taxonID	type	description	language	source
03CB87F0FF2AFFEBFF2A522AFADD1B2D.taxon	materials_examined	Type species: Interrhodeus brevicornus Karg, 2000, by original designation.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF2AFFEBFF2A522AFADD1B2D.taxon	diagnosis	Diagnosis (adult female). Arthrodial process of chelicera with a short coronet-like fringe; epistome with a serrate antero-medial projection of about uniform width, flanked on each side by a slightly longer and serrate projection; seta h 3 directly posterior to h 1 and distinctly postero-mediad to h 2; podonotal shield with 22 pairs of setae (setae s 1 and r 1 absent) and without scleronoduli; opisthonotal shield with 20 pairs of setae; all dorsal idiosomal setae smooth; laciniae of tritosternum separated from each other only in the distal half; presternal shields absent; ventrianal shield with five pairs of setae in addition to the circum-anal setae; unsclerotised integument around ventrianal shield with two pairs of setae; with a pair of elongate metapodal plates; peritreme extending anteriorly to mid-level of coxa II; pretarsus I present; seta pl 4 of tarsus IV absent. Numbers of setae on legs I – IV: coxa: 2, 2, 2, 1; trochanter: 6, 5, 5, 5; femur: 13, 11, 6, 6; genu: 13, 11, 9, 10; tibia: 14, 10, 8, 10; tarsus II – IV: 18, 18, 17.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF2AFFEEFF2A5050FBDD1A8C.taxon	materials_examined	Specimens examined. Holotype female: 6953 T, IB 1418905, La Selva, Heredia, Costa Rica (10 ° 26 ’ 0 ” N 84 ° 1 ’ 0 ” W), elev. 50 – 150 m, Coll. Alas, 6 - XI- 1993, ZMB Kat. Nr. 46010, [Costa Rica INBIO CRI 001418905]. Paratypes: one female, 6954, IB 1418902, ZMB Kat. Nr. 46011, [Costa Rica INBIO CRI 001418902]; one female, 6955, IB 1418922, ZMB Kat. Nr. 46012, [Costa Rica INBIO CRI 001418922]; one female, 6956, IB 1419397, Ex. Hernandia didymantha Hernandiaceae, 12 - X- 1994, ZMB Kat. Nr. 46013, [Costa Rica INBIO CRI 001419397]; one female, 6957, IB 1422099, Suampo experimental, 20 - I- 1995, ZMB Kat. Nr. 46014, [Costa Rica INBIO CRI 001422099]; one deutonymph, D-N 6958, IB 1419233, SSO 350 m, 3 - IX- 1993, ZMB Kat. Nr. 46015, [Costa Rica INBIO CRI 001419233]; one deutonymph, D-N 6959, IB 1418871, ZMB Kat. Nr. 46016, [Costa Rica INBIO CRI 001418871] (for all paratypes, other data as for holotype). (in MNHB).	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF2AFFEEFF2A5050FBDD1A8C.taxon	diagnosis	Diagnosis of adult. As for the genus. Adult female (Fig. 1 A – F) (holotype and four paratypes). Dorsal and ventral idiosomal, hypostomal, subcapitular and leg setae smooth. Gnathosoma: Fixed cheliceral digit 34 long, with five teeth in addition to apical tooth and a setiform pilus dentilis (Fig. 1 A); movable cheliceral digit 35 long, with four teeth in addition to apical tooth. Arthrodial process of chelicera with a short coronet-like fringe. Epistome with a serrate antero-medial projection of about uniform width and provided with two apical denticles, flanked on each side by a slightly longer, triangular and serrate projection (Fig. 1 B); margin of epistome smooth between antero-medial and antero-lateral projections, and serrate laterad to antero-lateral projections. Deutosternal denticles in eight rows, with seven to 13 denticles each; anterior row anteriorly convex, other rows straight and roughly transverse, second last row narrow (Fig. 1 C). Measurements of setae: h 1 14 – 17, h 2 13 – 15, h 3 10 – 12, sc 11 – 14. Dorsal idiosoma (Fig. 1 D). Podonotal shield ornamented with roundish markings, except for a punctate band along posterior margin; 158 – 163 long and 171 – 185 wide at widest point; with 22 pairs of setae (setae s 1 and r 1 absent); without scleronoduli and with five pairs of distinguishable lyrifissures. Opisthonotal shield imbricate, with roundish markings behind setae Z 4 and a punctate band along anterior margin; 131 – 146 long and 135 – 142 wide at widest point; with 20 pairs of setae and four pairs of distinguishable lyrifissures. Measurements of setae: j 1 17 – 19, j 2 18 – 19, j 3 15 – 16, j 4 13 – 15, j 5 14 – 15, j 6 13 – 14, z 1 18 – 19, z 2 16 – 18, z 3 14 – 15, z 4 14 – 15, z 5 12 – 14, z 6 12 – 13, s 2 15 – 16, s 3 14 – 15, s 4 14 – 15, s 5 13 – 15, s 6 12 – 13, r 2 14 – 15, r 3 18 – 20, r 4 13 – 14, r 5 13 – 14, r 6 14 – 15, J 1 12 – 13, J 2 12 – 13, J 3 12 – 14, J 4 13 – 15, J 5 15 – 16, Z 1 13 – 14, Z 2 13 – 14, Z 3 13 – 15, Z 4 15 – 16, Z 5 19 – 21, S 1 12 – 14, S 2 12 – 13, S 3 14 – 15, S 4 16 – 17, S 5 19 – 20, R 1 12 – 13, R 2 12 – 14, R 3 14 – 15, R 4 16 – 17, R 5 19 – 20. Ventral idiosoma (Fig. 1 E). Base of tritosternum 14 – 21 long and 12 – 13 wide proximally (Fig. 1 F); laciniae 43 – 51 long, separated for about half of their total length, pilose. Sternal shield lightly reticulate and with indistinct anterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; posterior margin straight; approximately 88 – 93 long, including the lightly sclerotised and punctate region, and 73 – 77 wide at widest point; with four pairs of setae and three pairs of lyrifissures. Triangular remnants of exopodal shield distinguishable between coxae II – III and III – IV. Elongate remnant of endopodal shield distinguishable adjacent to coxa IV and slightly anterior to it. Genital shield with light striae parallel to lateral margins, and a punctate band along most of its slightly convex posterior margin; longer than wide; extending posteriorly behind coxa IV; distance between st 5 – st 5 45 – 48. Unsclerotised integument posterolaterad to st 5 with a pair of lyrifissures. Ventrianal shield reticulate, except for a punctate band along anterior margin; 91 – 102 long and 90 – 102 wide at widest point, not fused to dorsal shield; with five pairs of setae (Jv 1 – Jv 3, Zv 1 and Zv 2) in addition to circum-anal setae; post-anal seta as long as para-anal setae; lyrifissures not distinguishable. Unsclerotised integument around ventrianal shield with two pairs of setae (Jv 5 and Zv 3) and a pair of elongate metapodal plates. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield fused with a section of exopodal shield next to coxa IV. Measurements of setae: st 1 18 – 20, st 2 17 – 18, st 3 16 – 18, st 4 14 – 16, st 5 14 – 15, Jv 1 14 – 15, Jv 2 15 – 16, Jv 3 16 – 17, Jv 5 14 – 16, Zv 1 13 – 14, Zv 2 15 – 16, Zv 3 14 – 15, para-anal 15 – 16, post-anal 16 – 18. Legs. Lengths: I: 303 – 312; II: 223 – 234; III: 203 – 220; IV: 281 – 295. Pretarsi I – IV similar in shape and length, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes. Deutonymph (Fig. 2 A – B) (two paratypes). Shape of setae as in adult female. Gnathosoma. Fixed cheliceral digit 28 – 29 long, with five teeth in addition to apical tooth, and a setiform pilus dentilis; movable cheliceral digit 28 – 30 long, with four teeth in addition to apical tooth. Arthrodial process of chelicera with a short coronet-like fringe. Epistome, deutosternum and position of hypostomal setae as in female. Measurements of setae: h 1 16, h 2 12 – 13, h 3 10, sc 14 – 15. Dorsal idiosoma (Fig. 2 A). Podonotal shield imbricate laterad to j setal series and with roundish markings between j series; 132 – 133 long and 143 – 147 wide at widest point; with 21 pairs of setae (setae s 1 and r 1 absent); without scleronoduli and with five pairs of distinguishable lyrifissures. Unsclerotised integument laterad to podonotal shield with a pair of setae (r 4). Opisthonotal shield mostly imbricate, except for a narrow band of roundish markings along anterior margin; 110 – 122 long and 104 wide at widest point; with 15 pairs of setae and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to opisthonotal shield with five pairs of setae (R 1 – R 5). Measurements of setae: j 1 16 – 17, j 2 18, j 3 15, j 4 13, j 5 13 – 14, j 6 11 – 13, z 1 15 – 16, z 2 15 – 16, z 3 12 – 13, z 4 12 – 13, z 5 11 – 12, z 6 11 – 12, s 2 13, s 3 12 – 13, s 4 13, s 5 12 – 13, s 6 12, r 2 13 – 14, r 3 16 – 17, r 4 11, r 5 11 – 12, r 6 11, J 1 11 – 12, J 2 11, J 3 10 – 11, J 4 11, J 5 12, Z 1 11 – 12, Z 2 10 – 11, Z 3 11 – 12, Z 4 13 – 14, Z 5 19 – 21, S 1 11 – 12, S 2 10 – 11, S 3 10 – 11, S 4 13, S 5 15, R 1 9 – 10, R 2 8, R 3 8 – 9, R 4 10, R 5 12 – 14. Ventral idiosoma (Fig. 2 B). Base of tritosternum 15 – 17 long and 11 – 13 wide proximally; laciniae 40 – 41 long, otherwise as in adult female. Sternal shield smooth, narrowing behind coxae II, with slightly rounded posterior margin; approximately 119 – 123 long and 58 – 59 wide at widest point; with four pairs of setae and three pairs of lyrifissures. Setae st 5 inserted on unsclerotised integument at level of posterior end of sternal shield. Ventrianal shield imbricate; 45 – 56 long along midline and 57 – 58 wide at widest point; with two pairs of setae (Jv 2 and Jv 3) in addition to circum-anal setae; post-anal seta about as long as para-anal setae. Unsclerotised integument around ventrianal shield with five pairs of setae (Jv 1, Jv 4 and Zv 1 - Zv 3). Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield indistinct. Measurements of setae: st 1 17 – 18, st 2 16 – 17, st 3 13 – 14, st 4 11 – 13, st 5 9 – 11, Jv 1 11 – 13, Jv 2 12, Jv 3 13, Jv 4 10 – 11, Zv 1 10 – 11, Zv 2 10 – 11, Zv 3 8 – 9, para – anal 13 – 15, post-anal 15 – 16. Legs. Lengths: I: 267 – 271; II: 198 – 201; III: 180 – 181; IV: 236 – 241. Leg chaetotaxy and pretarsi as in adult female.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF2AFFEEFF2A5050FBDD1A8C.taxon	discussion	Remarks. This species was described on the basis of the adult female holotype, 52 adult female paratypes and seven deutonymph paratypes. The following characteristics appear in the original description and illustrations of the species (Karg, 2000 a), but do not agree with our observations of female type specimens: movable cheliceral digit with three teeth in addition to apical tooth; most anterior row of deutosternal denticles roughly transverse; podonotal shield imbricate, with 21 pairs of setae, and with structures suggesting the presence of scleronoduli [Karg (2000 a) mentioned that scleronoduli were not clear]; unsclerotised integument laterad to podonotal shield with two pairs of setae; sternal shield smooth; exopodal shield consisting of a single structure between coxae II – IV; no endopodal shields; genital shield smooth; lyrifissures present only on podonotal (two pairs) and sternal shields (three pairs). No information was provided about the tritosternum. The only measurements provided in the original description referred to the length of the idiosoma (270 – 290), width of the idiosoma (140 – 180), length of seta J 5 (cited as I 5) (18), range of lengths of other dorsal idiosomal setae (15 – 16) and length of all st setae (18). Leg setal counts were given only for tibia IV (10 setae) and genu IV (10 setae). The deutonymph was illustrated in the original description of the species, but not described (Figure 23 in Karg, 2000 a). The following characteristics are present in the original illustrations of the deutonymph, but do not agree with our observations of the paratype specimens: podonotal shield smooth and with 23 pairs of setae; opisthonotal shield reticulate; ventrianal shield smooth; with a longitudinal line extending from the anterior end of the peritreme to the level of coxa I; lyrifissures present only on the sternal shield (two pairs).	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF2FFFEEFF2A51F6FDB71E38.taxon	materials_examined	Type species: Pennarhodeus pennatus Karg, 2000, by original designation.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF2FFFEEFF2A51F6FDB71E38.taxon	diagnosis	Diagnosis (adult female and male). Arthrodial process of chelicera with a short coronet-like fringe (except P. decoris, arthrodial process elongate, with three points); epistome with a smooth antero-medial projection either of about uniform width or slightly wider proximally, flanked on each side by a shorter projection whose lateral margins are smooth or lightly serrate; seta h 3 directly posterior to h 1 and mediad to h 2; setae dorsal, circumanal and Jv 4 usually pilose and other setae smooth; podonotal shield with 22 – 23 pairs of setae and without scleronoduli (except male of P. brevipennatus, with two scleronoduli); unsclerotised integument laterad to podonotal shield with zero or one pair of setae; opisthonotal shield with 15 – 17 pairs of setae; with four or five pairs of R setae on unsclerotised integument laterad to opisthonotal shield in female and two to five pairs of R setae on unsclerotised integument laterad to opisthonotal shield or on ventrianal shield in male; most dorsal idiosomal setae pilose; presternal shields absent; ventrianal shield with five pairs of setae in female and seven to nine pairs of setae in male; unsclerotised integument along margins of ventrianal shield with two pairs of setae in female and without setae in male; with a pair of rounded metapodal plates in female and without metapodal plates in male; peritreme extending anteriorly to level between anterior margin of coxa III and median region of coxa II; pretarsus I, a single, sessile, curved claw; seta pl 4 of tarsus IV absent. Numbers of setae on legs I – IV: coxa: 2, 2, 2, 1; trochanter: 6, 5, 5, 5; femur: 13, 11, 6, 6; genu: 13, 11, 9, 10 (9 in P. turris Karg); tibia: 14, 10, 8, 10 (9 in P. brevipennatus Karg); tarsus II – IV: 18, 18, 17. Adult male with spermatodactyl abruptly bent dorsally or ventrally and with one or two spur-like ventral setae on each of femur, genu and tibia II.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF2FFFECFF2A5562FC9A1D5C.taxon	diagnosis	Diagnosis (adult male). Arthrodial process of chelicera with a short coronet-like fringe; epistome with anteromedial projection about 1.2 times as long as antero-lateral projections, and with margin serrate laterad to antero-lateral projections; corniculi rather straight; dorsal idiosomal setae pilose, except r 3, Z 5 and S 5, feather-like; podonotal and opisthonotal shields imbricate; with two crescent-shaped scleronoduli between setae j 5 and j 6; posterior margin of opisthonotal shield slightly concave between setae Z 5; seta R 5 absent; sternogenital shield lightly reticulate; unsclerotised integument between sternogenital and ventrianal shields with a subtriangular accessory platelet; peritreme extending anteriorly to level of posterior margin of coxa II; with two spur-like ventral setae on each of femur, genu and tibia II. Adult male (Fig. 3 A – G) (holotype). Dorsal idiosomal setae (except r 3, Z 5 and S 5), para-anal and post-anal setae pilose; setae r 3, Z 5, S 5 and Jv 5 feather-like; hypostomal, subcapitular, ventral idiosomal (except Jv 5, paraanal and post-anal) and leg setae smooth. Gnathosoma. Fixed cheliceral digit 36 long; movable digit 37 long; teeth and pilus dentilis not visible because of position of chelicera. Spermatodactyl 77 long, abruptly bent ventrally at an acute angle, distal portion narrower than proximal portion (Fig. 3 A). Arthrodial process of chelicera with a short coronet-like fringe. Epistome with a smooth antero-medial projection slightly wider at the base and provided with three apical denticles, flanked on each side by shorter, smooth projection with four or five apical denticles (Fig. 3 B); margin of epistome with two or three large teeth between antero-medial and antero-lateral projections and serrate laterad to antero-lateral projections. Deutosternal denticles in eight rows, each with seven to 13 denticles; anterior row “ V shaped, other rows straight and transverse (Fig. 3 C). Corniculi rather straight. Measurements of setae: h 1 14, h 2 10, h 3 11, sc 12. Dorsal idiosoma (Fig. 3 D). Podonotal shield imbricate, except for a punctate band along posterior margin; 160 long and 173 wide at widest point; with 22 pairs of setae (seta r 1 absent); with two crescent-shaped scleronoduli between setae j 5 and j 6 and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to podonotal shield with a pair of setae (r 4). Opisthonotal shield imbricate, except for a punctate band along anterior margin; posterior margin slightly concave between setae Z 5; 125 long and 141 wide at widest point; with 18 pairs of setae and three pairs of distinguishable lyrifissures. Seta R 1 on ventrianal shield; seta R 5 absent. Measurements of setae: j 1 9, j 2 8, j 3 8, j 4 10, j 5 10, j 6 11, z 1 8, z 2 not seen (broken), z 3 10, z 4 10, z 5 not seen (broken), z 6 not seen (broken), s 1 11, s 2 9, s 3 9, s 4 11, s 5 10, s 6 10, r 2 11, r 3 15, r 4 11, r 5 not seen (broken), r 6 14, J 1 not seen (broken), J 2 not seen (broken), J 3 not seen (broken), J 4 11, J 5 12, Z 1 12, Z 2 13, Z 3 13, Z 4 13, Z 5 14, S 1 12, S 2 11, S 3 12, S 4 12, S 5 13, R 1 8, R 2 9, R 3 10, R 4 11. Ventral idiosoma (Fig. 3 E). Base of tritosternum 15 long and 12 wide proximally (Fig. 3 F); laciniae 46 long, separated for about 90 % of their total length, pilose. Sternogenital shield lightly reticulate, with indistinct anterior margin and punctate band along straight posterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; approximately 132 long, including the lightly sclerotised and punctate region, and 65 wide at widest point; with five pairs of setae and three pairs of lyrifissures; distance between st 5 - st 5 28; genital opening on anterior margin of shield. Elongate remnant of endopodal shield adjacent to coxa IV. Unsclerotised integument between sternogenital and ventrianal shields with a subtriangular accessory platelet and punctate near anterior margin of ventrianal shield. Ventrianal shield imbricate; 95 long and 170 wide at widest point and extending to dorso-lateral region of idiosoma, but not fused to dorsal shield; with nine pairs of setae (R 1, Jv 1 - Jv 5 and Zv 1 - Zv 3) in addition to circum-anal setae, a distinguishable pore and two distinguishable lyrifissures; post-anal seta about as long as para-anal setae. Peritreme extending anteriorly to level of posterior margin of coxa II. Peritrematal shield extending posteriorly beyond mid-level of coxa IV. Measurements of setae: st 1 13, st 2 11, st 3 13, st 4 11, st 5 11, Jv 1 11, Jv 2 12, Jv 3 12, Jv 4 11, Jv 5 12, Zv 1 11, Zv 2 11, Zv 3 9, para-anal 13, post-anal 13. Legs. Lengths: I: 240; II: 204; III: 177; IV: 230. Genu IV with ten setae and tibia IV with nine setae. With two spur-like ventral setae on each of femur, genu and tibia II (Fig. 3 G). Pretarsus I, a single, sessile, curved claw; pretarsi II – IV, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF2FFFECFF2A5562FC9A1D5C.taxon	discussion	Remarks. This species is known only from the adult male holotype. The following characteristics appear in the original description and illustrations of the species, but do not agree with our observations of the holotype: margin of epistome smooth between antero-medial and antero-lateral projections; podonotal shield with 22 pairs of setae and four crescent-shaped scleronoduli between setae j 5 and j 6; unsclerotised integument laterad to podonotal shield without setae; opisthonotal shield with 17 pairs of setae; unsclerotised integument laterad to opisthonotal shield with a pair of setae. Karg (2000 b) provided no information about the hypostome, tritosternum, ventral idiosoma, lyrifissures and leg setal counts. The only measurements provided in the original description referred to the length of the idiosoma (290), width of idiosoma (180), length of seta Z 5 (13), range of other dorsal idiosomal setae (8 – 10), length of legs I (220), II (190), III (170) and IV (220).	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF2DFFE2FF2A56C1FD5E1CBB.taxon	diagnosis	Diagnosis (adult female). Arthrodial process of chelicera elongate, three-pointed; epistome with three, well separated projections, the antero-medial about 1.2 times as long as the antero-lateral ones; epistome margin irregularly serrate; corniculi rather straight; dorsal idiosomal setae pilose; podonotal and opisthonotal shields imbricate; without scleronoduli; seta R 5 present; genital shield lightly reticulate; unsclerotised integument between genital and ventrianal shields with a transverse elongate accessory platelet; peritreme extending anteriorly to level of anterior margin of coxa III. Adult female (Fig. 4 A – F) (holotype). Dorsal idiosomal setae, Jv 5, para-anal and post-anal setae pilose; hypostomal, subcapitular, ventral idiosomal setae (except Jv 5, para-anal and post-anal) and leg setae smooth. Gnathosoma. Chelicerae narrow and elongate (Fig. 4 A); fixed cheliceral digit 88 long, with three teeth in addition to apical tooth and a setiform pilus dentilis; movable cheliceral digit 90 long, with three teeth in addition to apical tooth. Arthrodial process of chelicerae elongate, three-pointed. Epistome with three well separated projections; antero-medial projection smooth of about uniform width and provided with two apical denticles; antero-lateral projections slightly shorter, each provided with two or three lateral and three apical denticles (Fig. 4 B); epistome margin irregularly serrate. Deutosternal denticles in eight roughly transverse rows, each with 11 – 16 denticles (Fig. 4 C). With two pairs of rows of denticles outside of the deutosternal groove, one between deutosternal denticle rows fifth and sixth and the other in level with basal most row. Corniculi rather straight. Measurements of setae: h 1 14, h 2 10, h 3 11, sc 12. Dorsal idiosoma (Fig. 4 D). Podonotal shield imbricate, except for a punctate band along posterior margin; 217 long and 207 wide at widest point; with 22 pairs of setae (setae r 1 and r 4 absent); without scleronoduli and with six pairs of distinguishable lyrifissures. Opisthonotal shield imbricate, except for a punctate band along anterior margin; 150 long and 208 wide at widest point; with 15 pairs of setae and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to opisthonotal shield with four pairs of setae (R 2 – R 5); seta R 1 absent. Measurements of setae: j 1 18, j 2 25, j 3 24, j 4 26, j 5 25, j 6 24, z 1 21, z 2 27, z 3 26, z 4 25, z 5 25, z 6 26, s 1 17, s 2 21, s 3 25, s 4 31, s 5 31, s 6 28, r 2 29, r 3 31, r 5 27, r 6 30, J 1 26, J 2 27, J 3 27, J 4 32, J 5 16, Z 1 28, Z 2 30, Z 3 32, Z 4 46, Z 5 59, S 1 32, S 2 31, S 3 32, S 4 39, S 5 48, R 2 10, R 3 7, R 4 9, R 5 8. Ventral idiosoma (Fig. 4 E). Base of tritosternum 19 long and 12 wide proximally (Fig. 4 F); laciniae 45, separated for about 90 % of their total length, pilose. Sternal shield smooth, except for a few light longitudinal lines along margin of posterior half, anterior margin indistinct; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; posterior margin slightly concave; approximately 115 long, including the lightly sclerotised punctate region, 73 wide at widest point; with four pairs of setae and three pairs of lyrifissures. Subtriangular remnants of exopodal shield present between coxae II – III and III – IV. Genital shield lightly reticulate; longer than wide; extending posteriorly behind coxa IV; distance between st 5 – st 5 47; posterior margin straight. Unsclerotised integument between genital and ventrianal shields with a transverse elongate accessory platelet, with setae Jv 1 and Zv 1 and a pair of rounded metapodal plates. Ventrianal shield reticulate; 108 long and 158 wide at widest point, not fused to dorsal shield; with five pairs of setae (Jv 2, Jv 3, Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae and three pairs of distinguishable lyrifissures; post-anal seta longer than para-anal setae. Peritreme extending anteriorly to level of anterior margin of coxa III. Peritrematal shield narrow, not extending beyond peritreme. Measurements of setae: st 1 12, st 2 15, st 3 15, st 4 11, st 5 14, Jv 1 12, Jv 2 13, Jv 3 17, Jv 5 44, Zv 1 12, Zv 2 17, Zv 3 23, para-anal 24, post-anal 35. Legs: Lengths: I: 280; II: 220; III: 201; IV: 279. Genu and tibia IV each with ten setae. Pretarsus I, a single, sessile, curved claw; pretarsi II – IV, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF2DFFE2FF2A56C1FD5E1CBB.taxon	discussion	Remarks. This species is known only from the adult female holotype. The following characteristics appear in the original description and illustrations of the species (Karg, 2000 a), but do not agree with our observations of the holotype: fixed cheliceral digit with two teeth in addition to apical tooth; epistome with antero-medial projection about as long as antero-lateral projections; podonotal shield without punctate band along posterior margin and with 20 pairs of setae; opisthonotal shield without punctate band along anterior margin and with 15 pairs of setae; unsclerotised integument laterad to opisthonotal shield with three pairs of setae; genital shield smooth; peritrematal shield fused to podonotal shield; lyrifissures present only on sternal shield (one pair). No information was provided about the tritosternum and leg setal counts. The measurements provided in the original description referred to the length of the movable cheliceral digit (90), length of the idiosoma (340), width of idiosoma (200), length of setae j 1 (cited as i 1) (20), j 5 (cited as i 4) (25), r 3 (cited as r 5) (25), J 1 (cited as I 1) (25), Z 4 (45), Z 5 (53), S 5 (50), range of all st (12 – 15), Jv 5 (cited as V 8) (45), post-anal (cited as Ps) (35), range of other ventrianal setae (13 – 20), length of legs I (280), II (220), III (200) and IV (280).	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF23FFE1FF2A57F9FE001B78.taxon	diagnosis	Diagnosis (adult female and male). Adult female and male with arthrodial process of chelicera with a short coronet-like fringe; epistome with antero-medial projection about 1.2 times as long as antero-lateral projections, and with margin smooth laterad to anterolateral extensions; podonotal shield mostly with roundish markings except for reticulate antero-lateral region, without scleronoduli; dorsal idiosomal setae pilose, except seta z 1, smooth; opisthonotal shield reticulate; seta R 5 present, inserted on unsclerotised integument outside lateral margins of opisthonotal shield; no accessory platelet between genital / sternogenital and ventrianal shields; peritreme extending anteriorly to mid-level of coxa II. Adult female with genital shield smooth. Adult male with sternogenital shield lightly reticulate; with a spur-like ventral seta on each of femur, genu and tibia II. Adult female (Fig. 5 A – E) (holotype). Dorsal idiosomal setae (except z 1), Jv 5, para-anal and post-anal setae pilose; setae z 1, hypostomal, subcapitular, ventral idiosomal (except Jv 5, para-anal and post-anal) and leg setae smooth. Gnathosoma. Fixed cheliceral digit 35 long; movable cheliceral digit 36 long; teeth and pilus dentilis not visible because of position of chelicera. Arthrodial process of chelicera with a short coronet-like fringe. Epistome with a smooth antero-medial projection of about uniform width and provided with five apical denticles, flanked on each side by a shorter, smooth projection with three apical denticles (Fig. 5 A); margin of epistome with one or two denticles between antero-medial and antero-lateral projections, and smooth laterad to antero-lateral projections. Deutosternal denticles in eight transverse rows, each with eight to 12 denticles (Fig. 5 B). With two pairs of rows of denticles outside of the deutosternal groove, slightly anterior to the sixth and to the seventh deutosternal denticle rows, respectively. Measurements of setae: h 1 11, h 2 8, h 3 9, sc 9. Dorsal idiosoma (Fig. 5 C). Podonotal shield mostly with roundish markings, except for reticulate antero-lateral region and a punctate band along posterior margin; 156 long and 161 wide at widest point; with 22 pairs of setae (setae r 1 and r 4 absent); without scleronoduli and with five pairs of distinguishable lyrifissures. Opisthonotal shield reticulate, except for a punctate band along anterior margin; 146 long and 145 wide at widest point; with 15 pairs of setae and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to opisthonotal shield with five pairs of setae (R 1 – R 5). Measurements of setae: j 1 13, j 2 14, j 3 12, j 4 11, j 5 12, j 6 13, z 1 13, z 2 12, z 3 12, z 4 13, z 5 12, z 6 14, s 1 10, s 2 13, s 3 13, s 4 14, s 5 13, s 6 12, r 2 14, r 3 20, r 5 14, r 6 15, J 1 17, J 2 17, J 3 not seen (broken), J 4 17, J 5 9, Z 1 18, Z 2 18, Z 3 18, Z 4 17, Z 5 16, S 1 16, S 2 15, S 3 16, S 4 17, S 5 15, R 1 6, R 2 7, R 3 7, R 4 8, R 5 9. Ventral idiosoma (Fig. 5 D). Base of tritosternum 15 long and 10 wide proximally (Fig. 5 E); laciniae 31 long, separated for about 90 % of their total length, pilose. Sternal shield smooth and with indistinct anterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; posterior margin straight; approximately 87 long, including the lightly sclerotised punctate region, 70 wide at widest point; with four pairs of setae and three pairs of lyrifissures. Genital shield smooth; slightly shorter than wide; extending posteriorly behind coxae IV; distance between st 5 – st 5 37; posterior margin straight. Unsclerotised integument between genital and ventrianal shields with setae Jv 1 and Zv 1 and a pair of rounded metapodal plates. Ventrianal shield reticulate; 88 long and 121 wide at widest point, not fused to dorsal shield; with five pairs of setae (Jv 2, Jv 3, Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae and three pairs of distinguishable lyrifissures; post-anal seta longer than para-anal setae. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield narrow, not extending beyond peritreme. Measurements of setae: st 1 13, st 2 14, st 3 12, st 4 13, st 5 12, Jv 1 11, Jv 2 12, Jv 3 not seen (broken), Jv 5 11, Zv 1 11, Zv 2 12, Zv 3 12, para-anal 15, post-anal 25. Legs. Lengths: I: 242; II: 191; III: 161; IV: 232. Genu and tibia IV each with ten setae. Pretarsus I, a single, sessile, curved claw; pretarsi II-IV, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes. Adult male (Fig. 6 A – C) (paratype). Shape of setae as in adult female. Gnathosoma. Fixed cheliceral digit 33 long; movable cheliceral digit 32 long; teeth and pilus dentilis not visible because of position of chelicera. Spermatodactyl 48 long, abruptly bent ventrally at an acute angle, tapering distally (Fig. 6 A). Arthrodial process of chelicera with a short coronet-like fringe. Epistome, deutosternum and position of hypostomal setae as in female. Measurements of setae: h 1 15, h 2 8, h 3 11, sc 9. Dorsal idiosoma. Podonotal shield mostly with roundish markings, except for reticulate antero-lateral region and a punctate band along posterior margin; 150 long and 162 wide at widest point; with 22 pairs of setae (setae r 1 and r 4 absent); without scleronoduli and with five pairs of distinguishable lyrifissures. Opisthonotal shield reticulate, except for a punctate band along anterior margin; 139 long and 146 wide at widest point; with 15 pairs of setae and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to opisthonotal shield with five pairs of setae (R 1 – R 5). Measurements of setae: j 1 12, j 2 14, j 3 11, j 4 10, j 5 12, j 6 13, z 1 12, z 2 11, z 3 11, z 4 12, z 5 13, z 6 14, s 1 9, s 2 11, s 3 12, s 4 12, s 5 13, s 6 13, r 2 12, r 3 19, r 5 12, r 6 14, J 1 16, J 2 14, J 3 15, J 4 16, J 5 10, Z 1 16, Z 2 17, Z 3 17, Z 4 16, Z 5 14, S 1 15, S 2 14, S 3 15, S 4 15, S 5 13, R 1 7, R 2 7, R 3 7, R 4 8, R 5 9. Ventral idiosoma (Fig. 6 B). Base of tritosternum 13 long and 10 wide proximally; laciniae 30 long, separated for about 90 % of their total length, pilose. Sternogenital shield lightly reticulate, with indistinct anterior margin, a small punctate region posterolaterad to seta st 4 and a punctate band along straight posterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; approximately 129 long, including the lightly sclerotised and punctate region, and 69 wide at widest point; with five pairs of setae and three pairs of lyrifissures; distance between st 5 – st 5 22; genital opening on anterior margin of shield. Ventrianal shield reticulate, except for a punctate band along anterior margin; approximately 110 long and 126 wide at widest point, not fused to dorsal shield; with seven pairs of setae (Jv 1, Jv 3, Jv 5 and Zv 1 – Zv 3) in addition to circum-anal setae and two pairs of distinguishable lyrifissures; post-anal seta longer than para-anal setae. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield narrow, not extending beyond peritreme. Measurements of setae: st 1 11, st 2 12, st 3 11, st 4 10, st 5 10, Jv 1 10, Jv 2 not seen (broken), Jv 3 12, Jv 5 11, Zv 1 9, Zv 2 10, Zv 3 10, para-anal 15, post-anal 21. Legs. Lengths: I: 247; II: 171; III: 168; IV: 225. Leg chaetotaxy as in adult female. With a spur-like ventral seta on each of femur, genu and tibia II (Fig. 6 C). Pretarsi as in adult female.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF23FFE1FF2A57F9FE001B78.taxon	discussion	Remarks. This species was described on the basis of the adult female holotype, two adult female paratypes and two adult male paratypes. The following characteristics appear in the original description and illustrations of the species, but do not agree with our observations of the female holotype; podonotal shield ornamented only with roundish markings, with 24 pairs of setae and structures resembling scleronoduli (illustrated by Karg, 2000 a, but not mentioned in the text); seta z 1 pilose; opisthonotal shield imbricate; unsclerotised integument laterad to opisthonotal shield with three pairs of setae; genital shield longer than wide; peritrematal shield fused with a section of exopodal shield next to coxa IV; lyrifissures present only on sternal shield (three pairs). No information was provided about the tritosternum and leg setal counts. The measurements provided in the original description referred to the length of the idiosoma (280 – 300), width of idiosoma (130 – 140), length of setae j 1 (cited as i 1) (12), j 5 (cited as i 4) (10), r 3 (cited as r 5) (18), J 1 (cited as I 1) (17), J 3 (cited as I 3) (16), J 5 (cited as I 5) (14), Z 5 (13), S 5 (16), range of hypostomal setae (12 – 16), length of legs I (250), II (200), III (160) and IV (250). For the adult male paratype, the only characteristics provided in the original description referred to details of the spermatodactyl and ventral setae of femur, genu and tibia II, as well as measurements for length (250 – 280) and width (140) of the idiosoma.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF20FFE7FF2A50A2FC461A7B.taxon	diagnosis	Diagnosis (adult male). Arthrodial process of chelicera with a short coronet-like fringe; epistome with smooth margin, and with antero-medial projection about twice as long as antero-lateral projections; podonotal shield mostly with roundish markings except for reticulate antero-lateral region, without scleronoduli; with a narrow band projecting posteriorly from region lateral to seta r 3, bearing seta r 4; dorsal idiosomal setae pilose, except setae z 1, z 2, s 1 and R 1 - R 4, smooth; opisthonotal shield with polygons outlined by aligned granules; seta R 5 absent; sternogenital shield smooth; unsclerotised integument between sternogenital and ventrianal shields without accessory platelet; peritreme extending anteriorly to mid-level of coxa II; with a spur-like ventral seta on each of femur, genu and tibia II. Adult male (Fig. 7 A – G) (holotype). Dorsal idiosomal setae (except z 1, z 2, s 1 and R 1 – R 4), Jv 5, para-anal and post-anal setae pilose; setae z 1, z 2, s 1, R 1 – R 4, hypostomal, subcapitular, ventral idiosomal (except Jv 5, para-anal and post-anal) and leg setae smooth. Gnathosoma. Fixed cheliceral digit 52 long, with three teeth in addition to apical tooth and a setiform pilus dentilis (Fig. 7 A); movable cheliceral digit 51 long, with one tooth in addition to apical tooth. Spermatodactyl 110 long, abruptly bent dorsally, distal portion approximately parallel to proximal portion, and almost twice as long. Arthrodial process of chelicera with a short coronet-like fringe. Epistome with a smooth antero-medial projection, narrowed in anterior one-third, with four apical denticles; flanked on each side by shorter, smooth projection with five apical denticles (Fig. 7 B); margin of epistome smooth between antero-medial and antero-lateral projections and laterad to antero-lateral projections. Deutosternal denticles in eight rows, each with 10 – 15 denticles; most anterior row shallow U-shaped, seventh row convex, other rows straight and transverse (Fig. 7 C). Measurements of setae: h 1 18, h 2 11, h 3 15, sc 15. Dorsal idiosoma (Fig. 7 D). Podonotal shield mostly with roundish markings, except for reticulate antero-lateral region, and a punctate band along its posterior margin, between setae s 6; 188 long and 222 wide at widest point; with 23 pairs of setae (seta r 1 absent); without scleronoduli and with four pairs of distinguishable lyrifissures; with a narrow band projecting posteriorly from region lateral to seta r 3, bearing seta r 4. Opisthonotal shield with polygons defined by aligned granules, except for a punctate band along anterior margin; 177 long and 154 wide at widest point; with 15 pairs of setae and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to opisthonotal shield with four pairs of setae (R 1 - R 4); seta R 5 absent. Measurements of setae: j 1 20, j 2 17, j 3 16, j 4 15, j 5 15, j 6 16, z 1 15, z 2 14, z 3 15, z 4 16, z 5 20, z 6 16, s 1 12, s 2 17, s 3 17, s 4 16, s 5 18, s 6 19, r 2 18, r 3 28, r 4 12, r 5 20, r 6 20, J 1 21, J 2 20, J 3 18, J 4 20, J 5 13, Z 1 21, Z 2 21, Z 3 20, Z 4 20, Z 5 25, S 1 22, S 2 21, S 3 22, S 4 22, S 5 21, R 1 10, R 2 10, R 3 11, R 4 10. Ventral idiosoma (Fig. 7 E). Base of tritosternum 20 long and 14 wide proximally (Fig. 7 F); laciniae 63 long, separated for about 90 % of their total length, pilose. Sternogenital shield smooth and with indistinct anterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; posterior margin straight; approximately 148 long, including the lightly sclerotised and punctate region, 74 wide at widest point; with five pairs of setae and three pairs of lyrifissures; distance between st 5 - st 5 34; genital opening on anterior margin of shield. Ventrianal shield reticulate, except for a punctate band along anterior margin, its anterior margin with a rounded protrusion, shield 125 long and 157 wide at widest point, not fused to dorsal shield; with seven pairs of setae (Jv 1, Jv 3, Jv 5 and Zv 1 - Zv 3) in addition to circum-anal setae and four pairs of distinguishable lyrifissures; post-anal seta longer than para-anal setae. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield narrow, not extending beyond peritreme. Measurements of setae: st 1 15, st 2 15, st 3 15, st 4 13, st 5 12, Jv 1 16, Jv 2 15, Jv 3 16, Jv 5 23, Zv 1 14, Zv 2 15, Zv 3 15, para-anal 16, post-anal 22. Legs. Lengths: I: 331; II: 267; III: 225; IV: 320. Genu IV with nine setae and tibia IV with ten setae. With a spur-like ventral seta on each of femur, genu and tibia II (Fig. 7 G). Pretarsus I, a single, sessile, curved claw; pretarsi II – IV, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF20FFE7FF2A50A2FC461A7B.taxon	discussion	Remarks. This species is known only from the adult male holotype. The following characteristics appear in the original description and illustrations of the species, but do not agree with our observations of the holotype: podonotal shield ornamented only with roundish markings, with 24 pairs of setae and structures resembling scleronoduli (illustrated by Karg, 2000 a but not mentioned in the text). No information was provided about the hypostome, tritosternum, ventral idiosoma, lyrifissures and leg setal counts. The measurements provided in the original description referred to the length of the idiosoma (350), width of idiosoma (200), length of setae j 1 (cited as i 1) (18), j 5 (cited as i 4) (15), z 6 (cited as i 5) (16), J 1 (cited as I 1) (20), J 2 (cited as I 2) (20), J 4 (cited as I 4) (20), Z 5 (25), S 5 (20), length of legs I (320), II (250), III (220) and IV (300).	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF26FFE4FF2A56F3FD7E1988.taxon	materials_examined	Type species: Poropodalius hexapennatus Karg, 2000, by original designation.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF26FFE4FF2A56F3FD7E1988.taxon	diagnosis	Diagnosis (adult female and male). Arthrodial process of chelicera with a short coronet-like fringe; epistome with a smooth antero-medial projection, about uniform width or wider at the base, flanked on each side by shorter projection, these either uniform width and smooth or triangular and serrate; seta h 3 directly posterior to h 1 and mediad or slightly postero-mediad to h 2; dorsal idiosomal setae smooth or with distal half pilose; podonotal shield with 21 – 22 pairs of setae and four scleronoduli between setae j 5 and j 6 (except P. basisetae, where scleronoduli are between setae j 6 and z 6); unsclerotised integument laterad to podonotal shield with one or two pairs of setae; opisthonotal shield with 15 pairs of setae; with four or five pairs of R setae on unsclerotised integument laterad to opisthonotal shield in female and on ventrianal shield in male; presternal shields absent; ventrianal shield with five or six pairs of setae in female and 12 pairs of setae in male; unsclerotised integument along anterior margin of ventrianal shield with two pairs of setae in female and without setae in male; with zero or one pair of metapodal plates in female and without metapodal plates in male; peritreme extending anteriorly to level of median region of coxa II; pretarsus I either a single, sessile, curved claw or an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes, with the same length or shorter than pretarsi of legs II – IV; seta pl 4 of tarsus IV absent. Numbers of setae on legs I – IV: coxa: 2, 2, 2, 1; trochanter: 6, 5, 5, 5; femur: 13, 11, 6, 6; genu: 13, 11, 9, 10; tibia: 14, 10, 8, 10; tarsus II – IV: 18, 18, 17. Adult male with spermatodactyl abruptly bent dorsally and with a spur-like ventral seta on each of femur, genu and tibia II.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF25FFE5FF2A52F2FCE61A7B.taxon	diagnosis	Diagnosis (adult female). Epistome with smooth antero-lateral projections of about uniform width; dorsal idiosomal setae smooth; podonotal shield lightly imbricate, with four crescent-shaped scleronoduli between setae j 5 and j 6; opisthonotal and sternal shields lightly imbricate; unsclerotised integument between sternal and ventrianal shields without accessory platelet; ventrianal shield with five pairs of setae (Jv 2, Jv 3, Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae; peritrematal shield narrow, not extending beyond peritreme; pretarsi I – IV similar in shape and length, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes. Adult female (Fig. 8 A – D) (holotype). Dorsal and ventral idiosomal, hypostomal, subcapitular and leg setae smooth. Gnathosoma. Fixed cheliceral digit 30 long; movable digit 31 long; teeth and pilus dentilis not visible because of position of chelicera. Arthrodial process of chelicera with a short coronet-like fringe. Epistome with a smooth antero-medial projection of about uniform width, with four apical denticles, flanked on each side by shorter, smooth projection with two to four apical denticles (Fig. 8 A); margin of epistome smooth between antero-medial and antero-lateral projections and serrate laterad to lateral projections. Deutosternal denticles in seven rows, each with eight to 12 denticles; fifth row anteriorly convex, other rows straight and transverse (Fig. 8 B). Seta h 3 posterior to h 1 and slightly postero-mediad to h 2. Measurements of setae: h 1 13, h 2 not seen (broken), h 3 not seen (broken), sc 10. Dorsal idiosoma (Fig. 8 C). Podonotal shield lightly imbricate, except for a punctate band along posterior margin; 137 long and 128 wide at widest point; with 21 pairs of setae (seta r 1 absent); with four crescent-shaped scleronoduli between setae j 5 and j 6 and five pairs of distinguishable lyrifissures. Unsclerotised integument laterad to podonotal shield with two pairs of setae (r 3 and r 4). Opisthonotal shield lightly imbricate, except for a punctate band along anterior margin; 132 long and 131 wide at widest point; with 15 pairs of setae and five pairs of distinguishable lyrifissures. Unsclerotised integument laterad to opisthonotal shield with five pairs of setae (R 1 - R 5). Measurements of setae: j 1 11, j 2 10, j 3 9, j 4 8, j 5 8, j 6 not seen (broken), z 1 9, z 2 7, z 3 9, z 4 8, z 5 not seen (broken), z 6 8, s 1 6, s 2 5, s 3 8, s 4 8, s 5 9, s 6 8, r 2 7, r 3 13, r 4 7, r 5 8, r 6 10, J 1 8, J 2 8, J 3 8, J 4 10, J 5 12, Z 1 9, Z 2 9, Z 3 10, Z 4 13, Z 5 19, S 1 9, S 2 8, S 3 10, S 4 14, S 5 18, R 1 6, R 2 not seen (broken), R 3 5, R 4 5, R 5 5. Ventral idiosoma (Fig. 8 D). Tritosternum not seen (broken). Sternal shield lightly imbricate and with indistinct anterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; posterior margin straight; approximately 85 long, including the lightly sclerotised and punctate region, 79 wide at widest point; with four pairs of setae and three pairs of lyrifissures. Genital shield lightly reticulate, except for a punctate band along straight posterior margin; longer than wide; extending posteriorly behind coxae IV; distance between st 5 - st 5 42. Unsclerotised integument between genital and ventrianal shields with setae Jv 1 and Zv 1. Metapodal plates not distinguishable. Ventrianal shield reticulate; 88 long and 101 wide at widest point, not fused to dorsal shield; with five pairs of setae (Jv 2, Jv 3, Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae and four pairs of distinguishable lyrifissures. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield narrow, not extending beyond peritreme. Measurements of setae: st 1 14, st 2 not seen (broken), st 3 not seen (broken), st 4 not seen (broken), st 5 not seen (broken), Jv 1 10, Jv 2 not seen (broken), Jv 3 not seen (broken), Jv 5 17, Zv 1 8, Zv 2 not seen (broken), Zv 3 11, para-anal 14, post-anal not seen (broken). Legs. Lengths: I: 248; II: 177; III: 153; IV not suitable for measurement. Pretarsi I – IV similar in shape and length, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF25FFE5FF2A52F2FCE61A7B.taxon	discussion	Remarks. This species was described on the basis of the adult female holotype and an adult male paratype. The following characteristics appear in the original description and illustrations of the species (Karg, 2000 a), but do not agree with our observations of the female holotype (the paratype male was not re-examined in this study): unsclerotised integument laterad to opisthonotal shield with two pairs of setae; genital shield smooth; a pair of rounded metapodal plates, containing a circular structure; lyrifissures present only on podonotal shield (a pair) and sternal shield (two pairs). No information was provided about the chelicera, hypostome, tritosternum and leg setal counts. The only measurements provided for female in the original description referred to the length of the idiosoma (260), width of idiosoma (140), range of dorsal idiosomal setae (10 – 16), length of ventral idiosomal setae (10), length of legs I (250), II (170), III (150) and IV (220).	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF24FFFBFF2A5479FD811DC0.taxon	diagnosis	Diagnosis of adult. Epistome with serrate, triangular antero-lateral projections; dorsal idiosomal setae smooth; podonotal shield lightly imbricate, with four crescent-shaped scleronoduli between setae j 6 and z 6; opisthonotal shield with anterior half lightly imbricate and posterior half with indistinct roundish markings; sternal shield smooth; unsclerotised integument between sternal and ventrianal shields with a transverse elongate accessory platelet; ventrianal shield with five pairs of setae (Jv 2, Jv 3, Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae; peritrematal shield wider than peritreme, anterior and posterior ends pointed; pretarsi I – IV similar in shape, an elongate ambulacral stalk, a pair of strong claws, and three rounded pulvillar lobes; pretarsus I about half as long as other pretarsi. Adult female (Fig. 9 A – F) (holotype). Dorsal and ventral idiosomal, hypostomal, subcapitular and leg setae smooth. Gnathosoma. Fixed cheliceral digit 31 long, with five teeth in addition to apical tooth (pilus dentilis indistinguishable) (Fig. 9 A); movable cheliceral digit 32 long, with three teeth in addition to apical tooth. Arthrodial process of chelicera with a short coronet-like fringe. Epistome with a smooth antero-medial projection of about uniform width, with two apical denticles, flanked on each side by shorter, triangular and serrate projection (Fig. 9 B); margin of epistome smooth between antero-medial and antero-lateral projections and serrate laterad to anterolateral projections. Deutosternal denticles in seven straight transverse rows, each with eight to 12 denticles (Fig. 9 C). Seta h 3 directly posterior to h 1 and mediad to h 2. Measurements of setae: h 1 15, h 2 12, h 3 13, sc 13. Dorsal idiosoma (Fig. 9 D). Podonotal shield lightly imbricate, except for a punctate band along its posterior margin, between setae s 6; 132 long and 129 wide at widest point; with 22 pairs of setae (seta r 1 absent); with four crescent-shaped scleronoduli between setae j 6 and z 6 and five pairs of distinguishable lyrifissures. Unsclerotised integument laterad to podonotal shield with a pair of setae (r 4). Opisthonotal shield with anterior half lightly imbricate, a punctate band along anterior margin, and posterior half with indistinct roundish markings; 133 long and 126 wide at widest point; with 15 pairs of setae and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to opisthonotal shield with four pairs of setae (R 1 – R 4); seta R 5 absent. Measurements of setae: j 1 14, j 2 18, j 3 15, j 4 16, j 5 14, j 6 14, z 1 15, z 2 15, z 3 16, z 4 17, z 5 15, z 6 15, s 1 12, s 2 13, s 3 15, s 4 16, s 5 15, s 6 16, r 2 9, r 3 21, r 4 9, r 5 17, r 6 21, J 1 16, J 2 18, J 3 20, J 4 24, J 5 21, Z 1 20, Z 2 20, Z 3 25, Z 4 30, Z 5 32, S 1 20, S 2 21, S 3 22, S 4 27, S 5 29, R 1 8, R 2 8, R 3 10, R 4 10. Ventral idiosoma (Fig. 9 E). Base of tritosternum 12 long and 9 wide proximally (Fig. 9 F); laciniae 28 long, separated for about 90 % of their total length, pilose. Sternal shield smooth and with indistinct anterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; posterior margin straight; approximately 92 long, including the lightly sclerotised and punctate region, 58 wide at widest point; with four pairs of setae and three pairs of lyrifissures. Genital shield lightly reticulate; shorter than wide; extending posteriorly behind coxae IV; distance between st 5 - st 5 35; posterior margin straight. Unsclerotised integument between genital and ventrianal shields with a transverse elongate accessory platelet and with setae Jv 1 and Zv 1; with a pair of elongate, curved metapodal plates, each with a small median lobe on the convex paraxial margin. Ventrianal shield reticulate; 84 long and 102 wide at widest point, not fused to dorsal shield; with five pairs of setae (Jv 2, Jv 3, Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae and four pairs of distinguishable lyrifissures; post-anal seta as long as para-anal setae. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield wider than peritreme, with pointed anterior and posterior ends. Measurements of setae: st 1 15, st 2 16, st 3 15, st 4 13, st 5 12, Jv 1 11, Jv 2 not seen (broken), Jv 3 17, Jv 5 26, Zv 1 12, Zv 2 16, Zv 3 21, para-anal 17, post-anal 17. Legs. Lengths: I: 231; II: 158; III: 156; IV: 210. Pretarsi I – IV similar in shape, an elongate ambulacral stalk, a pair of strong claws, and three rounded pulvillar lobes; pretarsus I about half as long as other pretarsi.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF24FFFBFF2A5479FD811DC0.taxon	discussion	Remarks. This species is known only from the adult female holotype. The following characteristics appear in the original description and illustrations of the species, but do not agree with our observations of the holotype: podonotal shield reticulate, with 21 pairs of setae, scleronoduli anterior to setae j 6 (i 5 in Karg's notation); unsclerotised integument laterad to podonotal shield with a pair of setae; opisthonotal shield reticulate; genital shield smooth; metapodal plates very large and with a median constriction on the paraxial margin; lyrifissures present only on sternal shield (two pairs). No information was provided about the chelicera, hypostome, tritosternum and leg setal counts. The measurements provided in the original description referred to the length of the idiosoma (220 – 250), width of idiosoma (100 – 110), length of setae j 1 (cited as i 1) (13), j 3 (cited as i 2) (16), j 5 (cited as i 4) (15), r 3 (cited as r 5) (16), J 1 (cited as I 1) (16), J 2 (cited as I 2) (18), J 3 (cited as I 3) (18), J 5 (cited as I 5) (20), Z 3 (22), Z 4 (25), Z 5 (27), S 1 – S 5 (20), Jv 5 (cited as V 8) (25), post-anal (15), length of other ventral idiosomal setae (20) and length of metapodal plate (17).	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF3AFFFEFF2A553AFA4A1DA4.taxon	diagnosis	Diagnosis (adult female and male). Epistome with smooth antero-lateral projections, widest at the base; setae j 1, z 2 - z 6, s 5, r 3, r 6, J 2 - J 5, Z 1 - Z 5, S 2 - S 5 pilose in distal half; other dorsal idiosomal setae smooth; podonotal shield with four small circular scleronoduli between setae j 5 and j 6; peritrematal shield broad and long, extending anteriorly onto the dorsum but not fused to podonotal shield; pretarsus I, a single, sessile, curved claw. Adult female with podonotal shield mostly with series of short, wavy lines mostly aligned transversely or diagonally; opisthonotal shield imbricate with many imbrications containing series of short longitudinal lines; sternal shield lightly reticulate; unsclerotised integument between genital and ventrianal shields with three transverse elongate accessory platelets; ventrianal shield with six pairs of setae (Jv 2 - Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae. Adult male with podonotal shield imbricate; opisthonotal shield imbricate with imbrications without series of short, longitudinal lines; sternogenital shield lightly reticulate; unsclerotised integument between sternogenital and ventrianal shields with an ellipsoidal accessory platelet; ventrianal shield with 12 pairs of setae (R 1 - R 4, Jv 1 - Jv 5 and Zv 1 - Zv 3) in addition to circum-anal setae; seta r 4 on peritrematal shield. Adult female (Fig. 10 A – E) (holotype). Setae j 2 - j 6, z 1, s 1 - s 4, s 6, r 2, r 4, r 5, J 1, S 1, R 1 - R 4, hypostomal, subcapitular, ventral idiosomal (except Jv 5) and leg setae smooth; setae j 1, z 2 - z 6, s 5, r 3, r 6, J 2 - J 5, Z 1 - Z 5, S 2 - S 5 and Jv 5 pilose in distal half. Gnathosoma. Fixed cheliceral digit 66 long; movable cheliceral digit 68 long; teeth and pilus dentilis not visible because of position of chelicera. Arthrodial process of chelicera with a short coronet-like fringe. Epistome with a smooth antero-medial projection, widest at the base, with two apical denticles, flanked on each side by shorter, smooth projection with three apical denticles (Fig. 10 A); margin of epistome with two denticles between anteromedial and antero-lateral projections and smooth laterad to antero-lateral projections. Deutosternal denticles in eight rows, each with eight to 12 denticles; anterior row convex, posterior row V-shaped, other rows straight and transverse (Fig. 10 B). Seta h 3 directly posterior to h 1 and slightly postero-mediad to h 2. Measurements of setae: h 1 24, h 2 13, h 3 not seen (broken), sc 22. Dorsal idiosoma (Fig. 10 C). Podonotal shield with series of very short, wavy lines mostly transversely or diagonally aligned, except for a punctate band along posterior margin; 224 long and 260 wide at widest point; with 22 pairs of setae (seta r 1 absent); with four small circular scleronoduli between setae j 5 and j 6 and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to podonotal shield with a pair of short setae (r 4). Opisthonotal shield imbricate, except for a punctate band along its anterior margin, between setae Z 1; many imbrications containing series of short, longitudinal lines; shield 195 long and 258 wide at widest point; with 15 pairs of setae and five pairs of distinguishable lyrifissures. Unsclerotised integument laterad to opisthonotal shield with four pairs of setae (R 1 - R 4); seta R 5 absent. Measurements of setae: j 1 23, j 2 23, j 3 21, j 4 23, j 5 17, j 6 20, z 1 16, z 2 21, z 3 20, z 4 18, z 5 20, z 6 22, s 1 14, s 2 14, s 3 15, s 4 18, s 5 20, s 6 23, r 2 20, r 3 35, r 4 10, r 5 18, r 6 27, J 1 19, J 2 23, J 3 22, J 4 24, J 5 17, Z 1 23, Z 2 23, Z 3 30, Z 4 32, Z 5 35, S 1 23, S 2 23, S 3 26, S 4 30, S 5 32, R 1 10, R 2 10, R 3 13, R 4 16. Ventral idiosoma (Fig. 10 D). Base of tritosternum 28 long and 15 wide proximally (Fig. 10 E); laciniae 57 long, separated for about 90 % of their total length, pilose. Sternal shield lightly reticulate, with indistinct anterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; posterior margin straight; approximately 123 long, including the lightly sclerotised and punctate region, and 103 wide at widest point; with four pairs of setae and three pairs of lyrifissures. Genital shield lightly reticulate; longer than wide; extending posteriorly behind coxae IV; distance between st 5 - st 5 62; posterior margin straight. Unsclerotised integument between genital and ventrianal shields with three transverse elongate accessory platelets, and with setae Jv 1 and Zv 1; with a pair of roundish metapodal plates, each with a median constriction on the paraxial margin, and containing a circular marking. Ventrianal shield reticulate; 124 long and 243 wide at widest point, not fused to dorsal shield; with six pairs of setae (Jv 2 - Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae and four pairs of distinguishable lyrifissures; post-anal seta as long as para-anal setae. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield smooth, broad and long, extending anteriorly onto the dorsum but not fused to podonotal shield. Measurements of setae: st 1 16, st 2 22, st 3 not seen (broken), st 4 16, st 5 15, Jv 1 not seen (broken), Jv 2 19, Jv 3 25, Jv 4 25, Jv 5 30, Zv 1 16, Zv 2 18, Zv 3 16, para-anal 23, post-anal 24. Legs: Lengths: I: 354; II: 298; III: 279; IV: 357. Pretarsus I, a sessile, curved claw; pretarsi II – IV, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes. Adult male (Fig. 11 A – B) (two paratypes). Shape of setae as in adult female. Gnathosoma. A ll structures damaged and unavailable for description. Dorsal idiosoma: Podonotal shield imbricate, except for a punctate band along posterior margin; 188 long and 195 wide at widest point; with 22 pairs of setae (seta r 1 absent); with four circular scleronoduli between setae j 5 and j 6 and four pairs of distinguishable lyrifissures. Seta r 4 on distal end of peritrematal shield. Opisthonotal shield imbricate, except for a punctate band along its anterior margin, between setae Z 1; imbrications without series of short, longitudinal lines; 146 – 161 long and 171 – 189 wide at widest point; with 15 pairs of setae and five pairs of distinguishable lyrifissures. Setae R 1 - R 4 inserted on ventrianal shield; seta R 5 absent. Measurements of setae: j 1 16, j 2 not seen (broken), j 3 16, j 4 15 – 16, j 5 15, j 6 16, z 1 13 – 14, z 2 16, z 3 not seen (broken), z 4 16 – 17, z 5 not seen (broken), z 6 16 – 17, s 1 not seen (broken), s 2 14, s 3 16, s 4 16, s 5 18, s 6 17, r 2 14, r 3 not seen (broken), r 4 8 – 9, r 5 15, r 6 18, J 1 18 – 19, J 2 20, J 3 19 – 20, J 4 21, J 5 14 – 15, Z 1 21, Z 2 20, Z 3 23 – 24, Z 4 26, Z 5 28, S 1 20, S 2 20, S 3 22 – 23, S 4 22, S 5 24, R 1 9 – 10, R 2 11, R 3 15, R 4 14. Ventral idiosoma (Fig. 11 A): Tritosternum not seen (broken). Sternogenital shield lightly reticulate and with indistinct anterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; posterior margin straight; approximately 158 long, including the lightly sclerotised and punctate region, and 81 wide at widest point; with five pairs of setae and three pairs of lyrifissures; distance between st 5 - st 5 33 – 36; genital opening on anterior margin of shield. Unsclerotised integument between sternogenital and ventrianal shields with an ellipsoidal accessory platelet, punctate near anterior margin of ventrianal shield. Ventrianal shield imbricate; 104 – 110 long and 192 – 201 wide at widest point, not fused to dorsal shield; with 12 pairs of setae (R 1 – R 4, Jv 1 - Jv 5 and Zv 1 - Zv 3) in addition to circum-anal setae and four pairs of distinguishable lyrifissures; post-anal seta as long as paraanal setae. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield broad and long, extending anteriorly onto the dorsum but not fused to podonotal shield. Measurements of setae: st 1 not seen (broken), st 2 not seen (broken), st 3 not seen (broken), st 4 13, st 5 12, Jv 1 14, Jv 2 15, Jv 3 16 – 18, Jv 4 16, Jv 5 22, Zv 1 not seen (broken), Zv 2 15, Zv 3 15 – 16, para-anal 16 – 18, post-anal 20. Legs: Lengths: I: 311; II: 220; III: 203 – 207; IV: 270. With a spur-like ventral seta on each of femur, genu and tibia II (Fig. 11 B). Pretarsi as in adult female.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF3AFFFEFF2A553AFA4A1DA4.taxon	discussion	Remarks. This species was described on the basis of the adult female holotype, one adult female paratype and two adult male paratypes. The following characteristics appear in the original description and illustrations of the species, but do not agree with our observations of the female holotype: margin of epistome smooth between anteromedial and antero-lateral projections; setae j 1, j 6, z 4 - z 6, s 4 - s 5, r 3, r 5 - r 6, J 1 - J 5, Z 2 - Z 5, S 1 - S 5 and Jv 5 totally pilose, other setae smooth; genital shield smooth; peritrematal shield ornamented; lyrifissures present only on podonotal shield (one pair) and sternal shield (three pairs). No information was provided about the chelicera, hypostome, tritosternum and leg setal counts. The measurements provided in the original description referred to the length of the idiosoma (320 – 400), width of idiosoma (200 – 250), length of setae j 1 (cited as i 1) (20), j 4 (cited as i 3) (20), j 5 (cited as i 4) (20), r 3 (cited as r 5) (35), J 1 (cited as I 1) (17), J 3 (cited as I 3) (25), J 4 (cited as I 4) (25), Z 3 (25), Z 4 (30), Z 5 (30), S 4 (27), S 5 (32), Jv 5 (cited as V 8) (30) and length of other ventral idiosomal setae (25). For the adult male paratypes, the only characteristics provided referred to details of the spermatodactyl and ventral setae of femur, genu and tibia II, as well as measurements for length (380) and width (230) of the idiosoma.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF3FFFFDFF2A56D9FED21DF1.taxon	diagnosis	Diagnosis (adult female and male). Epistome with smooth antero-lateral projections of about uniform width; setae r 3, J 4, J 5, Z 4, Z 5, S 5 pilose in distal half; other dorsal idiosomal setae smooth; podonotal shield with indistinct roundish markings and four small circular scleronoduli between setae j 5 and j 6; opisthonotal shield lightly reticulate; pretarsus I, a single, sessile, curved claw. Adult female with sternal shield lightly imbricate; unsclerotised integument between sternal and ventrianal shields with a transverse elongate accessory platelet; ventrianal shield with five pairs of setae (Jv 2, Jv 3, Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae; peritrematal shield broad and long, extending anteriorly onto the dorsum but not fused to podonotal shield. Adult male with sternogenital shield with indistinct roundish markings; no accessory platelet between sternogenital and ventrianal shields; ventrianal shield with 12 pairs of setae (R 1 - R 5, Jv 1 - Jv 3, Jv 5 and Zv 1 - Zv 3) in addition to circum-anal setae; peritrematal shield narrow, not extending beyond peritreme. Adult female (Fig. 12 A – E) (holotype). Dorsal idiosomal setae (except r 3, J 4, J 5, Z 4, Z 5, S 5), hypostomal, subcapitular, ventral idiosomal (except post-anal) and leg setae smooth; setae r 3, J 4 - J 5, Z 4 - Z 5, S 5 and post-anal setae pilose in distal half. Gnathosoma. Fixed cheliceral digit 67 long, with five teeth in addition to apical tooth, and a setiform pilus dentilis (Fig. 12 A); movable cheliceral digit 66 long, with three teeth in addition to apical tooth. Arthrodial process of chelicera with a short coronet-like fringe. Epistome with a smooth antero-medial projection slightly wider at the base and provided with two apical denticles, flanked on each side by a shorter, smooth projection with three apical denticles (Fig. 12 B); margin of epistome with two denticles between antero-medial and antero-lateral projections and smooth laterad to antero-lateral projections. Deutosternum, hypostomal and subcapitular setae not visible. Dorsal idiosoma (Fig. 12 C). Podonotal shield with indistinct roundish markings, except for a punctate band along posterior margin; 201 long and 210 wide at widest point; with 22 pairs of setae (seta r 1 absent); with four small circular scleronoduli between setae j 5 and j 6 and five pairs of distinguishable lyrifissures. Unsclerotised integument laterad to podonotal shield with a pair of setae (r 4). Opisthonotal shield lightly reticulate, except for a punctate band along anterior margin; 167 long and 180 wide at widest point; with 15 pairs of setae and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to opisthonotal shield with five pairs of setae (R 1 - R 5). Measurements of setae: j 1 20, j 2 20, j 3 18, j 4 17, j 5 16, j 6 19, z 1 16, z 2 18, z 3 18, z 4 20, z 5 19, z 6 20, s 1 16, s 2 18, s 3 18, s 4 19, s 5 20, s 6 21, r 2 18, r 3 25, r 4 12, r 5 20, r 6 22, J 1 20, J 2 21, J 3 22, J 4 23, J 5 21, Z 1 21, Z 2 22, Z 3 25, Z 4 30, Z 5 33, S 1 20, S 2 20, S 3 22, S 4 26, S 5 29, R 1 11, R 2 11, R 3 12, R 4 13, R 5 12. Ventral idiosoma (Fig. 12 D). Base of tritosternum 24 long and 14 wide proximally (Fig. 12 E); laciniae 62 long, separated for about 90 % of their total length, pilose. Sternal shield lightly imbricate and with indistinct anterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; posterior margin concave; approximately 103 long, including the lightly sclerotised and punctate region, and 96 wide at widest point; with four pairs of setae and three pairs of lyrifissures. Genital shield lightly reticulate, except for a punctate band along straight posterior margin; longer than wide; extending posteriorly behind coxae IV; distance between st 5 - st 5 50. Unsclerotised integument between genital and ventrianal shields with a transverse elongate accessory platelet and with setae Jv 1 and Zv 1. Metapodal plates irregular in shape, with a median constriction on the paraxial margin, containing a circular marking. Ventrianal shield reticulate; 125 long and 152 wide at widest point, not fused to dorsal shield; with five pairs of setae (Jv 2, Jv 3, Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae and three pairs of distinguishable lyrifissures; post-anal seta as long as para-anal setae. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield broad and long, extending anteriorly onto the dorsum but not fused to podonotal shield. Measurements of setae: st 1 17, st 2 not seen (broken), st 3 not seen (broken), st 4 not seen (broken), st 5 17, Jv 1 15, Jv 2 not seen (broken), Jv 3 20, Jv 5 27, Zv 1 14, Zv 2 16, Zv 3 15, para-anal 20, post-anal 22. Legs. Lengths: I: 332; II: 275; III: 226; IV: 310. Pretarsus I, a single, sessile, curved claw; pretarsi II – IV, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes. Adult male (Fig. 13 A – C) (three paratypes). Shape of setae as in adult female. Gnathosoma. Fixed cheliceral digit 42 long, with two teeth in addition to apical tooth and a setiform pilus dentilis (Fig. 13 A); movable cheliceral digit 41 long, with three teeth in addition to apical tooth. Spermatodactyl 83 long, abruptly bent dorsally, distal portion approximately parallel to proximal portion and double its length. Arthrodial process of chelicera with a short coronet-like fringe. Epistome as in female. Deutosternum, hypostomal and subcapitular setae not visible. Dorsal idiosoma. Podonotal shield with indistinct roundish markings, except for a punctate band along posterior margin; 176 – 192 long and 203 – 212 wide at widest point; with 22 pairs of setae (seta r 1 absent); with four circular scleronoduli between setae j 5 and j 6 and five pairs of distinguishable lyrifissures. Unsclerotised integument laterad to podonotal shield with a pair of setae (r 4). Opisthonotal shield lightly reticulate, except for a punctate band along anterior margin; 168 – 179 long and 179 – 187 wide at widest point; with 15 pairs of setae and four pairs of distinguishable lyrifissures. Setae R 1 - R 5 inserted on ventrianal shield. Measurements of setae: j 1 17, j 2 17, j 3 16 – 18, j 4 15 – 17, j 5 13 – 15, j 6 not seen (broken), z 1 14, z 2 14 – 16, z 3 15 – 18, z 4 16 – 18, z 5 14 – 18, z 6 16, s 1 14, s 2 16 – 18, s 3 17, s 4 17 – 20, s 5 16 – 20, s 6 20, r 2 16, r 3 21 – 27, r 4 11 – 13, r 5 16 – 18, r 6 20 – 22, J 1 15 – 20, J 2 18 – 20, J 3 14 – 18, J 4 16 – 21, J 5 15 – 18, Z 1 15 – 21, Z 2 15 – 21, Z 3 18 – 22, Z 4 20 – 26, Z 5 23 – 28, S 1 15 – 20, S 2 17 – 20, S 3 18 – 22, S 4 20 – 24, S 5 21 – 25, R 1 8 – 12, R 2 9 – 12, R 3 12 – 14, R 4 10 – 14, R 5 11 – 15. Ventral idiosoma (Fig. 13 B). Base of tritosternum 17 long and 11 wide proximally; laciniae 51 long, separated for about 90 % of their total length, pilose. Sternogenital shield with indistinct roundish markings, indistinct anterior margin and a punctate band along straight posterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; approximately 136 – 154 long, including the lightly sclerotised and punctate region, and 73 – 84 wide at widest point; with five pairs of setae and three pairs of lyrifissures; distance between st 5 - st 5 29 – 37; genital opening on anterior margin of shield. Unsclerotised integument between sternogenital and ventrianal shields without accessory platelet. Ventrianal shield reticulate, except for a punctate band along anterior margin; 135 – 155 long and 201 – 230 wide at widest point, not fused to dorsal shield; with 12 pairs of setae (R 1 - R 5, Jv 1 - Jv 3, Jv 5 and Zv 1 - Zv 3) in addition to circum-anal setae and three pairs of distinguishable lyrifissures; post-anal seta as long as para-anal setae. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield narrow, not extending beyond peritreme. Measurements of setae: st 1 15 – 16, st 2 14 – 16, st 3 16 – 17, st 4 13 – 15, st 5 13 – 15, Jv 1 15, Jv 2 12 – 15, Jv 3 13 – 16, Jv 5 20 – 23, Zv 1 15, Zv 2 11 – 13, Zv 3 11 – 14, para-anal 15 – 20, post-anal 16 – 21. Legs. Lengths: I: 311; II: 270; III: 210 – 235; IV: 265 – 305. With a spur-like ventral seta on each of femur, genu and tibia II (Fig. 13 C). Pretarsi as in adult female.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF3FFFFDFF2A56D9FED21DF1.taxon	discussion	Remarks. This species was described on the basis of the adult female holotype, 12 adult female paratypes and 15 adult male paratypes. The following characteristics appear in the original description and illustrations of the species, but do not agree with our observations of the holotype: epistome with antero-medial projection as long as antero-lateral projections; podonotal shield ornamented with few irregular lines; unsclerotised integument laterad to opisthonotal shield with only two pairs of setae; setae S 5 and post-anal smooth and setae j 1, r 3, J 4, J 5, Z 4, Z 5 and Jv 5 totally pilose; unsclerotised integument between genital and ventrianal shields with five accessory platelets; unsclerotised integument laterad to ventrianal shield with four pairs of setae; peritrematal shield narrow, not extending beyond peritreme; lyrifissures only on podonotal shield (one pair) and sternal shield (three pairs). No information was provided about the hypostome, tritosternum and leg setal counts. The measurements provided in the original description referred to the length of the idiosoma (310 – 360), width of idiosoma (170 – 190), length of setae r 3 (cited as r 5) (28), J 4 (cited as I 4) (25), Z 4 (28), Z 5 (30), range of other dorsal idiosomal setae (18 – 20), length of all st 17, Jv 5 (cited as V 8) (25), range of other ventrianal idiosomal setae (16 – 20), length of legs I (290), II (250), III (230) and IV (310). For the male paratypes, the only characteristics provided referred to details of the spermatodactyl and ventral setae of femur, genu and tibia II, as well as measurements for the length (270 – 300) and width (150 – 190) of the idiosoma.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF3CFFF3FF2A5524FD5F1C2B.taxon	diagnosis	Diagnosis (adult female). Epistome with smooth antero-lateral projections of about uniform width; dorsal idiosomal setae smooth; podonotal shield with indistinct roundish markings and four crescent-shaped scleronoduli between setae j 5 and j 6; opisthonotal shield lightly imbricate; sternal shield with light reticulation along antero-lateral margin; no accessory platelet between sternal and ventrianal shields; ventrianal shield with six pairs of setae (Jv 2, Jv 3, Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae; peritrematal shield narrow, not extending beyond peritreme; pretarsi I – IV similar in shape, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes; pretarsus I about half as long as other pretarsi. Adult female (Fig. 14 A – C) (holotype). Dorsal and ventral idiosomal, hypostomal, subcapitular and leg setae smooth. Gnathosoma. Fixed cheliceral digit 31 long; movable cheliceral digit 32 long; teeth and pilus dentilis not visible because of position of chelicera. Arthrodial process of chelicera with a short coronet-like fringe. Epistome with a smooth antero-medial projection of about uniform width, with two apical denticles, flanked on each side by shorter, smooth projection with three apical denticles (Fig. 14 A); margin of epistome smooth between anteromedial and antero-lateral projections and serrate laterad to antero-lateral projections. Deutosternum, hypostomal and subcapitular setae not visible. Dorsal idiosoma (Fig. 14 B): Podonotal shield with indistinct roundish markings, except for a punctate band along posterior margin; 130 long and 119 wide at widest point; with 22 pairs of setae (seta r 1 absent); with four crescent-shaped scleronoduli between setae j 5 and j 6 and five pairs of distinguishable lyrifissures. Unsclerotised integument laterad to podonotal shield with a pair of setae (r 4). Opisthonotal shield lightly imbricate, except for a punctate band along anterior margin; 132 long and 125 wide at widest point; with 15 pairs of setae and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to opisthonotal shield with four pairs of setae (R 1 - R 4); seta R 5 absent. Measurements of setae: j 1 17, j 2 not seen (broken), j 3 20, j 4 not seen (broken), j 5 20, j 6 not seen (broken), z 1 16, z 2 18, z 3 21, z 4 not seen (broken), z 5 18, z 6 23, s 1 12, s 2 not seen (broken), s 3 22, s 4 22, s 5 21, s 6 23, r 2 21, r 3 29, r 4 12, r 5 21, r 6 24, J 1 25, J 2 not seen (broken), J 3 not seen (broken), J 4 25, J 5 22, Z 1 25, Z 2 not seen (broken), Z 3 24, Z 4 25, Z 5 27, S 1 25, S 2 24, S 3 25, S 4 25, S 5 25, R 1 10, R 2 10, R 3 14, R 4 16. Ventral idiosoma (Fig. 14 C). Tritosternum not seen (broken). Sternal shield with light reticulation along antero-lateral margin and with indistinct anterior margin; region anterior to the first pair of lyrifissures (iv 1) lightly sclerotised and punctate; posterior margin slightly concave; approximately 77 long, including the lightly sclerotised and punctate region, and 61 wide at widest point; with four pairs of setae and three pairs of lyrifissures. Genital shield with scanty reticulation and a punctate band along straight posterior margin; longer than wide; extending posteriorly behind coxae IV; distance between st 5 - st 5 38. Unsclerotised integument between genital and ventrianal shields with setae Jv 1 and Zv 1. Metapodal plates kidney-shaped, with narrow posterior end, containing a circular marking. Ventrianal shield reticulate; 87 long and 101 wide at widest point, not fused to dorsal shield; with five pairs of setae (Jv 2, Jv 3, Jv 5, Zv 2 and Zv 3) in addition to circum-anal setae and three pairs of distinguishable lyrifissures. Peritreme extending anteriorly to mid-level of coxa II. Peritrematal shield narrow, not extending beyond peritreme. Measurements of setae: st 1 not seen (broken), st 2 not seen (broken), st 3 not seen (broken), st 4 not seen (broken), st 5 not seen (broken), Jv 1 not seen (broken), Jv 2 not seen (broken), Jv 3 not seen (broken), Jv 5 24, Zv 1 not seen (broken), Zv 2 not seen (broken), Zv 3 not seen (broken), para-anal not seen (broken), post-anal not seen (broken). Legs. Lengths: I: 197; II: 152; III: 140; IV: 181. Pretarsi I – IV similar in shape, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes; pretarsus I about half as long as other pretarsi.	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
03CB87F0FF3CFFF3FF2A5524FD5F1C2B.taxon	discussion	Remarks. This species is known only from the adult female holotype. The following characteristics appear in the original description and illustrations of the species, but do not agree with our observations of the holotype: epistome with serrate anterior margin, a long flagellum-like antero-medial projection and without antero-lateral projections; podonotal shield reticulate; unsclerotised integument laterad to podonotal shield with two pairs of setae; opisthonotal shield ornamented with transverse lines; genital shield smooth; ventrianal shield with only a few weak transverse lines; lyrifissures present only on sternal shield (two pairs). No information was provided about the chelicera, hypostome, tritosternum and leg setal counts. The measurements provided in the original description referred to the length of the idiosoma (249), width of idiosoma (140), length of setae j 1 (cited as i 1) (17), j 2 (cited as z 1) (20), j 3 (cited as i 2) (19), j 5 (cited as i 4) (20), z 1 (cited as s 1) (15), z 2 (17), z 5 (cited as z 4) (17), s 2 (cited as r 2) (10), r 2 (cited as r 3) (20), r 6 (cited as r 5) (25), J 1 (cited as I 1) (22), J 4 (cited as I 4) (25), J 5 (cited as I 5) (22), Z 1 - Z 5 (24), S 1 (23), S 5 (25), length of all st (17), para-anal (10), Jv 5 (cited as lateral caudal setae) (22), length of other ventrianal idiosomal setae (17), length of the ventrianal shield (81), width of ventrianal shield (102), length of legs I (203), II (182), III (133) and IV (193).	en	Castilho, Raphael C., De, Gilberto J., Halliday, Bruce (2012): Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae). Zootaxa 3335: 1-28, DOI: 10.11646/zootaxa.3335.1.1
