05 ′ S, 34 ° 50 ′ W, 640 m. Material examined. The types and: USA: Florida state: off West Dry Rocks Reef, Eolis sta. 320; ~ 24 ° 26 ′ 46 ″ N, 81 ° 55 ′ 37 ″ W, 146 m; Yacht Eolis col., 1916: USNM 503753 [5]; off Fowey Rocks Lighthouse, Eolis sta. 355; ~ 25 ° 35 ′ 27 ″ N, 80 ° 05 ′ 48 ″ W, 128 m; Yacht Eolis col., 1916: USNM 503763 [11]; off Fowey Rocks Lighthouse, Eolis sta. 364; ~ 25 ° 35 ′ 27 ″ N, 80 ° 05 ′ 48 ″ W, 137 – 165 m; Yacht Eolis col., 1917: USNM 503771 [11]; Brazil: Pernambuco state: Revizee Nordeste; 08 ° 46 ′ 00 ″ S, 34 ° 44 ′ 00 ″ W, 690 m; R / V Natureza col., 18 / xi / 2000: MORG 49199 [4 †]; Sergipe state: MARSEAL SED 3 sta. BN 6: CZUFS GAS- 00102 [1]; MARSEAL SED 3 sta. DN 4: CZUFS GAS- 00103 [1]; MARSEAL SED sta. GN 4: CZUFS GAS- 00104 [1]. Revised description. Shell vitreous or whitish, tall, conical, with an obtuse apex, reaching about 3.2 mm long, 1.1 mm wide. Protoconch globose, subcylindrical, about 2.0 whorls, about 300 μm wide, smooth, transition to teleoconch marked by a gentle incremental scar. Teleoconch reaching eight whorls, strongly convex outline, faster increase in diameter from the fourth whorl; suture deep, well impressed, sloping; surface glossy, presenting faint microscopic spiral threads in subsutural zone of each whorl; incremental scars slightly demarcated appearing at irregular intervals; growth lines slightly demarcated; surface smooth or with up to three faint spiral keels. Last whorl about 40 % of total length, broad; base rounded or slightly crenulated by spiral keels. Aperture round to elliptical in shape, about 60 % of body whorl length, strongly expanded laterally, rounded and slightly advanced anteriorly, acute posteriorly; outer lip thin, slightly opisthocline, gradually advancing forward, rectilinear; inner lip thin, reflected. Umbilical fissure present.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA69DE37FF4FFF4075D6FC3A.taxon	description	Measurements. Lectotype (NHMUK 1887.2.9.1576): 8 whorls; SL = 3.21 mm; BWL = 1.38 mm; AL = 0.81 mm; SW = 1.14 mm; AW = 0.66 mm. NMW 1955.158.26255: 8 whorls; SL = 4.29 mm; BWL = 1.67 mm; AL = 1.02 mm; SW = 1.67 mm; AW = 0.77 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA69DE37FF4FFF4075D6FC3A.taxon	discussion	Remarks. Watson (1881) did not state the exact number of individuals on which the description was based. There are two shells housed at the NHMUK collection, which are considered as syntypes (Fig. 2 A – E). Other shells from the same locality and labeled as A. sarissa by J. C. Melvill are housed at the NMW. According to Lingwood (1981), part of the material from the Challenger expedition was distributed to collections in other museums. By the experience of H. Wood and G. Oliver (curators at the NMW; pers. Comm) on the Melvill-Tomlin collection, the Challenger material from this institution usually belongs to the type series of Watson’s species, as Melvill and Watson were contemporaries. Thus, we consider the three shells of NMW 1955.158.26255 as syntypes (one illustrated in Fig. 2 F – I). The shell of NHMUK 1887.2.9.1576 (Fig. 2 A – C, E) is undoubtedly the specimen figured by Watson (1886: 503, pl. 34, Fig. 3) and is here selected the lectotype to stabilize the concept of the species. Watson (1881) described a faint spiral sculpture in A. sarissa, which seems variable like in the type species Aclis minor (Fretter & Graham 1982: Figs. 282, 283). The paralectotype from the NMW (Fig. 2 F – H) have more pronounced spiral keels than the lectotype (Fig. 2 A – C). This variation in the expression of the spiral keels was also observed in the non-type material examined here (Fig. 3 A – G), which can be possibly accentuated by the state of preservation of the shell, since in eroded shells, the keels become vary faint. The spiral keels are more frequent and more notable in the anterior whorls (Figs 2 G, H, 3 E – H), a pattern which is also common in A. minor (Gofas et al. 2011: 225, text-fig). Since the original description, there was no published record of new material of A. sarissa. In the present study, additional records from the northeast of Brazil are provided, and the geographic distribution of the species is extended to the southeast of Brazil. These additional specimens of A. sarissa present a similar shell shape (Fig. 3 A – F) to the lectotype and have a small variation of the teleoconch sculpture (Fig. 3 A – H). The aperture varies from circular (Fig. 3 A, C) to oval and more advanced anteriorly (Fig. 3 E, G). The bathymetric range of the species is also extended upward (225 m) and downward (1000 m) but based exclusively on empty shells. Barros et al. (2001) commented that A. sarissa would be a possible synonym of Costaclis hyalina (cited as “ Hemiaclis hyalina ”) based on shells from the northeast Brazil. However, Barros et al. (2001, 2003) presented confused ideas about these species. During the examination of the LMUFRPE collection, no voucher material studied by Barros et al. (2001; 2003) could be found, but the shell figured by Barros et al. (2001: Fig. 2 D) is a misidentification of a newly described species of the present study (see Aclis pulchra nov. sp.), while the shell figured by Barros et al. (2003: Fig. 1) is a young specimen and it is not possible to infer a precise determination. Aclis sarissa and C. hyalina are distinct species because A. sarissa does not present axial ribs like C. hyalina, it has a more globular protoconch and more convex teleoconch whorls. Aclis sarissa is herein considered as a senior synonym of Aclis carolinensis Bartsch, 1911, from North Carolina, USA. The holotype of A. carolinensis (USNM 83743; Fig. 2 J – K) has a similar shape, spire angle, broad aperture expanded laterally, a quite smooth teleoconch and an umbilical fissure; additional shells from Florida, USA, has a variation from smooth to three faint spiral keels at the body whorl. It is hardly discernible from the material examined of A. sarissa, except for the more sinuous outline of outer lip. However, the latter feature is difficult to compare since most specimens have a partially broken outer lip. Considering the variation of sculpture in A. sarissa, the quite smooth surface also fits the taxon. Bartsch (1947) transferred both Aclis sarissa and Aclis carolinensis to Hemiaclis G. O. Sars, 1878, which is the current classification in MolluscaBase (2021 a). Bartsch (1947) did the same for several other species previously assigned in Aclis, but as noted by Bouchet & Warén (1986), many of them do not belong with certainty in Hemiaclis. Hemiaclis embraces species with a broadly conical shell with whorls increasing in diameter regularly, sutures relatively flat and a straight outer lip seen in lateral view (Bouchet & Warén 1986). Furthermore, the species of Hemiaclis have a smooth teleoconch, lacking keels, ribs and microscopic spiral striae. Geographic Distribution. USA: North Carolina, Florida; Brazil: Pernambuco, Sergipe. From 115 to 1000 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA64DE36FF4FF92573A5FC56.taxon	description	Fig. 4	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA64DE36FF4FF92573A5FC56.taxon	materials_examined	Type material. Holotype (herein examined): MZSP 77991; R / V Natureza coll., 28 / x / 2000. Paratypes (herein examined): Brazil: Sergipe state: type locality; MZSP 77992 [3]; Revizee Nordeste; 11 ° 35 ′ 30 ″, 37 ° 12 ′ 18 ″ W, 510 m; R / V Natureza col., 30 / x / 2000: MZSP 77993 [4]. Type locality. Brazil: Sergipe state; Revizee Nordeste, 10 ° 41 ′ 24 ″ S, 36 ° 18 ′ 42 ″ W, 365 m. Material examined. The types and: Brazil: Sergipe state: MARSEAL SED 3 sta. HN 4; CZUFS GAS- 00105 [1].	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA64DE36FF4FF92573A5FC56.taxon	description	Measurements. Holotype, MZSP 77991: 7 whorls; SL = 4.29 mm; BWL = 1.72 mm; AL = 1.02 mm; SW = 1.44 mm; AW = 0.86 mm. Paratype MZSP 77993: 7 whorls; SL = 4.26 mm; BWL = 1.73 mm; AL = 1.02 mm; SW = 1.5 mm; AW = 0.89 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA64DE36FF4FF92573A5FC56.taxon	discussion	Remarks. Aclis watsoni has been reported only on the coast of northeastern Brazil, based on few specimens. We report new material from the same region, which consists of a single shell with a worn surface but still exhibiting the strong spiral keels. Aclis watsoni closely resembles A. sarissa (Figs 2 – 3) in the general shape and dimensions of the shell. However, the holotype and paratype figured by Barros et al. (2007: Fig. 7 – 9) have a remarkable spiral sculpture since the third teleoconch whorl, which gives a truncated outline in the teleoconch whorls. The types of A. sarissa (Fig 2) have a more rounded outline, with a weaker spiral sculpture usually notable in the last teleoconch whorls. In view of the preservation status of the type series of A. sarissa and the possible variation in the expression of the spiral keels, one could suspect a possible synonymy of A. sarissa and A. watsoni, in which the types of A. sarissa would be eroded specimens. Additional specimens would be necessary to confirm the synonymy. Aclis watsoni resembles Aclis eolis Bartsch, 1947 (Fig. 19 A – D), from Florida, but differs by the more truncated outline of the teleoconch whorls. Furthermore, A. watsoni can be distinguished from A. eolis by the pattern of the teleoconch sculpture, the former has a single and pronounced spiral keel in the posterior whorls and reaches three pronounced spiral keels at the anterior whorls. All spiral keels are restricted to the lower half region of the whorls. In contrast, A. eolis has three weak spiral keels since the initial teleoconch whorls that are distributed through the whole surface of each whorl, equally spaced (Bartsch 1947) (Fig. 19 A – D). Geographic Distribution. Brazil: Sergipe state. From 365 m to 510 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA65DE31FF4FFBFA71A6FDA2.taxon	description	Fig. 5	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA65DE31FF4FFBFA71A6FDA2.taxon	materials_examined	Type material. Holotype (not found by Oliveira et al. 2018): IBUFRJ 18056. Paratypes (MNRJ and IBUFRJ not found by Pimenta et al. 2014 and Oliveira et al. 2018): OP I # 80; MNRJ 12852; OP II # 75; IBUFRJ 18057; OP II # 84; MNHN; OP I # 45; IBUFRJ 18058. Type locality. Brazil: Rio de Janeiro state: OP I sta. 75; 22 ° 31 ′ 28 ″ S, 40 ° 03 ′ 50 ″ W, 1050 m. Material examined. Brazil: Rio de Janeiro state, Campos Basin: OP I sta. 44: IBUFRJ 17191 [1]; OP I sta. 49: IBUFRJ 15431 [1 †]; OP I sta. 54: IBUFRJ 17107 [7]; OP I sta. 64: IBUFRJ 16108 [2]; OP I sta. 79: IBUFRJ 16897 [3]; OP I sta. 85: MNRJ 26714 [1 †]; OP II sta. 44: MNRJ 26715 [2 †]; OP I sta. 74: IBUFRJ 15441 [2]; OP II sta. 59: MNRJ 26713 [5 †]; OP II sta. 69: IBUFRJ 15520 [8]; OP II sta. 64: IBUFRJ 19664 [1]; OP II sta. 74: IBUFRJ 17258 [3]; OP II sta. 75: IBUFRJ 19665 [2]; OP II sta. 79: IBUFRJ 16769 [10]; Albacr sta. 18: IBUFRJ 17463 [1]; 21 ° 53 ′ 37 ″ S, 39 ° 50 ′ 19 ″ W, 1120 m; 24 / viii / 2001: MNRJ 26731 [1 †]; São Paulo state: Revizee Sul sta. 6705: MNRJ 28293 [1]; PADCT sta. 6554: MNRJ 36034 [1 †].	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA65DE31FF4FFBFA71A6FDA2.taxon	description	Measurements. Holotype (IBUFRJ 18056): 5 whorls; SL = 1.90 mm; BWL = 0.98 mm; AL = 0.6 mm; SW = 0.75 mm; AW = 0.41 mm. MNRJ 28293: 7 whorls; SL = 2.98 mm; BWL = 1.40 mm; AL = 0.85 mm; SW = 1.20 mm; AW = 0.70 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA65DE31FF4FFBFA71A6FDA2.taxon	discussion	Remarks. Besides the holotype, Absalão (2009) designated paratypes from different sampling sites in Campos Basin; however, they were not located by Pimenta (et al. 2014) and Oliveira (et al. 2018), during the curatorial revision of the types deposited in MNRJ and IBUFRJ, respectively, and they are presumably lost. The morphological comparisons of the present study were based on the original description and illustrations. The material of A. kanela examined here consists mostly of shells collected from nearby areas of the type locality. Additionally, we also extend the geographic distribution of the species southwards to the state of São Paulo. Aclis kanela resembles A. sarissa (Figs 2, 3) in the shape of the protoconch, aperture and by the convexity of the teleoconch whorls. Absalão (2009) distinguished these species based on the increase in the diameter of the teleoconch. According to him, A. sarissa has a “ typical elongated-turreted Aclis shell profile ”, with the first whorls markedly smaller than the last ones. In contrast, A. kanela would have a more regular increase in diameter. However, based on our comparison of the holotype illustration of A. kanela (Absalão 2009: fig. 2 I) with the lectotype of A. sarissa (Fig. 2 A – C, E.), the increase in diameter is very similar in both shells, and in the additional material we examined. The teleoconch surface of A. kanela has a completely smooth surface (except for microscopic spiral striae) in all specimens examined, while A. sarissa usually presents spiral keels. Even in specimens with lower keel elevation, it is possible to observe some degree of elevation (Figs 2 H, 3 A, D) in the last whorls of A. sarissa. Thus, based on the current knowledge we still consider A. kanela as a valid species, pending further comparisons with samples in better conditions of preservation. Geographic Distribution. Brazil: Rio de Janeiro (Absalão 2009), São Paulo (this study). From 424 m to 1941 m (Absalão 2009; this study).	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA63DE33FF4FFF40719AFE16.taxon	description	Fig. 6	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA63DE33FF4FFF40719AFE16.taxon	materials_examined	Material examined. Brazil: Rio de Janeiro state: OP II sta. 44; MNRJ 33505 [2 †]. Characterization. Shell tall, conical, reaching about 2.0 mm long, 0.9 mm wide, eight whorls, whitish. Protoconch conical, about 3.5 whorls, first whorl (embryonic) smooth, remaining whorls sculptured by microscopic sigmoid axial lines at small intervals. Teleoconch with slightly convex whorls, surface glossy presenting faint microscopical spiral striae and strong incremental scars with irregular intervals. Body whorl half the total length, base rounded; aperture wide, 60 % of body whorl length, strongly expanded laterally, rounded anteriorly, acute posteriorly; outer lip thin, sinuous, strongly advanced close to distal area; inner lip thin, reflected. Umbilical fissure present.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA63DE33FF4FFF40719AFE16.taxon	description	Measurements. MNRJ 33505: 7 whorls; SL = 2.0 mm; BWL = 1.0 mm; AL = 0.6 mm; SW = 0.9 mm; AW = 0.5 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA63DE33FF4FFF40719AFE16.taxon	discussion	Remarks. Aclis sp. 1 has a multispiral protoconch, with the first whorl (embryonic / protoconch I) smooth and the remaining whorls ornamented by microscopic sigmoid axial lines. This feature is not common in Aclis, but Aclis walleri Jeffreys, 1867 known from the North Atlantic and Aclis aurisparva Bertolaso & Garilli, 2009 from Pliocene deposits of Italy present a similar pattern of sculpture in the larval shell (Bouchet & Warén 1986: fig. 736; Bertolaso & Garilli 2009: figs 1 – 5). This pattern of sculpture is more common in other eulimids like species of Batheulima F. Nordsieck, 1968, Fuscapex Warén, 1981 and Niso Risso, 1826, which also presents some patterns of brownish coloration in the larval shell (Bouchet & Warén 1986). In Batheulima and Fuscapex, the presence of the sigmoid axial lines in the protoconch is variable, with species of the same genus showing a smooth surface (Bouchet & Warén 1986). This type of sculpture is possibly rare in Aclis. Despite the atypical protoconch, Aclis sp. 1 fits the genus by the shell shape with convex whorls (Fig. 6 A), presence of microscopic spiral striae at the teleoconch surface (Fig. 6 B), shape of the aperture and presence of an umbilical fissure (Fig. 6 C). A distinction between protoconch I and II is indicative of planktotrophy (Warén 1983). The remaining species studied here are probably non planktotrophic because, in general, they have a paucispiral protoconch with smooth surface, without the distinction between protoconch I and II (Warén 1983). The paucity of informative characters in the larval shells of Aclis species limits the inference of the type of larval development. The general shape of the species closely resembles A. sarissa, A. kanela and A. walleri. It can be distinguished from A. sarissa and A. kanela by the presence of microscopic sculpture in the larval shell. The teleoconch whorls of Aclis sp. 1 seems less convex than in A. walleri but based on limited material it is difficult to assess variation, pending further comparisons to distinguish Aclis sp. 1 as a new species or determining the occurrence of A. walleri in the southwestern Atlantic.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA60DE2CFF4FFE0D74DBFF66.taxon	description	Figs 7, 8	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA60DE2CFF4FFE0D74DBFF66.taxon	materials_examined	Type material. Holotype. MNRJ 16825 [†]; R / V Ocean Stalwart coll., 25 / ii / 2019. Paratypes. Brazil: Rio de Janeiro state: HAB 16 sta. D 5: MNRJ 17713 [1 †]; HAB 13 sta. H 4: MNRJ 16323 [1 †]; Campos Basin; 23 ° 05 ′ 00 ″ S, 40 ° 58 ′ 00 ″ W, 100 m; vi / 2004: MNRJ 26601 [1 †]; Campos Basin; 23 ° 05 ′ 23 ″ S, 40 ° 58 ′ 56 ″ W, 100 m; 16 / ix / 2004: MNRJ 30872 [1 †]; Espírito Santo state: off Vitória; 50 m; 1994; R / V Victor Hensen col.: MORG 39469 [1 †]. Type locality. Brazil: Rio de Janeiro state; Campos Basin, HAB 11 sta. G 3 (22 ° 03 ′ 41 ″, 40 ° 10 ′ 06 ″, 76 m). Material examined. The types and: Brazil: Rio de Janeiro state: OP II sta. 49; MNRJ 26716 [1 †]; Campos Basin; 22 ° 42 ′ 34 ″ S, 40 ° 41 ′ 24 ″ W, 110 m; 19 / ix / 2003; MNRJ 26588 [3 †]; MNRJ 33252 [1 †]; Campos Basin; 22 ° 42 ′ 34 ″ S, 40 ° 41 ′ 24 ″ W, 110 m; 19 / ix / 2009; MNRJ 31892 [6 †]; Campos Basin; 23 ° 05 ′ 23 ″ S, 40 ° 58 ′ 56 ″ W, 100 m; 16 / ix / 2004; MNRJ 33081 [13 †]; HAB 11 sta. A 5; MNRJ 33418 [1 †].	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA60DE2CFF4FFE0D74DBFF66.taxon	etymology	Etymology. The epithet means beautiful, pretty, fine, lovely, a Latin singular adjective in the nominative case, and refers to the beauty of the shell.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA60DE2CFF4FFE0D74DBFF66.taxon	description	Description. Shell tall, conical, reaching about 2.7 mm long, 1.2 mm wide, seven whorls. Protoconch globose, subcylindrical, nucleus elevated, about 2.0 whorls, 390 μm high, surface smooth. Teleoconch with strongly convex whorls, surface glossy presenting strong microscopical spiral striae and strong incremental scars with irregular intervals. Body whorl about half the total length, base strongly rounded; aperture broad, width about 80 % of length, elliptical in shape, about 60 % of body whorl length, strongly expanded laterally, rounded and slightly channeled anteriorly, acute posteriorly; outer lip thin, strongly opisthocline, gradually advancing forward, rectilinear; inner lip thin, reflected. Umbilicus wide. Measurements. MNRJ 16825: 6 whorls; SL = 2.4 mm; BWL = 1.2 mm; AL = 0.7 mm; SW = 1.0 mm; AW = 0.6 mm. MNRJ 26601: 6 whorls; SL = 2.6 mm; BWL = 1.3 mm; AL = 0.8 mm; SW = 1.1 mm; AW = 0.6 mm. MNRJ 26601: 7 whorls; SL = 2.7 mm; BWL = 1.2 mm; AL = 0.8 mm; SW = 1.2 mm; AW = 0.6 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA60DE2CFF4FFE0D74DBFF66.taxon	discussion	Remarks. Aclis pulchra sp. nov was misidentified as “ Aclis hyalina ” by Rios (2009: 197). We could not reassess the specimen studied by Rios (2009), but the shell figured fits the specimens studied here. Rios (2009) did not cite a precise locality for that record, but the geographic distribution was limited to Northeast Brazil. During our revision, we did not find any other specimen from this area Aclis pulchra sp. nov can be clearly distinguished from C. hyalina by the absence of axial sculpture, which is present in the latter species, and by having more convex teleoconch whorls and a wider umbilicus. Most of the shells studied have no trace of spiral keels, but they show the characteristic microscopic striae, specially visible below suture (Fig. 7 F – G). The shell of A. pulchra sp. nov has, in general, a rapid increase in diameter and may vary from less (Figs 7 A, 8 A, B, D) to more abrupt (Figs 7 D, E, 8 C). Aclis pulchra sp. nov resembles A. sarissa (Figs 2 – 3), but differs in having more convex teleoconch whorls, a wider umbilicus, more profound and better demarcated incremental scars. Besides that, the false suture is not conspicuous as in A. sarissa, and A. pulchra sp. nov has a more advanced and straighter outer lip. Aclis pulchra sp. nov resembles A. rushi Bartsch, 1911 (Fig. 17 C – D) and A. tenuis Verrill, 1882 (Fig. 18 A – B), however the former has a noticeably more conical shape with a faster increase in diameter, a wider umbilicus and an opisthocline outer lip, which is orthocline in A. rushi and prosocline in A. tenuis. Aclis pulchra sp. nov differs from A. carolinensis (Fig. 2 J – K) because the newly described species has a faster increase in diameter despite not so evident like in the species compared above, more convex teleoconch whorls, a wider umbilicus, and an aperture more expanded anteriorly. Aclis pulchra sp. nov. differs from A. costai sp. nov. (Fig. 12) and Aclis georgiana Dall, 1927 (Fig. 19 G – H) mainly by having a more conical shape, a teleoconch with more convex whorls and a wider umbilicus. Geographic Distribution. Brazil: Espírito Santo, Rio de Janeiro. From 50 m to 720 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7FDE2FFF4FF9627503FDA2.taxon	description	Fig. 9	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7FDE2FFF4FF9627503FDA2.taxon	materials_examined	Material examined. Brazil: Rio de Janeiro state: HAB 13 sta. H 3; MNRJ 16811 [1 †]; HAB 16 sta. H 3; MNRJ 34072 [2 †]. Characterization. Shell tall, conical, spire angle 15 °, reaching 3.0 mm long, 1.0 mm wide, eight whorls, colorless. Protoconch subcylindrical, dome-shaped, about 2.0 whorls, 320 μm high, surface smooth. Teleoconch with strongly convex whorls, surface glossy presenting faint microscopical spiral striae and strong incremental scars with irregular intervals. Body whorl about 40 % of the total length, base rounded; aperture high, about 67 % of body whorl length, strongly expanded laterally, rounded anteriorly, acute posteriorly; outer lip thin, strongly orthocline, sinuous, moderately advanced close to distal area; inner lip thin, reflected. Wide umbilicus present, bordered by a rounded edge.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7FDE2FFF4FF9627503FDA2.taxon	description	Measurements. MNRJ 34072: 6 whorls; SL = 3.0 mm; BWL = 1.2 mm; AL = 0.8 mm; SW = 1.0 mm; AW = 0.6 mm. Geographic Distribution. Brazil: Rio de Janeiro. From 71 m to 73 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7FDE2FFF4FF9627503FDA2.taxon	discussion	Remarks. These shells (Fig. 9) are similar to A. pulchra sp. nov (Figs 7, 8) described herein in relation to shell shape, dimensions of the protoconch and convexity of teleoconch whorls. However, shells with this tentative identification (‘ cf. ’) reach a higher number of whorls, has a more regular increase in diameter, which results in a more turriform shape, besides a less advanced outer lip, a slightly smaller aperture compared to the total length (0.25 vs. 0.30) and a slightly narrower umbilicus. We avoid describing these specimens as a new species due to the scarcity of studied material that hampers a proper comparison of shape variability with A. pulchra sp. nov.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7CDE2FFF4FFD59741FF922.taxon	description	Fig. 10	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7CDE2FFF4FFD59741FF922.taxon	materials_examined	Type material: Holotype. MNRJ 33425 [†]; R / V Ocean Stalwart coll., 17 / vii / 2009. Paratypes. Brazil: Rio de Janeiro state: HAB 17 sta. E 2. MNRJ 34073 [1 †]. Type locality. Brazil: Rio de Janeiro state; Campos Basin, HAB 11 sta. E 2 (22 ° 06 ′ 52 ″, 40 ° 39 ′ 06 ″, 53 m). Material examined. The types and: Brazil: Rio de Janeiro state: HAB 11 sta. E 2; MNRJ 16934 [1 †]; São Paulo state: off São Sebastião Island; 24 ° 12 ′ 45 ″ S, 44 ° 59 ″ W, 79 m; R / V Professor Wladimir Besnard col., xi / 1997; MNRJ 36009 [2 †]; MZSP 89179 [1 †].	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7CDE2FFF4FFD59741FF922.taxon	etymology	Etymology. The epithet means horn-shaped, a Latin singular adjective in the nominative case and refers to the shell shape.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7CDE2FFF4FFD59741FF922.taxon	description	Description. Shell tall, subcylindrical, reaching about 2.9 mm long, 0.9 mm wide, eight whorls. Protoconch globose, subcylindrical, nucleus elevated, about 1.75 whorls, surface smooth. Teleoconch with convex whorls, surface glossy presenting faint microscopical spiral striae and strong incremental scars with irregular intervals. Body whorl about half the total length, base rounded; aperture short, elliptical in shape, about 65 % of body whorl length, slightly expanded laterally, rounded anteriorly, acute posteriorly; outer lip thin, almost orthocline, slightly advanced at distal area; inner lip thin, reflected. Umbilicus lacking. Measurements. Holotype, MNRJ 33425: 7 whorls; SL = 2.9 mm; BWL = 1.3 mm; AL = 0.7 mm; SW = 0.9 mm; AW = 0.5 mm. Paratype, MNRJ 34073: 7 whorls; SL = 2.9 mm; BWL = 1.5 mm; AL = 0.6 mm; SW = 0.7 mm; AW = 0.4 mm. MNRJ 36009: 7 whorls; SL = 2.6 mm; BWL = 1.1 mm; AL = 0.6 mm; SW = 0.8 mm; AW = 0.4 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7CDE2FFF4FFD59741FF922.taxon	discussion	Remarks. Aclis corniculata sp. nov. is noticeably different from most of his congeners because this newly described species has a subcylindrical shape, lack of spiral keels, a small aperture and straight outer lip. Other specific features can be mentioned to distinguish the present species. Aclis corniculata sp. nov. differs from A. sarissa (Figs 2 – 3) by the more acute spire angle, narrower and shorter aperture, narrower umbilical fissure, straighter outer lip and by having no trace of spiral keels. Aclis corniculata sp. nov. differs from A. pulchra sp. nov. (Figs 7 – 8) by having a narrower aperture anteriorly, lacking a flattened expansion, and an orthocline or slightly prosocline outer lip, which is strongly opisthocline in A. pulchra sp. nov. Moreover, A. corniculata has a small umbilical fissure while A. pulchra sp. nov. has a broader umbilicus. Geographic Distribution. Brazil: Rio de Janeiro, São Paulo. From 53 m to 79 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7CDE28FF4FF8D97508FA92.taxon	description	Fig. 11	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7CDE28FF4FF8D97508FA92.taxon	materials_examined	Material examined. Brazil: Espírito Santo state: off Piúma; 1993; F. Pitombo leg.; IBUFRJ 8497 [1 †]; Rio de Janeiro state; HAB 16 sta. H 3; MNRJ 17688 [3 †]; Geomar XII sta. 90; IBUFRJ 7584 [1 †]; HAB 16 sta. G 3; MNRJ 16712 [4 †]. Characterization. Shell tall, conical, spire angle 23 ° – 31 °, reaching about 4.0 mm long, 1.5 mm wide, eight whorls, yellowish-brown in color. Protoconch conical, tip narrow, about 2.0 whorls, 340 μm wide, surface smooth. Teleoconch with moderately convex whorls, surface glossy presenting faint microscopical spiral striae and strong incremental scars with irregular intervals. Body whorl about half the total length, base rounded; aperture high, about 60 % of body whorl length, strongly expanded laterally, rounded anteriorly, acute posteriorly; outer lip thin, strongly opisthocline, sinuous, strongly advanced close to distal area; inner lip thin, reflected. Umbilicus present, bordered by a sharp edge.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7CDE28FF4FF8D97508FA92.taxon	description	Measurements. IBUFRJ 7584: 7 whorls; SL = 2.94 mm; BWL = 1.59 mm; AL = 1.03 mm; SW = 1.34 mm; AW = 0.72 mm. IBUFRJ 8497: 8 whorls; SL = 4.04 mm; BWL = 1.99 mm; AL = 1.18 mm; SW = 1.50 mm; AW = 0.80 mm. Geographic Distribution. Brazil: Espírito Santo, Rio de Janeiro. From 71 m to 76 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7CDE28FF4FF8D97508FA92.taxon	discussion	Remarks. Aclis aff. conula is noticeably different from the other species of Aclis from surrounding areas by the yellowish-brown color, a markedly oval aperture and inflated teleoconch whorls (Fig. 11 A, F). The umbilicus is also notable because it is broad and bordered by a sharp edge (Fig. 11 C). The teleoconch has the spiral micro striae typical of the genus but is weaker than commonly found in Aclis and more spread along the teleoconch surface (Fig. 11 D, E). The species presents a variation in the spire angle; the shell of IBUFRJ 7584 (Fig. 11 A) is more inflated than the shell of IBUFRJ 8497 (Fig. 11 F) (spire angle: 31 ° vs. 23 °). Aclis aff. conula closely resembles Aclis conula Dall, 1927 (Fig. 20 A – C), from the continental slope off Georgia, USA. Both taxa have about 2.0 protoconch whorls and the maximum diameter is slightly similar (IBUFRJ 7584 and holotype of A. conula, respectively: 340 μm vs. 310 μm). However, the protoconch of Aclis aff. conula is more pointed and has a considerably narrower tip (first whorl diameter: 180 μm vs. 255 μm). Moreover, Aclis aff. conula has a wider umbilicus which is bordered by a sharp edge while it is more rounded in A. conula. We can also add that Aclis aff. conula is restricted to the continental shelf, while A. conula is known only from the continental slope. These taxa possibly differ in color since Dall (1927) described A. conula as whitish. However, if the shell of A. conula was found in a worn condition it remains doubtful. Aclis aff. conula can be distinguished from Aclis pyramida Dall, 1927 (Fig. 20 D – F), from the continental slope off Georgia, USA, by having a less globular shape, a slower increase in diameter and a narrower protoconch. Both species reach about 2.0 protoconch whorls, but Aclis aff. conula has a smaller tip (first whorl: 180 μm vs. 284 μm) and maximum diameter (340 μm vs. 415 μm). Aclis aff. conula differs from A. rushi (Fig. 17 C – E) and A. tenuis (Fig. 18 A, B) because it has a broader shape, a wider umbilicus, an opisthocline and more advanced outer lip. We refrain from describing Aclis aff. conula as a new species because of the lack of sufficient material.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7BDE2AFF4FFA89716EFB76.taxon	description	Fig. 12	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7BDE2AFF4FFA89716EFB76.taxon	materials_examined	Type material. Holotype. MNRJ 23777; R / V Ocean Stalwart coll., 28 / vii / 2019. Paratypes: Brazil: Espírito Santo state: South of Abrolhos Bank, JOPS-II- 8 sta. 3228; 19 ° 45 ′ 30 ″ S, 38 ° 45 ′ 48 ″ W, 1100 m; 14 / iv / 1995; R / V Victor Hensen; MNRJ 22634 [1]; Rio de Janeiro state: Campos Basin; 22 ° 18 ′ 53 ″ S, 40 ° 11 ′ 59 ″ W, 200 m; 18 / xii / 2004; Supply Boat Astro Garoupa; MNRJ 33559 [1 †]; Santa Catarina state: PADCT 6611; W, MNRJ 60249 [2]; MNRJ 23603 [1]; off Itajaí; 27 ° 02 ′ 14 ″ S, 46 ° 53 ′ 51 ″ W, 200 m; vii / 2004; MNRJ 26730 [1 †]. Type locality. Brazil: Rio de Janeiro state: PCR BS 4 sta. G 6; 23 ° 48 ′ 57.96 ″ S, 42 ° 40 ′ 31.8 ″; 404 m. Material examined: The types and: Brazil: Rio de Janeiro state: Campos Basin; 22 ° 18 ′ 53 ″ S, 40 ° 11 ′ 59 ″ W, 200 m; 18 / xii / 2004; Supply Boat Astro Garoupa col.; MNRJ 35546 [17 †]; Revizee Sul sta. 6660; MNRJ 28256 [1 †]; Santos Basin; 24 ° 20 ′ 40 ″ S, 43 ° 50 ′ 21 ″ W, 610 m; 29 / i / 2009; MNRJ 35926 [4 †]; São Paulo state: Revizee Sul sta. 6671; MNRJ 28257 [1 †]; PADCT sta. 6587; MNRJ 28246 [1 †]; MNRJ 28255 [2 †]; Paraná state: Revizee Sul sta. 6703; MNRJ 28260 [1 †]; Revizee Sul sta. 6685; MNRJ 28258 [4 †]; PADCT sta. 6642; MNRJ 28252 [1 †]; Revizee Sul sta. 6653; MNRJ 28254 [1 †]; Revizee Sul sta. 6652; MNRJ 28253 [4 †]; Santa Catarina state: Revizee Sul sta. 6695; MNRJ 28259 [3 †]; PADCT sta. 6595; MNRJ 28247 [3 †]; off Itajaí; 150 m; 28 / i / 2005; MNRJ 28843 [3 †]; PADCT sta. 6635; MNRJ 28251 [9 †]; PADCT sta. 6601; MNRJ 28248 [5 †]; PADCT sta. 6608; MNRJ 28249 [23 †]; type locality; MNRJ 28250 [9 †].	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7BDE2AFF4FFA89716EFB76.taxon	etymology	Etymology. Named after our dear friend, Dr. Paulo Márcio Santos Costa, who donated part of the specimens studied here.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7BDE2AFF4FFA89716EFB76.taxon	description	Description. Shell vitreous or whitish, tall, subcylindrical, with an obtuse apex, reaching about 5.1 mm long, 1.7 mm wide. Protoconch globose, subcylindrical, about 2.0 whorls, 580 μm wide, smooth, transition to teleoconch marked by a gentle incremental scar. Teleoconch reaching about five whorls, strongly convex outline; suture deep, well impressed, sloping; surface glossy, presenting faint microscopic spiral threads in subsutural zone of each whorl, growth lines well-impressed, incremental scars strongly demarcated, appearing at irregular intervals. Last whorl about 45 % of total length, broad; base rounded. Aperture high, elliptical in shape, about 65 % of body whorl length, strongly expanded laterally, rounded and with a concavity in the anterior edge, acute posteriorly; outer lip thickened, raised suture near adapical margin, strongly opisthocline, gradually advancing forward, rectilinear; inner lip thick, reflected. Umbilical fissure usually present. Measurements. Holotype, MNRJ 23776: 7 whorls; SL = 4.8 mm; BWL = 2.2 mm; AL = 1.4 mm; SW = 1.6 mm; AW = 1.0 mm. Paratype, MNRJ 33559: 7 whorls; SL = 5.1 mm; BWL = 2.4 mm; AL = 1.6 mm; SW = 1.7 mm; AW = 1.1 mm. Paratype, MNRJ 22634: 6 whorls; SL = 3.8 mm; BWL = 2.0 mm; AL = 1.3 mm; SW = 1.5 mm; AW = 0.9 mm. Paratype, MNRJ 60249: 4 whorls; SL = 2.7 mm; BWL = 1.6 mm; AL = 0.9 mm; SW = 1.1 mm; AW = 0.6 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA7BDE2AFF4FFA89716EFB76.taxon	discussion	Remarks. This species is known only from empty shells, which are frequent in depths close to the continental shelf break and upper slope. The deepest record was found in 1100 m, off the coast of Espírito Santo state, but the single shell (paratype MNRJ 22634) was probably carried out by turbidity currents. Aclis costai sp. nov. has faint microscopic spiral striae in the subsutural region of each whorl (Fig. 12 I) like other Aclis species. However, the axial growth lines are more conspicuous than the spiral sculpture. The general shape of the shell, protoconch and aperture also suggests the present generic classification. The shell shape (Fig. 12 A – F) and the concavity, with a flattened surface in the anterior area of the aperture (Fig. 12 G) are diagnostic, differing the present species from most of his congeners. The anterior part of the aperture is commonly damaged, giving the appearance of a siphonal canal (e. g., Fig. 12 A, C, D), but it is a flattened area that could be recognized in shells with an undamaged aperture in the lower part (Fig. 12 E). Aclis costai sp. nov. resembles Aclis georgiana (Fig. 19 G – H) but can be distinguished by having a much broader length and width, a faster increase in diameter and a wider aperture, which is more expanded laterally and anteriorly than in A. georgiana. Geographic Distribution. Brazil: Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina. From 70 m to 1100 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA79DE2AFF4FFAED754BF82B.taxon	materials_examined	Type material. Holotype: MZSP 77988; R / V Natureza col., 26 / xi / 2001. Paratypes: Brazil: Rio Grande do Norte: Revizee Nordeste; 04 ° 51 ′ 00 ″ S, 35 ° 06 ′ 46 ″ W, 375 m; 24 / xi / 2001; R / V Natureza; MZSP 77989 [2]; Revizee Nordeste; 06 ° 14 ′ 24 ″ S, 34 ° 52 ′ 06 ″ W, 500 m; 26 / xi / 2001; R / V Natureza; MZSP 77990 [1]: LMUFRPE 022 [1]. Type locality. Brazil: Rio Grande do Norte; Revizee Nordeste; 06 ° 14 ′ 04 ″ S, 34 ° 52 ′ 33 ″ W, 510 m. Material examined. Only the holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA79DE2AFF4FFAED754BF82B.taxon	description	Measurements. Holotype, MZSP 77988: 3 whorls; SL = 1.08 mm; BWL = 0.80 mm; AL = 0.54 mm; SW = 0.74 mm; AW = 0.50 mm. Paratype LMUFRPE 022: 3 whorls; SL = 1.08 mm; BWL = 0.81 mm; AL = 0.54 mm; SW = 0.71 mm; AW = 0.50 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA79DE2AFF4FFAED754BF82B.taxon	discussion	Remarks. Aclis macrostoma is known only by the type series and has conspicuous characteristics among the taxa of the present study, such as the broad elliptical aperture, having about half the total size of the shell, and the outer lip vastly expanded in comparison to the other species (Barros et al. 2007: Figs. 1, 5). Although type specimens have a tiny size, the expanded aperture, well-formed inner lip and the relatively sizeable umbilical fissure indicate that this may represent an adult specimen. Geographic Distribution. Brazil: Rio Grande do Norte (Barros et al. 2007). From 375 to 510 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA77DE27FF4FFF407501FD86.taxon	description	Fig. 13	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA77DE27FF4FFF407501FD86.taxon	materials_examined	Type material. (see Signorelli et al. 2015 for notes and illustrations of the type material). Holotype: MACN-In 28924. Paratypes: Argentina: from the type locality: MACN-In 28294 - 1 [1]. Type locality. Argentina: Buenos Aires province: Mar del Plata, Chapadmalal; N. Landoni leg., 1976. Material examined. Brazil: Bahia state: Royal Charlote Bank Revizee Central V sta. 07 r: IBUFRJ 14050 [1 †]; Bank, Revizee Central V sta. 07 r: IBUFRJ 14050 [1 †]. Espírito Santo state: South edge of Abrolhos Shelf, Revizee Central V sta. 25 a: IBUFRJ 14008 [1 †]; Vitória-Trindade Chain: Jaseur Bank, Revizee Central V sta. 23 r: IBUFRJ 13998 [1 †]; Rio de Janeiro state: HAB 11 sta. D 2: MNRJ 15962 [3 †]; MNRJ 16969 [1 †]; HAB 11 sta. E 3: MNRJ 15963 [3 †]; MNRJ 16851 [1 †]; HAB 11 sta. E 2: MNRJ 15964 [1 †]; MNRJ 16583 [3 †]; HAB 11 sta. A 1: MNRJ 15965 [1 †]; HAB 11 sta. F 2: MNRJ 15966 [1 †]; MNRJ 16582 [1 †]; MNRJ 16966 [1 †]; HAB 11 sta. G 3: MNRJ 15968 [1 †]; Macaé, Santana Archipelago; 22 ° 24 ′ 30 ″ S, 41 ° 42 ′ 30 ″ W, 38 m; S / B Astro Garoupa col., v / 1993: MNRJ 22141 [5 †]: off Rio de Janeiro; 23 ° 49 ′ 04.20 ″ S, 42 ° 46 ′ 48.60 ″ W, 225 m; R / V Professor Wladimir Besnard col. Xi / 1997; MNRJ 28195 [7 †]: Campos Basin; MNRJ 28198 [1 †]; Revizee Sul sta. 6653: MNRJ 60185 [1]; PADCT sta. 6627: MNRJ 60254 [1]; Cabo Frio VII sta. 6147;: IBUFRJ 2428 [1 †]; Cabo Frio VII sta. 6165;: IBUFRJ 2429 [6 †]; IBUFRJ 14813 [4 †]; Cabo Frio VII sta. 6199; IBUFRJ 2430 [2 †]; Cabo Frio VII sta. 6198; IBUFRJ 2431 [1 †]; Macaé, Cabiúnas; S / B Astro Garoupa col., 20 / v / 1993: IBUFRJ 7223 [1 †]; Macaé, Santana Archipelago, sta. 9; 22 ° 21 ′ 30 ″ S, 41 ° 41 ′ W, 15 m; S / B Astro Garoupa col., v / 1993: IBUFRJ 7410 [11 †]; Cabo Frio VII sta. 6194; IBUFRJ 7463 [1 †]; Geomar XII sta. 76; IBUFRJ 7570 [1 †]; Geomar XII sta. 125; IBUFRJ 7693 [1 †]; Revizee V sta. 52 f: IBUFRJ 13344 [2 †]; São Paulo state: Revizee sul sta. 6656: MNRJ 28196 [1 †]; Revizee sul sta. 6680: MNRJ 28197 [5 †]; PADCT sta. 6631: MNRJ 60260 [1]; Revizee sul sta. 6652: MNRJ 60268 [2]; Caraguatatuba; 23 ° 53 ′ 097 ″ S, 45 ° 30 ′ 843 ″ W, 15 / ii / 2001: MZSP 42037 [2]; Ubatuba, sta. 4854; 23 ° 47 ′ S, 44 ° 58 ′ W, 47 m: MZSP 85269 [3]; Ubatuba; 23 ° 53 ′ S, 45 ° 09 ′ W, 38 m; R / V Véliger II col., 10 / x / 1986: MZSP 85401 [18]; MZSP 88368 [40]. Revised description. Shell vitreous or whitish, tall, conical, with an obtuse apex, reaching about 4.1 mm long, 1.2 mm wide. Protoconch globose, subcylindrical, about 2.0 whorls, 400 μm high, smooth, transition to teleoconch marked by a gentle incremental scar. Teleoconch reaching about eight whorls, regularly convex outline; suture slightly deep, well impressed, sloping; surface glossy, presenting faint microscopic spiral striae in subsutural zone of each whorl; incremental scars strongly demarcated, appearing at irregular intervals; presence of spiral keels, emerging faintly in the first teleoconch whorl, in number of two in the initial whorls, reaching five in the body whorl of holotype. Last whorl about 40 % of total length, broad; base rounded. Aperture high, elliptical in shape, about 60 % of body whorl length, slightly expanded laterally, rounded and slightly advanced anteriorly, acute posteriorly; outer lip thin, orthocline, rectilinear; inner lip thin, reflected. Umbilical fissure present.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA77DE27FF4FFF407501FD86.taxon	description	Measurements. Holotype (MACN-In 28924): 8 whorls; SL = 4.13 mm; BWL = 1.59 mm; AL = 0.91 mm; SW = 1.19 mm; AW = 0.67 mm. MNRJ 15964: 8 whorls; SL = 3.62 mm; BWL = 1.41 mm; AL = 0.83 mm; SW = 1.03 mm; AW = 0.64 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA77DE27FF4FFF407501FD86.taxon	discussion	Remarks. The present species was originally described in the genus Turritellopsis G. O. Sars, 1878 (Castellanos & Landoni 1984), which was included in Turritellidae until Warén (1996) transferred it to Mathildidae based on anatomical characteristics studied by him and by Haszprunar (1988). The type species of Turritellopsis, Turritellopsis stimpsoni, Dall, 1919 [= Turritellopsis acicula (Stimpson, 1851)] was revised and figured by Warén (1996). The shell has convex whorls with sharp spiral keels, similar to those present in Aclis marplatensis comb. nov. (Fig. 13 A – E). However, T. stimpsoni presents the intervals between the spiral keels sculptured by microscopic axial threads (Warén 1996: Fig. 28 A – C) and lacks the microscopic spiral striae adjacent to the sutures. Aclis marplatensis has no axial sculpture and has the microscopic spiral striae in the intervals between the suture and the subsutural spiral cord (Fig. 13 H, I), which leads to the new combination proposed here. The holotype of the species was recently illustrated with SEM images (Signorelli et al. 2015: fig. 6 K – L). Based on these images, it is possible to assure that there are no axial threads in the holotype compared to what was described and figured by Castellanos & Landoni (1984). The first record of the present species in Brazil was reported by Absalão et al. (1996) (as “ Turritellopsis marplatensis ”). These authors described that the shells from Brazil do not present microscopic axial sculpture, but only microscopic spiral striae as described above. A revised description of the shell is provided here to elucidate the shell morphology of the species. The development of the spiral keels is very homogeneous in A. marplatensis, with all of them showing a similar degree of elevation from the teleoconch surface and does not vary among the specimens studied. This pattern seems robust, taking into consideration that this is one of the most abundant species studied. Aclis marplatensis can be distinguished from Aclis eolis (Fig. 19 A – D), Aclis floridana Bartsch, 1911 (see Peñas & Rolán 2013: pl. 2, Fig. 2 A, B for illustrations of the type material), Aclis hendersoni Dall, 1927 (Fig. 19 E – F), all from Florida, USA, and from Aclis trilineata Watson, 1897 (see Bogi et al. 2016: Fig. 1 A – F, for illustrations), from Madeira and western Mediterranean, by the more pronounced spiral keels of the teleoconch and by the less expanded aperture anteriorly and laterally. Geographic Distribution. Brazil: Bahia, Espirito Santo, Vitória-Trindade Chain, Rio de Janeiro, São Paulo (Absalão 1996; present study); Argentina: Mar del plata (Castellanos & Landoni 1984). From 29 to 450 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA74DE21FF4FFDBD75C7FE16.taxon	description	Fig. 14	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA74DE21FF4FFDBD75C7FE16.taxon	materials_examined	Type material. Holotype: MNRJ 23777, R / V Ocean Stalwart col. 28 / ix / 2019. Paratypes. Brazil: Rio de Janeiro state: PADCT sta. 6541; MNRJ 28199 [5 †]. Type locality. Brazil: Santos Basin, off Rio de Janeiro state; PCR BS 4 sta. G 6; 23 ° 48 ′ 57.96 ″ S, 42 ° 40 ′ 31.8 ″ W, 404 m. Material examined. The types and: Brazil: Rio de Janeiro state: PADCT sta. 6541: MNRJ 28205 [10 †]; PADCT sta. 6627: MNRJ 28203 [1 †]; São Paulo state: Revizee Sul sta. 6662: MNRJ 28210 [2 †]; Revizee Sul sta. 6669: MNRJ 28211 [1 †]; Revizee Sul sta. 6660: MNRJ 28209 [1 †]; PADCT sta. 6625: MNRJ 28208 [2 †]; Revizee Sul sta. 6674: MNRJ 28212 [1 †]; Paraná state: Revizee Sul sta. 6703: MNRJ 28213 [2 †]; PADCT sta. 6641: MNRJ 28214 [1 †]; Santa Catarina state: PADCT sta. 6595: MNRJ 28207 [14 †]; Revizee Sul sta. 6672: MNRJ 28206 [1 †]; PADCT sta. 6635: MNRJ 28204 [1 †]; PADCT sta. 6601: MNRJ 28200 [2 †]; PADCT sta. 6606: MNRJ 28201 [2 †]; PADCT sta. 6608: MNRJ 28202 [2 †].	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA74DE21FF4FFDBD75C7FE16.taxon	etymology	Etymology. The specific epithet is a Latin singular feminine adjective in the nominative case meaning carved or engraved in bas-relief, which refers to the teleoconch sculpture.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA74DE21FF4FFDBD75C7FE16.taxon	description	Description. Shell whitish, tall, conical, with an obtuse apex, spire angle about 16 °, reaching about 4.3 mm long, 1.4 mm wide. Protoconch globose, subcylindrical, about 2.0 whorls, about 320 μm high and 260 μm wide, presence of microscopic spiral lines regularly spaced, transition to teleoconch marked by an abrupt incremental scar and onset of spiral keels. Teleoconch up to seven whorls, moderately convex outline, crenulated; suture slightly deep, well impressed, sloping; presence of microscopic spiral striae in whole surface of each whorl; presence of sharp, pronounced spiral keels, two since the onset of teleoconch, reaching three at the body whorl; incremental scars strongly demarcated, appearing at irregular intervals. Last whorl about 40 % of total length; base carinated, flattened below. Aperture elliptical in shape, about 55 % of body whorl length, slightly expanded laterally, rounded or pointed anteriorly, acute posteriorly; outer lip thin, orthocline in adults, prosocline in juveniles, gradually advancing forward, slightly sinuous, maximum projection close to distal region, retracted at distal region; inner lip thin, reflected. Umbilical fissure present. Measurements. Holotype, MNRJ 23777: 9 whorls; SL = 4.3 mm; BWL = 1.7 mm; AL = 1.0 mm; SW = 1.4 mm; AW = 0.8 mm. Paratype, MNRJ 28199: 7 whorls; SL = 3.0 mm; BWL = 1.2 mm; AL = 0.7 mm; SW = 1.0 mm; AW = 0.6 mm. Paratype, MNRJ 28199: 6 whorls; SL = 2.4 mm; BWL = 1.0 mm; AL = 0.5 mm; SW = 0.9 mm; AW = 0.5 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA74DE21FF4FFDBD75C7FE16.taxon	discussion	Remarks. Aclis anaglyptica sp. nov. presents the microscopic spiral striae, which are very numerous and thin (Fig. 14 G, H). The aperture is relatively short and slightly expanded laterally (Fig. 14 B) like in other Aclis species. Aclis anaglyptica can be easily recognized by its short, globular protoconch with microscopic spiral lines (Fig. 14 I – K), presence of sharp and strongly pronounced spiral keels beginning since the onset of teleoconch in all shells examined (Fig. 14 A – J). These features are the most conspicuous to recognize this species. Aclis anaglyptica sp. nov. differs from A. marplatensis (Fig. 13 A – I) by the fewer number of spiral keels in the body whorl (3 vs. 5), which are more pronounced and sharper than in A. marplatensis. Furthermore, the spiral keels of A. anaglyptica sp. nov. starts at the onset of teleoconch whorls, earlier than in A. marplatensis. Aclis anaglyptica sp. nov. differs from A. eolis (Fig. 19 A – D) and A. hendersoni (Fig. 19 E – F) by the less advanced aperture anteriorly and laterally. Additionally, A. anaglyptica sp. nov. differs from A. eolis by the more pronounced spiral keels and A. hendersoni by the homogeneous development of the spiral keels, while the latter has a single well pronounced spiral keel in the upper region of each whorl and the remaining ones are weaker. Aclis anaglyptica sp. nov. resembles Aclis maoria Powell, 1937, from New Zealand, but the former is more elongated and with a more advanced outer lip. Geographic Distribution. Brazil: Rio de Janeiro, São Paulo, Paraná, Santa Catarina. From 101 m to 980 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA73DE23FF4FFF407414FEDE.taxon	description	Fig. 15	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA73DE23FF4FFF407414FEDE.taxon	materials_examined	Type material. Holotype: MNRJ 28215 [†]; R / V Professor Wladimir Besnard coll., xi / 1997 - i / 1998. Paratypes: Brazil: Paraná state: Revizee Sul sta. 6699: MNRJ 2821 7 [1 †]; PADCT sta. 6641 (23 ° 35 ′ 49 ″, 41 ° 42 ′ 25 ″, 143 m): MNRJ 28216 [1 †]. off Itajaí; 26 ° 38 ′ 45 ″ S, 46 ° 51 ′ 54 ″ W, 150 m; ii / 2004: MNRJ 36004 [1 †]. Type locality. Brazil: Santa Catarina state: PADCT sta. 6635; 27 ° 10 ′ 23 ″ S, 47 ° 27 ′ 33 ″ W, 129 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA73DE23FF4FFF407414FEDE.taxon	etymology	Etymology. From the Latin adjective multus, meaning many, and carinatus, meaning keeled; referring to the plethora of spiral keels.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA73DE23FF4FFF407414FEDE.taxon	description	Description. Shell tall, conical, spire angle 14 °, reaching about 4.2 mm long, 1.2 mm wide, eight whorls, whitish. Protoconch globose, apex dome-shaped, about 3.0 whorls, 315 μm high, surface smooth. Teleoconch with moderately convex whorls, moderately convex outline, crenulated; suture slightly deep; presence of microscopic spiral striae near suture; presence of sharp, pronounced spiral keels, first teleoconch whorl smooth, two spiral keels emerging at second whorl, remaining whorls with a progressive number of keels, reaching seven at the body whorl; incremental scars strongly demarcated, appearing at irregular intervals. Body whorl about 40 % of the total length, base crenulated; aperture high, about 55 % of body whorl length, rounded anteriorly, acute posteriorly; outer lip thin, sinuous, slightly advanced close to distal area; inner lip thin, reflected. Umbilical fissure present. Measurements. Holotype, MNRJ 28215: 8 whorls; SL = 4.2 mm; BWL = 1.6 mm; AL = 0.9 mm; SW = 1.2 mm; AW = 0.7 mm.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA73DE23FF4FFF407414FEDE.taxon	discussion	Remarks. Aclis multicarinata sp. nov. is conspicuous by the late development of the spiral keels that starts around the third teleoconch whorl as faint elevations (Fig. 15 E). In the last whorls the spiral keels are more numerous and more pronounced (Fig. 15 A−B). The body whorl of the largest specimen has six spiral keels and one beginning at the basal region (Fig. 15 C, H). In addition to the features cited above, A. multicarinata sp. nov. can be distinguished from A. marplatensis comb. nov. by having a higher number of microscopic spiral striae, a more circular aperture, and a more advanced outer lip in the distal region; from A. anaglyptica sp. nov., by having a smaller interval between the spiral cords, a more rounded outline of the teleoconch whorls and a less expanded aperture anteriorly. Aclis multicarinata sp. nov. resembles A. maestratii Poppe & Tagaro, 2016, from the Philippines, but it differs by the late development of the spiral keels, which starts consistently since the beginning of the teleoconch in A. maestratii. Besides that, A. multicarinata sp. nov. has less pronounced spiral keels and a wider interval between the suture and the first spiral keel. Geographic Distribution. Brazil: Santa Catarina, Paraná. From 129 m to 150 m.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA70DE23FF4FFE0D70B2FC26.taxon	description	Fig. 16	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA70DE23FF4FFE0D70B2FC26.taxon	materials_examined	Type material. Holotype (herein examined): USNM 94288. Type locality. Cuba: Blake Collection; 1426 m; W. H. Rush coll. Material examined. The holotype and: USA: off Georgia: USFC sta. 2668, 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ W, 538 m, Steamer Albatross coll.: USNM 108042.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA70DE23FF4FFE0D70B2FC26.taxon	discussion	Remarks. MolluscaBase (2021 a) currently considers Aclis dalli Bartsch, 1911, from Cuba, (Fig. 16) as a junior synonym of A. sarissa, but there is no reference to the publication in which such synonymy is discussed. These species resemble each other by the convexity of teleoconch whorls and by the slightly oval aperture. However, A. sarissa is considerably shorter, narrower and with a wider apex than A. dalli, which should be considered as a valid species.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE22FF4FFB007211F9DB.taxon	description	Figs 18 C – D	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE22FF4FFB007211F9DB.taxon	materials_examined	Type material. Holotype (herein examined): USNM 108047. Type locality. USA: Georgia state Off Georgia, USFC sta. 2668; 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ W, 538 m; Steamer Albatross col, 05 / v / 1886. Not “ off Fernandina ” as stated in the original description (vide station number of the original label). Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE22FF4FFF407211FE5E.taxon	description	Figs 17 A – B	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE22FF4FFF407211FE5E.taxon	materials_examined	Type material. Holotype (herein examined): USNM 108019. Type locality. USA: Georgia state: USFC sta. 2668; 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ WW, 538 m; Steamer Albatross col.; 05 / v / 1886. Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE22FF4FFDC47211FC86.taxon	description	Figs 17 C – E	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE22FF4FFDC47211FC86.taxon	materials_examined	Type material. Holotype (herein examined): USNM 82973. Type locality. USA: off Fowey Rocks; 274 to 366 m; W. H. Rush coll. Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE22FF4FFCBC71BCFB1A.taxon	description	Figs 18 A – B	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE22FF4FFCBC71BCFB1A.taxon	materials_examined	Type material. Holotype (herein examined): USNM 44821. Type locality. USA: New Jersey state: off Martha’s Vineyard, USFC sta. 873; 40 ° 1 ’ 48.00 ” N, 70 ° 57 ’ W, 183 m; R / V Fish Hawk col., 13 / ix / 1880. Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE1CFF4FF94274B7FE32.taxon	description	Figs 18 E – I	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE1CFF4FF94274B7FE32.taxon	materials_examined	Type material. Holotype (herein examined): USNM 44811. Type locality. USA: New Jersey state: off Martha’s Vineyard, USFC sta. 894; 39 ° 53 ′ N, 70 ° 58 ′ W, 668 m; R / V Fish Hawk col., 02 / x / 1880. Material examined. The holotype and: holotype of Aclis tanneri: USNM 417444: USA: New Jersey state: Hudson Canyon, USFC sta. 2234; 39.15 N, 70.0542 W, 1481 m; R / V Fish Hawk col., 13 / ix / 1884.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA71DE1CFF4FF94274B7FE32.taxon	discussion	Remarks. MolluscaBase (2021 a) considers Aclis tenuis as the senior synonymy of a group of very similar species described from material of the USFC from New Jersey and Georgia localities: Aclis verrilli Bartsch, 1911, Aclis fernandinae Dall, 1927 and Aclis tanneri Bartsch, 1947. Such synonymy was proposed by Bouchet & Waren (1983: 306), who also speculated about the possible synonymy with A. walleri; however, the authors were somewhat dubious about the occurrence of A. walleri in West Atlantic and stated that the gap of distribution should be filled for a better comprehension. The occurrence of A. walleri in the western Atlantic (recorded by Verrill, 1880: 391; 1882: 528) was also contested by Bartsch (1911: 438) who considered that the material studied by Verrill (1880, 1882: 377), from the USFC are distinct from A. walleri and he described Aclis verrilli. In the present study, all types of the above names were examined, and some conclusions could be taken about the validity of them. Aclis verrilli (Fig. 18 E – I) is a valid species and can be distinguished from A. tenuis (Fig. 18 A – B) because it has a considerably narrower and multispiral protoconch, besides a faster increase in diameter in the teleoconch. Aclis tanneri (Fig. 18 H – I), by the other hand, cannot be distinguished from A. verrilli. The shell of Aclis tenuis and Aclis verrilli are considerably wider and conical, while A. fernandinae (Fig. 18 C – D) has a more cylindrical shape. Sexual dimorphism is known in Eulimidae with females usually bigger than males (Warén 1983), but until we have evidence for this in Aclis we consider both as valid species.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4CDE1FFF4FF9B47211F82E.taxon	description	Figs 20 A – C	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4CDE1FFF4FF9B47211F82E.taxon	materials_examined	Type material. Holotype (herein examined): USNM 108022. Type locality. USA: Georgia state: Off Georgia, USFC sta. 2668; 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ W, 538 m; Steamer Albatross col., 05 / v / 1886. Not “ off Fernandina ” as stated in the original description (vide station number of the original label). Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4CDE1FFF4FFF407211FE5E.taxon	description	Figs 19 A – D	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4CDE1FFF4FFF407211FE5E.taxon	materials_examined	Type material. Holotype (herein examined): USNM 573617. Type locality. USA: Florida state: Key West, Sand Key; 25 ° 30 ’ N, 80 ° 10 ’ 48.00 ” W, 119 m; 1913; R / V Eolis col. Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4CDE1FFF4FFDC47211FD6A.taxon	materials_examined	Type material. Holotype (herein examined): USNM 82973 A. Type locality. USA: Florida state, Florida Straits, off Fowey Rocks; 274 to 366 m; W. H. Rush coll. Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4CDE1FFF4FFB147211FA72.taxon	description	Figs 19 G – H	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4CDE1FFF4FFB147211FA72.taxon	materials_examined	Type material. Holotype (herein examined): USNM 108020. Type locality. USA: Georgia state: USFC sta. 2668; 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ W, 538 m; Steamer Albatross col., 05 / v / 1886. Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4CDE1FFF4FFC907211FBEE.taxon	description	Figs 19 E – F	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4CDE1FFF4FFC907211FBEE.taxon	materials_examined	Type material. Holotype (herein examined): USNM 333465. Type locality. USA: Gerogia state: USFC sta. 2668; 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ W, 538 m; Steamer Albatross col., 05 / v / 1886. Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4DDE1EFF4FF95B7211F8E1.taxon	description	Figs 20 D – F	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4DDE1EFF4FF95B7211F8E1.taxon	materials_examined	Type material. Holotype (herein examined): USNM 108223. Type locality. USA: Georgia state: Off Georgia, USFC sta. 2668; 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ W, 538 m; Steamer Albatross col., 05 / v / 1886. Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4ADE18FF4FF9187144FE16.taxon	description	Figs 21 A – G	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4ADE18FF4FF9187144FE16.taxon	materials_examined	Type material. Holotype (herein examined): USNM 108024. Type locality. USA: Georgia state: Off Georgia, USFC sta. 2668; 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ W, 538 m; Steamer Albatross col., 05 / v / 1886. Material examined. The holotype and: holotype of A. pendata: USNM 108021: USA: Georgia state: Off Georgia, USFC sta. 2668; 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ W, 538 m; Steamer Albatross col., 05 / v / 1886.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4ADE18FF4FF9187144FE16.taxon	discussion	Remarks. MolluscaBase (2021 a) considers A. stilifer as a synonym of A. pendata Dall, 1927. The holotype of A. stilifer (USNM 108024) (Figure 21 A – D) and the holotype of A. pendata (USNM 108021) (Figure 21 E – G) are proportionally similar in size, including the width of the first protoconch whorl (ca. 250 μm). The transition between protoconch-teleoconch is not clear (Figure. 21 D, G). We follow this synonymy and keep the priority with the name A. stilifer.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4BDE1BFF4FFA3874ECFE16.taxon	description	Fig. 22	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4BDE1BFF4FFA3874ECFE16.taxon	materials_examined	Type material. Lectotype (herein designated): NHMUK 1889.10. 1.612. Paralectotypes: Saint Helena: NHMUK 1889.10.1.613 – 616 [4]. Type locality. Saint Helena. Material examined. Saint Helena: ANSP 67898 [1].	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA4BDE1BFF4FFA3874ECFE16.taxon	discussion	Remarks. The current classification of this species in the Microstelma A. Adams, 1863 (Zebinidae) (MolluscaBase 2023 c) should be revised. It was originally described in the genus Amaurella A. Adams, 1867, which was demonstrated by Ponder (1985) to be a junior synonym of Microstelma. Ponder (1985) described the presence of weak to moderate axial ribs that do not extend over the base of the teleoconch whorls in Microstelma, which are lacking in Aclis. The original description, observations on the type series (Fig. 22 A, B) and of one topotype clearly indicate that the species has a completely smooth teleoconch, except for strong incremental scars typical of Eulimidae. Therefore, we propose a new combination of the species in Aclis. The shell herein designated as the lectotype is the figured specimen by E. A. Smith (1890: pl. 23, Fig. 27) (Fig. 22 A). All shells of the type series are glued to a cardboard (Fig. 22 A, B). The topotype of A. canaliculata comb. nov. from the ANSP (Fig. 22 C, D) contains a label by A. Warén (10 Feb. 2003) indicating that this shell is a type and most probably a syntype. This shell was donated to the ANSP by G. B. Sowerby and there is no evidence that E. A. Smith studied this specimen. Thus, we do not consider the shell of ANSP 67898 as a syntype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA48DE1BFF4FFD2D71CBFB0E.taxon	description	Figs 23 A, B	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA48DE1BFF4FFD2D71CBFB0E.taxon	materials_examined	Type material. Syntypes: NHMUK 1889.10.1.749 – 767 [19]. Type locality. Saint Helena.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA48DE1BFF4FFD2D71CBFB0E.taxon	discussion	Remarks. The presence of numerous axial ribs through the teleoconch surface does not fit in Aclis, but a precise classification in another group remains dubious. This name is a junior homonym of Aclis angulata P. Fischer in de Folin & Périer, 1869, which is currently considered a taxon inquirendum by MolluscaBase (2021 b). As the species of E. A. Smith is not an Aclis, we refrain from giving a new name for this species that should be placed in another genus or even family.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA48DE1BFF4FFB357179F9AE.taxon	description	Figs 23 C, D	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA48DE1BFF4FFB357179F9AE.taxon	materials_examined	Type material. Syntypes: NHMUK 1889.10.1.775 – 779 [5]. Type locality. Sanit Helena.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA48DE1BFF4FFB357179F9AE.taxon	discussion	Remarks. Aclis didyma is very similar to ‘ Aclis angulata E. A. Smith, 1890 ’ (see comments above) and has several axial ribs at small intervals in the teleoconch, a feature not known in Aclis. This taxon needs relocation to another group.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA48DE1BFF4FF9557450F86A.taxon	description	Figs 23 E – G	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA48DE1BFF4FF9557450F86A.taxon	materials_examined	Type material. Syntypes: NHMUK 1889.10.1.768 – 770 [3]. Type locality. Saint Helena.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA48DE1BFF4FF9557450F86A.taxon	discussion	Remarks. The heterostrophic protoconch indicates that this species is not a member of Aclis. A tentative placement in Ebala Gray, 1847 is here proposed. Ebala similima (E. A. Smith, 1890) comb. nov. is probably related to other species from West Africa treated by Peñas & Rolán (2001) under Anisocycla Monterosato, 1880 but now placed in Ebala (see Warén 2013 for further details about these generic names).	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA46DE15FF4FFC8673CDFACB.taxon	description	Figs 23 J, K	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA46DE15FF4FFC8673CDFACB.taxon	materials_examined	Type material. Holotype (herein examined): USNM 108037. Type locality. USA: Georgia state: off Georgia, USFC sta. 2668; 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ W, 538 m; Steamer Albatross col., 05 / v / 1886. The locality is not “ off Fernandina ” as stated in the original description (vide station number of the original label). Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA46DE15FF4FFC8673CDFACB.taxon	discussion	Remarks. The present species has inconspicuous shell features, but the almost flat teleoconch whorls and the relatively shorter aperture, not advanced anteriorly contrast with other Aclis species. Besides that, SEM images available for study indicate that the teleoconch does not present the microscopic spiral striae typical of Aclis. A new classification in the genus Fusceulima Laseron, 1955 is here proposed, since the shell morphology of the present species follows the diagnosis provided by Souza & Pimenta (2014).	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA46DE15FF4FFF40715DFC9F.taxon	description	Figs 23 H, I	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA46DE15FF4FFF40715DFC9F.taxon	materials_examined	Type material. Holotype (herein examined): USNM 94290. Type locality. Cuba: off Cuba, sta. 34; 1426 m; Rush leg. Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA46DE15FF4FFF40715DFC9F.taxon	discussion	Remarks. Bartsch (1911) introduced the species in Aclis and later transferred it to Costaclis by the presence of strong axial ribs in the teleoconch (Bartsch 1947). Bouchet & Warén (1986) did not adopt the classification in Costaclis. The present species has a subcylindrical shell shape, a bulbous protoconch with less than 2.0 whorls and the absence of an umbilical fissure, which does not fit Aclis and Costaclis. Based on the previous shell features, a new classification in the genus Graphis is proposed here, despite lacking spiral cords at the lower half of each teleoconch whorl as in Graphis albida (Kanmacher, 1798) (= Turbo unicus Montagu, 1803), the type species of Graphis. Other species without spiral cords and with a very different protoconch morphology are currently considered in Graphis, for example, Graphis gracilis (Monterosato, 1874) (see Gofas et al. 2014: Fig. 7 H, I, J, K).	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA46DE15FF4FFA7270B2F83D.taxon	description	Figs 23 L, M	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA46DE15FF4FFA7270B2F83D.taxon	materials_examined	Type material. Holotype (herein examined): USNM 108044. Type locality. USA: Georgia state: off Georgia, USFC sta. 2668; 30 ° 58 ′ 30 ″ N, 79 ° 38 ′ 30 ″ W, 538 m; Steamer Albatross col., 05 / v / 1886. The locality is not “ off Fernandina ” as stated in the original description (vide station number of the original label). Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA46DE15FF4FFA7270B2F83D.taxon	discussion	Remarks. The present species has almost flat teleoconch whorls, a relatively thicker inner lip without an umbilical fissure, and the SEM images available for study indicate that the teleoconch does not present the microscopic spiral striae typical of Aclis. We propose a tentative classification in the genus Melanella Bowdich, 1822 despite uncertainties about the diagnosis of the latter genus (Bouchet & Warén 1986; Souza & Pimenta 2019 b).	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA47DE14FF4FFBEA7144F934.taxon	description	Figs 24 A – C	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA47DE14FF4FFBEA7144F934.taxon	materials_examined	Type material. Holotype (herein examined): USNM 108389. Type locality. USA: Florida state: off Fernandina, USFC sta. 2415; 30 ° 44 ′ N, 79 ° 26 ′ W, 805 m; Steamer Albatross col., 01 / iv / 1885. Material examined. The holotype.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA47DE14FF4FFBEA7144F934.taxon	discussion	Remarks. Dall (1927) clearly stated that the shell of USNM 108389 is the type specimen. Thus, it should be considered the holotype and the type locality the USFC sta. 2415 (Off Fernandina), as written in the original label. The author also refers to several other specimens from USFC sta. 2668 (Off Georgia). Bartsch (1947) included the present species in Costaclis, which was not adopted later by Bouchet & Warén (1986). A new combination of the present species in Graphis is proposed here, similarly to the case of Graphis cubana commented above. Graphis rhyssa (Dall, 1927) comb. nov. also has strong axial ribs but lacks the spiral sculpture.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA47DE16FF4FF92A724DFEFB.taxon	description	Figs 24 D – E	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA47DE16FF4FF92A724DFEFB.taxon	materials_examined	Type material. Holotype: ZMA. MOLL. 138263. Paratype: Aruba; West coast of Aruba; ZMA. MOLL. 138275 [1]. Type locality. Aruba: West coast of Aruba.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
03CA878ABA47DE16FF4FF92A724DFEFB.taxon	discussion	Remarks. This species was originally described in Bermudaclis Bartsch, 1947, which is a junior synonym of Murchisonella Mörch, 1875 (Peñas & Rolán 2013). Peñas & Rolán (2013: 61) noted that this species is not a Murchisonellidae, but possibly belongs to Aclis. However, the presence of low axial ribs and fine spiral striae in the whole teleoconch surface (Fig. 24 D, E) does not fit in Aclis. The protoconch of this species seems slightly deviated. The study of the type material under SEM would be necessary for better classification, but we here suggest that this species is a ‘ lower heterobranch’.	en	Souza, Leonardo Santos De, Araújo, Tarciso Almeida De, Pimenta, Alexandre Dias (2024): Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena. Zootaxa 5433 (4): 451-499, DOI: 10.11646/zootaxa.5433.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5433.4.1
