taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037338/files/figure.png	https://doi.org/10.5281/zenodo.5037338	Fig. 4. Bryothinusa minuta (Sawada, 1955).A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – male tergite X; I – female tergite X; J – medin lobe, ventral view; K – medin lobe, lateral view; L – paramere; M – spermatheca. Scale bars: 0.1 mm.	Fig. 4. Bryothinusa minuta (Sawada, 1955).A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – male tergite X; I – female tergite X; J – medin lobe, ventral view; K – medin lobe, lateral view; L – paramere; M – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037372/files/figure.png	https://doi.org/10.5281/zenodo.5037372	Fig. 20. Collection localities of Bryothinusa species in Japan:A – B. algarum Sawada, 1971 in Nemoto, Chiba; B – B. minuta (Sawada, 1955), B. nakanei (Sawada, 1955) and B. sakishimana Sawada, 1991 in Takara-jima (Ôgomori beach), Tokara-rettô; C – B. minuta in Nakano-shima (Yoriki), Tokara-rettô; D – B. minuta, B. nakanei and B. sakishimana in Iriomote-jima (mouth of Honera-gawa); E – B. tsutsuii (Sawada, 1955) in Nakano-shima (Amadomari), Tokara-rettô; F – B. nakanei and B. tsutsuii in Iriomote-jima (Haemida); G – B. tsutsuii in Iriomote-jima (mouth of Nakama-gawa); H – B. itsuroi sp. nov. and B. koreana Ahn & Jeon, 2004 in Kakise, Taka-shima, Nagasaki.	Fig. 20. Collection localities of Bryothinusa species in Japan:A – B. algarum Sawada, 1971 in Nemoto, Chiba; B – B. minuta (Sawada, 1955), B. nakanei (Sawada, 1955) and B. sakishimana Sawada, 1991 in Takara-jima (Ôgomori beach), Tokara-rettô; C – B. minuta in Nakano-shima (Yoriki), Tokara-rettô; D – B. minuta, B. nakanei and B. sakishimana in Iriomote-jima (mouth of Honera-gawa); E – B. tsutsuii (Sawada, 1955) in Nakano-shima (Amadomari), Tokara-rettô; F – B. nakanei and B. tsutsuii in Iriomote-jima (Haemida); G – B. tsutsuii in Iriomote-jima (mouth of Nakama-gawa); H – B. itsuroi sp. nov. and B. koreana Ahn & Jeon, 2004 in Kakise, Taka-shima, Nagasaki.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037374/files/figure.png	https://doi.org/10.5281/zenodo.5037374	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037340/files/figure.png	https://doi.org/10.5281/zenodo.5037340	Fig. 5. Bryothinusa tsutsuii (Sawada, 1955).A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 5. Bryothinusa tsutsuii (Sawada, 1955).A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC00FF884CC6FF4612C9F997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC00FF884CC6FF4612C9F997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC00FF884CC6FF4612C9F997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037344/files/figure.png	https://doi.org/10.5281/zenodo.5037344	Fig. 6. Bryothinusa koreana Ahn & Jeon, 2004. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm. Modified from Lංඎ et al. (2020).	Fig. 6. Bryothinusa koreana Ahn & Jeon, 2004. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm. Modified from Lංඎ et al. (2020).	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC00FF884CC6FF4612C9F997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037372/files/figure.png	https://doi.org/10.5281/zenodo.5037372	Fig. 20. Collection localities of Bryothinusa species in Japan:A – B. algarum Sawada, 1971 in Nemoto, Chiba; B – B. minuta (Sawada, 1955), B. nakanei (Sawada, 1955) and B. sakishimana Sawada, 1991 in Takara-jima (Ôgomori beach), Tokara-rettô; C – B. minuta in Nakano-shima (Yoriki), Tokara-rettô; D – B. minuta, B. nakanei and B. sakishimana in Iriomote-jima (mouth of Honera-gawa); E – B. tsutsuii (Sawada, 1955) in Nakano-shima (Amadomari), Tokara-rettô; F – B. nakanei and B. tsutsuii in Iriomote-jima (Haemida); G – B. tsutsuii in Iriomote-jima (mouth of Nakama-gawa); H – B. itsuroi sp. nov. and B. koreana Ahn & Jeon, 2004 in Kakise, Taka-shima, Nagasaki.	Fig. 20. Collection localities of Bryothinusa species in Japan:A – B. algarum Sawada, 1971 in Nemoto, Chiba; B – B. minuta (Sawada, 1955), B. nakanei (Sawada, 1955) and B. sakishimana Sawada, 1991 in Takara-jima (Ôgomori beach), Tokara-rettô; C – B. minuta in Nakano-shima (Yoriki), Tokara-rettô; D – B. minuta, B. nakanei and B. sakishimana in Iriomote-jima (mouth of Honera-gawa); E – B. tsutsuii (Sawada, 1955) in Nakano-shima (Amadomari), Tokara-rettô; F – B. nakanei and B. tsutsuii in Iriomote-jima (Haemida); G – B. tsutsuii in Iriomote-jima (mouth of Nakama-gawa); H – B. itsuroi sp. nov. and B. koreana Ahn & Jeon, 2004 in Kakise, Taka-shima, Nagasaki.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC00FF884CC6FF4612C9F997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037374/files/figure.png	https://doi.org/10.5281/zenodo.5037374	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC00FF884CC6FF4612C9F997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC00FF8F4C89F9791502F8D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC00FF8F4C89F9791502F8D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC00FF8F4C89F9791502F8D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037346/files/figure.png	https://doi.org/10.5281/zenodo.5037346	Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 7. Bryothinusa yoshigoui sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC00FF8F4C89F9791502F8D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC07FF8F4FEFF8CF13F1FA6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC07FF8F4FEFF8CF13F1FA6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC07FF8F4FEFF8CF13F1FA6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037348/files/figure.png	https://doi.org/10.5281/zenodo.5037348	Fig. 8. Bryothinusa japonica Liu, Ono & Maruyama, 2020. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm. Modified from Lංඎ et al. (2020).	Fig. 8. Bryothinusa japonica Liu, Ono & Maruyama, 2020. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm. Modified from Lංඎ et al. (2020).	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC07FF8F4FEFF8CF13F1FA6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037374/files/figure.png	https://doi.org/10.5281/zenodo.5037374	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC07FF8F4FEFF8CF13F1FA6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC07FF924CC6FA321502F9B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC07FF924CC6FA321502F9B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC07FF924CC6FA321502F9B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037350/files/figure.png	https://doi.org/10.5281/zenodo.5037350	Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC07FF924CC6FA321502F9B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037374/files/figure.png	https://doi.org/10.5281/zenodo.5037374	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC07FF924CC6FA321502F9B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1AFF924E54F94F13F1F918.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1AFF924E54F94F13F1F918.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1AFF924E54F94F13F1F918.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037352/files/figure.png	https://doi.org/10.5281/zenodo.5037352	Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1AFF924E54F94F13F1F918.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037354/files/figure.png	https://doi.org/10.5281/zenodo.5037354	Fig. 11. Bryothinusa hokkaidensis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 11. Bryothinusa hokkaidensis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037374/files/figure.png	https://doi.org/10.5281/zenodo.5037374	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC19FF974E58F88015D5FD98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC19FF974E58F88015D5FD98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC19FF974E58F88015D5FD98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037356/files/figure.png	https://doi.org/10.5281/zenodo.5037356	Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC19FF974E58F88015D5FD98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037374/files/figure.png	https://doi.org/10.5281/zenodo.5037374	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC19FF974E58F88015D5FD98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1FFF974E66FD861368FCB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1FFF974E66FD861368FCB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1FFF974E66FD861368FCB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037358/files/figure.png	https://doi.org/10.5281/zenodo.5037358	Fig. 13. Bryothinusa itsuroi sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 13. Bryothinusa itsuroi sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1FFF974E66FD861368FCB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037372/files/figure.png	https://doi.org/10.5281/zenodo.5037372	Fig. 20. Collection localities of Bryothinusa species in Japan:A – B. algarum Sawada, 1971 in Nemoto, Chiba; B – B. minuta (Sawada, 1955), B. nakanei (Sawada, 1955) and B. sakishimana Sawada, 1991 in Takara-jima (Ôgomori beach), Tokara-rettô; C – B. minuta in Nakano-shima (Yoriki), Tokara-rettô; D – B. minuta, B. nakanei and B. sakishimana in Iriomote-jima (mouth of Honera-gawa); E – B. tsutsuii (Sawada, 1955) in Nakano-shima (Amadomari), Tokara-rettô; F – B. nakanei and B. tsutsuii in Iriomote-jima (Haemida); G – B. tsutsuii in Iriomote-jima (mouth of Nakama-gawa); H – B. itsuroi sp. nov. and B. koreana Ahn & Jeon, 2004 in Kakise, Taka-shima, Nagasaki.	Fig. 20. Collection localities of Bryothinusa species in Japan:A – B. algarum Sawada, 1971 in Nemoto, Chiba; B – B. minuta (Sawada, 1955), B. nakanei (Sawada, 1955) and B. sakishimana Sawada, 1991 in Takara-jima (Ôgomori beach), Tokara-rettô; C – B. minuta in Nakano-shima (Yoriki), Tokara-rettô; D – B. minuta, B. nakanei and B. sakishimana in Iriomote-jima (mouth of Honera-gawa); E – B. tsutsuii (Sawada, 1955) in Nakano-shima (Amadomari), Tokara-rettô; F – B. nakanei and B. tsutsuii in Iriomote-jima (Haemida); G – B. tsutsuii in Iriomote-jima (mouth of Nakama-gawa); H – B. itsuroi sp. nov. and B. koreana Ahn & Jeon, 2004 in Kakise, Taka-shima, Nagasaki.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1FFF974E66FD861368FCB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1FFF9A4C9AFC66155FFC18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1FFF9A4C9AFC66155FFC18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1FFF9A4C9AFC66155FFC18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037360/files/figure.png	https://doi.org/10.5281/zenodo.5037360	Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm.	Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1FFF9A4C9AFC66155FFC18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037374/files/figure.png	https://doi.org/10.5281/zenodo.5037374	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC1FFF9A4C9AFC66155FFC18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037362/files/figure.png	https://doi.org/10.5281/zenodo.5037362	Fig. 15. Bryothinusa okinawana sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 15. Bryothinusa okinawana sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037364/files/figure.png	https://doi.org/10.5281/zenodo.5037364	Fig. 16. Bryothinusa nigra sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 16. Bryothinusa nigra sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037374/files/figure.png	https://doi.org/10.5281/zenodo.5037374	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037336/files/figure.png	https://doi.org/10.5281/zenodo.5037336	Fig. 3. Bryothinusa algarum Sawada, 1971.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 3. Bryothinusa algarum Sawada, 1971.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037368/files/figure.png	https://doi.org/10.5281/zenodo.5037368	Fig. 18. Bryothinusa moriguchii sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – male tergite X; I – female tergite X; J – median lobe, ventral view; K – median lobe, lateral view; L – paramere; M – spermatheca. Scale bars: 0.1 mm.	Fig. 18. Bryothinusa moriguchii sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – male tergite X; I – female tergite X; J – median lobe, ventral view; K – median lobe, lateral view; L – paramere; M – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037366/files/figure.png	https://doi.org/10.5281/zenodo.5037366	Fig. 17. Bryothinusa nakanei (Sawada, 1955).A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – male tergite X; I – female tergite X; J – median lobe, ventral view; K – median lobe, lateral view; L – paramere; M – spermatheca.. Scale bars: 0.1 mm.	Fig. 17. Bryothinusa nakanei (Sawada, 1955).A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – male tergite X; I – female tergite X; J – median lobe, ventral view; K – median lobe, lateral view; L – paramere; M – spermatheca.. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037368/files/figure.png	https://doi.org/10.5281/zenodo.5037368	Fig. 18. Bryothinusa moriguchii sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – male tergite X; I – female tergite X; J – median lobe, ventral view; K – median lobe, lateral view; L – paramere; M – spermatheca. Scale bars: 0.1 mm.	Fig. 18. Bryothinusa moriguchii sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – male tergite X; I – female tergite X; J – median lobe, ventral view; K – median lobe, lateral view; L – paramere; M – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037332/files/figure.png	https://doi.org/10.5281/zenodo.5037332	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037370/files/figure.png	https://doi.org/10.5281/zenodo.5037370	Fig. 19. Bryothinusa gangjinensis Ahn & Jeon, 2004.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	Fig. 19. Bryothinusa gangjinensis Ahn & Jeon, 2004.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037374/files/figure.png	https://doi.org/10.5281/zenodo.5037374	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037376/files/figure.png	https://doi.org/10.5281/zenodo.5037376	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037334/files/figure.png	https://doi.org/10.5281/zenodo.5037334	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm.	2021-05-07	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi		Zenodo	biologists	Liu, Tian-tian;Ono, Hiroki;Maruyama, Munetoshi			
