identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03CBD358FF96FFABF3D7574FFD44FECB.text	03CBD358FF96FFABF3D7574FFD44FECB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis striata subsp. striata striata Ikeda 1936	<div><p>Cobitis striata striata Ikeda, 1936</p> <p>(Figs. 3A, 4A, B, 5A, 6A)</p> <p>Cobitis taenia striata Ikeda, 1936: 984, figs. 10, 11, 12, 13 (original description; type locality: near Takamatsu, Kagawa Pref., Shikoku, Japan); Cobitis taenia striata middle race: Minamori 1952: 201, fig. 2A; Cobitis taenia striata: Aoyagi 1957: 170, fig. 143; Cobitis taenia striata middle race: Saitoh and Aizawa 1987: 336, fig. 3G; Cobitis sp. M: Saitoh 1989: 390; middle form: Saitoh 1990: 240, figs. 3f, g, h, i, j: 241, fig. 4 (lower four); Cobitis taenia striata: Sezaki et al. 1994: 684, fig. 1B; Cobitis cf. striata: Kim et al. 1999: 388, fig. 9; Cobitis sp. 3: Hosoya 2002: 275; Cobitis striata complex middle race: Kitagawa et al. 2005: 112, table 1; Cobitis striata (middle race): Nakajima et al. 2008: 13, fig. 2E; normal type (Setouchi form) of middle race in Cobitis striata complex: Kitagawa et al. 2009: 12, fig. 2C; Cobitis striata complex middle race: Saitoh et al. 2010: 1003, table 1; Cobitis sp. 3 subsp. 1: Nakajima et al. 2012: 92, fig. 3a.</p> <p>Specimens examined. 1 male, 53.3 mm SL, Kizu River, Yodo River system, Yawata, Kyoto Pref., Honshu, 16. V. 2009, J. Nakajima; 1 male, 59.1 mm SL, Kino R., Wakayama, Wakayama Pref., Honshu, 29. XII. 2007, M. Nakatani; MPM-FI1500, 1 male, 58.6 mm SL, Muko R., Sanda, Hyogo Pref., Honshu, 7. VI. 2009, K. Tominaga; 3 males, 50.8–55.9 mm SL, Kagato R., Yoshii R. s., Setouchi, Okayama Pref., Honshu, 6. VI. 2007, J. Nakajima; FRLM24922, 1 male, 68.4 mm SL, Ota R., Hiroshima, Hiroshima Pref., Honshu, 29. IX. 1996, M. Watanabe; FAKU55767, 2 males, 58.9, 66.6 mm SL, Shimata R., Shuto, Yamaguchi Pref., Honshu, 4. V. 1982, K. Saitoh; TKPM-P17340, 1 male, 55.6 mm SL, Koto R., Takamatsu, Kagawa Pref., Shikoku, 3. XI. 2007, K. Tominaga; 1 male and 1 female, 49.6, 47.7 mm SL, Kasuga R., Takamatsu, Kagawa Pref., Shikoku, 28. IX. 2008, Y. Hashimoto; 2 males and 2 females, 55.3–73.7 mm SL, Doki R., Man-nou, Kagawa Pref., Shikoku, 3. XI. 2007, K. Tominaga; TKPM-P2283, 1 male and 2 females, 58.2–69.2 mm SL, Miyagouchitani R., Yoshino R. s., Kamiita, Tokushima Pref., Shikoku, 18. VII. 1995, Y. Sato; KPM-NI29502, 1 male and 1 female, 56.8, 61.5 mm SL, Kitara R., Ima R. s., Miyako, Fukuoka Pref., Kyushu, 12. XII. 2010, J. Nakajima; 1 male, 66.8 mm SL, Harai R., Yukuhashi, Fukuoka Pref., Kyushu, 25. X. 2007, J. Nakajima.</p> <p>Abbreviations used; N, individuals number; SL, standard length; HL, lateral head length; LPP, length of between pectoral-fin base and pelvic-fin origin; LPA, length of between pelvic-fin base and anal-fin origin; DCP, depth of caudal peduncle; PMN, prepelvic myotome number.</p> <p>Diagnosis. This species is distinguishable from other Japanese striated spined loaches by the following characteristics: body size moderate, the mature size about 50–60 mm SL in males, 60–80 mm SL in females; lamina circularis at the base of the pectoral fin of adult male simple roundish plate, the upper segments of the first branched soft ray narrow and weak; PMN commonly 13; lines L3 and L5 well developed with broad stripes in all season; line L4 faint; caudal fin and dorsal fin with 3–4 arcuate bars; upper spot at the caudal base jet-black, approximately eye diameter; lower spot at caudal base faint or missing; egg yolk diameter approximately 1.0 mm; karyotype diploid (2n = 50).</p> <p>Description. Lateral view in Figure 3A illustrate body shape, form and position of fins. Morphometric and meristic data for 15 males and 6 females are summarized in Table 2. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7–8; pelvic-fin rays ii, 6; caudal-fin rays 8+8. Body elongate, laterally compressed. Head and snout elongated. Interorbital space narrow, convex. Caudal peduncle relatively compressed. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well-developed lobes; upper lip with transverse wrinkles on surface. Barbels, three pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 13 (range, 13–14). Very small cycloid scales on the trunk. Lamina circularis at the base of the pectoral fin of adult male simple roundish plate (Fig. 6A). The first branched soft ray of pectoral fin longer than the others; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin narrow and weak. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens: 93.0 mm SL male, 98.0 mm SL female (Minamori 1952).</p> <p>Coloration. Male in the non-spawning season (Figs. 3A, 4A). Body yellowish white with light brown pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Upper part of head covered with some amorphous spots, these spots often stringed. Opercle and snout covered with large amorphous patterns. Body pigmentation organized in one middorsal and four lateral zones. Line L1 consisting of a series of 10–20 saddles or oval-shaped blotches. Line L2 formed by few small angular spots, only present on the predorsal region, reaching dorsally to interspaces of L1, barely distinct from L1 occasionally. Line L3 formed by sharp longitudinal line, reaching to caudal base. The posterior part of L3 often intermissive. Line L4 formed by weak dots or a narrow line, reaching beyond dorsal fin, sometimes nonexistent. Line L5 formed by broad longitudinal line from upper part of pectoral fin to the caudal-fin base. Caudal fin and dorsal fin with 3–4 arcuate bars. Anal fin pigmented along the fin rays. Upper spot at the caudal base jet-black comparable in size to eye diameter, lower spot at caudal base faint or missing.</p> <p>Male in the spawning season (Fig. 4B). Lines L2 and L4 not visible, L3 and L5 well developed with broad stripes from the upper part of the pectoral-fin base to the caudal-fin base.</p> <p>Female (Fig. 5A). Appearance similar to males in the non-spawning season.</p> <p>Sexual dimorphism. Males have roundish lamina circularis at the base of pectoral fins, but females do not. Generally, the body size of females is larger than that of males.</p> <p>Egg diameter. 0.98 ± 0.05 mm (females, N = 2; collected from the Yoshii River and the Asahi River, Okayama Prefecture)</p> <p>Karyotype. Diploid (2n = 50) (Ueno &amp; Ojima 1976; Ueno et al. 1980; Ueno 1981; Saitoh et al. 1984, 2000; Kimizuka 1987)</p> <p>Distribution. Rivers flowing into the Seto Inland Sea in Honshu, Shikoku, and Kyushu, and rivers flowing into the Japan Sea in Honshu: Kyoto, Osaka, Wakayama, Hyogo, Okayama, Hiroshima, Yamaguchi, Kagawa, Tokushima, Ehime, and Fukuoka Prefectures (Saitoh &amp; Aizawa 1987; Nakajima et al. 2008).</p> <p>Habitat and biology. This species inhabits sandy-mud bottoms of the middle and lower reaches of rivers. Saitoh (1990) described the spawning ecology of this species as the middle form of Cobitis striata.</p> <p>Remarks. Although this loach had been previously described as a subspecies of C. taenia Linnaeus, 1758, some recent genetic analysis refuted this designation (Šlechtová et al. 2008; Nakajima et al. 2011a). It has been confused what is the true C. striata described by Ikeda (1936) because the original description of this loach has ambiguous information on the type locality, and the type series is missing and feared lost (Saitoh &amp; Aizawa 1987). However, the author, Dr. Hyoji Ikeda, had stated that the type locality of C. striata is ‘near Takamatsu, Kagawa’ in his other literature (Okada &amp; Ikeda 1939; Aoyagi 1957) (Ikeda and Aoyagi are names of the same person). The results obtained in my field and literature surveys confirm that there is only one species, Cobitis sp. 3 subsp. 1 (sensu Nakajima et al. 2012), near this type locality. In addition, the description of C. striata is consistent with the morphological characters of Cobitis sp. 3 subsp. 1. Therefore, I conclude that C. striata Ikeda, 1936 is identical to Cobitis sp. 3 subsp. 1. The genetic features have been already reported by Kitagawa et al. (2005) and Saitoh et al. (2010).</p> <p>Japanese name. Chûgata-suji-shima-dojyô.</p></div> 	https://treatment.plazi.org/id/03CBD358FF96FFABF3D7574FFD44FECB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
03CBD358FF98FFADF3D750AFFD64FCE0.text	03CBD358FF98FFADF3D750AFFD64FCE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis striata subsp. fuchigamii Nakajima 2012	<div><p>Cobitis striata fuchigamii Nakajima, subsp. nov.</p> <p>(Figs. 3B, 4C, D, 5B, 6B)</p> <p>Onga form of Cobitis striata (middle race): Nakajima et al. 2008: 13, fig. 2F; Onga form of middle race of Cobitis striata complex: Kitagawa et al. 2009: 12, fig. 2D; Cobitis striata (the Onga form of the middle race): Nakajima et al. 2011b: 320, fig. 1C; Cobitis sp. 3 subsp. 2: Nakajima et al. 2012: 92, fig. 3b.</p> <p>Holotype. TKPM-P17341, male, 64.8 mm SL, Japan: Kakenouma River, Onga River system, Iizuka, Fukuoka Pref., Kyushu, 12. XII. 2010, J. Nakajima.</p> <p>Paratypes. MPM-FI1501, 1 male, 55.6 mm SL, same data as holotype; KPM-NI29503, 1 male, 57.2 mm SL, Kuro R., Onga R. s., Yahatanishi-ku, Fukuoka Pref., Kyushu, 9. VI. 2005, J. Nakajima; JNC042, 1 male, 56.0 mm SL, Hikosan R., Onga R. s., Oto, Fukuoka Pref., Kyushu, 24. VI. 2008, J. Nakajima; FKUN33734, 1 female, 63.4 mm SL, Chuganji R., Onga R. s., Kawasaki, Fukuoka Pref., Kyushu, 24. IV. 2004, J. Nakajima.</p> <p>Non-type specimens. 3 males, 47.4–59.5 mm SL, Kakenouma R., Onga R. s., Iizuka, Fukuoka Pref., Kyushu, 17. V. 2008, J. Nakajima; 2 females, 51.2, 61.1 mm SL, Kakenouma R., Onga R. s., Iizuka, Fukuoka Pref., Kyushu, 10. XII. 2003, J. Nakajima; 2 males and 2 females, 49.6–59.5 mm SL, Hikosan R., Onga R. s., Oto, Fukuoka Pref., Kyushu, 24. VI. 2008, J. Nakajima; 1 male and 1 female, 59.5, 79.5 mm SL, Kuro R., Onga R. s., Yahatanishi-ku, Fukuoka Pref., Kyushu, 16. V. 2005, J. Kawahara.</p> <p>Diagnosis. This subspecies is distinguishable from other Japanese striated spined loaches by the following characteristics: body size moderate, the mature size about 50–60 mm SL in males, 50–70 mm SL in females; lamina circularis at the base of the pectoral fin of adult male simple roundish plate, the upper segments of the first branched soft ray narrow and weak (Fig. 6B); PMN commonly 13; line L3 formed by sharp longitudinal line, reaching to caudal base; line L4 formed by narrow longitudinal line, reaching beyond dorsal fin, narrower than L 3 in male of non-spawning season; line L5 organized in 11–14 roundish, oblong or ovoid blotches in non-spawning season; caudal fin and dorsal fin with 3–4 arcuate bars; upper spot at the caudal base jet-black comparable in size to eye diameter; lower spot at caudal base faint or missing; egg yolk diameter approximately 1.0 mm; karyotype diploid.</p> <p>Description. Lateral view in Figure 3B illustrate body shape, form and position of fins. Morphometric and meristic data for 10 males and 5 females are summarized in Table 2. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7–8; pelvic-fin rays ii, 6; caudal-fin rays 8+8. Body elongate, laterally compressed. Head and snout elongated. Interorbital space narrow, convex. Caudal peduncle relatively compressed. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well-developed lobes; upper lip with transverse wrinkles on surface. Barbels, 3 pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 13 (range, 13–14). Very small cycloid scales on the trunk. Lamina circularis at the base of the pectoral fin of adult male simple roundish plate (Fig. 6B). The first branched soft ray of pectoral fin longer than the other rays; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin narrow and weak. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens: 64.8 mm SL male, 79.5 mm SL female.</p> <p>Coloration. Male in the non-spawning season (Figs. 3B, 4C). Body yellowish white with light brown pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Upper part of head covered with some amorphous spots, these spots often stringed. Opercle and snout covered with some large amorphous patterns. Body pigmentation organized in one middorsal and four lateral zones. Line L1 consisting of a series of 10–16 saddles or oval-shaped blotches. Line L2 formed by convex semicircular spots, only present on the middorsal body, reaching dorsally to interspaces of L1. Line L3 formed by sharp longitudinal line, reaching to caudal base. The posterior part of L3 often intermissive. Line L4 formed by narrow longitudinal line, reaching beyond dorsal fin, narrower than L3. Latter part of L4 often interrupted. Line L5 organized in 11–14 blotches from upper part of the pectoral fin to the caudal-fin base; blotches roundish, frequently oblong or ovoid. Caudal fin and dorsal fin with 3–4 arcuate bars. Anal fin pigmented along the fin rays. Upper spot at the caudal base jet-black comparable in size to eye diameter, lower spot at caudal base faint or missing.</p> <p>Male in the spawning season (Fig. 4D). Line L4 not visible or formed by faint longitudinal line, L3 and L5 well developed with broad stripes from the upper part of the pectoral-fin base to the caudal-fin base.</p> <p>Female (Fig. 5B). Appearance similar to males in the non-spawning season. But line L4 often formed by longitudinal jagged line, reaching anterior anal-fin base, broader or as wide as L3. Latter part of L4 often interrupted. Lines L3 and L5 tend to be developed with broad stripes in spawning season in large individual.</p> <p>Sexual dimorphism. Males have roundish lamina circularis at the base of the pectoral fin, but females do not. Generally the body size of the female is larger than that of the male. The L4 of the male tends narrower than that of the female.</p> <p>Egg diameter. 0.96 ± 0.09 mm (females, N = 4; collected from the Onga River system, Fukuoka Prefecture)</p> <p>Karyotype. Diploid. (Kitagawa et al. 2009)</p> <p>Distribution. Onga River system, northern Kyushu: Fukuoka Prefecture. (Nakajima et al. 2008)</p> <p>Habitat and biology. This species inhabits sandy-mud bottoms of the middle and lower reach of rivers. Although the spawning ecology is unknown, it is suggested that this subspecies needs a well-vegetated zone as a spawning site (Nakajima et al. 2011b).</p> <p>Etymology. The subspecific name is dedicated to Mr. Nobuyoshi Fuchigami, discoverer of this spined loach in the Onga River system.</p> <p>Remarks. The genetic features have been already reported by Kitagawa et al. (2009). This is a subspecies endemic to the Onga River system.</p> <p>Japanese name. Onga-suji-shima-dojyô.</p></div> 	https://treatment.plazi.org/id/03CBD358FF98FFADF3D750AFFD64FCE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
03CBD358FF9EFFACF3D75297FD5AFAA0.text	03CBD358FF9EFFACF3D75297FD5AFAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis striata subsp. hakataensis Nakajima 2012	<div><p>Cobitis striata hakataensis Nakajima, subsp. nov.</p> <p>(Figs. 3C, 4E, F, 5C, 6C)</p> <p>Hakata form of Cobitis striata (middle race): Nakajima et al. 2008: 13, fig. 2G; Hakata form of middle race of Cobitis striata complex: Kitagawa et al. 2009: 12, fig. 2E, F; Cobitis sp. 3 subsp. 3: Nakajima et al. 2012: 92, fig. 3c.</p> <p>Holotype. TKPM-P17342, male, 58.0 mm SL, Japan: Tatara River, Kasuya, Fukuoka Pref., Kyushu, 12. XII. 2010, J. Nakajima.</p> <p>Paratypes. JNC005, 1 male, 54.4 mm SL, same data as holotype; JNC041, 1 male, 60.4 mm SL, Tatara R., Kasuya, Fukuoka Pref., Kyushu, 23. V. 2005, J. Nakajima; KPM-NI29504, male, 55.0 mm SL, Tatara R., Kasuya, Fukuoka Pref., Kyushu, 18. V. 2008, J. Nakajima; MPM-FI1502, 1 male, 48.8 mm SL, same data; FKUN33756, 1 female, 87.4 mm SL, Naka R., Minami-ku, Fukuoka, Fukuoka Pref., Kyushu, 20. IV. 2005, J. Nakajima; JNC006, 1 male, 59.1 mm SL, Muromi R., Nishi-ku, Fukuoka, Fukuoka Pref., Kyushu, 13. V. 2010. E. Miyamura.</p> <p>Non-type specimens. 1 male and 2 females, 56.4–63.4 mm SL, same data as holotype; 1 male and 2 females, 64.0– 69.7 mm SL, Muromi R., Nishi-ku, Fukuoka, Fukuoka Pref., Kyushu, 5. VI. 2006, J. Nakajima; 2 males, 65.0, 65.7 mm SL, Tatara R., Kasuya, Fukuoka Pref., Kyushu, 23. V. 2005, J. Nakajima; 1 male and 1 female, 52.6, 55.5 mm SL, Tatara R., Kasuya, Fukuoka Pref., Kyushu, 18. V. 2008, J. Nakajima.</p> <p>Diagnosis. This subspecies is distinguishable from other Japanese striated spined loaches by the following characteristics: body size moderate, the mature size about 50–60 mm SL in males, 55–80 mm SL in females; lamina circularis at the base of the pectoral fin of adult male simple roundish plate, the upper segments of the first branched soft ray narrow and weak (Fig. 6C); PMN commonly 13; line L3 formed by incomplete longitudinal line, reaching to caudal base; line L4 formed by longitudinal jagged weblike line, reaching to postanal body, broader than L 3 in male of non-spawning season; line L5 organized in 11–14 roundish or ovoid blotches in non-spawning season; caudal fin and dorsal fin with 3–4 arcuate bars; upper spot at the caudal base jet-black comparable in size to eye diameter; lower spot at caudal base faint or missing; egg yolk diameter approximately 1.0mm; karyotype diploid.</p> <p>Description. Lateral view in Figure 3C illustrate body shape, form and position of fins. Morphometric and meristic data for 11 males and 5 females are summarized in Table 2. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7–8; pelvic-fin rays ii, 6; caudal-fin rays 8+8. Body elongate, laterally compressed. Head and snout elongated. Interorbital space narrow, convex. Caudal peduncle relatively compressed. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well-developed lobes; upper lip with transverse wrinkles on surface. Barbels, 3 pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 13 (range, 13–14). Very small cycloid scales on the trunk. Lamina circularis at the base of the pectoral fin of adult male simple roundish plate (Fig. 6C). The first branched soft ray of pectoral fin longer than the others; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin narrow and weak. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens: 65.7 mm SL male, 69.7 mm SL female.</p> <p>Coloration. Male in the non-spawning season (Figs. 3C, 4E). Body yellowish white with dark brown pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Upper part of head, opercle and snout covered with oval or amorphous shape spots. Body pigmentation organized in one middorsal and four lateral zones. Line L1 consisting of a series of 14–16, saddles or oval-shaped blotches, irregularly chained to each other. Line L2 formed by longitudinal jagged line, reaching to middorsal region, often fused with L1. Line L3 formed by incomplete longitudinal line, reaching to caudal base. Line L4 formed by longitudinal jagged weblike line, reaching to postanal body, broader than L3. Line L5 organized in 11–14 blotches from upper part of the pectoral fin to the caudal-fin base; blotches roundish or ovoid. Caudal fin and dorsal fin with 3–4 arcuate bars. Anal fin pigmented along the fin rays. Upper spot at the caudal base jet-black comparable in size to eye diameter, lower spot at the caudal base faint or missing.</p> <p>Male in the spawning season (Fig. 4F). Line L4 not visible or formed by faint longitudinal line, present only in anterior half of body. Lines L3 and L5 well developed with broad stripes from the upper part of the pectoral-fin base to the caudal-fin base.</p> <p>Female (Fig. 5C). Appearance similar to males in the non-spawning season, but number of blotches of line L5 tends to be more than in the male, line L5 of female organized in 11–17 blotches.</p> <p>Sexual dimorphism. Males have roundish lamina circularis at the base of the pectoral fin, but females do not. Generally, the body size of females is larger than that of males.</p> <p>Egg diameter. 0.98 ± 0.05 mm (females, N = 3; collected from the Tatara River system, Fukuoka Prefecture).</p> <p>Karyotype. Diploid (Kitagawa et al. 2009).</p> <p>Distribution. Rivers flowing into Hakata Bay, northern Kyushu: Fukuoka Prefecture (Nakajima et al. 2008).</p> <p>Habitat and biology. This species inhabits sandy-mud bottoms of the middle and lower reach of rivers. Life histories are unknown.</p> <p>Etymology. The subspecific name is derived from the popular common name of the Fukuoka City area in which the type locality is situated.</p> <p>Remarks. The genetic features have been reported by Kitagawa et al. (2009).</p> <p>Japanese name. Hakata-suji-shima-dojyô.</p></div> 	https://treatment.plazi.org/id/03CBD358FF9EFFACF3D75297FD5AFAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
03CBD358FF9FFFAEF3D754D7FD57FC3B.text	03CBD358FF9FFFAEF3D754D7FD57FC3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis kaibarai Nakajima 2012	<div><p>Cobitis kaibarai Nakajima, sp. nov.</p> <p>(Figs. 3D, 5D, 6D, 7A, B)</p> <p>Cobitis sp. S Kyushu-gata: Saitoh 1989: 389; Cobitis sp. 2 subsp. 3: Hosoya 2002: 275; Cobitis striata complex Small race, Kyushu form: Kitagawa et al. 2005: 112, table 1; Kyushu form of Cobitis striata (small race): Nakajima et al. 2008: 13, fig. 2D; Cobitis striata complex Kyushu form: Saitoh et al. 2010; 1003, table 1; Cobitis sp. 4: Nakajima et al. 2012: 92, fig. 3d.</p> <p>Holotype. TKPM-P17343, 1 male, 54.5 mm SL, Japan: Mitsuru River, Chikugo River system, Ukiha, Fukuoka Pref., Kyushu, 15. XII. 2010, J. Nakajima.</p> <p>Paratypes. MPM-FI1503, 1 male, 52.3 mm SL, creek of Kikuchi R. s., Tamana, Kumamoto Pref., Kyushu, 19. III. 2008, J. Nakajima; JNC015, 1 male, 47.2 mm SL, same data; JNC016, 1, male, 44.4 mm SL, same data; KPM- NI29505, 1 male, 48.3 mm SL, creek of Rokkaku R. s., Taku, Saga Pref., Kyushu, 22. XII. 2010, E. J. Kim; JNC018, 1 male, 46.9 mm SL, same data; KPM-NI9231, 1 female, 47.4 mm SL, Sakai R., Tamana, Kumamoto Pref., Kyushu, 24. XII. 1997, M. Watanabe.</p> <p>Non-type specimens. 2 males and 2 females, 49.4–56.1 mm SL, creek of Kikuchi R. s., Tamana, Kumamoto Pref., Kyushu, 4. VI. 2007, J. Nakajima; 2 males, 51.9, 52.3 mm SL, Kose R., Chikugo R. s., Ukiha, Fukuoka Pref., Kyushu, 14. VIII. 2007, J. Nakajima; 1 female, 63.9 mm SL, same data; 1 male and 1 female, 47.9, 68.1 mm SL, creek of Kase R. s., Saga, Saga Pref., Kyushu, 16. V. 2008, J. Nakajima; 1 male and 1 female, 50.1, 65.8 mm SL, creek of Rokkaku R. s., Taku, Saga Pref., 23. XI. 2010, Y. Suzawa; 1 female, 56.7 mm SL, creek of Rokkaku R. s., Taku, Saga Pref., Kyushu, 22. XII. 2010, E. J. Kim.</p> <p>Diagnosis. This species is distinguishable from other Japanese striated spined loaches by the following characteristics: body size relatively small, the mature size about 45–55 mm SL in males, 55–65 mm SL in females; lamina circularis at the base of the pectoral fin of adult male simple roundish plate, the upper segments of the first branched soft ray narrow and weak (Fig. 6D); PMN commonly 13; line L3 formed by incomplete longitudinal line, reaching to caudal base, fused with L1 and L4 on posterior part of body; line L4 formed by longitudinal jagged line, reaching beyond dorsal fin in male of non-spawning season; line L5 organized in 10–16 roundish or ovoid blotches, fused with L4 on caudal body in non-spawning season; caudal fin and dorsal fin with 4–5 arcuate bars; upper spot at the caudal base jet-black approximately eye diameter; lower spot at caudal base faint or missing; egg yolk diameter approximately 0.8 mm; karyotype diploid (2n = 50).</p> <p>Description. Lateral view in Figure 3D illustrate body shape, form and position of fins. Morphometric and meristic data for 12 males and 6 females are summarized in Table 2. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7; pelvic-fin rays ii, 5–6; caudal-fin rays 8+8. Body elongate, laterally compressed. Head and snout elongated. Interorbital space narrow, convex. Caudal peduncle relatively compressed. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well-developed lobes; upper lip with transverse wrinkles on surface. Barbels, 3 pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 13 (range, 13–14). Very small cycloid scales on the trunk. Lamina circularis at the base of the pectoral fin of adult male simple roundish plate (Fig. 6D). The first branched soft ray of pectoral fin longer than the other rays; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin narrow and weak. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens: 54.5 mm SL male, 68.1 mm SL female.</p> <p>Coloration. Male in the non-spawning season (Figs. 3A, 7A). Body yellowish white with brown pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Upper part of head, opercle and snout covered with oval or amorphous shaped spots. Body pigmentation organized in one middorsal and four lateral zones. Line L1 consisting of a series of 12–25 blotches; blotches saddle or oval-shaped. Line L2 formed by longitudinal jagged line or small angular blotches, reaching to postdorsal region, often fused with L1. Line L3 formed by incomplete longitudinal line, reaching to caudal base, fused with L1 and L 4 in postdorsal region. Line L4 formed by longitudinal jagged line, reaching beyond dosal fin, width variable. Line L5 organized in 10–16 blotches from upper part of the pectoral fin to the caudal-fin base; blotches roundish or ovoid, fused with L4 on caudal body. Caudal fin and dorsal fin with 4–5 arcuate bars. Anal fin pigmented along the fin rays. Upper spot at the caudal base jet-black comparable in size to eye diameter, lower spot at caudal base faint or missing.</p> <p>Male in the spawning season (Fig. 7B). Line L4 not visible or formed by narrow longitudinal line, present only in anterior half of body. Lines L3 and L5 well developed with broad stripes from upper part of the pectoral-fin base to the caudal-fin base.</p> <p>Female (Fig. 5D). Appearance similar to males in the non-spawning season.</p> <p>Sexual dimorphism. Males have roundish lamina circularis at the base of the pectoral fin, but females do not. Generally, the body size of females is larger than that of males.</p> <p>Egg diameter. 0.83 ± 0.04 mm (females, N = 2; collected from the Kikuchi River system, Kumamoto Prefecture).</p> <p>Karyotype. Diploid (2n = 50) (Ueno &amp; Ojima 1976; Ueno et al. 1980; Ueno 1981; Saitoh et al. 2000).</p> <p>Distribution. Rivers flowing into Ariake estuary, northern Kyushu: Saga, Fukuoka, and Kumamoto Prefectures (Nakajima et al. 2008).</p> <p>Habitat and biology. This species inhabits sandy-mud bottoms of the middle and lower reaches of rivers and small streams. Life histories are unknown.</p> <p>Etymology. The specific name is dedicated to the late Mr. Ekiken Kaibara who was the first real naturalist and biologist in Japan. He recorded the distribution of spined loaches from Chikushi (modern-day Fukuoka Prefecture), Kyushu Island for the first time (Kaibara 1709).</p> <p>Remarks. The genetic features have been reported by Kitagawa et al. (2005) and Saitoh et al. (2010).</p> <p>Japanese name. Ariake-suji-shima-dojyô.</p></div> 	https://treatment.plazi.org/id/03CBD358FF9FFFAEF3D754D7FD57FC3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
03CBD358FF9DFFB1F3D75524FD6EF940.text	03CBD358FF9DFFB1F3D75524FD6EF940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis magnostriata Nakajima 2012	<div><p>Cobitis magnostriata Nakajima, sp. nov.</p> <p>(Figs. 3E, 5E, 6E, 7C, D)</p> <p>Cobitis taenia striata: Okada and Nakamura 1948:185, fig. 113; Cobitis taenia striata large race: Minamori 1956: 91, fig. 1;? Cobitis taenia striata: Aoyagi 1957: 169, fig. 142; Cobitis taenia striata: Okada 1960: 562, figs. 93a, 93b; Cobitis taenia striata: Nakamura 1963: 161, figs. 97a, b; Cobitis taenia f. striata: Miyadi et al. 1976: 247, pl. 31; Cobitis taenia striata large race: Saitoh and Aizawa 1987: 336, fig. 3A; Cobitis sp. L: Saitoh 1989: 386; Cobitis sp. L: Saitoh and Matsuda 1990: 19, fig. 1; Cobitis sp. 1: Hosoya 2002: 276; Cobitis striata complex large race: Kitagawa et al. 2005: 112, table 1; Cobitis striata complex large race: Saitoh et al. 2010: 1003, table 1; Cobitis sp. 1: Nakajima et al. 2012: 90, fig. 2a.</p> <p>Holotype. TKPM-P17344, male, 62.4 mm SL, Japan: small stream flowing into Biwa Lake, Adogawa, Shiga Pref., Honshu, 6. V. 2007, K. Tominaga.</p> <p>Paratypes. FRLM24920, 1 male, 63.3 mm SL, Chinai R., Takashima, Shiga Pref., Honshu, 16. VIII. 1997, Y. Fujioka; KPM-NI29506, 1 male, 64.3 mm SL, same data as holotype; MPM-FI1504, 1 male, 63.8 mm SL, same data as holotype; JNC049, 1 female, 93.3 mm SL, same data as holotype; JNC040, 1 male, 72.5 mm SL, small stream flowing into Biwa Lake, Takashima, Shiga Pref., Honshu, 3. VII. 2007, J. Nakajima.</p> <p>Non-type specimens. 5 males and 3 females, 60.4–84.0 mm SL, same data as holotype; 1 male and 2 females, 55.4–60.4 mm SL, small river of Yodo R.s., Otsu, Shiga Pref., Honshu, 22. VII. 2008, K. Tominaga.</p> <p>Diagnosis. This species is distinguishable from other Japanese striated spined loaches by the following characteristics: body size large, mature size about 60–70 mm SL in males, 70–90 mm SL in females; lamina circularis at the base of the pectoral fin of adult male simple roundish plate, the upper segments of the first branched soft ray broad and strong (Fig. 6E); PMN commonly 14; caudal peduncle relatively deep; line L1 organized in longitudinal line from snout tip to dorsal-fin base, and formed by some oval- or saddle-shaped blotches on postdorsal body; line L2 lacking commonly; line L3 formed by sharp longitudinal line, reaching beyond dorsal fin; line L4 lacking or formed by weak dotted line; line L5 well developed with broad stripe; caudal and dorsal fins margined by a broad black band; upper and lower spot at caudal base connected into dumbell-shape; egg yolk diameter approximately 1.1 mm; karyotype tetraploid (2n = 98).</p> <p>Description. Lateral view in Figure 3E illustrate body shape, form and position of fins. Morphometric and meristic data for 10 males and 6 females are summarized in Table 2. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7–8; pelvic-fin rays ii, 6; caudal-fin rays 8+8. Body elongate, laterally compressed. Head and snout elongated. Interorbital space narrow, convex. Eye diameter slightly small. Caudal peduncle relatively deep. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well- developed lobes; upper lip with transverse wrinkles on surface. Barbels, 3 pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 14 (range, 13–14). Very small cycloid scales on the trunk. Lamina circularis at the base of the pectoral fin of adult male simple roundish plate (Fig. 6E). The first branched soft ray of pectoral fin longer than the other rays; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin broad and strong. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens: 72.5 mm SL male, 93.3 mm SL female.</p> <p>Coloration. Male in the non-spawning season (Figs. 3E, 7C). Body yellowish white with black pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Opercle and snout covered with some large amorphous patterns. Line L1 organized in longitudinal line from the snout tip to the dorsal-fin base, and formed by some oval- or saddle-shaped blotches on postdorsal body; uncommonly formed by 4–6 predorsal and 4–6 postdorsal blotches. Line L2 lacking; uncommonly formed by tiny oval blotches, reaching to subdorsal body. Line L3 formed by sharp longitudinal line, reaching beyond dorsal fin. Line L4 lacking or formed by weak dotted line, present only in anterior half of body. Line L5 well developed with broad stripe from upper part of the pectoral fin to the caudal-fin base. Caudal fin and dorsal fin margined by a broad black band; various cloudy blotches or broken marks on other regions. Anal fin pigmented along the fin rays. Upper and lower spot at caudal base connected into dumbell-shape; upper spot jet-black, larger than eye diameter.</p> <p>Male in the spawning season (Fig. 7D). Line L3 well developed with broad stripe from upper part of the pectoralfin base to the posterior part of the dorsal fin. Line L5 well developed with broad stripe from upper part of the pectoral-fin base to the caudal-fin base. Lines L2 and L4 lacking.</p> <p>Female (Fig. 5E). Appearance similar to males in the non-spawning season.</p> <p>Sexual dimorphism. Males have roundish lamina circularis at the base of the pectoral fins, but females do not. Generally, the body size of females is larger than that of males.</p> <p>Egg diameter. 1.16 ± 0.05 mm (female, N = 1; collected from the Biwa Lake, Shiga Prefecture).</p> <p>Karyotype. Tetraploid (2n = 98) (Ueno &amp; Ojima 1976; Ueno et al. 1980; Ueno 1981; Saitoh et al. 1984, 2000; Kimizuka 1987).</p> <p>Distribution. Biwa Lake and tributary rivers, central Honshu: Shiga Prefecture (Saitoh &amp; Aizawa 1987).</p> <p>Habitat and biology. This species inhabits sandy-mud bottoms 1–3 meters below the surface of the lake, and migrates to small streams in the spawning season (Saitoh &amp; Matsuda 1990).</p> <p>Etymology. This species has the largest body and the most awesome appearance in the Japanese Cobitis striata complex; this is reflected by the specific epithet magno-, which in Latin means ‘large’ or ‘great’.</p> <p>Remarks. Although this is an endemic species of Biwa Lake and tributary rivers, it has been introduced in some rivers in Honshu Island (Kitahara 2007; Matsuzawa &amp; Senou 2008). The genetic features have been reported by Kitagawa et al. (2005) and Saitoh et al. (2010).</p> <p>Japanese name. Ôgata-suji-shima-dojyô.</p></div> 	https://treatment.plazi.org/id/03CBD358FF9DFFB1F3D75524FD6EF940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
03CBD358FF82FFB3F3D757F7FD01FDEB.text	03CBD358FF82FFB3F3D757F7FD01FDEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis minamorii Nakajima 2012	<div><p>Cobitis minamorii Nakajima, sp. nov.</p> <p>(Figs. 3F, 5F, 6F, 8A, B)</p> <p>Cobitis taenia striata small race: Minamori 1952: 201, fig. 2B; Cobitis taenia striata small race: Saitoh and Aizawa 1987: 336, fig. 3D; Cobitis sp. S (Sanyo-gata): Saitoh 1989: 388; small form: Saitoh 1990: 240, figs. 3a, b, c, d, e: 241, fig. 4 (upper four); Cobitis sp. 2 subsp. 1: Hosoya 2002: 275; Cobitis striata complex small race, Sanyo form: Kitagawa et al. 2005: 112, table 1; Cobitis striata complex small race: Saitoh et al. 2010: 1003, table 1; Cobitis sp. 2 subsp. 1: Nakajima et al. 2012: 90, fig. 2b.</p> <p>Holotype. TKPM-P17345, 1 male, 48.9 mm SL, Japan: Kagato River, Yoshii River system, Setouchi, Okayama Pref., Honshu, 6. VI. 2007, J. Nakajima.</p> <p>Paratypes. KPM-NI29507, 1 male, 42.0 mm SL, same data as holotype; MPM-FI1505, 1 male, 37.2 mm SL, same data as holotype; KPM-NI9225, female, 33.4 mm SL, Asahi R., Okayama, Okayama Pref., Honshu, 31. VII. 1997, T. Okazaki and M. Watanabe.</p> <p>Non-type specimens. FAKU055761, 3 males, 38.1–46.4 mm SL, Imizo R., Ashida R. s., Fukuyama, Hiroshima Pref., Honshu, 2. V. 1982, K. Saitoh; FAKU055675, 3 males, 40.1–47.6 mm SL, Omachi R., Asahi R. s., Okayama, Okayama Pref., Honshu, 23. IV. 1981, K. Saitoh; 2 males and 7 females, 40.0– 55.5 mm SL, same data as holotype.</p> <p>Diagnosis. This species is distinguishable from other Japanese striated spined loaches by the following characteristics: body size small, the mature size about 35–45 mm SL in males, 45–55 mm SL in females; lamina circularis at the base of the pectoral fin of adult male a roundish plate, somewhat narrowing toward the outer part; upper segments of the first branched soft ray of pectoral fin narrow and weak (Fig. 6F); PMN commonly 12; snout short; eye diameter relatively large; body depth and caudal peduncle relatively deep; line L1 consisting of a series of 15–25 blotches; blotches saddle or oval-shaped; line L2 formed by longitudinal jagged line or angular blotches, reaching to pre- or middorsal region, often fused with L1; line L3 formed by incomplete longitudinal narrow line, reaching beyond dorsal fin; line L5 formed by narrow incomplete longitudinal line or organized in chained tiny 15–20 blotches; caudal and dorsal fin with 3–4 irregular vertical bars and exterior bar weak margined; upper spot at the caudal base black, smaller than eye diameter; lower spot at caudal base existing but faint; each spots not connected; egg yolk diameter approximately 0.8 mm; karyotype diploid (2n = 49 in male, 50 in female).</p> <p>Description. Lateral view in Figure 3F illustrate body shape, form and position of fins. Morphometric and meristic data for 11 males and 7 females are summarized in Table 2. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7–8; pelvic-fin rays ii, 6; caudal-fin rays 8+8. Body some what waistless, laterally compressed. Snout short comparatively. Interorbital space broad, convex. Eye diameter relatively large. Body depth and caudal peduncle relatively deep. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well-developed lobes; upper lip with transverse wrinkles on surface. Barbels, 3 pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 12 (range, 11–14). Very small cycloid scales on the trunk. Lamina circularis at the base of the pectoral fin of adult male roundish plate, somewhat narrowing toward the outer part (Fig. 6F). The first branched soft ray of pectoral fin longer than the other rays; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin narrow and weak. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens: 48.9 mm SL male, 55.5 mm SL female.</p> <p>Coloration. Male in the non-spawning season (Figs. 3F, 8A). Body pearl white with dark brown pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Upper part of head covered with 3–5 roundish. Opercle and snout covered with oval and amorphous shaped spots. Body pigmentation organized in one middorsal and four lateral zones. Line L1 consisting of a series of 15–25 blotches; blotches saddle or oval-shaped. Line L2 formed by longitudinal jagged line or angular blotches, reaching to pre- or middorsal region, often fused with L1. Line L3 formed by incomplete longitudinal narrow line, reaching to beyond dorsal fin, fused with L1 and L4 on posterior part. Line L4 formed by tiny blotches or narrow longitudinal line, reaching to dorsal fin. Line L5 formed by narrow incomplete longitudinal line or organized in chained tiny 15–20 blotches from upper part of the pectoral fin to the caudal-fin base. Caudal fin and dorsal fin with 3–4 irregular vertical bars and exterior bar often margined. Anal fin pigmented along the fin rays. Upper spot at the caudal base black, smaller than eye diameter, lower spot at caudal base faint; spots not connected.</p> <p>Male in the spawning season (Fig. 8B). Line L4 not visible or formed by narrow longitudinal line, present only in anterior half of body. Lines L3 and L5 well developed with narrow stripes from upper part of the pectoral-fin base to the caudal-fin base; the posterior part often intermissive.</p> <p>Female (Fig. 5F). Appearance similar to males in the non-spawning season.</p> <p>Sexual dimorphism. Males have roundish lamina circularis at the base of the pectoral fin, but females do not. Generally, the body size of females is larger than that of males.</p> <p>Egg diameter. 0.85 ± 0.04 mm (females, N = 3; collected from the Yoshii River system, Okayama Prefecture).</p> <p>Karyotype. Diploid (2n = 49 in male, 2n = 50 in female) (Saitoh et al. 1984, 2000).</p> <p>Distribution. Rivers flowing into Seto Inland Sea, Sanyo district, western Honshu: Okayama and Hiroshima Prefectures (Saitoh &amp; Aizawa 1987).</p> <p>Habitat and biology. This species inhabits sandy-mud bottoms of the lower reach of rivers and small stream. Saitoh (1990) described the spawning ecology of this species as small form of C. striata.</p> <p>Etymology. The specific name is dedicated to Dr. Sumio Minamori who was the pioneer of the study of speciation of loaches in Japan.</p> <p>Remarks. The genetic features have been reported by Kitagawa et al. (2005) and Saitoh et al. (2010).</p> <p>Japanese name. Sanyô-kogata-suji-shima-dojyô.</p></div> 	https://treatment.plazi.org/id/03CBD358FF82FFB3F3D757F7FD01FDEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
03CBD358FF80FFB5F3D757E0FD1AFEB3.text	03CBD358FF80FFB5F3D757E0FD1AFEB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis minamorii subsp. oumiensis Nakajima 2012	<div><p>Cobitis minamorii oumiensis Nakajima, subsp. nov.</p> <p>(Figs. 3G, 5G, 6G, 8C, D)</p> <p>Cobitis taenia striata Biwa-small race: Minamori 1956: 91, fig. 1; Cobitis taenia striata Biwa small race: Saitoh and Aizawa 1987: 336, fig. 3B; Cobitis sp. S (Biwako-gata): Saitoh 1989: 387; Cobitis sp. S: Saitoh and Matsuda 1990: 19, fig. 1; Cobitis sp. 2 subsp. 4: Hosoya 2002: 276; Cobitis striata complex small race, Biwa form: Kitagawa et al. 2005: 112, table 1; Cobitis striata complex Biwa small race: Saitoh et al. 2010: 1003, table 1; Cobitis sp. 2 subsp. 4: Nakajima et al. 2012: 90, fig. 2e.</p> <p>Holotype. TKPM-P17346, 1 male, 51.7 mm SL, Japan: small stream flowing into Biwa Lake, Takashima, Shiga Pref., Honshu, 24. V. 2007, S. Yodo.</p> <p>Paratypes. JNC037, 1 male, 48.2 mm SL, small stream flowing into Biwa Lake, Takashima, Shiga Pref., Honshu, 15. V. 2009, J. Nakajima; KPM-NI29508, 1 male, 48.9 mm SL, same data as holotype; MPM-FI1506, 1 male, 48.6 mm SL, same data as holotype; KPM-NI8989, 1 female, 69.3 mm SL, Imazu, Takashima, Shiga Pref., Honshu, 19. V. 2000, T. Satou.</p> <p>Non-type specimens. FAKU55876, 5 males, 45.2–53.2 mm SL, Takashima, Shiga Pref., Honshu, 6. VI. 1983, K. Saitoh; 2 males and 3 females, 44.4–68.7 mm SL, same data as holotype; 2 females, 63.4, 74,5 mm SL, small stream flowing into Biwa Lake, Takashima, Shiga Pref., Honshu, 15. V. 2009, J. Nakajima.</p> <p>Diagnosis. This subspecies is distinguishable from other Japanese striated spined loaches by the following characteristics: body size small, the mature size about 45–55 mm SL in males, 50–60 mm SL in females; lamina circularis at the base of the pectoral fin of adult male roundish plate, somewhat narrowing toward the outer part; upper segments of the first branched soft ray of pectoral fin narrow and weak (Fig. 6G); PMN commonly 12; snout short; line L1 consisting of a series of 14–16 blotches; blotches oval-shaped, often chainlike longitudinal jagged line; line L2 lacking; line L3 formed by sharp longitudinal line, reaching beyond dorsal fin; line L4 lacking or formed by weak dotted line; line L5 well developed with broad stripe in all season; caudal and dorsal fins margined by a broad black band; Upper and lower spot at caudal base black, upper sharpen, lower somewhat pale; spots slightly connected; egg yolk diameter approximately 0.8 mm; karyotype diploid (2n = 50).</p> <p>Description. Lateral view in Figure 3G illustrate body shape, form and position of fins. Morphometric and meristic data for 11 males and 5 females are summarized in Table 2. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7; pelvic-fin rays ii, 5–6; caudal-fin rays 8+8. Body elongate, laterally compressed. Snout short comparatively. Interorbital space narrow, convex. Caudal peduncle relatively compressed. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well-developed lobes; upper lip with transverse wrinkles on surface. Barbels, 3 pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 12 (range, 11–13). Very small cycloid scales on the trunk. Lamina circularis at the base of the pectoral fin of adult male roundish plate, somewhat narrowing toward the outer part (Fig. 6G). The first branched soft ray of pectoral fin longer than the other rays; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin narrow and weak. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens: 53.2 mm SL male, 74.5 mm SL female.</p> <p>Coloration. Male in the non-spawning season (Figs. 3G, 8C). Body pearl white with black pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Head, opercle and snout covered with some large amorphous patterns. Line L1 consisting of a series of 14–16 blotches; blotches ovalshaped, often chainlike longitudinal jagged line. Line L2 lacking. Line L3 formed by sharp longitudinal line, reaching beyond dorsal fin. Line L4 lacking or formed by weak dotted line, present only in anterior half of body. Line L5 well developed with broad stripe from upper part of the pectoral fin to the caudal-fin base. Caudal fin and dorsal fin margined by a broad black band; various cloudy blotches or broken marks on other regions. Anal fin pigmented along the fin rays. Upper and lower spot at caudal base black, upper sharpen, lower somewhat pale; spots slightly connected.</p> <p>Male in the spawning season (Fig. 8D). Line L3 well developed with broad stripe from upper part of the pectoral-fin base to the posterior part of the dorsal fin. Line L5 well developed with broad stripe from upper part of the pectoral-fin base to the caudal-fin base. Lines L2 and L4 lacking.</p> <p>Female (Fig. 5G). Appearance similar to males in the non-spawning season.</p> <p>Sexual dimorphism. Males have roundish lamina circularis at the base of the pectoral fin, but females do not. Generally, the body size of females is larger than that of males.</p> <p>Egg diameter. 0.84 ± 0.03 mm (females, N = 2; collected from small stream flowing into the Biwa Lake, Shiga Prefecure).</p> <p>Karyotype. Diploid (2n = 50) (Ueno &amp; Ojima 1976; Ueno et al. 1980; Ueno 1981; Kimizuka 1987; Saitoh et al. 2000).</p> <p>Distribution. Lake Biwa, central Honshu: Shiga Prefecture (Saitoh &amp; Aizawa 1987).</p> <p>Habitat and biology. This subspecies inhabits sandy-mud bottoms of small streams, and spawns in paddy field (Saitoh &amp; Matsuda 1990).</p> <p>Etymology. The subspecific name is derived from Oumi, the old name of Lake Biwa, the type locality.</p> <p>Remarks. The genetic features have been reported by Kitagawa et al. (2005) and Saitoh et al. (2010).</p> <p>Japanese name. Biwa-kogata-suji-shima-dojyô.</p></div> 	https://treatment.plazi.org/id/03CBD358FF80FFB5F3D757E0FD1AFEB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
03CBD358FF86FFB4F3D750ACFD1EFBAB.text	03CBD358FF86FFB4F3D750ACFD1EFBAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis minamorii subsp. tokaiensis Nakajima 2012	<div><p>Cobitis minamorii tokaiensis Nakajima, subsp. nov.</p> <p>(Figs. 3H, 5H, 6H, 8E, F)</p> <p>Cobitis taenia: Aizawa 1981: 188, fig. 2A, B, C; Cobitis taenia striata Tokai small race: Saitoh and Aizawa 1987: 336, fig. 3E; Cobitis sp. S (Tokai-gata): Saitoh 1989: 389; Cobitis sp. 2 subsp. 2: Hosoya 2002: 275; Cobitis striata complex small race, Tokai form: Kitagawa et al. 2005: 112, table 1; Cobitis striata complex Tokai small race: Saitoh et al. 2010: 1003, table 1; Cobitis sp. 2 subsp. 2: Nakajima et al. 2012: 90, fig. 2c.</p> <p>Holotype. MPM-FI1507, 1 male, 47.9 mm SL, Japan: creek of Kumozu River system, Tsu, Mie Pref., Honshu, 24. IV. 2011, J. Kitamura.</p> <p>Paratypes. KPM-NI29509, 1 male, 43.2 mm SL, creek of Ibi R. s., Tado, Mie Pref., Honshu, 21. X. 2007, J. Nakajima; MPM-FI1508, 1 male, 39.3 mm SL, creek of Kushida R. s., Matsusaka, Mie Pref., Honshu, 11. IV. 2011, J. Kitamura; TKPM-P17347, 1 male, 39.0 mm SL, creek of Iride-ota R. s., Kosai, Shizuoka Pref., Honshu, 9. VI. 2008, J. Nakajima; FRLM24921, 1 male, 52.4 mm SL, Ohtani R., Nagara R. s., Gifu, Gifu Pref., Honshu, 19. VII. 1997, T. Okazaki.</p> <p>Non-type specimens. 3 males and 4 females, 42.5–55.5 mm SL, creek of Ibi R. s., Tado, Mie Pref., Honshu, 21. X. 2007, J. Nakajima; 2 males, 41.2, 42.3 mm SL, Harai R., Kushida R. s., Matsusaka, Mie Pref., Honshu, 7. III. 2008, K. Tominaga; 1 male and 1 female, 37.5, 50.3 mm SL, creek of Iride-ota R. s., Kosai, Shizuoka Pref., Honshu, 9. VI. 2008, J. Nakajima; 2 males and 1 female, 34.1–41.6 mm SL, Shonai R., Kasugai, Aichi Pref., Honshu, 2. VI. 2006, T. Oonaka.</p> <p>Diagnosis. This subspecies is distinguishable from other Japanese striated spined loaches by the following characteristics: body size small, the mature size about 35–45 mm SL in males, 40–50 mm SL in females; lamina circularis at the base of the pectoral fin of adult male roundish plate, somewhat narrowing toward the outer part; upper segments of the first branched soft ray of pectoral fin narrow and weak (Fig. 6H); PMN commonly 12; snout short; line L1 consisting of a series of 15–25 blotches; line L2 formed by sparse angular blotches, reaching to pre- or middorsal region, often fused with L1; line L3 formed by a longitudinal line, reaching to caudal base, fused with L1 and L4 on postdorsal body; line L5 organized in 10–16 blotches in non-spawning season; caudal and dorsal fin with thin 3–4 arcuate bars; upper spot at the caudal base black, smaller than eye diameter; lower spot at caudal base faint; spots not connected; egg yolk diameter approximately 0.8 mm; karyotype diploid (2n = 50).</p> <p>Description. Lateral view in Figure 3H illustrate body shape, form and position of fins. Morphometric and meristic data for 12 males and 6 females are summarized in Table 2. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7–8; pelvic-fin rays ii, 5–6; caudal-fin rays 8+8. Body somewhat waistless, laterally compressed. Snout short comparatively. Interorbital space broad, convex. Eye diameter relatively large. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well-developed lobes; upper lip with transverse wrinkles on surface. Barbels, 3 pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 12 (range, 11–13). Very small cycloid scales on the trunk. Lamina circularis at the base of pectoral fin of adult male roundish plate, somewhat narrowing toward the outer part (Fig. 6H). The first branched soft ray of pectoral fin longer than the other rays; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin narrow and weak. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens: 47.9 mm SL in male, 55.5 mm SL in female.</p> <p>Coloration. Male in the non-spawning season (Figs. 3H, 8E). Body pearl white with dark brown pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Upper part of head covered with some roundish blotches or by longitudinal line. Opercle and snout covered with oval and amorphous shaped spots. Body pigmentation organized in one middorsal and four lateral zones. Line L1 consisting of a series of 15–25 blotches; blotches saddle or oval-shaped. Line L2 formed by sparse angular blotches, reaching to pre- or middorsal region, often fused with L1. Line L3 formed by longitudinal line, reaching to caudal base, fused with L1 and L4 on postdorsal body. Line L4 formed by tiny blotches or narrow incomplete longitudinal line. Line L5 organized in 10–16 blotches from upper part of the pectoral fin to caudal-fin base; blotches roundish or ovoid. Caudal fin and dorsal fin with thin 3–4 arcuate bars. Anal fin pigmented along the fin rays. Upper spot at the caudal base black, smaller than eye diameter; lower spot at caudal base faint; spots not connected.</p> <p>Male in the spawning season (Fig. 8F). Lines L2 and L4 not visible. Line L3 well developed with broad stripe from upper part of the pectoral-fin base to posterior part of body, fused with L1 on postdorsal body. Line L5 well developed with broad stripe from upper part of the pectoral-fin base to the caudal-fin base.</p> <p>Female (Fig. 5H). Appearance similar to males in the non-spawning season, but zone L4 tends to be better developed than in males.</p> <p>Sexual dimorphism. Males have roundish lamina circularis at the base of the pectoral fin, but females do not. Generally, the body size of females is larger than that of males.</p> <p>Egg diameter. 0.84 ± 0.04 mm (female, N = 1; collected from the Iride-ota River system, Shizuoka Prefecture).</p> <p>Karyotype. Diploid (2n = 50) (Ueno et al. 1980; Kimizuka 1987; Saitoh et al. 2000).</p> <p>Distribution. Rivers flowing into Ise Bay, Mikawa Bay, and western part of Enshu-nada coast, Tokai District, central Honshu: Shizuoka, Gifu, Aichi, and Mie Prefectures (Saitoh &amp; Aizawa 1987).</p> <p>Habitat and biology. This subspecies inhabits sandy-mud bottoms of the lower reaches of rivers and small streams. Life histories are unknown.</p> <p>Etymology. The subspecific name is derived from the Tokai District of Central Honshu, which is the main distribution area of this subspecies.</p> <p>Remarks. The genetic features have been reported by Kitagawa et al. (2005) and Saitoh et al. (2010).</p> <p>Japanese name. Tokai-kogata-suji-shima-dojyô.</p></div> 	https://treatment.plazi.org/id/03CBD358FF86FFB4F3D750ACFD1EFBAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
03CBD358FF87FFB7F3D755D4FD04F8B8.text	03CBD358FF87FFB7F3D755D4FD04F8B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis minamorii subsp. saninensis Nakajima 2012	<div><p>Cobitis minamorii saninensis Nakajima, subsp. nov.</p> <p>(Figs. 3I, 5I, 6I, 8G, H)</p> <p>Cobitis taenia striata spotted-small race: Minamori 1956: 91, fig. 1; Cobitis taenia striata spotted small race: Saitoh and Aizawa 1987: 336, fig. 3F; Cobitis sp. S (San-in-gata): Saitoh 1989: 389; Cobitis sp. 2 subsp. 3: Hosoya 2002: 275; Cobitis striata complex small race, San-in form: Kitagawa et al. 2005: 112, table 1; Cobitis striata complex spotted small race: Saitoh et al. 2010: 1003, table 1; Cobitis sp. 2 subsp. 3: Nakajima et al. 2012: 90, fig. 2d.</p> <p>Holotype. TKPM-P17348, 1 male, 62.1 mm SL, Japan: creek of Hii River system, Izumo, Shimane Pref., Honshu, 10. IV. 2011, J. Nakajima.</p> <p>Paratypes. KPM-NI29510, 1 male, 53.8 mm SL, Hiwa R., Hokuei, Tottori Pref., Honshu, 9. IV. 2011, J. Nakajima; MPM-FI1509, 1 male, 43.9 mm SL, same data; JNC034, 1 male, 47.6 mm SL, same data; JNC036, 1 male, 48.4 mm SL, Terauchi R., Hino R. s., Nanbu, Tottori Pref., Honshu, 10. IV. 2011, J. Nakajima; KPM-NI9218, 1 male, 59.3 mm SL, Tai R., Kishida R. s., Shin-onsen, Hyogo Pref., Honshu, 17. VIII. 1998, T. Kitagawa.</p> <p>Non-type specimens. FAKU55788, 3 males, 44.1–52.1 mm SL, Shimizu R., Sendai R. s., Tottori, Tottori Pref., Honshu, 31. V. 1981, K. Saitoh; FAKU055797, 2 females, 50.9, 73.0 mm SL, Kouchi R., Kedaka, Tottori Pref., Honshu, 16. X. 1983, K. Saitoh; 3 males and 2 females, 49.0– 52.1 mm SL, Tai R., Kishida R. s., Shin-onsen, Hyogo Pref., Honshu, 23. V. 2008, K. Tominaga; 3 males, 50.4–54.0 mm SL, creek of Hii R. s., Izumo, Shimane Pref., Honshu, 31. V. 2008, J. Nakajima; 2 females, 50.9, 73.0 mm SL, Hiwa R., Hokuei, Tottori Pref., Honshu, 9. IV. 2011, J. Nakajima.</p> <p>Diagnosis. This subspecies is distinguishable from other Japanese striated spined loaches by the following characteristics: body size relatively small, the mature size about 40–60 mm SL in males, 50–70 mm SL in females; lamina circularis at the base of the pectoral fin of adult male roundish plate, somewhat narrowing toward the outer part; upper segments of the first branched soft ray of pectoral fin narrow and weak (Fig. 6I); PMN commonly 12; snout elongated; line L1 consisting of a series of 15–20 blotches; line L2 formed by longitudinal jagged line, reaching to postdorsal region, fused with L1; line L3 formed by incomplete longitudinal line, reaching to caudal base, fused with L1 and L4 on postdorsal body in non-spawning season; line L5 organized in 11–13 blotches in non-spawning season; caudal and dorsal fins with thin 4–6 arcuate bars; upper spot at the caudal base black, approximately eye diameter; lower spot at caudal base faint; spots not connected; egg yolk diameter approximately 0.8 mm; karyotype diploid (2n = 50).</p> <p>Description. Lateral view in Figure 3I illustrates body shape, form and position of fins. Morphometric and meristic data for 14 males and 6 females are summarized in Table 2. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7–9; pelvic-fin rays ii, 6; caudal-fin rays 8+8. Body elongate, laterally compressed. Head and snout elongated. Interorbital space broad, convex. Snout elongated. Caudal peduncle relatively compressed. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well-developed lobes; upper lip with transverse wrinkles on surface. Barbels, 3 pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 12 (range, 11–14). Very small cycloid scales on the trunk. Lamina circularis at the base of the pectoral fin of adult male roundish plate, somewhat narrowing toward the outer part (Fig. 6I). The first branched soft ray of pectoral fin longer than the other rays; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin narrow and weak. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens: 62.1 mm SL male, 73.0 mm SL female.</p> <p>Coloration. Male in the non-spawning season (Figs. 3I, 8G). Body yellowish white with brown pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Upper part of head, opercle and snout covered with oval or amorphous shaped spots. Body pigmentation organized in one middorsal and four lateral zones. Line L1 consisting of a series of 15–20 blotches; blotches saddle or oval-shaped, often chained. Line L2 formed by longitudinal jagged line, reaching to postdorsal region, fused with L1. Line L3 formed by incomplete longitudinal line, reaching to caudal base, fused with L1 and L4 on postanal body. Line L4 formed by longitudinal jagged weblike line, reaching to postanal body. Line L5 organized in 11–13 blotches; blotches roundish or ovoid. Caudal fin and dorsal fin with 4–6 arcuate bars. Anal fin pigmented along the fin rays. Upper spot at the caudal base jet-black comparable in size to eye diameter, lower spot at caudal base faint or missing.</p> <p>Male in the spawning season (Fig. 8H). Line L4 not visible or formed by narrow longitudinal line, present only in anterior half of body. Lines L3 and L5 well developed with broad stripes from upper part of the pectoral-fin base to the caudal-fin base.</p> <p>Female (Fig. 5I). Appearance similar to males in the non-spawning season. But line L4 often formed by longitudinal jagged line, broader or as wide as L3.</p> <p>Sexual dimorphism. Males have roundish lamina circularis at the base of the pectoral fin, but females do not. Generally, the body size of females is larger than that of males.</p> <p>Egg diameter. 0.86 ± 0.08 mm (females, N = 4; collected from the Hii River system, Shimane Prefecture).</p> <p>Karyotype. Diploid (2n = 50) (Kimizuka 1987; Saitoh et al. 2000).</p> <p>Distribution. Rivers flowing into Japan sea, San-in District, western Honshu: Hyogo, Tottori, and Shimane Prefectures (Saitoh &amp; Aizawa 1987).</p> <p>Habitat and biology. This subspecies inhabits sandy-mud bottoms of lower reaches of rivers and small streams. Life histories are unknown.</p> <p>Etymology. The subspecific name is derived from the San-in District of eastern Honshu, which is the main distribution area of this subspecies.</p> <p>Remarks. The genetic features have been reported by Kitagawa et al. (2005) and Saitoh et al. (2010).</p> <p>Japanese name. San-in-kogata-suji-shima-dojyô.</p></div> 	https://treatment.plazi.org/id/03CBD358FF87FFB7F3D755D4FD04F8B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
03CBD358FF85FFB9F3D7518CFD29FD88.text	03CBD358FF85FFB9F3D7518CFD29FD88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis minamorii subsp. yodoensis Nakajima 2012	<div><p>Cobitis minamorii yodoensis Nakajima, subsp. nov.</p> <p>(Figs. 3J, 5J, 6J)</p> <p>Cobitis taenia striata Yodo small race: Saitoh and Aizawa 1987: 336, fig. 3C; Cobitis sp. S (Yodogawa-gata): Saitoh 1989: 388; Cobitis striata complex Yodo small race: Saitoh et al. 2010: 1003, table 1; Cobitis sp. 2 subsp. 5: Nakajima et al. 2012: 90, fig. 2f.</p> <p>Holotype. FAKU55719, 1 male, 45.1 mm SL, Japan: Yodo River, Asahi-ku, Osaka Pref., Honshu, 16. X. 1980, K. Saitoh.</p> <p>Paratypes. FAKU134643, 1 male, 43.0 mm SL, same data as holotype; FAKU134643, 1 male, 43.1 mm SL, same data as holotype; FAKU134643, 1 female, 70.6 mm SL, same data as holotype; FAKU55701, female, 64.9 mm SL, Uji R., Yodo R. s., Uji, Kyoto Pref., Honshu, 15. VII. 1976, K. Saitoh; FAKU55702, 1 male, 50.5 mm SL, Uji R., Yodo R. s., Uji, Kyoto Pref., Honshu, 12. V. 1978, K. Saitoh; KUN-P41321, 6 females, 59.9–73.9 mm SL, Yodo R., Asahi-ku, Osaka Pref., Honshu, 8. V. 1971, collector unknown.</p> <p>Diagnosis. This subspecies is distinguishable from other Japanese striated spined loaches by the following characteristics: body size small, the mature size about 50 mm SL in males, 60 mm SL in females; lamina circularis at the base of the pectoral fin of adult male roundish plate, somewhat narrowing toward the outer part; upper segments of the first branched soft ray of pectoral fin narrow and weak (Fig. 6J); PMN commonly 12; snout elongated; eye diameter relatively small; line L1 consisting of a series of 14–17 blotches; line L2 formed by tiny angular blotches, reaching to pre- or middorsal region, often fused with L1; line L3 formed by incomplete longitudinal narrow line, reaching beyond dorsal fin, fused with L1 and L 4 in posterior part; line L5 formed by narrow longitudinal line; caudal and dorsal fins margined by a narrow black band, two or three speckles on other regions; upper spot at the caudal base black, smaller than eye diameter; lower spot at caudal base existing but faint; spots not connected; karyotype diploid (2n = 50).</p> <p>Description. Lateral view in Figure 3J illustrate body shape, form and position of fins. Morphometric and meristic data for 4 males and 8 females are summarized in Table 2. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7; pelvic-fin rays ii, 5–6; caudal-fin rays 8+8. Body elongate, laterally compressed. Snout short. Interorbital space narrow, convex. Eye diameter relatively small. Caudal peduncle relatively compressed. Eye diameter small. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well-developed lobes; upper lip with transverse wrinkles on surface. Barbels, 3 pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 12 (range, 11–13). Very small cycloid scales on the trunk. Lamina circularis at the base of the pectoral fin of adult male roundish plate, somewhat narrowing toward the outer part (Fig. 6J). The first branched soft ray of pectoral fin longer than the other rays; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin narrow and weak. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens: 50.5 mm SL male, 73.9 mm SL female.</p> <p>Coloration. Male (Fig. 3J). Body pearl white with black pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Head, opercle and snout covered with some large amorphous patterns. Body pigmentation organized in one dorsal and four lateral zones. Line L1 consisting of a series of 14–17 blotches; blotches saddle shape. Line L2 formed by tiny angular blotches, reaching to pre- or middorsal region, often fused with L1. Line L3 formed by incomplete longitudinal narrow line, reaching beyond dorsal fin, fused with L1 and L 4 in posterior part. Line L4 formed by narrow longitudinal line, reaching near caudal base, fused with L1 and L 3 in posterior part. Line L5 formed by narrow longitudinal line from upper part of the pectoral fin to the caudal-fin base. Caudal fin and dorsal fin with 3–4 bars of small speckles, exterior bar often margined. Anal fin pigmented along the fin rays. Upper spot at the caudal base black, smaller than eye diameter; lower spot at caudal base faint; spots not connected.</p> <p>Female (Fig. 5J). Appearance similar to males. But line L4 more developed than males.</p> <p>Sexual dimorphism. Males have roundish lamina circularis at the base of the pectoral fin, but females do not. Generally, the body size of females is larger than that of males.</p> <p>Egg diameter. Unknown.</p> <p>Karyotype. Diploid (2n = 50) (Ueno &amp; Ojima 1976; Ueno et al. 1980; Saitoh et al. 2000).</p> <p>Distribution. Middle and lower reaches of Yodo River system, central Honshu: Kyoto and Osaka Prefectures (Saitoh &amp; Aizawa 1987).</p> <p>Habitat and biology. This subspecies inhabits sandy-mud bottoms of lower reaches of rivers and wando-pools (Saitoh &amp; Aizawa 1987; Saitoh 1989).</p> <p>Etymology. The subspecific name is derived from the type locality.</p> <p>Remarks. The subspecies is endemic to the middle and lower reaches of the Yodo River system; however, the last known individual of this subspecies was collected in August 1996. It points out that the decline of the subspecies has been attributed to river improvement and urbanization (Saitoh 1989, 2005). The morphology of this subspecies has been well studied by Saitoh and Aizawa (1987). The genetic features have been reported by Saitoh et al. (2010).</p> <p>Japanese name. Yodo-kogata-suji-shima-dojyô.</p></div> 	https://treatment.plazi.org/id/03CBD358FF85FFB9F3D7518CFD29FD88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
03CBD358FF8AFFB9F3D753A8FA6FF8B3.text	03CBD358FF8AFFB9F3D753A8FA6FF8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cobitis striata	<div><p>Key to 10 species and subspecies of Japanese Cobitis striata complex</p> <p>1. Upper segments of first branched soft ray of pectoral fin broad and strong; prepelvic myotome number commonly 14; egg diameter over 1.1 mm................................................................ C. magnostriata sp. nov.</p> <p>- Upper segments of first branched soft ray of pectoral fin narrow and weak; prepelvic myotome number commonly equal to or less than 13; egg diameter about or less than 1.0 mm.......................................................... 2</p> <p>2. Lamina circularis of adult male simple roundish plate; prepelvic myotome number commonly 13; black spot on lower caudalfin base inconspicuous or absent.......................................................................... 3</p> <p>- Lamina circularis of adult male roundish plate, somewhat narrowing toward outer part; prepelvic myotome number commonly 12; black spot on lower caudal-fin base small................................................................ 6</p> <p>3. Caudal fin and dorsal fin with 4–5 arcuate bars in adult; mature size small, 45–55 mm SL in male, 50–60 mm SL in female; egg diameter 0.8–0.9 mm................................................................. C. kaibarai sp. nov.</p> <p>- Caudal fin and dorsal fin with 3–4 arcuate bars in adult; mature size moderate, 60–70 mm SL in male, 60–90 mm SL in female; egg diameter 0.9–1.0 mm................................................................................ 4</p> <p>4. Line L4 formed by weak dots or narrow line in non-spawning season; line L5 formed by longitudinal line in all seasons.............................................................................................. C. striata striata</p> <p>- Line L4 formed by dots or weblike line in non-spawning season; line L5 organized in 11–14 roundish or ovoid blotches in nonspawning season....................................................................................... 5</p> <p>5. Line L4 formed by narrow longitudinal line, narrower than L 3 in male in non-spawning season; light brown pigmentation in fresh specimens.............................................................. C. striata fuchigamii subsp. nov.</p> <p>- Line L4 formed by longitudinal, jagged weblike line, broader than L 3 in male in non-spawning season; dark brown pigmentation in fresh specimens................................................ C. striata hakataensis subsp. nov.</p> <p>6. Lines L3 and L5 well-developed broad stripes in all seasons........................ C. minamorii oumiensis subsp. nov.</p> <p>- Lines L3 and L5 formed by narrow longitudinal line or organized blotches in non-spawning season.................... 7</p> <p>7. Line L5 formed by narrow longitudinal line or organized in connected tiny blotches in non-spawning season.............. 8</p> <p>- Line L5 organized in 10–16 blotches in non-spawning season................................................... 9</p> <p>8. Line L5 formed by narrow incomplete longitudinal line or organized in 15–20 connected tiny blotches; body depth 15.5–20.0% in SL............................................................. C. minamorii minamorii sp. nov.</p> <p>- Line L5 formed by narrow longitudinal line; body slender, body depth 12.7–19.2% in SL. C. minamorii yodoensis subsp. nov.</p> <p>9. Line L2 formed by sparse angular blotches; L4 formed by tiny blotches or narrow incomplete longitudinal line in male in nonspawning season; caudal and dorsal fins with 3–4 thin arcuate bars in adult............ C. minamorii tokaiensis subsp. nov.</p> <p>- Line L2 formed by longitudinal jagged line; L4 formed by longitudinal jagged weblike line, reaching to postanal body in male in non-spawning season; caudal and dorsal fins with 4–6 thin arcuate bars in adult....... C. minamorii saninensis subsp. nov.</p> </div>	https://treatment.plazi.org/id/03CBD358FF8AFFB9F3D753A8FA6FF8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakajima, Jun	Nakajima, Jun (2012): Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan. Zootaxa 3586: 103-130
