taxonID	type	description	language	source
03CB0C7FFFA1FFCF9D77C06DA135F90A.taxon	description	Figs 1 – 8	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA1FFCF9D77C06DA135F90A.taxon	type_taxon	Type species Dictyophara jocosa Gerstaecker, 1895 (original designation).	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA1FFCF9D77C06DA135F90A.taxon	diagnosis	Emended diagnosis The genus can be distinguished by the following combination of characters: cephalic process moderately long and slender, conical and strongly curved upward; vertex with lateral carinae abruptly constricted and strongly upturned in front of eyes, then gradually convergent anteriad and acuminate at apex; frons with median carina robust and strongly convex, intermediate carinae distinctly expanded outward in apical third, their apical portion being distinctly visible in posterodorsal view; genae with a longitudinal carina above eyes; pronotum with median carina sharp and high, intermediate carinae absent; forewings with sparse transverse veins, stigmal area small and quadrangular, with two or three cells; legs elongate and slender, fore femur not flattened and dilated, without spine; hind tibia with eight apical teeth; apical spines of tarsomeres with long setae; female abdominal sternite VII with a pair of large horn-shaped spines strongly produced ventrad near anterior margin and a pair of blunt triangular lobes on posterior margin; gonocoxae VIII with a pair of small triangular sclerotized plates on posterolateral margin of endogonocoxal lobes. Redescription HEAD (Figs 4 A – C, 6 A – C, 7 A – C, 8 A – C). Produced into a moderately long and slender cephalic process. Cephalic process (Figs 4 B, 6 B, 7 B, 8 B) conical, strongly curved upward, and gradually narrowed toward apex. Vertex (Figs 4 A, 6 A, 7 A, 8 A) broadest at base, basal width narrower than transverse diameter of eyes, posterior plane elevated above pronotum; lateral carinae strongly ridged, foliaceous, and sub-parallel in basal third, abruptly constricted and strongly upturned in front of eyes, then gradually convergent anteriad, and acuminate at apex; posterior margin of vertex ridged and angularly concave at about 90 – 100 °; median carina distinct on a bulge between eyes. Frons (Figs 4 C, 6 C, 7 C, 8 C) with lateral carinae ridged, nearly parallel, slightly expanded outward below antennae; intermediate carinae slightly converging posteriorly and nearly approaching to frontoclypeal suture, but only sharp and distinct in apical third where they are distinctly expanded outward in ventral view and curved anterodorsad in lateral view, so their apical portion is distinctly visible in posterodorsal view; median carina robust and strongly convex in lateral view. Postclypeus and anteclypeus (Figs 4 C, 6 C, 7 C, 8 C) convex medially, with distinct median carina. Rostrum very long, surpassing middle of hind femora; basal segment distinctly longer than distal one. Genae (Figs 4 B, 6 B, 7 B, 8 B) with a longitudinal carina above eyes, appearing as bifurcated from lateral carinae of vertex in dorsal view (Figs 4 A, 6 A, 7 A, 8 A). Compound eyes (Figs 4 A – C, 6 A – C, 7 A – C, 8 A – C) large and globose. Ocelli (Figs 4 B, 6 B, 7 B, 8 B) relatively large, reddish. Antenna (Figs 4 B, 6 B, 7 B, 8 B) with very small scape; pedicel large and subglobose, with more than 50 distinct sensory plaque organs distributed over entire surface; flagellum long, setuliform. THOARAX. Pronotum (Figs 4 A, 6 A, 7 A, 8 A) distinctly shorter than mesonotum medially, narrow anteriorly, broad posteriorly; anterior margin pointed medially, forming a sharp angle, lateral marginal areas straight and sloping with two long longitudinal carinae on each side between eyes and tegulae, lower lateral carinae expanded and visible in dorsal view; posterior margin angularly concave at about 100 – 110 °; median carina sharp and high, with a lateral pit on each side, intermediate carinae absent. Mesonotum (Figs 4 A, 6 A, 7 A, 8 A) tricarinate, lateral carinae incurved anteriorly toward median carina. Forewings (Fig. 3 A – D) hyaline, much longer than abdomen, with ratio of length to width about 3: 1; veins with short setae on ventral side; venation with sparse transverse veins; MP bifurcating MP 1 + 2 and MP 3 + 4 near middle and beyond CuA; number of apical cells between R and CuA equal to 14; stigmal area small and quadrangular, with two or three cells. Legs elongate and slender, fore and middle femora distinctly elongate, fore femur not flattened and dilated, without spine near apex; hind tibia with 5 – 7 lateral spines and eight apical teeth; hind tarsomeres I and II each with 7 – 8 apical spines; apical spines of tarsomeres with long setae instead of platellae. ABDOMEN. With pregenital segments elongate and broad, without distinct median and intermediate carinae dorsally. Female abdominal sternite VII with a pair of large horn-shaped spines directed ventrad near anterior margin and a pair of blunt triangular lobes on posterior margin (Fig. 5 A). MALE GENITALIA. Pygofer (Figs 4 D – F, 6 D – F, 8 D – F) in lateral view distinctly wider ventrally than dorsally, dorsal margin slightly excavated to accommodate segment X, dorsoposterior margins angular. Gonostyles (Figs 4 E, 6 G, 8 G) symmetrical, base narrow, expanded toward apex, broadest subapically; dorsal margin with a claw-like, sclerotised process at apex directed dorsad, outer dorsal edge with a hook-like sclerotised process near middle directed ventrad. Aedeagus (Figs 4 G – I, 6 H – J, 8 H – J) with one pair of long endosomal processes extended from phallotheca or lacking such processes; phallobase sclerotised basally and membranous and inflated apically, with paired lobes covered with numerous minute superficial spines (Figs 4 G – I, 6 H – J) or without spines (Fig. 8 H – J). Segment X (Figs 4 D, 6 D, 8 D) large, in dorsal view with apex deeply excavated to accommodate anal style; anal style elongate and large. FEMALE GENITALIA (Fig. 5 A – G). Gonocoxae VIII (Fig. 5 E) with two membranous and flattened endogonocoxal processes (Gxp) on endogonocoxal lobe: Gxp 1 large and elongate, with a long sclerotized plate in it; Gxp 2 smaller and shorter. A pair of small triangular sclerotized plates on posterolateral margin of endogonocoxal lobes (Fig. 5 A). Gonapophyses VIII (Fig. 5 E) with anterior connective lamina large and sclerotized, with seven teeth of varying sizes and shapes. Gonapophyses IX (Fig. 5 F) with posterior connective lamina triangular, symmetrical, fused with the intergonocoxal plate at base; intergonocoxal plate extended cephalad into genital cavity, forming wall of gonospiculum. Gonoplacs (Fig. 5 G) with two lobes homologous; lateral lobe large and moderately sclerotized, with long setae at apex; the posterior lobe membranous, containing long sclerotized plate. Segment X (Fig. 5 D) large and broad in dorsal view, apex deeply excavated to accommodate anal style; anal style large and elongate. Female ectodermal genital ducts ditrysian. Bursa copulatrix (Fig. 5 B) superficially membranous, regularly gridded, without sclerotized ornamentations. A pair of large digitiform glands (Fig. 5 B) branched at anterior extremity of the anterior vagina on each side of the spermatheca. Spermatheca (Fig. 5 B) divided clearly into five parts: orificium receptaculi, ductus receptaculi, diverticulum ductus, pars intermedialis, and glandula apicalis.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA1FFCF9D77C06DA135F90A.taxon	distribution	Diversity and distribution Centromeriana is comprised of four species being endemic to the Congolian region of the western tropical Africa as defined by Linder et al. (2012) and closely matching to the Guineo-Congolian region of White (1979, 1983) including the ‘ Dahomey gap’.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA1FFCF9D77C06DA135F90A.taxon	biology_ecology	Ecology and economic importance Unknown. Heinrichs & Barrion (2004) listed Centromeriana sp. among insects occurring on rice in Gambia. However, based on morphological characters they specify in their identification key (p. 155) and a schematical drawing of the head and thorax (fig. 355: 157), this is a misidentification; the record probably refers to some other dictyopharid genus. Oke et al. (2015) reported adult Centromeriana spp. as minor pests of leaves of Amaranthus spp. in Nigeria but the identification is uncertain.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA5FFC39D53C4D6A135FD7D.taxon	description	Figs 1 A, C, 2 C, 3 A, 4 A – I, 5 A – G	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA5FFC39D53C4D6A135FD7D.taxon	materials_examined	Type material examined Lectotype (here designated) CAMEROON: ♂, “ jocosa Gerst. *, Centromeria, Victoria. Buchh. [handwritten dark blue label pinned in the collection next to the specimen]; Zool. Mus. Greifswald, II 27382; Victoria (Camerun), Buchholz leg. (labelled by I. Malenovský, 2016) [newly added white printed label pinned to the specimen]; LECTOTYPUS ♂, Dictyophara jocosa Gerstaecker, 1895, desig. Z. S. Song, I. Malenovský & A. P. Liang, 2016 [newly added red label pinned to the specimen] ” (ZIMG; dry-mounted: pinned through mesonotum, left foreleg and apical half of left forewing missing). Other material examined CAMEROON: 1 ♀, “ Kamerun, Conradt; Centromeriana jocosa Gerst. [Melichar’s handwriting], det. Melichar. ” (MMBC); 1 ♀, “ N. Kamerun, Joh. Albrechtshöhe, 14. VII – 17. VIII. [18] 96, L. Conradt; 14 / 7 – 17 / 8. [18] 96; jocosa [Melichar’s handwriting], det. Melichar. ” (MMBC); 1 ♀, “ N. Kamerun, Johann Albrechtshöhe, L. Conradt; 21 / 10. [18] 95; Centromeriana jocosa Gerst. [Melichar’s handwriting] ” (MFNB); 2 ♂♂, “ N. Kamerun, Joh. Albrechtshöhe, I. [18] 96, L. Conradt ” (MFNB); 1 ♂, “ Kamerun, L. Conradt ” (MFNB). EQUATORIAL GUINEA: 1 ♂, “ Fernando Póo [= Bioko island], Isab. [= Santa Isabel = the city of Malabo]; Centromeriana jocosa Ger. [Melichar’s handwriting], det. Melichar ” (MMBC); 2 ♂♂, 2 ♀♀, “ Span. Guinea, Nkolentangan [near Alén in mainland Equatorial Guinea], 7. XI – 8. V [1907 – 1909], G. Tessmann; Centromeriana jocosa Gers. [Synave’s handwriting], H. Synave det., 1969 ” (MFNB). GABON: 1 ♀ “ NHRS-GULI 000009123; Gabun.; Staudgr. [= O. Staudinger] ” (NHRS).	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA5FFC39D53C4D6A135FD7D.taxon	description	Redescription LENGTH. Body length (from apex of head to tip of forewings): ♂ 14.1 – 16.9 mm, ♀ 16.5 – 17.7 mm; head length (including two portions: the former is from apex of cephalic process to constricted and curved part, and the latter is from curved part to base of eyes): ♂ (0.8 – 1.0) + (1.0 – 1.2) mm, ♀ (0.8 – 1.1) + (1.1 – 1.2) mm; head width (including eyes): ♂ 1.4 – 1.6 mm, ♀ 1.6 – 1.7 mm; forewing length: ♂ 11.7 – 12.6 mm, ♀ 12.3 – 13.0 mm. COLORATION. General color (Figs 1 A, C, 2 C) brownish ochraceous marked with pale green and purplish-red on head and thorax, and dark brown and black on abdomen in dorsal view. Head pale green or yellow, head apex (cephalic process) purplish-red to dark brown, frons between intermediate carinae including median carina purplish-red. Clypeus dark purplish red to dark brown. Pronotum and mesonotum greenish or yellowish ochraceous, median carina along with posterior margin, areas behind eyes including lateral carinae and apical margins of paranotal lobes on pronotum, and three broad stripes on mesonotum medially and laterally all purplish-red. Tegulae purplish red. Forewing and hindwing membrane hyaline, except an unsharply delimited brownish macula at apex of forewings; venation brown, stigmal area brown. Thorax greenish or yellowish ochraceous ventrolaterally with an orange or purplish red band adjacent to base of hind coxae. Legs yellowish green, basal sections of femora, apices of fore and middle tibiae and lateral and apical spines all fuscous, apices of all femora dark brown to black. Abdomen dorsally dark brown and black with a paler brownish midline and a longitudinal row of large round yellowish ochraceous spots on each side or abdomen dorsally more or less uniformly yellowish ochraceous and dark brown to black only laterally; ventrally mostly blackish brown, posterior margins of sternites yellowish ochraceous; male and female terminalia dark brown, segment X dorsally yellowish ochraceous. MALE GENITALIA. Pygofer in lateral view (Fig. 4 E) large and broad, dorso-posterior margin obtusely angular; in ventral view (Fig. 4 F) much longer than in dorsal view (Fig. 4 D) with ratio of ventral to dorsal length about 4.0: 1. Gonostyles (Fig. 4 E – F) symmetrical, sicle-shaped, relatively narrow basally, slightly expanding towards apex, broadest subapically, apex straight; dorsal claw-like process large, elongate, and acute apically. Aedeagus (Fig. 4 G – I) with endosomal processes (Fig. 4 G – H) elongate, membranous and acute apically, extended posteriad and strongly curved dorso-anteriad, apex with some minute spines; phallobase sclerotized and pigmented at base, membranous and inflated apically, with one pair of short and thumb-like dorsolateral lobes directed laterad, their apex bearing minute spines (Fig. 4 G), and one pair of robust, elongate, thumb-like (apically broad) ventral lobes, confluent, curved and directed dorso-posteriad, base with some minute spines (Fig. 4 I). Segment X, in lateral view, with apical lobes hook-shaped: narrow, sharp and strongly projecting ventrad (Fig. 4 E); in dorsal view, relatively narrow and elongate, ratio of length to width near middle about 2.2: 1 (Fig. 4 D). FEMALE GENITALIA (Fig. 5 A – G) as in generic description.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA5FFC39D53C4D6A135FD7D.taxon	distribution	Distribution Cameroon, Equatorial Guinea and Gabon; also reported Democratic Republic of the Congo (Schmidt 1915; Fennah 1958 a) and Guinea (Lallemand 1958) but these require verification. The record from Sierra Leone (Melichar 1912) is here referred to as C. lindbergae sp. nov.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA5FFC39D53C4D6A135FD7D.taxon	discussion	Remarks Gerstaecker (1895) described C. jocosa based on material from “ Victoria ” (= Limbe, 4 ° 01 ′ N, 9 ° 13 ′ E), Cameroon but did not provide any further details on the type series; he particularly did not state the number and sex of the specimens he used for the description nor did he designate a holotype. In the Gerstaecker collection in ZIMG, there is a single male of C. jocosa. It fully conforms the original description including the associated locality data. Following the Recommendation 73 F and Article 74 of ICZN (1999), we designate this specimen as the lectotype to stabilize the nomenclature in the genus.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA9FFC59D06C146A5C6F977.taxon	description	urn: lsid: zoobank. org: act: B 6 EE 7817 - C 8 D 3 - 413 F- 9301 - 251 C 6389 C 5 DB Figs 1 B, 3 B, 6 A – J	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA9FFC59D06C146A5C6F977.taxon	diagnosis	Diagnosis Centromeriana lindbergae sp. nov. is very similar to C. jocosa in size, coloration and body structure but can be differentiated from the latter by the structure of the male genitalia: the segment X (anal tube) which is broad in dorsal view and with broad and blunt apical lobes, weakly projecting ventrad in lateral view (in C. jocosa, the segment X is much narrower and elongate in dorsal view and with narrow, sharp, hook-like apical lobes strongly projecting ventrad in lateral view); and the aedeagus which has relatively short and only slightly curved endosomal processes, the apices of which are directed dorso-posteriad (in C. jocosa, the endosomal processes are longer and more strongly curved, with apices directed dorso-anteriad), and the ventral lobes of the phallobase are more slender, convergent and tapering apically, but divergent medially (in C. jocosa, they are robust and thumb-like, broad apically and confluent medially). Centromeriana lindbergae sp. nov. differs from C. rhinoceros sp. nov. in the presence of a brownish macula on the forewing apex and a less inclined cephalic process. Centromeriana lindbergae sp. nov. differs from C. simplex in a darker body coloration pattern, the presence of a brownish macula on the forewing apex, and the structure of the male genitalia, particularly a broader segment X (narrower in C. simplex), obtusely angled dorsoposterior margin of pygofer (right-angled in C. simplex), a more slender and sicle-shaped gonostyle (robust and triangular in C. simplex), and the aedeagus with long endosomal processes (absent in C. simplex) and differently shaped and sized lobes of the phallobase (C. simplex has large and broad triangular dorsal lobes, expanded laterally).	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA9FFC59D06C146A5C6F977.taxon	etymology	Etymology The new species is named after Ms. Gunvi Lindberg, manager of the Hemiptera collection and the microscopic slide collection at the Naturhistoriska riksmuseet, Stockholm, Sweden, in recognition of her kindest help and support to the first author when he visited NHRS in 2014.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA9FFC59D06C146A5C6F977.taxon	materials_examined	Type material examined Holotype SIERRA LEONE: ♂, “ Sierra Leone, no date and collector [white printed label]; NHRS-GULI 000009124 [white printed label]; HOLOTYPE [newly added red printed label]; ♂, Centromeriana lindbergae sp. nov., det. Z. S. Song, I. Malenovský & A. P. Liang, 2016 [newly added red printed label] ” (NHRS, dry-mounted: pinned through mesonotum, right wing apex missing, abdomen detached and stored in glycerine in a glass microvial). Paratype SIERRA LEONE: 1 ♂, “ Sierra Leone [white printed label]; jocosa Gerst, det. Melichar [white handwritten and printed label]; Hung. Nat. Hist. Mus., Budapest, coll. Hemiptera [yellow printed label]; PARATYPUS ♂, Centromeriana lindbergae sp. nov., det. Z. S. Song, I. Malenovský & A. P. Liang, 2016 [newly added red printed label] ” (HNHM, dry-mounted: pinned through mesonotum, abdomen detached and stored in glycerine in a plastic microvial).	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA9FFC59D06C146A5C6F977.taxon	description	Description LENGTH. Body length (from apex of head to tip of forewings): ♂ 14.3 mm; head length (including two portions: the former is from apex of cephalic process to constricted and curved part, and the latter is from curved part to base of eyes): ♂ (0.7 – 0.8) + (1.1 – 1.2); head width (including eyes): ♂ 1.6 mm; forewing length: ♂ 11.4 – 11.9 mm. COLORATION AND EXTERNAL MORPHOLOGY (Figs 1 B, 3 B, 6 A – C). Largely identical to C. jocosa. MALE GENITALIA. Pygofer in lateral view (Fig. 6 E) similar to C. jocosa, ventral margin much longer than dorsal one with ratio of ventral to dorsal length about 4.2: 1; dorso-posterior margin obtusely angular. Gonostyles (Fig. 6 E – G) identical to C. jocosa, sicle-shaped, narrow basally, slightly expanding towards apex, broadest subapically; dorsal claw-like process large, elongate, and acute apically. Aedeagus with endosomal processes (Fig. 6 H – J) relatively short, membranous, slightly inflated, acute apically, slightly curved with apices directed obliquely dorso-posteriad, apex with minute spines; phallobase sclerotized and pigmented at base, membranous and inflated apically, with one pair of thumb-like dorsolateral lobes, directed anteriad, apex with numerous minute spines (Fig. 6 H), a pair of long and relatively slender, curved ventral lobes, divergent medially, convergent and tapering apically with some minute superficial spines (Fig. 6 I – J). Segment X large and broad in dorsal view, ratio of length to width near middle about 1.3: 1 (Fig. 6 D); in lateral view, with broad and blunt apical lobes, weakly projecting ventrad (Fig. 6 E). Female unknown.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA9FFC59D06C146A5C6F977.taxon	distribution	Distribution Sierra Leone.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFA9FFC59D06C146A5C6F977.taxon	discussion	Remark The paratype specimen from HNHM was misidentified and erroneously reported under C. jocosa by Melichar (1912).	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFAEFFC79D07C36EA543FD03.taxon	description	urn: lsid: zoobank. org: act: 304 EABAF- 397 A- 401 C- 931 F-A 8 F 2 D 782 B 702 Figs 2 A, D, 3 C, 7 A – C	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFAEFFC79D07C36EA543FD03.taxon	diagnosis	Diagnosis The species is formally described here in the absence of males but it can be distinguished from all other Centromeriana species by asexual characters, particularly the backward inclination of the cephalic process which also has a relatively smaller size. From C. jocosa and C. lindbergae sp. nov. it also differs in the clear forewing apex and from C. simplex in the body coloration, having a dark pattern, particularly on the abdomen.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFAEFFC79D07C36EA543FD03.taxon	etymology	Etymology The specific epithet is a noun in apposition derived through Latin from the Ancient Greek word ρίνóκερως (= rhinoceros), which is composed of ρίνο- (= nose) and κέρας (= horn). The new species is named for its horn-shaped cephalic process.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFAEFFC79D07C36EA543FD03.taxon	materials_examined	Type material examined Holotype TOGO: ♀, “ Toǵo, Bismarckburǵ, L. Conradt [dark blue printed label]; HOLOTYPE [newly added red printed label]; ♀, Centromeriana rhinoceros sp. nov., det. Z. S. Song, I. Malenovský & A. P. Liang, 2016 [newly added red printed label] ” (MFNB, dry-mounted: pinned through mesonotum). Paratype TOGO: 1 ♀, “ Toǵo, Bismarckburǵ, L. Conradt [dark blue printed label]; PARATYPE [newly added yellow printed label]; ♀, Centromeriana rhinoceros sp. nov., det. Z. S. Song, I. Malenovský & A. P. Liang, 2016 [newly added red printed label] ” (MFNB, dry-mounted: pinned through mesonotum).	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFAEFFC79D07C36EA543FD03.taxon	description	Description LENGTH. Body length (from apex of head to tip of forewings): ♀ 14.4 mm; head length (including two portions: the former is from apex of cephalic process to constricted and curved part, and the latter is from curved part to base of eyes): ♀ 0.7 + 1.0; head width (including eyes): ♀ 1.4 mm; forewing length: ♀ 11.9 mm. COLORATION (Figs 2 A, D, 7 A – C) AND STRUCTURE. Largely identical to C. jocosa except for the forewing membrane which has clear apex (Fig. 3 C). Cephalic process in lateral view relatively short, strongly upturned and curved upward and slightly backward in more than 90 ° (Fig. 7 B). Male unknown.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFAEFFC79D07C36EA543FD03.taxon	distribution	Distribution Togo.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFADFFD99D60C0CCA0EBFDD1.taxon	description	Figs 2 B, E, 3 D, 8 A – J	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFADFFD99D60C0CCA0EBFDD1.taxon	materials_examined	Type material examined Lectotype (here designated) EQUATORIAL GUINEA: ♂, “ Is. Fernando Poo [= Bioko island], Basile, 400 – 600 m. s. m., VIII – IX. 1901, L. Fea; Centromeriana simplex M. [Melichar’s handwriting], det. Melichar.; Typus [dark red label]; Syn- typus [bright red label]; Collectio Dr. L. Melichar, Moravské museum Brno; Invent. č. 4940 / Ent., Mor. muzeum, Brno; LECTOTYPUS ♂, Centromeriana simplex Melichar, 1912, desig. Z. S. Song, I. Malenovský & A. P. Liang, 2016 [newly added red label] ” (MMBC; dry-mounted, pinned through mesonotum, abdomen detached and stored in glycerine in a glass microvial pinned under the specimen).	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFADFFD99D60C0CCA0EBFDD1.taxon	description	Redescription LENGTH. Body length (from apex of head to tip of forewings): ♂ 13.3 mm; head length (including two portions: the former is from apex of cephalic process to constricted and curved part, and the latter is from curved part to base of eyes): ♂ 0.7 + 1.0; head width (including eyes): ♂ 1.4 mm; forewing length: ♂ 10.4 mm. COLORATION (Fig. 2 B, E). Largely similar to C. jocosa, but much paler than the latter, without dark brown or black on abdomen in dorsal habitus. Head including cephalic process pale green, a longitudinal stripe in front of eyes on genae and median carina on frons reddish ochraceous (Fig. 8 A, C). Pronotum and mesonotum greenish ochraceous, median carina along with posterior margin, areas behind eyes including lower lateral carinae and apical margins of paranotal lobes on pronotum, and median carina and lateral areas on mesonotum reddish ochraceous. Forewings and hindwings with venation and stigmal area pale greenish ochraceous. Abdomen dorsally and ventrally greenish ochraceous. Apical lobes of male segment X black (Fig. 8 E). MALE GENITALIA. Pygofer in lateral view (Fig. 8 E) slightly shorter than in C. jocosa and C. lindbergae sp. nov., ventral margin longer than dorsal one with ratio of ventral to dorsal width about 3.0: 1; dorsoposterior margin right-angled. Gonostyles (Fig. 8 E – G) broad, strongly expanding towards apex, broadest and convex subapically; dorsal claw-like process narrow and elongate, apex acute. Aedeagus (Fig. 8 H – J) lacking distinct endosomal processes; phallobase sclerotized and pigmented at base, membranous and inflated apically, with one pair of large, robust, triangular dorsolateral lobes, directed laterad, without superficial spines (Fig. 8 H); and one pair of short and thumb-like ventral lobes, directed straightly posteriad (Fig. 8 I). Segment X, in dorsal view, elongate with ratio of length to width near middle about 1.7: 1 (Fig. 8 D); in lateral view, ventral margins straight, apical lobes blunt, rounded, weakly produced ventrad (Fig. 8 E). Female unknown.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFADFFD99D60C0CCA0EBFDD1.taxon	distribution	Distribution Equatorial Guinea.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
03CB0C7FFFADFFD99D60C0CCA0EBFDD1.taxon	discussion	Remarks This species was described by Melichar (1912) based on two male specimens from Fernando Poo [= Bioko], Equatorial Guinea, originally coming from the collection of the museum in Genoa, Italy. One of these two syntypes has been preserved in Melichar’s personal collection in MMBC. According to Article 74 of ICZN (1999) we designate this specimen as a lectotype for C. simplex to stabilize the nomenclature. We have redescribed and illustrated the species based on the lectotype.	en	Song, Zhi-Shun, Malenovský, Igor, Liang, Ai-Ping (2017): Revision of the Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Dictyopharidae). European Journal of Taxonomy 278: 1-21, DOI: 10.5852/ejt.2017.278
