taxonID	type	description	language	source
03CB87ABFFC590730E4F10E41BFFFCF2.taxon	diagnosis	Diagnosis. Males of the genus Lomachaeta can be distinguished from other Sphaeropthalmina by the presence of an apical row of stout, lanceolate bristles on metasomal segments 2 – 5, which are directed obliquely inwards (Fig. 1). Females of this genus can be distinguished from other Mutillidae by the following combination of characters: presence of subovate eyes, presence of a reticulated surface on the head and mesosoma (Fig. 5), absence of carinae defining the lateral margins of the pygidial area, presence of apically edentate mandibles, a petiolate first metasomal segment (Fig. 5), and presence of tuberculate propodeal spiracles (Fig. 5). The following additional characters will help differentiate Lomachaeta from other Sphaeropthalmina: males having a tridentate mandible with a ventral tooth or slight excision that is angulate medially, lacking parapsidal lines and notauli, having an inconspicuous pterostigma (Fig. 2), having a distinctly petiolate first metasomal segment, having a hypopygium that is rounded apically but bidentate medially, having reduced wing venation in comparison with other New World species (Fig. 2), and having the cuspis reduced to a knob­like structure (Figs. 14 – 22); females having a genal carina and a pyriform mesosoma (widest anteriorly and gradually narrowing posteriorly) (Fig. 5). The genitalia of male Lomachaeta are of limited taxonomic use because they are quite similar between species (Figs. 3,4, 14 – 22). The genitalia, however, along with the presence or absence of a ventral mandibular tooth, do suggest several relationships. The following three species groups are defined by male characters: L. crocopinna species­group, L. hicksi species­group and L. hyphantria species­group. The L. crocopinna species­group is defined by lack of the ventral mandibular tooth, presence of distinctly brachyplumose pubescence, and presence of long setae on the ventral margin of the parameres (Figs. 19, 20), and includes L. crocopinna, sp. nov. and L. ptilohyalus, sp. nov.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC590730E4F10E41BFFFCF2.taxon	description	The L. hyphantria species­group is defined by the lack of the ventral mandibular tooth, presence of indistinct brachyplumose pubescence (serrate), presence of confluent punctures on the mesonotum (Fig. 13), and absence of long setae on the ventral margin of the parameres (Fig. 21), and includes only L. hyphantria, sp. nov. The L. hicksi species­group is defined by presence of a ventral mandibular tooth, presence of indistinct brachyplumose pubescence (serrate), and absence of long setae on the ventral margin of the parameres (Figs. 14 – 18, 22) and includes L. chionothrix, L. cirrhomeris and L. hicksi. The males of L. argenta, L. ibarrai, and L. vianai remain unknown. As such, this species are not placed into species­groups, because the species­groups defined above are based on characters derived from males. Hosts. Host data suggests that Lomachaeta species are opportunistic parasitoids. Rearing parasitoids from twig and trap nests provided new host records for four more species of Lomachaeta. Host data suggests that the species may be limited to cavity nesting hosts of small size, including Solierella Spinola, Pisonopsis Fox and Trypoxylon Latreille. Specific hosts are listed under each Lomachaeta species. It is noteworthy that the females of the species­group lack a distinct pygidium, as do many twig nesting sphecoid wasps, bees, and their hymenopteran parasitoids, such as mutillid species of the Sphaeropthalma uro species­group.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC590730E4F10E41BFFFCF2.taxon	distribution	Distribution. Lomachaeta ranges throughout the United States, south to Guatemala, and is also found in Argentina and Bolivia. It is likely that Lomachaeta occurs along with its hosts throughout the Neotropics.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC390710E4F122C1B42FBCA.taxon	description	(Figs. 3, 6, 14)	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC390710E4F122C1B42FBCA.taxon	diagnosis	Diagnosis. Presence of large punctures on the medial portion of the pronotum, the totally black brown body with white setae, and black brown trochanters separate this species from all other Lomachaeta. This is the only species that has an aedeagus that is bidentate ventrally (Fig. 14). The punctures of the pronotum (Fig. 6) are similar to some forms of L. hicksi (Fig. 11), but the punctures of the mesonotum are much denser and deeper and the setae of the dorsum are much denser for L. chionothrix.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC390710E4F122C1B42FBCA.taxon	materials_examined	Holotype Male. Head. Width 0.95 mm, black, clothed throughout with sparse, long, erect, pale white setae. Clypeus with several black setae along apical margin. Compound eye maximum diameter 0.21 mm. Median ocellus 0.069 mm, lateral ocellus 0.054 mm, ocellocular distance 0.26 mm, and interocellular distance 0.077 mm. Malar space 0.092 mm. Mandible black, orange apically, distinctly emarginate beneath, with distinct tooth beneath. Antennal tubercle and scape black. Pedicel and flagellomeres brown. Antenna clothed with pale white setae throughout. Length of scape, pedicel and first three flagellomeres: 0.26, 0.077, 0.050, 0.093 and 0.11 mm, respectively. Width of first flagellomeres 0.088 mm. Front reticulate, punctures crowded. Vertex with small, distinct, separated punctures. Mesosoma. Black; clothed throughout with sparse, long, erect, pale white setae. Pronotum with moderately large, irregularly spaced punctures, some well separated, some confluent. Mesonotum with moderately large, close punctures throughout. Scutellum almost flat, with punctures similar to those on mesonotum. Dorsum and posterior face of propodeum broadly reticulate throughout. Tegula dark brown, glabrous, except punctate along margin, with long, pale, erect setae. Legs dark brown, clothed with sparse, long, erect, pale white setae. Tibia with pale white bristles. Tibial spurs pale. Wings with pterostigma 0.14 mm in length along costa. Marginal cell 0.31 mm in length. Second submarginal cell pentagonal, 0.23 mm in length.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC390710E4F122C1B42FBCA.taxon	description	Metasoma. Black, except metasomal segments 6 – 7 brown; clothed throughout with sparse, long, erect, pale white setae. T 5 – T 7 with additional dark setae. Bristles at posterior margins of T 2 – T 4, brown. T 1 mostly impunctate, posterior margin with row of small, contiguous punctures. T 2 with moderately small, distinct, widely separated punctures. T 3 – T 7 with progressively small to fine, separated punctures. S 2 with moderately small, distinct, widely separated punctures. Genitalia Figs. 3, 14; aedeagus bidentate. Length. Approximately 4.5 mm. Female. Unknown.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC390710E4F122C1B42FBCA.taxon	materials_examined	Type material. Holotype ɗ, GUATEMALA, Zacapa, Rio Hondo, 7. VI. 1987, unknown (CNCI). Paratypes, 5 ɗ, same data as holotype (JPPC, DGMC and UGAC). Other material examined. MEXICO, Nayarit, Mecatan, 800 ’, 1 ɗ, 23. V. 1949, G. M. Bradt (AMNH).	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC390710E4F122C1B42FBCA.taxon	description	Variation. The six male specimens vary from 3.6 mm to 4.7 mm and lack any noticeable difference in coloration or sculpture. Hosts. Unknown.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC390710E4F122C1B42FBCA.taxon	etymology	Etymology. From the Greek chion “ snow ” and thrix “ hair ”, in reference to the white setae of the metasoma.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC0907E0E4F153C1A52FD22.taxon	description	(Figs. 4, 7, 15, 28, 29)	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC0907E0E4F153C1A52FD22.taxon	diagnosis	Diagnosis. The male of this species is easily recognized by having a totally black body with orange femora, brown orange tegula and small, sparse punctures medially on the pronotum (Fig. 7). These characters distinguish the male of this species from all other Lomachaeta. The sculpture of the pronotum of the male is similar to that of L. hicksi (Fig. 8). The female can be identified by the totally orange body with a black brown pygidial area and a black fringe of setae on the fifth tergite.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC0907E0E4F153C1A52FD22.taxon	materials_examined	Holotype Male. Head. Width 0.98 mm, black, clothed throughout with sparse, long, erect, pale white setae. Compound eye maximum diameter 0.39 mm. Median ocellus 0.050 mm, lateral ocellus 0.046 mm, ocellocular distance 0.28 mm, and interocellular distance 0.13 mm. Malar space very short, 0.10 mm in length. Mandible black, brown orange apically, distinctly emarginated and toothed beneath. Antennal tubercle brown. Antenna black. Scape clothed with pale white setae throughout. Pedicel and flagellomeres with pale golden setae. Length of scape, pedicel and first three flagellomeres: 0.30, 0.069, 0.077, 0.10 and 0.085 mm, respectively; width of first flagellomeres 0.092 mm. Front reticulate, punctures large and crowded. Vertex with small, distinct, separated punctures. Mesosoma. Black, clothed throughout with sparse, long, erect, pale white and brown setae. Pronotum with moderately large, irregularly spaced punctures along lateral margin, confluent at posterior margin, small and sparsely spaced medially. Mesonotum with small, close punctures throughout. Scutellum almost flat, with punctures similar to those on mesonotum, more crowded than mesonotum. Dorsum and posterior face of propodeum broadly reticulate throughout. Metapleuron and propodeal side glabrous, dorsal margin of metapleuron finely rugose, and reticulations of dorsum and posterior face of propodeum extending slightly onto propodeal side. Tegula brown orange, glabrous, except entire margin punctate and with long, pale, erect setae. Legs brown orange with femora orange, clothed with sparse, long, erect, pale white setae. Tibia with pale white bristles. Tibial spurs pale. Wings with pterostigma 0.22 mm in length along costa. Marginal cell 0.26 mm in length. Second submarginal cell pentagonal, 0.31 mm in length.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC0907E0E4F153C1A52FD22.taxon	description	Metasoma. Black except metasomal segments 6 – 7 brown. Metasoma clothed throughout with sparse, long, erect, pale white setae. T 5 – T 7 with dark setae. Bristles at posterior margins of T 2 – T 4 black brown. T 1 punctate on lateral margins, glabrous medially, with row of very large, contiguous punctures along posterior margin. T 2 with small, distant, widely separated punctures. T 3 – T 7 with progressively small to fine, separated punctures. S 2 with large, close punctures throughout. Genitalia Fig. 4, 15. Length. Approximately 5.4 mm. Allotype Female. Head. Width is 0.77 mm, orange. Flagellum darkened orange brown. Head clothed with very sparse, erect, black setae, except additional short pale white setae on clypeus and gena. Antennae, maxillary and labial palps covered with very short pale silvery setae. Mandible edentate, not emarginate nor toothed beneath. Front and vertex broadly reticulate. Posterior margin of gena defined by carina, gena broadly reticulate to carina. Compound eye maximum diameter 0.31 mm. Malar space very short, 0.12 mm in length. Length of scape, pedicel and first three flagellomeres: 0.29, 0.050, 0.092, 0.062 and 0.069 mm, respectively. Width of first flagellomeres 0.054 mm. Mesosoma. Orange. Clothed with very sparse, black setae. Dorsum of mesosoma narrowly reticulate, posterior face of propodeum more shallowly so than elsewhere. Humeral angle slightly angulate. Posterior 0.75 of mesopleuron reticulate. Metapleuron and propodeal side glabrous. Coxa, trochanter, and femur orange. Tibia and tarsus orange brown. Legs sparsely clothed with pale white glittering setae. Tarsus with black brown spines. Tibial spurs pale. Metasoma. Orange. T 1 clothed with pale white setae and with small, sparse punctures. T 2 – T 5 clothed with sparse black setae. T 2 punctured and with marginal fringe of dense pale white setae. T 3 – T 5 indistinctly punctate. T 5 with apical fringe of dense black setae. Pygidium black brown, glabrous, pygidial area undefined. S 1 with median longitudinal carina. S 3 – S 5 indistinctly, finely punctate at posterior margins. S 2 – S 4 clothed with pale white setae. S 5 with black brown setae. Length. Approximately 3.0 mm.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC0907E0E4F153C1A52FD22.taxon	materials_examined	Type material. Holotype ɗ, USA, Arizona, Santa Cruz Co., Patagonia, 31 ° 53 ’ N, 110 ° 77 ’ W, 3. IX. 1995, B. Brown (CNCI). Paratypes, 10 ɗ, Arizona, Santa Cruz Co., Patagonia, 31 ° 53 ’ N, 110 ° 77 ’ W, 1 ɗ 16. VI. 1994, 3 ɗ 8. VII. 1995, 1 3. IX. 1995, 1 10. IX. 1995, 1 ɗ 24. IX. 1995, 3 ɗ 14. X. 1995, B. Brown (JPPC, DGMC, UGAC and CNCI). Allotype Ψ, USA, Arizona, Santa Cruz Co., Patagonia, Sonorita Creek Reserve, 31 ° 53 ’ N, 110 ° 77 ’ W, 14. VIII. 1994, B. Brown (CNCI). Other material examined. USA: Arizona: Cochise Co.: Chiricahua Mnts., 5400 ’, Hidden Terrace, 4.5 mi SW portal, 1 Ψ 16 – 20. VIII. 1982, M. A. Cazier (AMNH); Portal, 2 ɗ 8. IX. 1974, H. & M. Townes (AEIC); Portal, 11 ɗ reared by F. D. Parker (BBSL); Wilcox, 1 ɗ reared by F. D. Parker (BBSL); 8 m N Tombstone, 1 ɗ reared by F. D. Parker (BBSL); Coconino Co., 3 m S Sedona, 10 Ψ reared by F. D. Parker (BBSL); 5 m S Sedona, 4 Ψ 6 ɗ reared by F. D. Parker (BBSL); 6 m S Jacob’s Lake, 1 ɗ reared by F. D. Parker (BBSL); Gila Co.: 20 m N Claypool, 2 Ψ 2 ɗ reared by F. D. Parker (BBSL); 1 m S Jakes Corner, 10 Ψ reared by F. D. Parker (BBSL); 5 mi S Payson, 1 Ψ 4 ɗ reared by F. D. Parker (BBSL); 2 m S Pine, 1 Ψ reared by F. D. Parker (BBSL); 8 m S Punkin Cnt., 1 Ψ reared by F. D. Parker (BBSL); Mohave Co., 3 m S Oatman, 1 Ψ reared by F. D. Parker (BBSL); Pima Co.: 21 – 35 mi W Tucson, 1 ɗ 21. IV. 1969, J. Schuh (AMNH); 1 ɗ reared by F. D. Parker (BBSL); Santa Cruz Co., Madera Cyn., 2 Ψ reared by F. D. Parker (BBSL); Yavapai Co.: Clarkdale, 4 ɗ reared by F. D. Parker (BBSL), 6 m E Camp Verde, 2 ɗ reared by F. D. Parker (BBSL). California: Santa Barbara Co., Sedgewick Ranch Reserve, 45 km NW Santa Barbara, 1 Ψ 24. VI – 8. VII. 1997, (UCDC); Fresno Co., Lookout Mnt., 6 m NW Coalinga, 2 ɗ reared by F. D. Parker (BBSL); Santa Barbara Co., Sedgewick Reserve, 45 km NW Santa Barbara, 1 ɗ 19. IX. 1997, E. Schlinger (UCDC); San Diego Co., Lake Wohlford, 1 ɗ 30. IV. 1974, H. & M. Townes (AEIC); Santa Cruz Co., Felton, 1 ɗ reared by F. D. Parker (BBSL); Tulare Co., 9 km SE Wilsonia, 2380 m, 1 Ψ 1. VIII. 1993, P. Ward (UCDC). New Mexico: Hidalgo Co.: Rodeo, 2 ɗ reared by F. D. Parker (BBSL), 5 m E Granite Pass, 1 ɗ reared by F. D. Parker (BBSL); 10 m NE Lordsburg, 1 ɗ reared by F. D. Parker (BBSL); Valencia Co., 5 m N Fence Lake, 1 ɗ reared by F. D. Parker (BBSL). Nevada: Clark Co., Kyle Cyn, 3 ɗ reared by F. D. Parker (BBSL); Clark Co., Searchlight, 1 Ψ 3 ɗ reared by F. D. Parker (BBSL); Lincoln Co., Tule Desert 1 ɗ 10. IV – 9. V. 1984, 1 ɗ 17. VIII – 30. IX. 1983, R. Bechtel & J. Knight (BBSL); Washoe Co., Verdi, 1? reared by F. D. Parker (BBSL). Texas: Va l Ve rd e C o., Devil’s River, Dolan Falls, 2 Ψ 16 – 19. VI. 1995; 2 Ψ 24 – 26. VI. 1995, A. Hook and O. Hernandez (JPPC and DGMC). Utah: Emery Co., San Rafael Desert, 3 m SSE Temple Mt., 5300 ’, 1 Ψ F. D. Parker (BBSL); Washington Co.: Beaver Dam Wash, Lytle Ranch Preserve, 1 ɗ 10 – 24. VI. 1987, W. J. Hanson (BBSL); Duncan Flats, 1 ɗ 21 – 31. VII. 1981, C. R. Nelson (BBSL); Leeds Canyon, 2 ɗ 15 – 18. VII. 1980 (BBSL); Lytle Ranch, malaise trap, 1 ɗ 4 – 7. VIII. 1992, D. Judd & D. Feener (AEIC); Paradise Canyon, 1 ɗ 30. V – 4. VI. 1983, 1 ɗ 21 – 30. VIII. 1983, D. Beck (BBSL); Pintura, 1 ɗ reared by F. D. Parker (BBSL), South Pintura, 1 ɗ 10. VII – 14. VIII. 1986, W. J. Hanson (BBSL). MEXICO: Baja California, Rio San. Salv., Hwy 3, 1077 m, 1 ɗ 7 – 12. VI. 1997, E. Schlinger (UCDC). Baja California Sur: Puerto Escondito, 2 ɗ 10 – 14. VII. 1989, R. Shaver (UCDC); Puerto Escondito, Lorinto, 3 ɗ 23. VI. 1991, R. Shaver (UCDC).	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC0907E0E4F153C1A52FD22.taxon	description	Variation. The male specimens vary in length from 4.6 mm to 6.2 mm and the coloration of the femora and tegulae vary from stramineous to brown orange. The female specimens vary in length from 3.0 mm to 4.2 mm, but lack any noticeable difference in coloration or sculpture. Hosts. Both sexes were reared from Solierella blaisdelli, S. plenoculoides similis, and Pisonopsis birkmani Rohwer. Males were also reared from a Trypoxylon sp.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC0907E0E4F153C1A52FD22.taxon	etymology	Etymology. From the Greek kirrhos “ orange ” and meros “ thigh, ” in reference to the orange femora.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFC0907E0E4F153C1A52FD22.taxon	discussion	Remarks. Lomachaeta cirrhomeris occurs sympatrically with L. hicksi. Given the range of color variation of the latter, future studies may prove that L. cirrhomeris is just another color morph of L. hicksi. However, current rearing data from these areas suggests that these are actually two separate species.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCF907D0E4F126E1A61FA8A.taxon	description	(Fig. 20, 24)	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCF907D0E4F126E1A61FA8A.taxon	diagnosis	Diagnosis. The totally orange metasoma, the distinctly brachyplumose setae, and lack of a distinct ventral mandibular tooth separate this species from all other Lomachaeta. It is similar to L. ptilohyalus in lack of the ventral mandibular tooth, distinct brachyplumose setae, and long setae present on the ventral margin of the parameres (Fig. 20). The apical metasomal segments are orange, however, and the dorsum has more developed punctures than L. ptilohyalus. The pronotum has punctures similar to some forms of L. hicksi (Fig. 8).	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCF907D0E4F126E1A61FA8A.taxon	materials_examined	Holotype Male. Head. Width 0.86 mm, entirely black, clothed throughout with sparse, long, erect, pale white brachyplumose setae. Compound eye maximum diameter 0.40 mm. Median ocellus 0.069 mm, lateral ocellus 0.046 mm, ocellocular distance 0.25 mm, and interocellular distance 0.085 mm. Malar space very short, 0.065 mm in length. Mandible black except tip brown orange, slightly emarginated beneath, without distinct tooth beneath. Antennal tubercle dark brown. Antenna black. Scape clothed with both pale white and black brown setae throughout. Length of scape, pedicel and first three flagellomeres: 0.28, 0.054, 0.085, 0.092 and 0.10 mm, respectively. Width of first flagellomeres 0.85 mm. Front with punctures large and crowded. Vertex with small, distinct, separated punctures. Mesosoma. Black, clothed throughout with sparse, long, erect, pale white and brown brachyplumose setae. Pronotum with moderately large, irregularly spaced shallow punctures on lateral margin, confluent at posterior margin, small and sparsely spaced medially. Mesonotum with small, close punctures throughout. Scutellum almost flat, only very slightly convex, with punctures similar to mesonotum, more crowded. Dorsum and posterior face of propodeum broadly reticulate throughout. Tegula dark brown, glabrous, except entire margin punctate and with long, pale, erect setae. Legs brown. Femur clothed with sparse, long, erect, pale white brachyplumose setae. Tibia and tarsus with golden brown setae. Tibial spurs pale. Wings with pterostigma 0.12 mm in length along costa. Marginal cell 0.24 mm in length. Second submarginal cell pentagonal, 0.19 mm in length.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCF907D0E4F126E1A61FA8A.taxon	description	Metasoma. Orange except S 1 and metasomal segment 7 brown. Clothed throughout with sparse, long, erect, pale white setae except T 7 completely clothed with erect black brown setae. T 2 – T 6 also with single row of erect dark setae just anterior to apical margin. Bristles at posterior margins of T 2 – T 4, pale white. T 1 punctate on lateral margin and medially, posterior margin with row of very large, contiguous punctures. T 2 with small, widely separated punctures. T 3 – T 7 with progressively small to fine, separated punctures. S 2 with large, close punctures throughout. Genitalia Fig. 20. Length. Approximately 4.8 mm. Female. Unknown.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCF907D0E4F126E1A61FA8A.taxon	materials_examined	Type material. Holotype ɗ, USA, Texas, Val Verde Co., Devil’s River, Dolan Falls, 24 – 26. VI. 1995, A. Hook and O. Hernandez (UMIC). Paratypes, 2 ɗ, same data as holotype (BFLC). Other material examined. USA: Arizona: Coconino Co.: 20 m S Jacob’s Lake, 1 ɗ reared by F. D. Parker (BBSL); 7 m S Sedona, 3 ɗ reared by F. D. Parker (BBSL); Yavapai Co., Clarkdale, 1 ɗ reared by F. D. Parker (BBSL); 11 m W Jerome, 2 ɗ reared by F. D. Parker (BBSL). Nevada, Clark Co., Mormon Well N, 2 ɗ 16. VII. 1998 (BBSL); Mormon Well S, 2 ɗ 16. VII. 1998 (BBSL); White Pine Co., Charcoal Ovens State Park, 1 ɗ 30. VI. 1960, T. R. Haig (UCDC). Texas, Culberson Co., Choza Springs., Guatalupe Mts., 1610 m, 1 ɗ 8. IV. 1996 (BBSL).	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCF907D0E4F126E1A61FA8A.taxon	description	Variation. The specimens vary from 3.0 mm to 4.8 mm. The specimens from Arizona are the smallest. In some of the smaller specimens, coloration of the first metasomal segment differs slightly from that of the larger specimens by the first metasomal segment being darker in coloration and, for the smallest specimen (Clarkdale, AZ), the metasoma is more brown than orange. Host. Solierella plenoculoides similis.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCF907D0E4F126E1A61FA8A.taxon	etymology	Etymology. From the Greek krokos “ orange ” and pinna “ feather ”, in reference to the orange metasoma and the brachyplumose setae.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCC907C0E4F14FC18BEF9C2.taxon	description	(Figs. 13, 21)	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCC907C0E4F14FC18BEF9C2.taxon	diagnosis	Diagnosis. Lomachaeta hyphantria has several highly autapomorphic characters. It differs from all congeners by having a blunt tubercle present at the occipital and genal carinae junction, and by having elongate punctures present posteriorly on the mesonotum. This species also lacks a ventral basal tooth on the mandibles, and has yellow orange femora and tegulae.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCC907C0E4F14FC18BEF9C2.taxon	materials_examined	Holotype Male. Head. Width 0.71 mm, brown except red brown around compound eyes. Clothed throughout with sparse, long, erect, pale white setae. Compound eye maximum diameter 0.28 mm. Median ocellus 0.062 mm, lateral ocellus 0.050 mm, ocellocular distance 0.17 mm, and interocellular distance 0.069 mm. Malar space very short, 0.085 mm in length. Mandible yellow except tip red brown, slightly emarginate beneath. Scape red brown, clothed with pale white setae throughout. Pedicel red brown. Flagellomeres brown. Length of scape, pedicel and first three flagellomeres: 0.22, 0.054, 0.046, 0.077 and 0.065 mm, respectively; width of first flagellomeres 0.065 mm. Front reticulate, with pale brown setae. Vertex with small, distinct, separated punctures and pale white setae. Clypeus with dense, white setae. Blunt tubercle present at junction of occipital and genal carinae. Mesosoma. Brown, clothed throughout with sparse, long, erect, pale white setae. Pronotum with moderate, closely spaced punctures, confluent on posterior margin. Mesonotum with large punctures anteriorly, posteriorly with 2 reticulations with length equal to 3 punctures in same area. Scutellum almost flat, slightly convex, with small, shallow punctures similar to mesonotum but more crowded. Scutellum with anterior, median margin concave. Dorsum and posterior face of propodeum reticulate throughout. Tegula yellow brown, glabrous, margin with long, pale, erect setae. Legs orange brown, clothed with sparse, long, erect, and dense, short, decumbent white setae; spurs pale. Wings with pterostigma 0.14 mm in length along costa. Marginal cell 0.40 mm in length. Second submarginal cell pentagonal, 0.24 mm in length.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCC907C0E4F14FC18BEF9C2.taxon	description	Metasoma. Metasomal segment 1 and 5 – 7 red brown, apical segments lighter. Metasomal segments 2 – 4 dark brown. Metasoma clothed throughout with sparse, long, erect, pale white setae, except T 5 – T 7 also with setae brown. Bristles at posterior margins of T 2 – T 4, pale brown. T 1 punctate on lateral margin, glabrous medially, posterior margin with row of small, contiguous punctures. T 2 with small, distant, widely separated punctures. T 3 – T 7 with progressively small to fine, separated punctures. S 2 with large, close punctures throughout. Genitalia Fig. 21. Length. Approximately 3.3 mm. Female. Unknown.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCC907C0E4F14FC18BEF9C2.taxon	materials_examined	Type material. Holotype ɗ, BOLIVIA, Dep. Beni, Rio Itenez, 4 km above Costa Marque, Brazil, 12 – 18. IX. 1964, J. K. Bowseman and J. Lussenhop (AMNH). Paratype, 1 ɗ, same data as holotype. Hosts. Unknown.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCC907C0E4F14FC18BEF9C2.taxon	etymology	Etymology. From the Latin hyphantria “ weave ” in reference to the large reticulations on the mesonotum.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCD907B0E4F17161D28F962.taxon	description	(Figs. 10, 19, 26)	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCD907B0E4F17161D28F962.taxon	diagnosis	Diagnosis. This species can be distinguished from other Lomachaeta by lack of a distinct ventral mandibular tooth, distinctly brachyplumose setae, and by sparse, very small punctures on the pronotum and mesonotum (Fig. 10). It is most similar to L. crocopinna, especially regarding the presence of elongate setae on the parameres, but differs from this species by coloration of the metasoma.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCD907B0E4F17161D28F962.taxon	materials_examined	Holotype Male. Head. Width 0.92 mm, black, clothed throughout with sparse, long, erect, pale white setae. Compound eye maximum diameter 0.37 mm. Median ocellus 0.069 mm, lateral ocellus 0.062 mm, ocellocular distance 0.27 mm, and interocellular distance 0.10 mm. Malar space very short, 0.12 mm in length. Mandible black except tip brown orange, slightly emarginate beneath, without distinct tooth beneath. Antennal tubercle brown. Antenna black. Scape clothed with pale white and black brachyplumose setae throughout. Pedicel and flagellomeres with decumbent white setae. Length of scape, pedicel and first three flagellomeres: 0.30, 0.092, 0.077, 0.12 and 0.10 mm, respectively; width of first flagellomeres 0.088 mm; front with punctures small and dense; vertex with small, distinct, separated punctures and brown setae. Mesosoma. Black brown. Clothed throughout with sparse, long, erect, pale white and black brown setae. Pronotum with small, distantly spaced punctures, not confluent on posterior margin. Mesonotum with small, distant punctures throughout. Scutellum almost flat, only very slightly convex, with small, shallow punctures similar to mesonotum but more crowded. Dorsum and posterior face of propodeum reticulate throughout. Tegula black with brown margin, glabrous, except entire margin punctate and with long, pale, erect setae. Legs brown, clothed with sparse, long, erect, brachyplumose pale white setae. Tibia and tarsus with golden bristles. Tibial spurs pale. Wings with pterostigma 0.17 mm in length along costa. Marginal cell 0.23 mm in length. Second submarginal cell pentagonal, 0.22 mm in length.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCD907B0E4F17161D28F962.taxon	description	Metasoma. T 1 brown, orange laterally. Metasomal segment 2 and S 3 orange. T 3 orange, black brown medially. S 1 and metasomal segments 4 – 6 brown. Metasoma clothed throughout with sparse, long, erect, pale white setae. T 5 – T 7 also with brown setae anterior to apical margin, denser on apical segments. Bristles at posterior margins of T 2 – T 4, pale orange. T 1 punctate on lateral margins, glabrous medially, posterior margin with row of small, contiguous punctures. T 2 with small, distant, widely separated punctures. T 3 – T 7 with progressively small to fine, separated punctures. S 2 with large, close punctures throughout. Genitalia Fig. 19. Length. Approximately 5.2 mm.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCD907B0E4F17161D28F962.taxon	materials_examined	Type material. Holotype, ɗ, MEXICO, Oaxaca, 10 m North of Huajuapan de Leon, 7. III. 1985, L. Stange & R. Miller (CNCI). Paratypes, 4 ɗ, USA, California, San Bernardino Co., 8 – 9 m S Kelso, reared by F. D. Parker (BBSL).	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCD907B0E4F17161D28F962.taxon	description	Female. Unknown. Host. Solierella plenoculoides similis.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCD907B0E4F17161D28F962.taxon	etymology	Etymology. From the Greek ptilon “ feather ” and hyalos “ glass, ” in reference to the brachyplumose setae and sparse punctures on the dorsum of the mesosoma.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCB90670E4F10431B87FB02.taxon	description	(Figs. 5, 8 – 11, 16 – 18, 22, 25, 27)	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCB90670E4F10431B87FB02.taxon	diagnosis	Diagnosis. Males are defined by presence of a ventral mandibular tooth, indistinct brachyplumose pubescence (serrate), and absence of long setae on the ventral margin of the parameres (Figs. 16 – 18, 22). Females of this species do not have the sculpture of mesosomal dorsum concealed by stout decumbent orange white setae. However, the dorsum of the mesosoma is concolorous with head, T 2 lacks stout decumbent orange white setae, the thoracic dorsum varies from broadly reticulate throughout to the anterior third to half being distinctly more narrow than posteriorly, T 5 having sparse, black setae and the pygidial area orange in coloration.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCB90670E4F10431B87FB02.taxon	materials_examined	Material examined. USA: Arizona: Cochise Co.: Bisbee, 4 Ψ 2 ɗ reared by F. D. Parker (BBSL); Huachuca Mts., Bear Canyon, 1 ɗ 5. VII. 1966, Hanson (BBSL); Portal, 6 ɗ 10 – 13. VIII. 1974, 2 ɗ 19. VIII. 1987, 1 ɗ 31. VIII. 1987, 1 ɗ 12. IX. 1987, 1 ɗ 2. X. 1987, H. & M. Townes (AEIC); 5 mi West of Portal, 1 ɗ 7. VI. 1965 (UAIC); 5 mi W of Portal, South Western Research Station, 1 ɗ 10. VI. 1963, 2 ɗ 13. VII. 1963, 1 ɗ 7. X. 1962 (AMNH); 8.5 m N Portal, 1 Ψ 5 ɗ reared by F. D. Parker (BBSL); Cave Creek, Sunny Flat, 1 ɗ 13. IX. 1980 (BBSL); Chiricahua Mts., 2 Ψ 12 ɗ reared by F. D. Parker (BBSL); Coconino Co.: 6 m N Cameron, 2 ɗ reared by F. D. Parker (BBSL); Felton, 6 Ψ reared by F. D. Parker (BBSL); 4 m S Jacob’s Lake, 1 Ψ 6 ɗ reared by F. D. Parker (BBSL); 11 m S Jacob’s Lake, 9 Ψ 11 ɗ reared by F. D. Parker (BBSL); 20 m S Jacob’s Lake, 1 ɗ reared by F. D. Parker (BBSL); Gila Co.: Payson, 1 Ψ reared by F. D. Parker (BBSL); 2 m S Pine, 4 ɗ reared by F. D. Parker (BBSL); 19 m E Punken, 3 Ψ reared by F. D. Parker (BBSL); Pima Co., Madera Canyon, Santa Rita Mountains, 1 ɗ 18 – 23. VI. 1962, F. Werner (UAIC); Santa Cruz Co., Patagonia, 31 ° 53 ’ N, 110 ° 77 ’ W, 1 ɗ 14. VIII. 1994, 4 ɗ 3. IX. 1995, 1 ɗ 24. IX. 1995, 1 ɗ 14. X. 1995, B. Brown (JPPC, DGMC, and CNCI); Yavapai Co.: 15 m E Camp Verde, 1 Ψ 18 ɗ reared by F. D. Parker (BBSL); Clarkdale, 1 Ψ reared by F. D. Parker (BBSL); 11 m W Jerome, 4 Ψ reared by F. D. Parker (BBSL); Prescott, 14 Ψ 18 ɗ reared by F. D. Parker (BBSL). Arkansas: Pulaski Co., Pinnacle Mt. State Park, 3 ɗ 15 – 25. VI. 2002, J. P. Pitts (JPPC); Pulaski Co., Camp Robinson, 8 ɗ VII. 2002, B. Baldwin (JPPC). California: Alpine Co., Silver Creek, 1 ɗ 13. VII. 1948, R. M. Bohart (UCDC); Fresno Co., White Creek, 12 m W Coalinga, 1 Ψ 3 ɗ reared by F. D. Parker (BBSL); Inyo Co: 6 m NE Pine, 11 Ψ 10 ɗ reared by F. D. Parker (BBSL); 8 m NE Pine, 9 Ψ 6 ɗ reared by F. D. Parker (BBSL); 11 m NE Big Pine, 1 Ψ 10 ɗ reared by F. D. Parker (BBSL); Napa Co., Pope Valley, 1 / 3 km S Aetna Springs, 6 ɗ 3 – 14. IX. 1993, L. S. Kimsey (UCDC); Los Angeles Co. Tanbark Flat, 1 ɗ 17. VI. 1956, R. M. Bohart (UCDC); Riverside Co.: 3 m W Banning, 4 ɗ reared by F. D. Parker (BBSL); 9 m W Banning, 4 Ψ 3 ɗ reared by F. D. Parker (BBSL); 1 m W Jacumba, 2 Ψ 1 ɗ reared by F. D. Parker (BBSL); Fullmer Lake, 3 Ψ 1 ɗ reared by F. D. Parker (BBSL); Sacramento Co.: 1 Ψ 2 ɗ IV. 1947, unknown (UAIC); Nevada Co., Sagehen Creek 1920 m: 3 ɗ 2 – 5. VII. 1981, L. French & P. S. Ward (UCDC), 2 ɗ 21. V. 1997, E. Schlinger (UCDC), 1 ɗ 28. V. 1997, E. & M. Schlinger (UCDC); Sacramento, 11 Ψ 6 ɗ IV. 1947, reared by F. D. Parker (BBSL); San Bernardino Co.: Granite Mts., 4050 ’, 1 Ψ 22 – 25. V. 1983, T. L. Griswold (BBSL); Morongo Valley, 3 Ψ 3 ɗ reared by F. D. Parker (BBSL); Pinnacles Natl. Mon., 1 mi W Chaparral Campgrounds, 1 ɗ 31. V. 1998, O. Messinger (BBSL); Santa Cruz Co., Felton, 10 ɗ reared by F. D. Parker (BBSL); Shasta Co., 2 mi W. Shingletown, 2750 ’, malaise trap, 1 ɗ 1 – 9. VIII. 1986, R. Miller (JPPC); Solano Co., Cold Canyon Reserve, 11 km W Winters, 2 ɗ 18. IX – 2. X. 1991, 1 ɗ 13 – 24. IX. 1990, 1 ɗ 4 – 10. VII. 1990, D. Carmean (UCDC). Florida: Alachua Co., Gainesville, Picree Homestead, 1 ɗ 10. IX. 1976, Pierce (UCDC); Alachua Co., Paines Prairie, 1 ɗ 21. III. 1977, G. B. Fairchild (UCDC); Highlands Co., Lake Placid, Archbold Biological Station, 6 ɗ, 2. VI. 1986, 9. VI. 1986, 25. VII. 1986, 8. X. 1986, 28. IX. 1987, 1. XI. 1996, M. Deyrup (ABSC); Orange Co., Rock Springs Run, 1 ɗ 31. V. l 995, 1 ɗ 7. VI. 1995, 1 ɗ 14. VI. 1995, S. W. Fullerton (CNCI, UCFC); Orange Co., Wekiwa Springs State Park, 1 ɗ 2. IV. 2001, 1 ɗ 13. IV. 2001, 1 ɗ 28. IV. 2001, 1 ɗ 26. V. 2001, 3 ɗ 1 Ψ 9. V 1.2001, 1 Ψ 21. VII. 2001, 3 ɗ 23. V 1.2001, 1 ɗ 21. VII. 2001, burn zone 43 / 31, malaise trap, P. J. Russell and S. W. Fullerton (UCFC). Georgia: Clarke Co., Whitehall Forest, Malaise Trap, 1 ɗ 1. X. 1997, JPP (JPPC), 1 ɗ 5 – 9. V. 1988, 1 ɗ 12 – 16. V. 1988, 2 ɗ 23 – 26. V. 1988, 1 ɗ 13 – 16. VI. 1988, C. L. Smith (UGAC), 2 ɗ 20. V. 1967, 27. V. 1970, R. T. Franklin (UGAC); Jones Co., 1 ɗ 26. IV – 8. V. 1993, 1 ɗ 11 – 18. V. 1993, 1 ɗ 24 – 31. V. 1993, 1 ɗ 8 – 15. VI. 1993, 1 ɗ 22 – 29. VI. 1993, 1 ɗ 20 – 27. VII. 1993 (UGAC). Maryland: Prince George’s Co., Lauret, Patuxent Research Station, 2 ɗ 1 – 8. VI. 1986, 3 ɗ 8 – 15. VI. 1986, 1 ɗ 5 – 13. VII. 1986, D. B. Wahl (CNCI). Massachusetts: Norfolk Co., malaise trap, 1 ɗ 7. VII. 1966, B. and J. Matthews (JPPC). Missouri: Wayne Co., Williamsville, 1 ɗ 26. V – 16. VI. 1969, J. T. Becker (CNCI). North Carolina: Sampson Co., Clinton, 1 ɗ 24. IV. 1951, H. & M. Townes (AEIC); Wake Co., 1 ɗ 4. VII. 1984, S. Nolfo (UCDC). New Mexico: Catron Co., Datril, 1 Ψ reared by F. D. Parker (BBSL); 5 m W Datril, 1 ɗ reared by F. D. Parker (BBSL); Hidalgo Co.: 5 m E Granite, 2 Ψ 4 ɗ reared by F. D. Parker (BBSL); 10 m NE Lordsburg, 3 ɗ reared by F. D. Parker (BBSL); 15 m N Lordsburg, 2 ɗ reared by F. D. Parker (BBSL); Sierra Co., Kingston, 1 ɗ reared by F. D. Parker (BBSL); Socorro Co.: Bingham, 2 Ψ 6 ɗ reared by F. D. Parker (BBSL); 3 m NW Socorro, 1 Ψ 3 ɗ reared by F. D. Parker (BBSL). Nevada: Clark Co., 16 m W Searchlight, 1 ɗ reared by F. D. Parker (BBSL); Douglas Co., Topaz Lake, 1 ɗ 17. VIII. 1960, D. Q. Cavagnaro (UCDC); Washoe Co.: Patrick, 2 ɗ reared by F. D. Parker (BBSL); Verdi: 1 ɗ 4 Ψ no date, 1 Ψ 25. XII. 1960, 1 Ψ 1 ɗ from Queen Ann’s Lace, 25. XII. 1960, 6 ɗ from Queen Ann’s Lace, 16. XII. 1960, 1 ɗ from Wild Buckwheat, 25. XII. 1960, 3 Ψ from Elderberry, 28. I. 1961, reared by F. D. Parker (UCDC). Oklahoma, Latimer Co., 1 Ψ VII. 1988, Karl Stephan (UGAC). Oregon: Benton Co., Corvallis, 1 ɗ 29. VII. 1978, H. &. M. Townes (AEIC); Grant Co., Aldrich Mts., 1 ɗ 16. VI. 1987, H. &. M. Townes (AEIC); Hood River Co., Parkdale, 1 ɗ 20. VII. 1978, H. &. M. Townes (AEIC). Texas: Erath Co., 5 mi. N. Stephensville, 1 ɗ 16 – 17. V. 1980, P. T. Riherd; Gillespie Co., Fredericksburg, 2 ɗ 5. V. 1987, 1 ɗ 7. V. 1987, H. &. M. Townes (AEIC); Kerr Co., Kerrville, 1 ɗ 1. V. 1987, H. &. M. Townes (AEIC); Val Verde Co., Devil’s River, Dolan Falls, 1 ɗ 24 – 26. VI. 1995, A. Hook & O. Hernandez (DGMC). South Carolina, Aiken Co.: Savanah River Project, Funnel Trap, 1 Ψ 19. V. 1994, C. New (JPPC); Savanah River Project, water trap, 1 ɗ 22. IV. 1994, C. New (UGAC); Greenville Co., Greenville, 1 ɗ 9. III. 1955, L. & G. Townes (AEIC). Utah: Cache Co.: Black Smith Fork Canyon, 13 Ψ 17 ɗ reared by J. P. Pitts (BBSL); Crowley Canyon, 1 ɗ 27. VIII – 3. VIII. 989, Hanson (BBSL); Four Mile Canyon, 1 ɗ 3 – 11. VI. 1992, 1 ɗ 21 – 31. VII. 1992, Hanson & Kelly (BBSL); Logan Canyon, 132 Ψ 171 ɗ reared by F. D. Parker (BBSL); 9 mi W Mendon, 5 Ψ reared by F. D. Parker (BBSL); Tony Grove Junction, 1 ɗ 8 – 16. VIII. 1983 (BBSL); Garfield Co., Cass Creek Reserve, Mt. Hillers, 1 ɗ 26. VII. 1978 D. Vogt (BBSL); Kane Co.: Chamb Ranch, 1 ɗ 17 – 30. VIII. 1982, C. R. Neson (BBSL); 4 mi N Glendale, 1 ɗ 9 – 14. VIII. 1986, Hanson (BBSL); Uintah Co.: SW Bonanza, 5000 ’, 1 ɗ 12. VI. 1981, 1 ɗ 9. VII. 1981, 1 ɗ 11. VIII. 1981, Schwartz & Miller (BBSL); Utah Co., Hobble Creek Canyon, 5 Ψ 30 – 31. VII. 1982, C. R. Nelson (BBSL); Washington Co.: Leeds Canyon, 1 ɗ reared by F. D. Parker (BBSL); Upper Deep Creek, 1 ɗ 20 – 31. VIII. 1981, C. R. Nelson (BBSL). Wyoming: Lincoln Co., Kemmerer, 1 ɗ 13. VII. 1981, Meadows & Coombs (BBSL). MEX­ ICO: Baja California, Rio San. Salv., Hwy 3, 1077 m, 3 ɗ 2 – 7. VI. 1997, M. E. Irwin & E. I. Schlinger (UCDC). Puebla, 3 mi NW Petlailcingo, 1 ɗ 3. IV. 1962, L. Stange (UCDC); Ixtalhucan del Rio Jal, 1 ɗ 17. IX. 1970, R. M. Bohart (UCDC).	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCB90670E4F10431B87FB02.taxon	distribution	Distribution. Ranges from Oregon to Mexico and east to Florida and Massachusetts. Hosts. Solierella blaisdelli, S. plenoculoides similis, and Pisonopsis birkmani.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFCB90670E4F10431B87FB02.taxon	discussion	Remarks. Males of this species are geographically variable in coloration. Coloration of the male metasoma varies from being completely orange for individuals from Central and Southern California to completely black for individuals from the Northern extent of the range (Fig. 25), although a small number of specimens from Maryland have orange integument around the felt line of T 2 (e. g., Fig. 27). Furthermore, the color can vary from completely dark (brown to black) with some orange localized around the felt lines to having T 2 and S 2 completely orange (Fig. 27) in the remaining extent of the Southern range. Also for some individuals, the dark coloration of T 2 is sometimes so reduced that it appears as a medial dark maculation. This maculation can vary from a slight spot in the center of the tergite to a longitudinal band stretching the length of the tergite. The color variation is so apparent that it caused one author to create six synonyms. Color characters have been found to be inadequate for defining species. Males of L. coloradensis, L. punctinota, L. hicksi, and L. variegata and females of L. variegata and L. minutula have been reared out of the same nest from several areas multiple times. Also, the male genitalia of all species concerned are highly conservative (Figs. 16 – 18, 22). The only conclusion that can be drawn is that these species are synonymous. Furthermore, after studying the types of D. mickeli and L. formosula and many additional specimens of D. mickeli, they are concluded to be synonyms of L. hicksi as well. Punctation also tends to vary geographically. When punctuation is studied over the entire range, however, no species­level characters can be found. As with the color, it seems that one form of punctuation predominates over another in certain regions. For specimens that are darkly colored, the pronotum normally has large dense punctures laterally and has small, sparse punctures medially. This usually occurs in specimens from the Northwest (Fig. 8) with the punctures tending to be denser in individuals from the Northeast (Fig. 10). For these northeastern specimens, the pronotal sculpturing is often similar to that of L. cirrhomeris specimens (Fig. 7). For specimens in the Southern USA, which have orange tergal maculations or completely orange metasomas, the sculpturing tends to differ slightly in density compared to the Northern USA. The pronotal and mesonotal sculpturing vary from that of individuals from California (Fig. 9) to that of individuals in the remaining Southern USA (Fig. 11).	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD690660E4F14741D0CFDEA.taxon	diagnosis	Diagnosis. This species differs from North and Central American species by not having the pronotal spiracles tuberculate. It differs from L. vianai by having the scape as long as the following four flagellomeres combined, the propodeal spiracle prominently tuberculate, and the apical margin of metasomal tergites 2 – 4 with erect spines that are distinctly stouter than surrounding setae.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD690660E4F14741D0CFDEA.taxon	description	Male. Unknown.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD690660E4F14741D0CFDEA.taxon	distribution	Distribution. Known only from the holotype.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD690660E4F14741D0CFDEA.taxon	discussion	Remarks. Lomachaeta ibarrai and L. vianai are known only from the females. They differ from the North and Central American species in that the pronotal spiracles are not tuberculate. The tuberculate state of the thoracic spiracles, along with a combination of other characters, has also been used for delimiting Lomachaeta females from Smicromutilla females. The differences found between L. ibarrai and L. vianai and the remaining Lomachaeta species may be of generic importance. It is possible that after the males of these species are discovered, it may be found that L. ibarrai and L. vianai are not congeneric with the North and Central American Lomachaeta species. The holotype was the only material examined and it resides in the AMNH. It is damaged slightly with the metasoma being broken off and glued to the point.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD790660E4F132E1B74FB27.taxon	diagnosis	Diagnosis. This species differs from North and Central American species by not having the pronotal spiracles tuberculate. Differs from L. ibarrai by having the scape shorter than the following four flagellomeres combined, the propodeal spiracle not prominently tuberculate, apical margin of metasomal tergites 2 – 4 with decumbent spines that are only slightly stouter than surrounding setae.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD790660E4F132E1B74FB27.taxon	distribution	Distribution. Known only from the holotype.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD790660E4F132E1B74FB27.taxon	discussion	Remarks. The holotype was the only material examined. The holotype is located at the AMNH. The holotype is mounted on a card, such that the ventral surface of the specimen could not be examined. A label bearing the words “ s / termites ” (with termites) is located beneath the locality label. It is doubtful that there is any biological significance to this observation as stated by Casal (1969), because all of the known hosts for Mutillidae are holometabolous insects.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD590620E4F143E1D25FE31.taxon	diagnosis	Diagnosis. Its larger size, bidentate mandibles and laterally flattened parameres serve to distinguish this species from S. powelli.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD590620E4F143E1D25FE31.taxon	materials_examined	Holotype Male. Coloration. Head, mesosoma and antenna black. Mandible and antennal tubercle brown orange. Tegula and legs brown. Metasoma orange except sixth segment brown apically and seventh segment completely brown. Head. Head width 1.2 mm. Compound eye maximum diameter 0.49 mm. Median ocellus 0.08 mm, lateral ocellus 0.07 mm, ocellocular distance 0.42 mm, and interocellular distance 0.05 mm. Mandible bidentate with dorsal carina terminating ~ 0.33 from apex. Malar space 0.092 mm. Front, vertex and gena nitid, with sparse, shallow punctures and erect, pale black setae. Front more closely and coarsely punctate than vertex. Clypeus convex, highest along midline, closely punctate and with erect, black setae and shorter white setae. Gena with sparse, shallow punctures, weakly separated from postgena. Pedicel and flagellomere I approximately equal in length, flagellomere I distinctly shorter than flagellomere II. Length of scape, pedicel and first three flagellomeres: 0.37, 0.11, 0.11, 0.18 and 0.11 mm, respectively; width of first flagellomeres 0.15 mm.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD590620E4F143E1D25FE31.taxon	description	Mesosoma. Pronotum nitid, with sparse shallow punctures and erect, pale white setae. Humeral angle rounded. Mesonotum nitid, with sparse punctures and erect, pale black or white setae. Parapsidial furrows obsolete. Scutellum with large punctures more closely spaced than mesonotum and erect, pale black or white setae. Propleuron with large punctures, 3 – 4 X larger than pronotal punctures. Mesopleuron shallowly reticulate and with erect, pale white setae. Metapleuron and sides of propodeum nitid, glabrous. Posterior face of propodeum broadly, shallowly reticulate, some punctures confluent. Tegula disk nitid, impunctate medially, margins with sparse punctures. Wings hyaline with pterostigma 0.16 mm in length along costa. Marginal cell 0.42 mm in length. Second submarginal cell pentagonal, 0.30 mm in length. Legs clothed with pale setae. Metasoma. T 1 nitid, sparsely punctate throughout, with erect, pale white setae. T 2 nitid, with sparse, shallow punctures and erect, white pale setae, posterior margin with a band of small, distinct punctures and thin fringe of stouter pale white setae. Felt line of T 2 approximately 0.5 X length of tergum. T 3 – 5 nitid, with sparse, small punctures and erect, white setae. T 6 nitid with erect black setae. Pygidium sparsely punctate and with erect black setae, area undefined laterally, nitid and glabrous anteriorly. S 1 elevated into a carina midline; S 2 nitid, with sparse, shallow punctures and sparse, erect, pale white setae. S 3 – 6 with sparse punctures and erect, pale setae. Hypopygium with sparse, erect, black setae, deeply punctate, broadly emarginate medially. Genitalia (Fig. 23). Parameres laterally flattened. Length. Approximately 4.8 mm. Female. Unknown.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD590620E4F143E1D25FE31.taxon	materials_examined	Type material. Holotype ɗ, USA, California, San Bernardino Co., Granite Mts., 4050 ’, 9. VI. 1980, T. Griswold (BBSL). Paratypes 2 ɗ, San Bernardino Co., Granite Mts., 4050 ’, 1 ɗ 22 – 25. V. 1983, 1 ɗ 3. V. 1998, T. Griswold (BBSL). Other material. USA, California: Kern Co.: California City, 1 ɗ 6. V. 1982, 1 ɗ 14. V. 1995, P. F. Torchio (BBSL); 12 m E Mojave, 1 ɗ 30. IV. 1986, P. F. & D. M. Torchio (BBSL); San Bernardino Co.: Cotton Hills, 3300 ’, 1 ɗ 29. V. 1983, T. Griswold (BBSL); Cottonwood Wash, 2520 ’, 1 ɗ 25. V. 1983, T. Griswold (BBSL); Mid Hills, 5620 ’, 2 ɗ 6. VI. 1976, 1 ɗ 16. V 1.1980, T. Griswold (BBSL). Nevada, Mineral Co., Jct. 95 & 306, 1 ɗ 17. VI – 21. VII. 1988, Pan Trap, J. L. Carpenter (BBSL). Var ia t io n. Size varies from 3.5 – 5.3 mm.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD590620E4F143E1D25FE31.taxon	etymology	Etymology. From the Old English adjective beadugrim “ war­furious, ” in reference to the blade­like parameres.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD590620E4F143E1D25FE31.taxon	discussion	Remarks. Mickel (1964) stated that the reduced wing venation is an important character in distinguishing Smicromutilla from other genera. However, only the wing venation of S. powelli is reduced in comparison to Lomachaeta. When S. beadugrimi is compared to S. powelli, it has a much more complete wing venation that is not distinct from that of Lomachaeta. Due to the differences between these two species of Smicromutilla, which are as drastic as the differences between the genera Lomachaeta and Smicromutilla, this species may not be congeneric with S. powelli. However, rather than erecting another monotypic genus, we have chosen to place this species into Smicromutilla.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD390620E4F13041BB9FC07.taxon	diagnosis	Diagnosis. Its smaller size, tridentate mandibles, much reduced wing venation, and cylindrical tapering parameres (similar to Lomachaeta species) serve to distinguish this species from S. beadugrimi.	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
03CB87ABFFD390620E4F13041BB9FC07.taxon	materials_examined	Material Examined. No new material was examined other than the Holotype (CASC) and several paratypes (CNCI and BBSL).	en	Pitts, James P., Manley, Donald G. (2004): A Revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa 474: 1-27, DOI: 10.5281/zenodo.157428
