taxonID	type	description	language	source
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	description	(Figs 1 B; 2 B; 4; 20 B – D; 21 G; 22) (Japanese name: Hoso-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	materials_examined	Type material. Not examined. Material examined. JAPAN: H ඈκκൺ ං ൽô: 2 ♀♀, Usu-chô, Date-shi, Hokkaidô, 12. iv. 2020, Y. Tasaku. H ඈඇඌΗǕ: 44 unsexed spec., Sunosakikaigan, Sunosaki, Tateyama-shi, Chiba-ken, 6. ii. 2007, H. Ono; 1 unsexed spec., Kanaya, Futtsu-shi, Chiba-ken, 9. iv. 2007, H. Ono; 2 unsexed spec., Shibasaki-bashi, Kanaya, Futtsu-shi, Chiba-ken, 14. vi. 2009, H. Ono; 2 unsexed spec., Nagasakibana, Chôshi-shi, Chiba-ken, 16. ix. 2009, H. Ono; 2 unsexed spec., same locality, 13. x. 2009, H. Ono; 1 unsexed spec., Kuruwa-kaigan, Akiya, Yokosuka-shi, Kanagawa-ken, 14. ix. 2013, H. Ono; 3 JJ 5 unsexed spec., Mouth of Kô-gawa, Numazu-shi, Shizuoka-ken, 1. x. 2000, T. Kurihara; 3 ♀♀, 5 unsexed spec., Tsumeki-zaki, Shimoda-shi, Shizuoka-ken, 21. iv. 2003, M. Maruyama; 1 J, Wakamatsu-zaki, Numazu - shi, Shizuoka-ken, 1. x. 2000, T. Kurihara; 2 unsexed spec., Kusumibana, Ôbatake, Kojima, Kurashiki-shi, Okayama-ken, 21. iii. 2009, O. Yamaji; 3 unsexed spec., Ta-no-ura, Naga-shima, Kaminoseki-chô, Yamaguchiken, 15. x. 2005, Y. Nakase. SΗ ං κ ඈκඎ: 1 J, Kaigan-ji, Nishishirakata, Tadotsu-chô, Kagawa-ken, 31. iii. 2008, H. Fujimoto; 2 JJ 2 ♀♀, 3 unsexed spec., Ôyatomi-machi, Sakaide-shi, Kagawa-ken, 29. iii. 2008, H. Fujimoto; 1 J 2 ♀♀, 10 unsexed spec., Mouth of Umayado-gawa, Umayado, Higashi-kagawa-shi, Kagawa-ken, 11. v. 2014, T. Taki; 7 unsexed spec., Ado-ike, Hiketa, Higashi-kagawa-shi, Kagawa-ken, 28. vii. 2013, T. Taki; 17 unsexed spec., same locality, 11. v. 2014, T. Taki; 2 unsexed spec., Koura-kaigan, Koiso, Higashi-kagawa-shi, Kagawa-ken, 29. iv. 2014, T. Taki; 1 unsexed spec., same locality, 21. iv. 2014, T. Taki. K ඒǕඌΗǕ: 1 J, Suno-saki, Imazu, Nishi-ku, Fukuoka-shi, Fukuoka-ken, 26. vi. 2020, Y. Hisasue; 31 unsexed spec., same locality, 16. x. 2020, T. Liu & T. Nozaki; 1 unsexed spec., Miyaga-hama, Ibusuki-shi, Kagoshimaken, 19. iv. 2008, M. Maruyama; 2 unsexed spec., Ko-ura, Kasasa-machi, Minamisatsuma-shi, Kagoshima-ken, 20. iv. 2008, M. Maruyama; 3 unsexed spec., Aiboshi-gawa, Aiboshi, Minamisatsuma-shi, Kagoshima-ken, 20. iv. 2008, M. Maruyama. N ൺඇඌൾ ං - ඌΗඈඍත: 6 unsexed spec., Ôgomori, Takara-jima, Tokara-rettô, Kagoshima-ken, 6. vii. 2019, T. Liu, N. Tsuji & H. Ono; 3 JJ 1 ♀, 27 unsexed spec., Yoriki (a gravel beach), Nakano-shima, Tokara-rettô, Kagoshima-ken, 7. vii. 2019, T. Liu, N. Tsuji & H. Ono; 15 unsexed spec., same locality, 8. vii. 2019, T. Liu, N. Tsuji & H. Ono; 1 unsexed spec., You, Kasari-chô, Amami-Ôshima, Ôshima- gun, Kagoshima-ken, 27. ii. 2004, H. Kamezawa; 2 ♀♀, 2 unsexed spec., Kasari-wan, Kasari-chô, Amami-Ôshima, Kagoshima-ken, 4. vi. 2008, M. Maruyama; 4 unsexed spec., Yuwan, Uken-son, Amami-Ôshima, Kagoshima-ken, 5. vi. 2008, M. Maruyama; 1 J 1 ♀, 3 unsexed spec., Shokazu, Setouchi-chô, Kakeroma-jima, Kagoshima-ken, 6. vi. 2008, M. Maruyama; 1 ♀, Onna-son, Inbu-gun, Okinawa-ken, 11 – 13. v. 2005, M. Sugimoto; 3 JJ, Teruma, Yonagusuku-son, Okinawa-ken, 14. ix. 2003, M. Moriguchi; 2 unsexed spec., Fûne, Ishigaki-shi, Okinawa-ken, 7. iii. 2000, T. Kurihara; 6 JJ 8 unsexed spec., same locality, 8. iii. 2000, T. Kurihara; 3 unsexed spec., same locality, 12. iii. 2000, T. Kurihara; 10 unsexed spec., Nagura-Anparu, Nagura, Ishigaki-shi, Okinawa-ken, 4. ii. 2009, H. Ono; 2 JJ 2 ♀♀, 42 unsexed spec., Mouth of Honera-gawa, Iriomote-jima, Takana, Taketomi-chô, Okinawa-ken, 2. iv. 2019, T. Nozaki; 3 unsexed spec., Mouth of Honera-gawa, Takana, Taketomi-chô, Iriomote-jima, Okinawa-ken, 24. vi. 2019, T. Liu & N. Tsuji. OǤ ൺඌൺඐൺඋൺ-ඌΗඈඍත: 2 JJ 1 ♀, 6 unsexed spec., Ô-mura, Chichi-jima, Ogasawara-shotô, Tôkyô-to, 1. x. 1977, Y. Watanabe. Deposited at KUM but some will be distributed to NMPC and BMNH.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	description	Redescription. Body (Fig. 1 B) small, about 1.6 – 1.7 mm long, densely covered with minute setae. Head, pronotum and elytra light yellow; abdominal tergites VI – VII almost infuscate posteriorly; antenna, mouthparts and legs similarly coloured, lightly paler than head colour and paler toward apex. Head oval, wider than long, longer than pronotum and elytra; eyes (Fig. 2 B) small, reniform, shorter than 1 / 4 head length, with 3 or 4 facets; antennal length shorter than distance from anterior margin of head to posterior margin of elytra; all antennal segments longer than wide; segment I 1.5 times longer than II, almost 2.5 times longer than III, and shorter than XI. Mouthparts (Figs 4 A – E) with distribution of labral setae as in Figure 4 A; mandibles (Fig. 4 B) elongate triangle, asymmetrical, right mandible with median tooth; median tooth longer than 1 / 4 of mandibles; left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 4 C) with two spines interdigitating close; 2 – 3 setae on inner margin of lacinia; labial palpus (Fig. 4 D) with most setae located basally; mentum (Fig. 4 E) with antero- - lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” of mentum distant from apical setae. Pronotum longer than elytra, narrower than head. Elytra widened apically, wider than long, narrower than pronotum; hind wings absent. Male abdominal sternite VIII (Fig. 4 F) with posterior margin slightly serrate. Medial posterior margin protruded sharp in form on tergite X of male (Fig. 4 H). Median lobe of aedeagus as in Figs 4 J, K; paramere (Fig. 4 L) elongate. Female abdominal sternite VIII (Fig. 4 G) with posterior margin slightly serrate but rounder and smoother than in male. Medial posterior margin protruded sharp in form on tergite X of female (Fig. 4 I), and sharper than in male. Spermatheca (Fig. 4 M) small and apical part very short; basal part complicated in shape; several times coiled near base. Measurements. BL 1.60 – 1.70, FBL 0.70 – 0.80, HL 0.25 – 0.28, HW 0.25 – 0.34, PL 0.24 – 0.26, PW 0.25 – 0.28, EL 0.20 – 0.23, EW 0.24 – 0.25, HTL 0.23 – 0.26, AL 0.55 – 0.65, AS-I 0.08 – 0.09, AS-II 0.05 – 0.06, AS-III 0.03 – 0.04.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	diagnosis	Differential diagnosis. This species is easily distinguished from the other Japanese species of Bryothinusa by the smaller body and strongly reduced eyes, each consisting of 3 or 4 facets only.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	distribution	Distribution. Japan (Hokkaidô; Honshû: Chiba-ken, Tôkyô-to, Kanagawa-ken, Shizuoka-ken, Wakayama-ken, Okayama-ken, Yamaguchi-ken; Shikoku: Kagawa-ken; Kyûshû: Fukuoka-ken, Kagoshima-ken; Nansei-shotô: Kagoshima-ken: Takara-jima, Nakano-shima, Amami - Ôshima, Kakeroma-jima; Okinawa-ken: Okinawa-jima, Ishigaki-jima, Iriomote-jima; Ogasawara-shotô: Chichijima); Korea.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	discussion	Remarks. This species is very small, currently the smallest species in Japan, and the specimens were collected under stones. In the Japanese species of Bryothinusa, B. minuta is most widely distributed, from Hokkaidô to Iriomote-jima. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22. Bryothinusa tsutsuii (Sawada, 1955) (Figs 1 C; 2 C; 5; 20 E – G; 21 D; 22) (Japanese name: Kiiro-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	description	of article (characters); M ඈඈ * © ൾ et al. (1973): 74 (in key to species of Bryothinusa), 77 (nomenclatorial history; diagnosis); M ඈඈ * © ൾ & L ൾ * û ඇൾ * © (1975): 109 – 110 (in key to species of Bryothinusa), 111 (in tabular key to known species of Bryothinusa); A ඌ * ü ൾ (2004): 595 (in annotated catalogue of world species); S ආൾඍൺඇൺ (2004): 464 (in catalogue of Palearctic species); F * © ൺඇκ & A * ü ඇ (2011): 27 (in checklist); S ർ * ü * “ අκൾ & S ආൾඍൺඇൺ (2015): 676 (in catalogue of Palearctic species); L ං ඎ et al. (2020): 589 (in key of Japanese species), 592 (in checklist).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	materials_examined	Type material. Not examined. Material examined. JAPAN: N ൺඇඌൾ ං - ඌΗඈඍත: 3 JJ 2 ♀♀, 9 unsexed spec., Mage-shima (near Tanega-shima), Kagoshima-ken, 14. x. 2000, M. Maruyama (under stone on sandy beach, in a port); 5 JJ 3 ♀♀, 18 unsexed spec., Ama-domari (a gravel beach), Nakano-shima, Tokara-rettô, Kagoshima-ken, 7. vii. 2019, T. Liu & H. Ono; 1 J 6 ♀♀, 19 unsexed spec., Uken-son, Edateku-jima (near Amami-Ôshima), Kagoshima-ken, 5. vi. 2008, M. Maruyama; 6 unsexed spec., Izena, Izena-son, Shimajirigun, Okinawa-ken, 5. v. 2013, M. Asano; 4 JJ 5 ♀♀, 66 unsexed spec., Mihama (Sunset Beach), Chatan-chô, Okinawa-jima, Okinawa-ken, 27. iii. 2009, H. Suenaga; 1 ♀, 7 unsexed spec., Chatan-chô, Okinawa, Okinawa-ken, Nansei-shotô, 26. v. 2002, M. Moriguchi; 1 J 2 ♀♀, 7 unsexed spec., Teruma, Yonagusuku-son, Okinawa-ken, 14. ix. 2003, M. Moriguchi; 1 ♀, 3 unsexed spec., Ou-jima, Nago-shi, Okinawa-ken, 30. iv. 2006, M. Moriguchi; 2 JJ 1 ♀, 2 unsexed spec., Inbu-gun, Onnason, Okinawa-ken, 11 – 13. v. 2005, M. Sugimoto; 3 JJ 1 ♀, 7 unsexed spec., Kadena-Marina, Kadena-chô, Okinawa-ken, 25. iv. 2005, M. Moriguchi (under stone on sandy beach in estuary); 1 unsexed spec., Nakasuji, Ishigaki-shi, Ishigaki-jima, Okinawa-ken, 14. iii. 2000, T. Kurihara; 1 J 2 ♀♀, 8 unsexed spec., Inoda, Ishigaki-jima, Okinawa-ken, 31. v. 2003, M. Moriguchi; 2 JJ 1 ♀, 13 unsexed spec., Fûne, Ishigaki-shi, Ishigaki-jima, Okinawa-ken, 8. iii. 2000, T. Kurihara; 2 unsexed spec., Akaishi, Ishigaki - shi, Ishigaki-jima, Okinawa-ken, 30 – 31. iii. 2010, T. Hayama; 1 unsexed spec., Haemida, Taketomi-chô, Iriomote-jima, Okinawa-ken, 19. iii. 2000, T. Kurihara; 2 ♀♀, 7 unsexed spec., same locality, 24. x. 2020, T. Nozaki; 1 J 1 ♀, 7 unsexed spec., Mouth of Nakama-gawa, Haiminaka, Iriomote-jima, Yaeyama-gun, Okinawa-ken, 4. ix. 2020, S. Inoue & Y. Hisasue. Deposited at KUM but some will be distributed to NMPC and BMNH.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	description	Redescription. Body (Fig. 1 C) small about 1.9 – 2.2 mm long, covered with minute setae. Head, pronotum and elytra yellowish brown; abdominal tergites IV – VII infuscate posteriorly; antenna, mouthparts and legs similarly coloured, slightly paler than head colour and paler toward apex. Head oval, slightly wider than long, longer than pronotum and elytra; eyes (Fig. 2 C) small, reniform, shorter than tempora length, setae present between about 15 – 17 facets; antennal length little shorter than distance from anterior margin of head to posterior margin of elytra; all antennal segments longer than wide; segment I 1.3 times longer than II, almost 2 times longer than III, and as long as XI. Mouthparts (Figs 5 A – E) with distribution of labral setae as in Figure 5 A; mandibles (Fig. 5 B) elongate triangle, asymmetrical, right mandible with median tooth; median tooth longer than 1 / 4 of mandibles; left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 5 C) with three spines closely interdigitating and second spine very small, not obvious; 4 setae on inner margin of lacinia; labial palpus (Fig. 5 D) with most setae located basally; mentum (Fig. 5 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” of mentum lower than level of anterior margin. Pronotum almost as long as elytra, slightly wider than head. Elytra wider than long, almost as wide as pronotum; hind wings absent. Male abdominal sternite VIII (Fig. 5 F) with posterior margin slightly serrate. Median lobe of aedeagus as in Figures 5 H, I; paramere (Fig. 5 J) elongate. Female abdominal sternite VIII (Fig. 5 G) with posterior margin slightly serrate but smoother than in male. Spermatheca (Fig. 5 K) about 1.5 times coiled near base. Measurements. BL 1.90 – 2.20, FBL 1.00 – 1.20, HL 0.33 – 0.45, HW 0.43 – 0.45, PL 0.35 – 0.40, PW 0.40 – 0.46, EL 0.38 – 0.40, EW 0.45 – 0.46, HTL 0.33 – 0.38, AL 0.83 – 1.05, AS-I 0.11 – 0.14, AS-II 0.08 – 0.11, AS-III 0.05 – 0.07.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	diagnosis	Differential diagnosis. In general appearance the species is closely allied to B. algarum, B. japonica, B. koreana and B. yoshigoui sp. nov., but easily distinguished from B. japonica, B. koreana and B. yoshigoui sp. nov. by the shape of the spermatheca and the median lobe of the aedeagus without granule-like projections and from B. algarum by the simplified shape of the spermatheca.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	distribution	Distribution. Japan (Nansei-shotô: Kagoshima-ken: Tanega-shima, Takara-jima, Nakano-shima, Edateku-jima; Okinawa-ken: Izena-jima, Okinawa-jima, Ou-jima, Ishigaki-jima, Iriomote-jima).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC0CFF884CEBFD86167EF7D8.taxon	discussion	Remarks. The number of facets is different from the original description, but other morphological characters completely coincide with it. The type specimen was not found despite the efforts of Mr. Shigehiko Shiyake of Ôsaka Museum of Natural History, to which Dr. Kohei Sawada donated the collection. This species is mainly found in Nansei-shotô but probably found also in mainland Kyûshû. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC00FF884CC6FF4612C9F997.taxon	description	(Figs 1 D; 2 D; 6; 20 H; 21 A; 22) (Japanese name: Chôsen-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC00FF884CC6FF4612C9F997.taxon	materials_examined	Type material. Not examined. Material examined. JAPAN: H ඈඇඌΗǕ: 1 J 1 ♀, Kusumi-bana, Ôbatake, Ko-jima, Kurashiki-shi, Okayama-ken, 21. iii. 2009, O. Yamaji; 3 JJ 2 ♀♀, Miya-jima, Hiroshima-ken, 23. iv. 1997, K. Sawada & R. Sawada; 5 JJ 2 ♀♀, 6 unsexed spec., Tanoura, Naga-shima, Kaminoseki-chô, Yamaguchi-ken, 15. x. 2005, Y. Nakase; 1 ♀, same locality, 1. v. 2008, M. Moriguchi. SΗ ං κ ඈκඎ: 2 ♀♀, 7 unsexed spec., Koura-kaigan, Koiso, Higashi-kagawa-shi, Kagawa-ken, 21. iv. 2013, T. Taki; 1 ♀, 2 unsexed spec., same locality, 29. iv. 2014, T. Taki; 4 ♀♀, 6 unsexed spec., Kaburagoshi, Matsubara, Higashi-kagawa-shi, Kagawa-ken, 7. ix. 2015, T. Taki. K ඒǕඌΗǕ: 6 unsexed spec., Mouth of Sakurai-gawa, Shima-nogita, Itoshima-shi, Fukuoka-ken, 21. iv. 2020, Y. Hisasue; 2 ♀♀, 4 unsexed spec., Kakise, Takashima, Nagasaki-shi, Nagasaki-ken, 4. viii. 2020, Y. Hisasue. Deposited at KUM but some will be distributed to NMPC and BMNH. Additional description. Number of facets is 17 – 19 as in Figure 2 D. Antennal segment I 1.1 times longer than II, almost 2 times longer than III, and as long as XI. Mouthparts with median tooth longer than 1 / 4 of mandibles; 6 setae on inner margin of lacinia; male and female abdominal sternite VIII, spermatheca and aedeagus as in Figure 6 (edited from L ං ඎ et al. (2020 )). See also L ං ඎ et al. (2020). Differential diagnosis. In general appearance, the species is closely similar to B. algarum, B. japonica, B. tsutsuii and B. yoshigoui sp. nov., but is easily distinguished from B. japonica by the apical part of the spermatheca without granule-like projections, and from B. tsutsuii by the median lobe of the aedeagus with granule-like projections while that of B. tsutsuii and B. algarum median lobe of aedeagus without granule-like projections, and differs from B. yoshigoui sp. nov. in the number of facets and body colour.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC00FF884CC6FF4612C9F997.taxon	distribution	Distribution. Japan (Honshû: Okayama-ken, Hiroshimaken, Yamaguchi-ken; Shikoku: Kagawa-ken; Kyûshû: Fukuoka-ken, Nagasaki-ken); Korea (Jeonnam).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC00FF884CC6FF4612C9F997.taxon	discussion	Remarks. Presumably this species occurs also in other areas, especially Japan Sea coast of western Japan and Seto Inland Sea coast of Japan. A distribution map of examined specimens of this species in Japan is illustrated in Fig. 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC00FF8F4C89F9791502F8D8.taxon	description	(Figs 1 E; 2 E; 7; 22) (Japanese name: Hiroshima-nagisa-hanekakushi) Type material. H ඈඅඈඍඒඉൾ: J, JAPAN: H ඈඇඌΗǕ: Mitsuguchi-kaigan, Yasuura-chô, Kure, Hiroshima-ken, 28. vii. 2019, H. Yoshigou (KUM). P ൺ * © ൺඍඒඉൾඌ: H ඈඇඌΗǕ: 6 JJ 1 ♀, 2 unsexed spec., same data as holotype (KUM, NMPC). Description. Body (Fig. 1 E) small about 2.3 – 2.4 mm long, covered with minute setae; background colour deep reddish brown, head, pronotum and elytra reddish brown; posteriorly abdominal tergite VI infuscate; antenna and legs similarly reddish-brown coloured, lightly paler than body colour and paler toward apex. Head oval, wider than long, longer than pronotum and elytra; eyes (Fig. 2 E) small, reniform, shorter than 1 / 2 of head length, with 12 – 14 facets; antennal length a little shorter than distance from anterior margin of head to posterior margin of elytra; antennal segments all longer than wide; segment I as long as II, 1.2 times longer than III, and shorter than XI; posterior margin of antennal segment III narrower than anterior margin. Mouthparts (Figs 7 A-E) with distribution of labral setae as in Figure 7 A; mandibles (Fig. 7 B) elongate triangle, asymmetrical, right with median tooth; median tooth longer than 1 / 4 of mandibles; left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 7 C) with second spines very small and not obvious, at tip with two spines not closely interdigitating; 4 setae on inner margin of lacinia; labial palpus (Fig. 7 D) with most setae located basally; mentum (Fig. 7 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” of mentum near the level of anterior margin. Pronotum almost as long as elytra, slightly narrower than head. Elytra widened apically, wider than long, almost as wide as pronotum; hind wings absent. Male abdominal sternite VIII (Fig. 7 F) with posterior margin slightly serrate. Median lobe of aedeagus (Figs 7 H, I) with some granules on apical lobe, and not clearly visible in dorsal view; paramere elongate, as in Figure 7 J. Female abdominal sternite VIII (Fig. 7 G) with posterior margin slightly serrate but smoother than in male. Spermatheca (Fig. 7 K) small, simplified in shape, about half times coiled near base. Measurements. BL 2.30 – 2.40, FBL 1.00 – 1.20, HL 0.38 – 0.45, HW 0.43 – 0.48, PL 0.35 – 0.38, PW 0.40 – 0.43, EL 0.35 – 0.38, EW 0.41 – 0.43, HTL 0.34 – 0.39, AL 0.88 – 0.97, AS-I 0.09 – 0.10, AS-II 0.09 – 0.10, AS-III 0.07 – 0.08.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC00FF8F4C89F9791502F8D8.taxon	diagnosis	Differential diagnosis. In general appearance, the species is closely similar to B. tsutsuii, B. algarum, B. koreana and B. japonica, in but easily distinguished from B. tsutsuii and B. algarum by the spermathecal shape and the median lobe of the aedeagus with granule-like projections, from B. koreana at the number of facets and the darker body colour, and from B. japonica by the apical part of spermatheca without granule-like projections.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC00FF8F4C89F9791502F8D8.taxon	distribution	Distribution. Japan (Honshû: Hiroshima).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC00FF8F4C89F9791502F8D8.taxon	etymology	Etymology. This new species is named after Mr. Hidenori Yoshigou, a famous ichthyologist and the collector of the type series.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC00FF8F4C89F9791502F8D8.taxon	discussion	Remarks. The specimens were collected on the coastline of Setonaikai (inland sea between Honshû and Shikoku), Japan. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF8F4FEFF8CF13F1FA6C.taxon	description	(Figs 1 F; 2 F; 8; 21 A – B, G; 22) (Japanese name: Nippon-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF8F4FEFF8CF13F1FA6C.taxon	materials_examined	Material examined. JAPAN: H ඈඇඌΗǕ: 7 JJ 1 ♀, 22 unsexed spec., Matsumi-ga-ura, Ôchiba, Kosai-shi, Shizuoka-ken, 11. iv. 2004, Y. Tahira; 8 unsexed spec., Miya-gawa, Iso-chô, Ise-shi, Mie-ken, 8. vi. 2008, K. Aki- ta; 4 JJ 2 ♀♀, 67 unsexed spec., Kumozu-gawa, Karasu-chô, Mie-ken, 15. v. 2004, K. Akita; 2 JJ 1 ♀, 26 unsexed spec., Mouth of Kansa-gawa, Nansei-chô, Mie-ken, 5. v. 2008, K. Akita; 4 JJ 4 unsexed spec., Mouth of Yoshii-gawa, Nishikôzai, Okayama-shi, Okayama-ken, 24. x. 2004, O. Yamaji; 2 JJ 4 ♀♀, 6 unsexed spec., Mitsuguchi-kaigan, Yasuura-chô, Kure, Hiroshima-ken, 28. vii. 2019, H. Yoshigou. SΗ ං κ ඈκඎ: 2 JJ 5 ♀♀, 112 unsexed spec., Shimoda, Shimanto-shi, Kôchi-ken, 21. iv. 2012, M. Maruyama; 1 J 4 ♀♀, 2 unsexed spec., Kagami-gawa, Takajô-machi, Kôchi-shi, Kôchi-ken, 20. vi. 2008, H. Fujimoto. K ඒǕඌΗǕ: 3 JJ 1 ♀, 4 unsexed spec., Sunosaki, Imazu, Nishi-ku, Fukuoka-shi, Fukuoka-ken, 16. x. 2020, T. Liu & T. Nozaki; 8 unsexed spec., Mouth of Sakurai-gawa, Shima-Nogita, Itoshima-shi, Fukuoka-ken, 21. iv. 2020, Y. Hisasue; 1 ♀, same locality, 11. x. 2018, Liu et al. Deposited at KUM but some will be distributed to NMPC and BMNH. Additional description. Number of facets 23 – 25 as Fig. 2 F. Antennal segment I 1.2 times longer than II, almost 2.4 times longer than III, and longer than XI. Mouthparts with median tooth longer than 1 / 4 of mandibles; 6 setae on inner margin of lacinia; male and female abdominal sternite VIII, spermatheca and aedeagus as in Figure 8. See also L ං ඎ et al. (2020).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF8F4FEFF8CF13F1FA6C.taxon	diagnosis	Differential diagnosis. In general appearance, the species is closely allied to B. tsutsuii, B. algarum, B. koreana and B. yoshigoui sp. nov., but easily distinguished from them by the apical part of spermatheca without granule-like projections.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF8F4FEFF8CF13F1FA6C.taxon	distribution	Distribution. Japan (Honshû: Chiba-ken, Shizuoka-ken, Mie-ken, Ôsaka-fu, Hyôgo-ken, Wakayama-ken, Okayama-ken, Hiroshima-ken, Yamaguchi-ken; Shikoku: Kagawa-ken, Ehime-ken, Kôchi-ken; Kyûshû: Fukuoka-ken).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF8F4FEFF8CF13F1FA6C.taxon	discussion	Remarks. About 200 specimens were collected before the original description but they were not included in the type series due to the difficulty of identification. There was considerable variation in the colouration of the specimens, ranging from light brown to brown, but it is unclear whether this is due to geographical variation or to the methods in which the specimens were preserved. This species is quite common in sandy to mud flats in estuaries in Japan, mainly west of Kantô District. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF924CC6FA321502F9B8.taxon	description	(Figs 1 G; 2 G; 9; 21 B, D; 22) (Japanese name: Sakishima-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF924CC6FA321502F9B8.taxon	materials_examined	Type material. Not examined. Material examined. JAPAN: N ൺඇඌൾ ං - ඌΗඈඍත: 3 JJ 2 ♀♀, 11 unsexed spec., Ôgomori, Takara-jima, Tokara-rettô, Kagoshima-ken, 3. vii. 2019, T. Liu, N. Tsuji & H. Ono; 1 unsexed spec., Mouth of Honera-gawa, Takana, Taketomi-chô, Iriomote-jima, Okinawa-ken, 2. iv. 2019, T. Nozaki. Deposited at KUM but some will be distributed to NMPC and BMNH.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF924CC6FA321502F9B8.taxon	description	Redescription. Body (Fig. 1 G) small, about 1.6 – 1.9 mm in length, covered with minute setae. Abdomen almost black and darker than head and pronotum; elytra lighter than pronotum; antenna and legs yellowish brown. Head evenly rounded, more or less produced anteriorly, wider than long, shorter than pronotum and elytra; eyes (Fig. 2 G) large, setae present between about 21 – 23 facets, slightly longer than tempora length; antenna yellowish brown, longer than total length of head and pronotum; segment I 1.4 times longer than II, almost 3.8 times longer than III, and longer than XI; antennal segments V – X almost as long as wide. Mouthparts (Figs 9 A – E) with distribution of labral setae as in Fig. 9 A, with large and thick spiniform seta antero-laterally; mandibles (Fig. 9 B) elongate triangle, asymmetrical, right mandible with median tooth; median tooth shorter than 1 / 4 of mandibles; left without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 9 C) with second spine very small, not obvious; 3 setae on inner margin of lacinia; labial palpus (Fig. 9 D) with most setae located basally; mentum (Fig. 9 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” far away from apical setae. Pronotum shorter than elytra, wider than head. Elytra subquadrate, narrower than pronotum; almost unicolour and hind wings present but small, apparently flightless. Male abdominal sternite VIII (Fig. 9 F) with posterior margin slightly serrate. Median lobe of aedeagus as in Figures 9 H, I; paramere (Fig. 9 J) elongate. Female abdominal sternite VIII (Fig. 9 G) slightly serrate alike male on posterior margin. Spermatheca (Fig. 9 K) about 1.5 times coiled near base, apical part elongate. Measurements. BL 1.60 – 1.90, FBL 0.70 – 1.00, HL 0.28 – 0.33, HW 0.35 – 0.40, PL 0.33 – 0.38, PW 0.40 – 0.45, EL 0.35 – 0.40, EW 0.38 – 0.40, HTL 0.33 – 0.35, AL 0.70 – 0.83, AS-I 0.12 – 0.15, AS-II 0.09 – 0.10, AS-III 0.03 – 0.04.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF924CC6FA321502F9B8.taxon	diagnosis	Differential diagnosis. In Nansei-shotô, this species is especially similar to B. nakanei, but easily distinguished from it by the paler elytral colour, the number of facets, the spermathecal shape and the posterior margin of the tergite X rounded medially. This species is also similar to B. constricta sp. nov. in general appearance but differentiated from it by the elytral shape not being almost square and the shape of the spermatheca.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF924CC6FA321502F9B8.taxon	distribution	Distribution. Japan (Nansei-shotô: Kagoshima-ken: Takara-jima; Okinawa-ken: Iriomote-jima).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC07FF924CC6FA321502F9B8.taxon	discussion	Remarks. This species is distributed in Nansei-shotô of Japan. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF924E54F94F13F1F918.taxon	description	(Figs 1 H; 2 H; 10; 22) (Japanese name: Kubire-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF924E54F94F13F1F918.taxon	materials_examined	Type material. H ඈඅඈඍඒඉൾ: J, JAPAN: H ඈඇඌΗǕ: Futtsu-misaki, Futtsu-shi, Chiba-ken, 1. v. 2014, H. Ono (KUM). P ൺ * © ൺඍඒඉൾඌ: H ඈඇඌΗǕ: 3 ♀♀, 6 unsexed spec., same locality as holotype, 28. v. 2010, H. Ono (KUM); 2 JJ 5 ♀♀, 10 unsexed spec., same data as holotype (KUM, NMPC, BMNH); 1 ♀, 4 unsexed spec., Izumo, Koryô-chô, Shimane-ken, 3. ix. 2007, T. Hayama (KUM).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF924E54F94F13F1F918.taxon	description	Description. Body (Fig. 1 H) small, about 1.7 – 2.0 mm in length, covered with minute setae. Abdomen almost black and darker than head and pronotum; elytra lighter than pronotum; legs yellowish brown. Head evenly rounded, more or less produced anteriorly, wider than long, shorter than pronotum and as long as elytra; eyes (Fig. 2 H) large, setae present between about 20 – 22 facets, slightly longer than tempora length; antenna yellowish brown, equal to total length of head and pronotum; segment I 1.6 times longer than II, almost 3.5 times longer than III, and longer than XI; antennal segments III – X almost wider than long, and VI – VII wider than width of III – IV. Mouthparts (Figs 10 A – E) with distribution of labral setae as in Figure 10 A, with large and thick spiniform seta at antero-laterally; mandibles (Fig. 10 B) elongate triangle, asymmetrical, right mandible with median tooth; median tooth shorter than 1 / 4 of mandibles; left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 10 C) at right angle; at tip with two spines very small and not closely interdigitating; 3 setae on inner margin of lacinia; labial palpus (Fig. 10 D) with most setae located basally; mentum (Fig. 10 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” far away from apical setae. Pronotum longer than elytra, wider than head. Elytra widened apically, wider than long, almost as wide as pronotum; hind wings present but very small, apparently flightless. Male abdominal sternite VIII (Fig. 10 F) with posterior margin slightly serrate. Median lobe of aedeagus as in Figures 10 H, I; paramere (Fig. 10 J) elongate. Female abdominal sternite VIII (Fig. 10 G) slightly serrate alike male on posterior margin. Spermatheca (Fig. 10 K) simplified in shape, about 1.5 times coiled near base. Measurements. BL 1.70 – 2.00, FBL 0.80 – 1.00, HL 0.30 – 0.31, HW 0.35 – 0.39, PL 0.30 – 0.33, PW 0.38 – 0.40, EL 0.29 – 0.30, EW 0.36 – 0.40, HTL 0.30 – 0.31, AL 0.61 – 0.67, AS-I 0.11 – 0.14, AS-II 0.06 – 0.09, AS-III 0.03 – 0.04.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF924E54F94F13F1F918.taxon	diagnosis	Differential diagnosis. In general appearance, the species is similar to B. sakishimana, but easily distinguished from it by the elytral shape being narrowed anteriorly and the shape of the spermatheca.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF924E54F94F13F1F918.taxon	etymology	Etymology. Bringing attention to the constricted body shape; adjective.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF924E54F94F13F1F918.taxon	distribution	Distribution. Japan (Honshû: Chiba-ken, Shimane-ken).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF924E54F94F13F1F918.taxon	discussion	Remarks. This species is found mainly on the Japan Sea coast of western Japan and east of Tôkyô Bay, but it is probably occurring in other areas of mainland Japan. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	description	(Figs 1 I; 2 I; 11; 21 C; 22) (Japanese name: Hokkai-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	materials_examined	Type material. H ඈඅඈඍඒඉൾ: J, JAPAN: H ඈκκൺ ං ൽත: Shunkunitai, 43.2725 N, 145.472 E, Nemuro-shi, Hokkaidô, 11. ix. 1999, M. Maruyama (KUM). P ൺ * © ൺඍඒඉൾඌ: H ඈκκൺ ං ൽත: 3 JJ 4 ♀♀, 96 unsexed spec., same data as holotype (KUM, NMPC, BMNH); 1 ♀, 4 unsexed spec., Usu-chô, Date-shi, Hokkaidô, 12. iv. 2020, Y. Tasaku (KUM). K ඒǕඌΗǕ: 1 ♀, Nokono-shima, Fukuoka-shi, Fukuoka-ken, 21. iv. 2018, T. Hashizume (KUM).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	description	Description. Body (Fig. 1 I) small, about 1.8 – 2.2 mm in length, covered with minute setae. Abdomen almost dark brown and darker than head and pronotum; elytra lighter than pronotum; legs yellowish brown; antenna brown, similar coloured with mouthparts and legs, lightly paler than body colour. Head evenly oval, wider than long, shorter than pronotum and elytra; eyes (Fig. 2 I) large, setae present between about 22 – 25 facets, slightly longer than tempora length; antenna brown, equals to total length of head and pronotum; segment I as long as II, almost 2.3 times longer than III, and as long as XI; antennal segments III – IV subquadrate, V – X almost same length and wider than long. Mouthparts (Figs 11 A – E) with distribution of labral setae as in Figure 11 A, with large and thick spiniform seta at antero-laterally; mandibles (Fig. 11 B) elongate triangle, asymmetrical, right mandible with some median tooth and irregular; median tooth longer than 1 / 4 of mandibles; left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 11 C) at obtuse angle; at tip with two spines very small and not closely interdigitating; 2 – 3 setae on inner margin of lacinia; labial palpus (Fig. 11 D) with most setae located basally, small ligula; mentum (Fig. 11 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” far away from apical setae. Pronotum shorter than elytra, wider than head. Elytra subquadrate but wider than long, slightly narrower than pronotum; hind wings present but very small, apparently flightless. Male abdominal sternite VIII (Fig. 11 F) with posterior margin slightly serrate. Median lobe of aedeagus as in Figures 11 H, I; paramere (Fig. 11 J) elongate. Female abdominal sternite VIII (Fig. 11 G) slightly serrate similar male on posterior margin. Spermatheca (Fig. 11 K) simplified in shape, about 1 time coiled near base. Measurements. BL 1.80 – 2.20, FBL 0.90 – 1.00, HL 0.25 – 0.30, HW 0.38 – 0.40, PL 0.33 – 0.35, PW 0.40 – 0.43, EL 0.35 – 0.38, EW 0.35 – 0.40, HTL 0.25 – 0.35, AL 0.60 – 0.68, AS-I 0.10 – 0.11, AS-II 0.09 – 0.10, AS-III 0.04 – 0.05.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	diagnosis	Differential diagnosis. In general appearance, the species is similar to B. fulvipennis sp. nov. and B. itsuroi sp. nov. but distinguished from them by the combination of the following character states: the body colour (especially head and pronotum colour) and the spermathecal shape.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	etymology	Etymology. Toponym. Named after Hokkaidô, the type locality of this new species; adjective.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	distribution	Distribution. Japan (Hokkaidô; Kyûshû: Fukuoka-ken).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1AFF914CE7F8FE15C8F8F8.taxon	discussion	Remarks. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC19FF974E58F88015D5FD98.taxon	description	(Figs 1 J; 2 J; 12; 21 B; 22) (Japanese name: Usucha-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC19FF974E58F88015D5FD98.taxon	materials_examined	Type material. H ඈඅඈඍඒඉൾ: J, JAPAN: H ඈඇඌΗǕ: Mouth of Obitsugawa, Kuzuma, Kisarazu-shi, Chiba-ken, 2. iii. 2007, H. Ono (KUM). P ൺ * © ൺඍඒඉൾඌ: H ඈඇඌΗǕ: 1 ♀, same data as holotype (KUM); 1 unsexed spec., same locality as holotype, 19. ii. 2007, H. Ono (KUM); 1 J 2 unsexed spec., Mouth of Obitsu-gawa, Kuroto, Kisarazu-shi, Chiba-ken, 19. ii. 2007, H. Ono (KUM); 1 unsexed spec., same locality, 12. iii. 2009, H. Ono (KUM); 1 ♀, 3 unsexed spec., same locality, 16. iii. 2009, H. Ono (KUM); 1 J, same locality, 4. i. 2010, H. Ono (KUM); 1 unsexed spec., same locality, 12. i. 2011, H. Ono (KUM); 1 unsexed spec., same locality, 14. i. 2011, H. Ono (KUM); 1 unsexed spec., Kushidagawa, Matsuzaka-shi, Mie-ken, 28. iv. 2002, Y. Kawakami (KUM); 10 unsexed spec., Himeshima, Yodo-gawa, Nishiyodo-gawa-ku, Ôsaka-fu, 12. iv. 2002, Y. Kawakami (KUM, MNPC, BMNH); 3 unsexed spec., Hanakawa Yodo-gawa, Nishiyodo-gawa-ku, Ôsaka-fu, 25. iv. 2002, Y. Kawakami (KUM); 4 unsexed spec., same locality, 28. iii. 2002, Y. Kawakami (KUM); 1 unsexed spec., Kinburo, Kitagi-shima, Kasaoka-shi, Okayama-ken, 11. x. 2008, O. Yamaji (KUM); 3 unsexed spec., Kurahashi-jima (Umikoshi), Kurahashi-chô, Kure-shi, Hiroshima-ken, 1. ii. 2009, K. Nakano (KUM). SΗ ං κ ඈκඎ: 11 unsexed spec., Kaigan-ji, Nishishirakata, Tadotsu-chô, Kagawa-ken, 31. iii. 2008, H. Fujimoto (KUM); 1 unsexed spec., Ôyatomi-machi, Sakaide-shi, Kagawa-ken, 29. iii. 2008, H. Fujimoto (KUM); 1 unsexed spec., Kôtô-gawa, Gôtô - chô, Takamatsu-shi, Kagawa-ken, 12. iv. 2008, H. Fujimoto (KUM).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC19FF974E58F88015D5FD98.taxon	description	Description. Body (Fig. 1 J) small, about 1.6 – 2.0 mm in length, covered with minute setae. Abdomen almost black and darker than head and pronotum; elytra lighter than pronotum; antenna and legs yellowish brown. Head evenly rounded, more or less produced anteriorly, wider than long, shorter than pronotum and elytra; eyes (Fig. 2 J) large, setae present between about 22 – 24 facets, slightly longer than tempora length; antenna yellowish brown, almost equals to total length of head and pronotum; segment I 1.7 times longer than II, almost 4 times longer than III, and longer than XI; antennal segments IV – X almost as long as wide. Mouthparts (Figs 12 A – E) with distribution of labral setae as in Figure 12 A, with large and thick spiniform seta antero-laterally; mandibles (Fig. 12 B) elongate triangle, asymmetrical, right with median tooth; median tooth shorter than 1 / 4 of mandibles; left without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 12 C) second spine very small, not obvious; 3 setae on inner margin of lacinia; labial palpus (Fig. 12 D) with most setae located basally; mentum (Fig. 12 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” far away from apical setae. Pronotum almost as long as elytra, wider than head. Elytra subquadrate but wider than long, almost as wide as pronotum; hind wings present but very small, apparently flightless. Male abdominal sternite VIII (Fig. 12 F) with posterior margin slightly serrate. Median lobe of aedeagus (Fig. 12 H) and dorsal view as in Figure 12 I; paramere (Fig. 12 J) elongate. Female abdominal sternite VIII (Fig. 12 G) slightly serrate alike male on posterior margin. Spermatheca (Fig. 12 K) about 2 times coiled near base. Measurements. BL 1.60 – 2.00, FBL 0.90 – 1.00, HL 0.28 – 0.30, HW 0.36 – 0.43, PL 0.36 – 0.40, PW 0.40 – 0.45, EL 0.35 – 0.41, EW 0.36 – 0.45, HTL 0.23 – 0.33, AL 0.61 – 0.76, AS-I 0.13 – 0.14, AS-II 0.07 – 0.08, AS-III 0.03 – 0.04.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC19FF974E58F88015D5FD98.taxon	diagnosis	Differential diagnosis. In general appearance the species is very similar to B. hokkaidensis sp. nov. and B. itsuroi sp. nov., but easily distinguished from B. hokkaidensis sp. nov. by the paler body colour and the shape of the spermatheca, and from B. itsuroi sp. nov. by the number of facets and the shape of the spermatheca.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC19FF974E58F88015D5FD98.taxon	etymology	Etymology. The species epithet refers to the yellowish - brown elytra; adjective.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC19FF974E58F88015D5FD98.taxon	distribution	Distribution. Japan (Honshû: Chiba-ken, Mie-ken, Ôsaka-fu, Okayama-ken, Hiroshima-ken; Shikoku: Kagawa-ken).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC19FF974E58F88015D5FD98.taxon	discussion	Remarks. This species has been found mainly in coasts of bay and inland sea such as Tôkyô-wan, Ise-wan and Setonaikai. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF974E66FD861368FCB8.taxon	description	(Figs 1 K; 2 K; 13; 20 H; 22) (Japanese name: Niseusucha-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF974E66FD861368FCB8.taxon	materials_examined	Type material. H ඈඅඈඍඒඉൾ: J, JAPAN: H ඈඇඌΗǕ: Kuruwa-kaigan, Akiya, Yokosuka-shi, Kanagawa-ken, 14. ix. 2013, H. Ono (KUM). P ൺ * © ൺඍඒඉൾඌ: H ඈඇඌΗǕ: 1 ♀, 18 unsexed spec., Uraga-wan, Nishi-uraga, Yokosuka-shi, Kanagawa-ken, 4. iii. 2013, H. Ono & I. Kawashima (KUM, NMPC, BMNH); 7 JJ 3 ♀♀, 9 unsexed spec., same data as holotype (KUM). K ඒǕඌΗǕ: 1 J 1 ♀, 16 unsexed spec., Kakise, Takashima, Nagasaki-shi, Nagasaki-ken, 4. iii. 2020, Y. Hisasue (KUM). N ൺඇඌൾ ං - ඌΗඈඍත: 1 J 1 ♀, 1 unsexed spec., Mage-shima (near Tanega-shima), Kagoshima-ken, 14. x. 2000, M. Maruyama (under stone on sandy beach, in a port) (KUM).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF974E66FD861368FCB8.taxon	description	Description. Body (Fig. 1 K) small, about 1.7 – 1.9 mm in length, covered with minute setae. Abdomen almost black and darker than head and pronotum; elytra lighter than pronotum; legs yellowish brown. Head evenly rounded, more or less produced anteriorly, wider than long, shorter than pronotum and elytra; eyes (Fig. 2 K) large, setae present between about 29 – 31 facets, longer than tempora; antenna yellowish brown, almost equals to total length of head and pronotum; segment I as long as II, almost 3 times longer than III, and longer than XI; antennal segments VII – X almost wider than long. Mouthparts (Figs 13 A – E) with distribution of labral setae as in Figure 13 A, with large and thick spiniform seta antero-laterally; mandibles (Fig. 13 B) elongate triangle, asymmetrical, right with median tooth; median tooth shorter than 1 / 4 of mandibles; left without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 13 C) at right angle; at tip second spine very small, not obvious; 3 setae on inner margin of lacinia; labial palpus (Fig. 13 D) with most setae located basally; mentum (Fig. 13 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” far away from apical setae. Pronotum shorter than elytra, as wide as head. Elytra subquadrate, wider than pronotum; hind wings present but small, apparently flightless. Male abdominal sternite VIII (Fig. 13 F) with posterior margin slightly serrate. Median lobe of aedeagus (Fig. 13 H) and dorsal view as in Figure 13 I; paramere (Fig. 13 J) elongate. Female abdominal sternite VIII (Fig. 13 G) slightly serrate alike male on posterior margin. Spermatheca (Fig. 13 K) of complicated shape, very small, shorter than 0.25 mm and several times coiled near base. Measurements. BL 1.70 – 1.90, FBL 0.80 – 1.00, HL 0.31 – 0.33, HW 0.38 – 0.41, PL 0.33 – 0.38, PW 0.40 – 0.41, EL 0.42 – 0.43, EW 0.41 – 0.43, HTL 0.33 – 0.39, AL 0.61 – 0.66, AS-I 0.10 – 0.11, AS-II 0.09 – 0.10, AS-III 0.03 – 0.04.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF974E66FD861368FCB8.taxon	diagnosis	Differential diagnosis. In general appearance the species is very similar to B. fulvipennis sp. nov. and B. hokkaidensis sp. nov., but can be easily distinguished from them by the number of facets, the shape of the spermatheca, and the antennal segment I as long as II.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF974E66FD861368FCB8.taxon	etymology	Etymology. The new species is named after Mr. Itsuro Kawashima, a scientific illustrator and researcher of fireflies, who helped collect the type series of this species in Kanagawa.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF974E66FD861368FCB8.taxon	distribution	Distribution. Japan (Honshû: Kanagawa-ken; Kyûshû: Nagasaki-ken; Nansei-shotô: Kagoshima-ken: Tanega - shima).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF974E66FD861368FCB8.taxon	discussion	Remarks. Unfortunately, one of the habitats in Uraga-wan, Kanagawa Prefecture has likely disappeared due to recent seawall construction. This species has a wide distribution but is found very locally where it occurs. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF9A4C9AFC66155FFC18.taxon	description	(Figs 1 L; 2 L; 14; 21 B; 22) (Japanese name: Hime-nagisa-hanekakushi) Type material. H ඈඅඈඍඒඉൾ: ♀, JAPAN: H ඈඇඌΗǕ: Mouth of Obitsu-gawa, Kuroto, Kisarazu-shi, Chiba-ken, 12. iii. 2009, H. Ono (KUM). P ൺ * © ൺඍඒඉൾඌ: H ඈඇඌΗǕ: 3 ♀♀, same data as holotype (KUM); 1 ♀, same locality as holotype, 16. iii. 2009, H. Ono (KUM). Description. Body (Fig. 1 L) small, about 1.5 – 1.8 mm in length, covered with minute setae. Basically brown, head darker than pronotum and elytra, pronotum and elytra similarly coloured, from lower part of posteriorly abdominal tergite V to upper part of tergite VII infuscate; antenna, mouthparts and legs similarly coloured, lightly paler than head colour and paler toward apex. Head oval, wider than long, shorter than elytra; eyes (Fig. 2 L) large, reniform, longer than 1 / 2 of head length, with 22 – 24 facets; antennal length little shorter than distance from anterior margin of head to posterior margin of elytra; segment I 1.4 times longer than II, almost 3 times longer than III, and longer than XI; antennal segments I – III and XI longer than wide, antennal segments IV – VI almost as long as wide, antennal segments VII – X wider than long. Mouthparts (Figs 14 A – E) with distribution of labral setae as in Figure 14 A, with large and thick spiniform seta at antero-laterally; mandibles (Fig. 14 B) elongate triangle, asymmetrical, right with median tooth; median tooth shorter than 1 / 4 of mandibles; left without tooth, blunt at apex; maxilla with apical apex of lacinia (Fig. 14 C) with second spines small, at tip with two spines not closely interdigitating; 3 setae on inner margin of lacinia; labial palpus (Fig. 14 D) with most setae located basally; mentum (Fig. 14 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” of mentum slightly lower than level of anterior margin. Pronotum shorter than elytra, wider than head. Elytra subquadrate, almost as wide as pronotum; hind wings fully developed. Male unknown. Female abdominal sternite VIII in Figure 14 F. Spermatheca (Fig. 14 G) very small, simplified in shape, about one time coiled near base. Measurements. BL 1.50 – 1.80, FBL 0.83 – 0.85, HL 0.23 – 0.28, HW 0.31 – 0.32, PL 0.29 – 0.30, PW 0.34 – 0.35, EL 0.35 – 0.36, EW 0.35 – 0.36, HTL 0.28 – 0.30, AL 0.58 – 0.63, AS-I 0.10 – 0.11, AS-II 0.07 – 0.08, AS-III 0.03 – 0.04.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF9A4C9AFC66155FFC18.taxon	diagnosis	Differential diagnosis. The species is distinct based on the body colour and shape. In body size this species is close to B. minuta but easily distinguished from it by the number of facets and by the fully developed hind wings.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF9A4C9AFC66155FFC18.taxon	etymology	Etymology. This new species is named after Mrs. Aiko Ono for supporting the entomological life of her husband H. Ono, the second author of this paper.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF9A4C9AFC66155FFC18.taxon	distribution	Distribution. Japan (Honshû: Chiba-ken).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC1FFF9A4C9AFC66155FFC18.taxon	discussion	Remarks. The type series consists of only female specimens. The second author repeatedly searched for this species in the type locality, but no additional specimens were found. Nevertheless, we describe it because of its very distinct general appearance. This species was only found in Chiba-ken. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	description	(Figs 1 M; 2 M; 15; 22) (Japanese name: Okinawa-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	materials_examined	Type material. H ඈඅඈඍඒඉൾ: J, JAPAN: N ൺඇඌൾ ං - ඌΗඈඍත: Gesashi, Higashison, Okinawa-jima, Okinawa-ken, 15. viii. 2004, M. Moriguchi (KUM). P ൺ * © ൺඍඒඉൾඌ: 1 ♀, Mage-shima (near Tanega-shima), Kagoshima-ken, 14. x. 2000, M. Maruyama (under stone on sandy beach, in a port) (KUM); 3 ♀♀, 3 unsexed spec., same data as holotype. (KUM).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	description	Description. Body (Fig. 1 M) small, about 2.1 – 2.2 mm in length, covered with minute setae. Basically, yellowish brown. head and pronotum similarly coloured, elytra lightly paler than body colour, abdomen darkest; antenna, mouthparts and legs similarly coloured, lightly paler than head colour and paler toward apex. Head oval, wider than long, shorter than pronotum and elytra; eyes (Fig. 2 M) small, reniform, longer than 1 / 2 of head length, with 29 – 31 facets; antennal length shorter than distance from anterior margin of head to posterior margin of elytra; segment I as long as II, almost 2 times longer than III, and as long as XI; antennal segments I – IV and XI longer than wide; antennal segments V – X almost as long as wide. Mouthparts (Figs 15 A – E) with distribution of labral setae as in Figure 15 A; mandibles (Fig. 15 B) elongate triangle, asymmetrical, right mandible with median tooth; median tooth shorter than 1 / 4 of mandibles; left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 15 C) at right angle and second spines not very small, at tip with two spines not closely interdigitating; 4 setae on inner margin of lacinia; labial palpus (Fig. 15 D) with most setae located basally; mentum (Fig. 15 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” of mentum lower than level of anterior margin. Pronotum shorter than elytra, wider than head. Elytra subquadrate, wider than pronotum; hind wings fully developed. Male abdominal sternite VIII (Fig. 15 F) with posterior margin obviously serrate. Median lobe of aedeagus (Fig. 15 H) and dorsal view in Figure 15 I; paramere (Fig. 15 J) elongate. Female abdominal sternite VIII (Fig. 15 G) with posterior margin slightly serrate but obviously smoother than in male. Spermatheca (Fig. 15 K) simplified in shape, about half times coiled near base. Measurements. BL 2.10 – 2.20, FBL 1.00 – 1.10, HL 0.33 – 0.38, HW 0.40 – 0.43, PL 0.38 – 0.40, PW 0.43 – 0.45, EL 0.48 – 0.50, EW 0.50 – 0.51, HTL 0.33 – 0.36, AL 0.78 – 0.85, AS-I 0.10 – 0.13, AS-II 0.09 – 0.11, AS-III 0.05 – 0.06.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	diagnosis	Differential diagnosis. In general appearance the species in is closely allied to B. moriguchii sp. nov., but easily distinguished from it by the shape of the spermatheca, and lack of large, thick spiniform seta at the antero-lateral part of the labrum, and the posterior margin of the tergite X being rounded medially.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	etymology	Etymology. Toponym. Named after Okinawa, the type locality of this new species; adjective.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	distribution	Distribution. Japan (Nansei-shotô: Kagoshima-ken: Tanega-shima; Okinawa-ken: Okinawa-jima).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	discussion	Remarks. This species probably occurs in other areas, especially the coastline of southwest Japan. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22. Bryothinusa nigra sp. nov. (Figs 1 N; 16; 21 B, D; 22) (Japanese name: Kuro-nagisa-hanekakushi) “ Bryothinusa nakanei ”: S ൺඐൺൽൺ (1971): 88 (misidentification); A * ü ඇ & J ൾඈඇ (2004): 34 (misidentification; illustration of spermatheca); A * ü ඇ et al. (2017): 307 (misidentification; in checklist of Staphylinidae in Korea). Type material. H ඈඅඈඍඒඉൾ: J, JAPAN: H ඈඇඌΗǕ: Mouth of Obitsu-gawa, Kuroto, Kisarazu-shi, Chiba-ken, 24. i. 2011, H. Ono (KUM). P ൺ * © ൺඍඒඉൾඌ: H ඈඇඌΗǕ: 1 unsexed spec., same locality as holotype, 19. ii. 2007, H. Ono (KUM); 2 JJ 4 ♀♀, 15 unsexed spec., same locality as holotype, 12. iii. 2009, H. Ono (KUM, NMPC, BMNH); 1 J 1 ♀, 5 unsexed spec., same locality as holotype, 16. iii. 2009, H. Ono (KUM); 1 J 5 unsexed spec., same locality as holotype, 4. i. 2010, H. Ono (KUM); 2 unsexed spec., same locality as holotype, 14. i. 2011, H. Ono (KUM); 3 JJ 2 ♀♀, 17 unsexed spec., same data as holotype (KUM); 1 ♀, 3 unsexed spec., Edo-gawa, Gyôtoku, Ichikawa-shi, Chiba-ken, 18. vi. 2012, H. Ono (KUM); 1 unsexed spec., Kushida-gawa, Matsunase-chô, Matsuzaka-shi, Mie-ken, 2. iv. 2002, T. Wada (KUM). SΗ ං κ ඈκඎ: 1 ♀, 7 unsexed spec., Nio, Nio-chô, Kagawa-ken, 20. iv. 2008, H. Fujimoto (KUM); 2 JJ 1 ♀, Kaigan-ji, Nishishirakata, Tadotsu-chô, Kagawa-ken, 31. iii. 2008, H. Fujimoto (KUM). K ඒǕඌΗǕ: 2 JJ 1 ♀, 7 unsexed spec., Aiboshi-gawa, Aiboshi, Minamisatsuma-shi, Kagoshima-ken, 20. iv. 2008, M. Maruyama (KUM). N ൺඇඌൾ ං - ඌΗඈඍත: 1 J 6 ♀♀, 19 unsexed spec., Uken-son, Edateku-jima (near Amami-Ôshima), Kagoshima-ken, 5. vi. 2008, M. Maruyama (KUM).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	description	Description. Body (Fig. 1 N) small about 2.0 – 2.5 mm in length, covered with minute setae. Body almost black; elytra lighter than other parts; antenna and legs brown. Head evenly rounded, more or less produced anteriorly, wider than long, almost as long as pronotum, shorter than elytra; eyes (Fig. 2 N) large, setae present between about 32 – 33 facets, longer than 2 / 3 head length; antenna almost as long as distance from anterior margin of head to posterior margin of pronotum; segment I 1.3 times longer than II, almost 2.7 times longer than III, and longer than XI; antennal segments V – X almost as long as wide. Mouthparts (Figs 16 A – E) with distribution of labral setae as in Figure 16 A, with large and thick spiniform seta at antero-laterally; mandibles (Fig. 16 B) elongate triangle, asymmetrical, right mandible with median tooth; median tooth almost as long as 1 / 4 of mandibles; left without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 16 C) at tip with two spines very small and not closely interdigitating; 4 setae on inner margin of lacinia; labial palpus (Fig. 16 D) with most setae located basally; mentum (Fig. 16 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” far away from apical setae. Pronotum shorter than elytra, wider than head. Elytra subquadrate but wider than long, wider than pronotum; hind wings fully developed. Male abdominal sternite VIII (Fig. 16 F) with posterior margin slightly serrate. Median lobe of aedeagus (Fig. 16 H) and dorsal view as in Figure 16 I; paramere (Fig. 16 J) elongate. Female abdominal sternite VIII (Fig. 16 G) with posterior margin slightly serrate alike male. Spermatheca (Fig. 16 K) very small, shorter than 0.25 mm. Measurements. BL 2.00 – 2.50, FBL 1.00 – 1.30, HL 0.35 – 0.40, HW 0.41 – 0.44, PL 0.31 – 0.42, PW 0.40 – 0.54, EL 0.41 – 0.46, EW 0.50 – 0.56, HTL 0.37 – 0.39, AL 0.65 – 0.70, AS-I 0.11 – 0.14, AS-II 0.08 – 0.10, AS-III 0.04 – 0.05. Differential diagnosis. In general appearance, the species is closely similar to B. nakanei and B. gangjinensis, but it is easily distinguished from them by a different shape and size of spermatheca. It differs from B. nakanei also by the shape of the posterior margin of tergite X (rounded medially in the new species).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	etymology	Etymology. The species epithet, the Latin adjective niger (- a, - um), refers to the blackish colour of the body.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	distribution	Distribution. Japan (Honshû: Chiba-ken, Kanagawa-ken, Mie-ken; Shikoku: Kagawa-ken; Kyûshû: Kagoshima-ken; Nansei-shotô: Kagoshima-ken: Edateku-jima); Korea.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	discussion	Remarks. S ൺඐൺൽൺ (1971) illustrated the mouthparts and sexual organs of “ B. nakanei ” but the large, thick spiniform seta at the antero-lateral part of the labrum and the shape of aedeagus is very similar to those of B. nigra sp. nov. Therefore, we consider that the Figure 3 (S ൺඐൺൽൺ 1971) is of B. nigra sp. nov. A * ü ඇ & J ൾඈඇ (2004) illustrated the spermatheca of “ B. nakanei ” based on the Korean specimens but its shape agrees well with that of B. nigra sp. nov .. Therefore, we consider that their figure (A * ü ඇ & J ൾඈඇ 2004: Fig. 18) represents the spermatheca of B. nigra sp. nov. The misidentification by A * ü ඇ & J ൾඈඇ (2004) was probably caused by the redescription of S ൺඐൺൽൺ (1971). See also the Remarks of B. nakanei below. This species is found mainly west of Kantô District. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22. Bryothinusa nakanei (Sawada, 1955) (Figs 1 O; 2 O; 17; 21 B, D-F; 22) (Japanese name: Nakane-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	description	key to known species of Bryothinusa); A ඌ * ü ൾ (2004): 592 (in annotated catalogue of world species); S ආൾඍൺඇൺ (2004): 464 (in catalogue of Palearctic species); F * © ൺඇκ & A * ü ඇ (2011): 27 (in checklist); S ർ * ü * “ අκൾ & S ආൾඍൺඇൺ (2015): 676 (in catalogue of Palearctic species); L ං ඎ et al. (2020): 589 (in key to Japanese species), 592 (in checklist).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	materials_examined	Type material examined. H ඈඅඈඍඒඉൾ: unsexed specimen (not dissected), JAPAN: N ൺඇඌൾ ං - ඌΗඈඍත: “ Halesthenus / nakanei / Sawada. // TAKARA- JIMA / (TOKARA Is.) / 29. v. 1953 / coll. S. Uéno / Holotype / 49 - 16 ”. Deposited in the Hokkaido University Museum. Additional material examined. JAPAN: N ൺඇඌൾ ං - ඌΗඈඍත: 1 J, Mage - shima (near Tanega-shima), Kagoshima-ken, 14. x. 2000, M. Maruyama (under stone on sandy beach, in a port); 8 JJ 6 ♀♀, 57 unsexed spec., Ôgomori, Takara-jima, Tokara-rettô, Kagoshima-ken, 3. vii. 2019, T. Liu, N. Tsuji & H. Ono; 1 unsexed spec., same locality, 6. vii. 2019, T. Liu, N. Tsuji & H. Ono; 2 ♀♀, 9 unsexed spec., Tean, Setouchi-chô, Amami - Ôshima, Kagoshima-ken, 19. vii. 2007, M. Moriguchi; 2 JJ 2 unsexed spec., Chatan-chô, Okinawa-jima, Nansei-shôtô, Okinawa-ken, 26. v. 2002, M. Moriguchi; 3 ♀♀, 9 unsexed spec., Senaha, Yomitani-son, Okinawajima, Okinawa-ken, 21. v. 2004, M. Moriguchi; 1 J 1 ♀, 2 unsexed spec., Fuchaku, Onna-son, Okinawa-ken, 8. iv. 2000, T. Watanabe; 2 JJ 3 ♀♀, 31 unsexed spec., Todoroki-gawa, Shiraho, Ishigaki-shi, Okinawa-ken, 29. iii. 2010, T. Hayama; 2 ♀♀, 10 unsexed spec., Hira-kawa, Ishigaki-shi, Ishigaki-jima, Okinawa-ken, 29. iii. 2010, T. Hayama; 2 ♀♀, 5 unsexed spec., Kabira-wan, Ôtake, Ishigaki-jima, Okinawa-ken, 3 - 11. vi. 2000, S. Hori; 2 ♀♀, 11 unsexed spec., same locality, 1. vi. 2003, M. Moriguchi; 2 JJ 1 ♀, 9 unsexed spec., same locality, 26. iii. 2015, Y. Hisasue; 1 ♀, 2 unsexed spec., same locality, 11. iii. 2000, T. Kurihara; 2 JJ 1 ♀, 50 unsexed spec., Nagura, Ishigaki-jima, Okinawa-ken, 21. iv. 2009, M. Maruyama; 3 ♀♀, 5 unsexed spec., Nagura-anparu, Nagura, Ishigaki-shi, Ishigaki-jima, Okinawa-ken, 4. ii. 2009, H. Ono; 1 J 1 unsexed spec., Nakasuji, Ishigaki-shi, Ishigaki-jima, Okinawa-ken, 12. iii. 2000, T. Kurihara; 4 ♀♀, 8 unsexed spec., Nakama-gawa, Ôtomi, Iriomote-jima, Okinawa-ken, 19 – 20. iv. 2009, M. Maruyama; 1 ♀, Mouth of Honeragawa, Takana, Taketomi-chô, Iriomote-jima, Okinawa-ken, 24. vi. 2019, T. Liu & N. Tsuji; 2 unsexed spec., Mouth of Maira-gawa, Taketomi-chô, Iriomote-jima, Okinawa-ken, 21. iii. 2000, T. Kurihara; 2 ♀♀, 53 unsexed spec., Haemida, Taketomi-chô, Iriomote-jima, Okinawa-ken, 19. iii. 2000, T. Kurihara. Deposited at KUM but some will be distributed to NMPC and BMNH.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	description	Redescription. Body (Fig. 1 O) small, about 2.0 – 2.2 mm in length, covered with minute setae. Abdomen almost black and darker than head and pronotum; elytra lighter than pronotum, clearly bicolour, base darker than apex; legs yellowish brown. Head evenly rounded, more or less produced anteriorly, wider than long, shorter than pronotum and elytra; eyes (Fig. 2 O) large, setae present between about 34 – 36 facets, slightly longer than 2 / 3 of head length; antenna almost as long as distance from anterior margin of head to posterior margin of pronotum; segment I 1.3 times longer than II, almost 2.8 times longer than III, and longer than XI; antennal segments VII – X slightly wider than long. Mouthparts (Figs 17 A – E) with distribution of labral setae as in Figure 17 A, with large and thick spiniform seta at antero-laterally; mandibles (Fig. 17 B) elongate triangle, asymmetrical, right with median tooth; median tooth shorter than 1 / 4 of mandibles; left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 17 C) at right angle; at tip with two spines not closely and second spine very small, not obvious; 6 setae on inner margin of lacinia; labial palpus (Fig. 17 D) with most setae located basally; mentum (Fig. 17 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” far away from apical setae. Pronotum shorter than elytra, almost as wide as head. Elytra subquadrate but wider than long, wider than pronotum; hind wings fully developed. Male abdominal sternite VIII (Fig. 17 F) with posterior margin slightly serrate; medial posterior margin of tergite X protruded (Fig. 17 H). Median lobe of aedeagus and dorsal view as in Figures 17 J, K; paramere (Fig. 17 L) elongate. Female abdominal sternite VIII (Fig. 17 G) slightly sunken at median posterior margin but wider and smoother than in male; medial posterior margin of tergite X more protruded than in male (Fig. 17 I). Spermatheca (Fig. 17 M) of complicated shape, about 2.5 times coiled near base. Measurements. BL 2.00 – 2.20, FBL 1.10 – 1.20, HL 0.35 – 0.38, HW 0.45 – 0.50, PL 0.40 – 0.41, PW 0.45 – 0.50, EL 0.48 – 0.50, EW 0.45 – 0.54, HTL 0.35 – 0.38, AL 0.75 – 0.80, AS-I 0.11 – 0.15, AS-II 0.09 – 0.11, AS-III 0.04 – 0.05.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	diagnosis	Differential diagnosis. In general appearance the species is very similar to B. nigra sp. nov. and B. gangjinensis. In the shape of the spermatheca, the species is closely similar to B. moriguchii sp. nov. However, the new species is easily distinguished from B. nigra sp. nov. and B. gangjinensis by the medial posterior margin of tergite X being protruded and is different from B. moriguchii sp. nov. in the body colour being darker.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	distribution	Distribution. Japan (Nansei-shotô: Kagoshima-ken: Tanega-shima, Takara-jima, Amami-Ôshima; Okinawa-ken: Okinawa-jima, Ishigaki-jima, Iriomote-jima).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC12FF9F4E51FC0615F8F7D8.taxon	discussion	Remarks. S ൺඐൺൽൺ (1955) originally described B. nakanei based on a single specimen from Takara-jima, Tokara-rettô, Nansei-shotô, without illustrating any sexual characters. S ൺඐൺൽൺ (1971) redescribed “ B. nakanei ” in detail including the male aedeagus based on the “ holotype. ” However, we examined the holotype of B. nakanei deposited in the late Dr. Takehiko Nakane collection of the Hokkaido University Museum and confirmed it was not dissected. S ൺඐൺ- ൽൺ (1971) also examined a male specimen of this species from Aburatsubo of Kanagawa, Honshû. The redescription is probably based on the specimen from Kanagawa. Moreover, the redescription does not coincide with the holotype but actually agrees well with the characteristics of B. nigra sp. nov. Bryothinusa nakanei is very similar to B. nigra sp. nov. in general appearance and S ൺඐൺൽൺ (1971) probably confused them. As discussed in the Remarks of B. nigra sp. nov., A * ü ඇ & J ൾඈඇ (2004) described the spermatheca of “ B. nakanei ” from Korea but it was of the same shape as that of B. nigra sp. nov. Bryothinusa nakanei may not occur in Korea because it has not been found in temperate zones in Japan, but in Nansei-shotô, a subtropical zone. This species likely occurs in other areas, mainly in Nansei-shotô. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	description	(Figs 1 P; 2 P; 18; 22) (Japanese name: Moriguchi-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	materials_examined	Type material. H ඈඅඈඍඒඉൾ: J, JAPAN: N ൺඇඌൾ ං - ඌΗඈඍත: Yagaji-jima, Nago-shi, Okinawa-ken, 21. iii. 2005, M. Moriguchi (KUM). P ൺ * © ൺඍඒඉൾඌ: 3 JJ 3 ♀♀, 4 unsexed spec., same data as holotype (KUM, NMPC, BNMH); 2 ♀♀, 4 unsexed spec., Uehara, Iriomote-jima, 2. iv. 2004, M. Moriguchi (KUM).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	description	Description. Body (Fig. 1 P) small, about 1.9 – 2.2 mm in length, covered with minute setae. Basically, dark brown, elytra brown palest, head and pronotum similarly coloured, paler than abdomen; antenna, mouthparts and legs similarly coloured, lightly paler than head colour and paler toward apex. Head oval, wider than long, almost as long as pronotum, shorter than elytra; eyes (Fig. 2 P) small, reniform, longer than 2 / 3 of head length, with 32 – 34 facets; antennal length slightly longer than distance from anterior margin of head to posterior margin of pronotum; segment I as long as II, almost 2.5 times longer than III, and as long as XI; antennal segments I – III and XI longer than wide, antennal segments IV – X almost as long as wide. Mouthparts (Figs 18 A-E) with distribution of labral setae as in Figure 18 A, with large and thick spiniform seta antero-laterally; mandibles (Fig. 18 B) elongate triangle, asymmetrical, right with median tooth; median tooth shorter than 1 / 4 of mandibles; left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 18 C) with second spines very small and not obvious, at tip with two spines not closely interdigitating; 3 setae on inner margin of lacinia; labial palpus (Fig. 18 D) with most setae located basally; mentum (Fig. 18 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” of mentum lower than level of anterior margin. Pronotum shorter than elytra, almost as wide as head. Elytra subquadrate but wider than long, wider than pronotum; hind wings fully developed. Male abdominal sternite VIII (Fig. 18 F) with posterior margin slightly serrate. Medial posterior margin not protruded, sharp in form on tergite X of male (Fig. 18 H). Median lobe of aedeagus (Fig. 18 J) and dorsal view in Figure 18 K; paramere (Fig. 18 L) elongate. Female abdominal sternite VIII (Fig. 18 G) with posterior margin slightly sunken but smoother than in male. Medial posterior margin protruded sharp in form on tergite X of female (Fig. 18 I). Spermatheca (Fig. 18 M) of complicated shape, about 2.5 times coiled near base. Measurements. BL 1.90 – 2.20, FBL 1.00 – 1.10, HL 0.35 – 0.40, HW 0.43 – 0.45, PL 0.38 – 0.40, PW 0.45 – 0.46, EL 0.45 – 0.48, EW 0.50 – 0.55, HTL 0.33 – 0.38, AL 0.75 – 0.85, AS-I 0.10 – 0.13, AS-II 0.10 – 0.13, AS-III 0.04 – 0.05.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	diagnosis	Differential diagnosis. In general appearance the species is very similar to B. okinawana sp. nov., but easily distinguished from it by having large, thick spiniform seta at the antero-lateral part of the labrum, and the coil number of the spermathecal basal part and of the posterior margin of the tergite X being protruded.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	etymology	Etymology. This new species is named after Mr. Mitsuru Moriguchi, the collector of the type series.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	distribution	Distribution. Japan (Nansei-shotô: Okinawa-ken: Yagajijima, Iriomote-jima).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC17FFA24CF2FF46167EFEB8.taxon	discussion	Remarks. This species may occur in other areas, especially the coastline of western Japan. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	description	(Figs 1 Q; 1 Q; 19; 21 F; 22) (Japanese name: Kanjin-nagisa-hanekakushi)	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	description	talogue of world species); A ඌൺඇඈ & K ඈ * ° ං ආൺ (2010): 297 (new record from Japan: Tanega-shima); F * © ൺඇκ & A * ü ඇ (2011): 26 (in checklist); S ർ * ü * “ අκൾ & S ආൾඍൺඇൺ (2015): 676 (in catalogue of Palearctic species); A * ü ඇ et al. (2017): 307 (in checklist of the Staphylinidae in Korea); L ං ඎ et al. (2020): 589 (in key of Japanese species), 592 (in checklist).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	materials_examined	Type material. Not examined. Material examined. JAPAN: H ඈඇඌΗǕ: 2 JJ 17 unsexed spec., Mouth of Minato-gawa, Minato, Futtsu-shi, Chiba-ken, 27. v. 2013, H. Ono; 4 JJ 1 ♀, 24 unsexed spec., same locality, 24. viii. 2009 H. Ono; 2 JJ 2 ♀♀, 7 unsexed spec., Koajiro-wan, Koajiro, Misaki-machi, Miura-shi, Kanagawa-ken, 23. v. 2013, H. Ono; 1 J 2 ♀♀, 67 unsexed spec., Miya-gawa, Iso-chô, Ise-shi, Mie-ken, 8. vi. 2008, K. Akita; 2 ♀♀, 6 unsexed spec., Hannan 2 - ku, Kishiwada-shi, Ôsaka-fu, 11. vi. 2010, Y. Kawakami; 1 ♀, 5 unsexed spec., same locality, 7. x. 2010, Y. Kawakami; 1 ♀, Arida-gawa, Miyazaki-chô, Arida-shi, Wakayama-ken, 28. v. 2002, Y. Kawakami. SΗ ං κ ඈκඎ: 1 ♀, 1 unsexed spec., Kabe-gawa, Kamoshô, Sanuki-shi, Kagawa-ken, 13. iv. 2008, H. Fujimoto; 1 ♀, 2 unsexed spec., Kôtô-gawa, Gôtô-chô, Takamatsu-shi, Kagawa-ken, 12. iv. 2008, H. Fujimoto; 1 J 3 unsexed spec., Kamobe-gawa, Kamoshô, Sanuki - shi, Kagawa-ken, 12. iv. 2008, H. Fujimoto; 1 J, Aya-gawa, Ejiri-chô, Sakaide-shi, Kagawa-ken, 12. iv. 2008, H. Fujimoto; 2 unsexed spec., Nishihabu-machi, Matsuyama-shi, Ehime-ken, 19. iv. 2009, H. Suenaga. K ඒǕඌΗǕ: 1 J, Iki-no-matsubara, Nishi-ku, Fukuoka-shi, Fukuoka-ken, 1. vii. 2018, Y. Hisasue; 1 J 1 ♀, 13 unsexed spec., Chichibu-ga-ura, Shimabara, Nagasaki-ken, 10. v. 2001, T. Watanabe; 1 ♀, 3 unsexed spec., Kawashima, Nobeoka-shi, Miyazaki-ken, 24. vii. 2006, J. Nakajima; 1 ♀, 4 unsexed spec., Nagayoshi-kawa, Fukiage-chô, Hioki-shi, Kagoshima-ken, 9. xii. 2007, K. Nakamura; 1 ♀, 6 unsexed spec., Manose-gawa, Masukawa, Kaseda, Minamisatsuma-shi, Kagoshima-ken, 20. iv. 2008, M. Maruyama; 1 J 2 ♀♀, 11 unsexed spec., Aiboshi-gawa, Aiboshi, Minamisatsuma-shi, Kagoshima-ken, 20. iv. 2008, M. Maruyama. N ൺඇඌൾ ං - ඌΗඈඍත: 3 JJ 6 unsexed spec., Ou-jima, Nago-shi, Okinawa-ken, 30. iv. 2006, M. Moriguchi; 2 unsexed spec., Awase-Higata, Okinawa-shi, Okinawa-ken, 30. iv. 2006, M. Moriguchi; 11 unsexed spec., Makiya, Nago-shi, Okinawa-jima, Okinawa-ken, 4. vii. 2010, M. Moriguchi. Deposited at KUM but some will be distributed to NMPC and BMNH.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	description	Redescription. Body (Fig. 1 Q) small, about 1.9 – 2.4 mm in length, covered with minute setae. Abdomen almost black, darker than head and pronotum; elytra darker than pronotum, bicoloured but not clearly, base darker than apex; antenna and legs yellowish brown. Head evenly rounded, more or less produced anteriorly, wider than long, shorter than elytra; eyes (Fig. 2 Q) large, setae present between about 32 – 34 facets, slightly longer than 2 / 3 of head length; antennal length slightly longer than distance from anterior margin of head to posterior margin of pronotum; segment I 1.5 times longer than II, almost 2.8 times longer than III, and longer than XI; antennal segments V – X almost as long as wide. Mouthparts (Figs 19 A – E) with distribution of labral setae as in Figure 19 A; mandibles (Fig. 19 B) elongate triangle, asymmetrical, right with densely median tooth, left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (Fig. 19 C) at right angle; at tip with two spines not clearly; 7 setae on inner margin of lacinia; labial palpus (Fig. 19 D) with most setae located basally; mentum (Fig. 19 E) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “ v seta ” far away from apical setae. Pronotum shorter than elytra, wider than head. Elytra subquadrate, wider than pronotum; hind wings fully developed. Male abdominal sternite VIII (Fig. 19 F) with posterior margin slightly serrate. Median lobe of aedeagus and dorsal view as in Figures 19 H, I; paramere (Fig. 19 J) with apical lobe elongate. Female abdominal sternite VIII (Fig. 19 G) slightly sunken at median posterior margin but wider and smoother than in male. Spermatheca (Fig. 19 K) large, about 1.5 times coiled near base and larger than 0.25 mm. Measurements. BL 1.90 – 2.40, FBL 0.90 – 1.20, HL 0.29 – 0.30, HW 0.36 – 0.45, PL 0.31 – 0.45, PW 0.37 – 0.49, EL 0.36 – 0.55, EW 0.43 – 0.56, HTL 0.28 – 0.43, AL 0.60 – 0.75, AS-I 0.11 – 0.14, AS-II 0.06 – 0.11, AS-III 0.04 – 0.05.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	diagnosis	Differential diagnosis. In general appearance, the species closely resembles B. nigra sp. nov. and B. nakanei but is easily distinguished from B. nigra sp. nov. in the shape and size of the spermatheca and is different from B. nakanei in having the posterior margin of the tergite X rounded medially.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	distribution	Distribution. Japan (Honshû: Chiba, Kanagawa, Mie, Ôsaka, Wakayama; Shikoku: Kagawa, Ehime; Kyûshû: Fukuoka, Nagasaki, Miyazaki, Kagoshima; Nansei-shotô: Tanega-shima, Okinawa: Ou-jima, Okinawa-jima); Korea (Jeonnam).	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
03CB023BAC2AFFA24F99FE6614F5FA98.taxon	discussion	Remarks. This species is mainly found west of Kantô District and Okinawa Island. A distribution map of examined specimens of this species in Japan is illustrated in Figure 22.	en	Liu, Tian-tian, Ono, Hiroki, Maruyama, Munetoshi (2021): Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1): 163-201, DOI: 10.37520/aemnp.2021.009, URL: http://dx.doi.org/10.37520/aemnp.2021.009
