identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C887F4FFA2FFE2D7B6FD68B7840F4A.text	03C887F4FFA2FFE2D7B6FD68B7840F4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scapheremaeus subiasi Palacios-Vargas & Jaimes Murillo & Ojeda 2024	<div><p>Scapheremaeus subiasi sp. nov.</p><p>(Figs. 1–12)</p><p>Diagnosis. Body size: 434 × 217. Body surface foveolate-reticulate. Costulae well developed and lamellar setae associated with a tubercle. Costulae ridge-like, fully developed, interrupted medially in some of the specimens. Sensillum clavate, with a pigmented capitulum, sometimes very dark. Humeral process present. Ten pairs of notogastral setae of two different types. Six pairs of genital setae. Semi-round lenticule (33 × 36). Monodactylous.</p><p>Description. Measurements. Body length: 434, width 217 (holotype, female), paratypes (five); length 433–305 (394) and width 236–207 (221).</p><p>Integument. Body color light to dark brown. Body surface densely foveolate (Figs. 1 and 2), lateral sides with parallel ridges and troughs which are also shown in the femora of all legs.</p><p>Prodorsum (Figs. 1, 3). Rostrum rounded. Costulae well developed, terminated by tubercles bearing lamellar setae. Costular ridge sometimes incomplete. Rostral setae (ro, 20) setiform, smooth, directed medially. Lamellar setae minute (le, 10) setiform, straight (Fig. 4). Interlamellar and exobothridial setae and alveoli absent. Bothridium (bo) irregular shape. Sensillum (ss, 53) club-shape, with dark pigmentation, and a short stalk (Fig. 4).</p><p>Gnatosoma (Figs. 5–6). Subcapitulum longer than wide. Subcapitular setae thin, smooth; m shorter than a and h (Fig. 5). Palps with setation 2-1-3-7 (+ w). Chelicerae with two pectinate setae; cha slightly shorter than chb. Trägårdh´s organ long, tapered (Fig. 6).</p><p>Notogaster (Figs. 1, 3 and 8). Length (n= 6): 364–315 (333); width: 256–216 (232). General shape irregularly elliptical, almost square in the most anterior region. Lenticulus (len; 36 x 30) distinctly round. Humeral region with a well–developed process. Centrodorsal region (247 x 153) with foveolate microsculpture, and well-developed diagonal extensions towards the humeral region close to setae c2. Circumdorsal scissure complete. Ten pairs of notogastral setae, c2, la, h1, h2, h3, p1, p2 and p3 rod-shaped (Fig. 3). Setae lm and lp are small, spiniform and located in the centrodorsal region. Lyrifissures im, ip, ih visible, located in the circumdorsal border and lyrifissure ips and opisthonotal gland opening gla are not visible (Figs. 3,8).</p><p>Epimeral region (Fig. 7). Microsculpture predominantly foveolate, including the epimeral region and the genital and anal plates, and parallel ridges on the sides from the epimeral region onwards (Fig. 2). Epimeral chaetotaxy: 3-1-2-2, setae setiform and minute.</p><p>Anogenital region (Figs. 2, 7–8). Five pairs of genital setae (g1-g6), g1, g2 and g3 located adjacently on the anterior margin, followed by a pair of aggenital setae. Anal plate is longer than wider, with two pairs of setae, three pairs of adanal setae and lyrifissure iad in paranal position. Genitoanal chaetotaxy: 6-1-2-3 (Fig. 7). Microsculpture in the region foveolate in the center, to the sides consisting of parallel ridges and troughs (Fig. 2).</p><p>Legs (Figs. 9–12). All legs with tuberculate microsculpture and a series of parallel ridges on anti-axial surface. Porose areas (p.a.) clearly visible and oval shape in femur of all legs. Formula of leg setation and solenidia: femur-genua-tibia-tarsus (solenidia): leg I: 4-2(1)-4(2)-16(2); leg II: 4-2(1)-3(1)-14(1); leg III: 2-1(1)-3(1)-14; leg IV: 2-1(1)-3(1)-12. Length of leg segments I and II (femur-genua-tibia-tarsus): leg I: 85-24-58-23, leg II: 79-22-43-87; legs III and IV (trochanter-femur-genua-tibia-tarsus): III: 39-69-22-43-87, IV: 41-65-23-52-28. Total length of legs (from femur to tarsus): 190, 231, 221, 168. Monodactylous.</p><p>Type locality. MEXICO: Morelos: Sierra de Huautla, El Zapote, ex. Epiphytic moss and lichens. 1-X-2021. V. Granados, coll.</p><p>Type material. Female holotype and 7 paratypes in semipermanent slides (one stored in alcohol). All deposited in the collection of Laboratorio de Ecología y Sistemática de Microartrópodos, UNAM .</p><p>Etymology. The species name subiasi is dedicated to Dr. Luis Subías, in homage to the tireless oribatologist who left us an important legacy with his Catalogue of Oribatid Species.</p><p>Remarks. Scapheremaeus subiasi sp. nov. is morphologically most similar to S. magdalenae Palacios-Vargas &amp; Ríos, 1998 and S. hectorperezius Palacios-Vargas &amp; Ríos, 1998 from Cedros Island, Baja California and Los Tuxtlas, Veracruz respectively. They share the presence of a humeral process, ten notogastral setae, monodactylus legs and microsculpture of the body represented by a foveolar pattern, as well as an anal plate that is longer than wider. However, the new species differs from the two mentioned species by the larger body size 439 x 222 and 365 × 180, respectively. The localization of lamellar setae on well-developed lamellar tubercules (vs. absent or inconspicuous), and type and size of setae ro and le (smooth vs basiliform, barbulate; le smaller than ro, vs same size). The most evident difference between these species in the general appearance is the posterior end of notogaster, Scapheremaeus subiasi sp. nov. has a smooth contour vs. a constriction and crest on the border shown in the other species.</p></div>	https://treatment.plazi.org/id/03C887F4FFA2FFE2D7B6FD68B7840F4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Palacios-Vargas, José G.;Jaimes Murillo, Kevin A.;Ojeda, Margarita	Palacios-Vargas, José G., Jaimes Murillo, Kevin A., Ojeda, Margarita (2024): New species of Scapheremaeus (Acari: Oribatida: Cymbaeremaeidae) from an epiphytic environment in Morelos, Mexico. Zootaxa 5556 (1): 97-105, DOI: 10.11646/zootaxa.5556.1.9, URL: https://doi.org/10.11646/zootaxa.5556.1.9
03C887F4FFA5FFE3D7B6FE71B2E90C4A.text	03C887F4FFA5FFE3D7B6FE71B2E90C4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scapheremaeus Berlese 1910	<div><p>Key for all known species of Scapheremaeus from Mexico (modified from Palacios-Vargas et al. 2022)</p><p>1. Notogaster without a centrodorsal region; total body length more than two times its width; notogaster rectangular, with a transverse line separating the lenticule... Scapheremaeus alvarezius Ríos and Palacios-Vargas, 1998 (Veracruz: Los Tuxtlas).</p><p>- Notogaster with a well-defined centrodorsal region; sensillum (ss) variously shaped................................ 2</p><p>2. Notogaster pyriform, almost round, with only seven pairs of setae and strongly contrasting microsculpture between the centrodorsal region and the circumgastric scissure, tridactylous, genitoanal chaetotaxy 4-1-1-2.............................................................. Scapheremaeus colloffi Palacios-Vargas et al. 2022 (Veracruz: Los Tuxtlas).</p><p>- Notogaster usually elongate with eight or more pairs of setae, genitoanal chaetotaxy distinct......................... 3</p><p>3. Eight pairs of notogastral setae, genitoanal chaetotaxy 6-1-2-3; humeral process very long, pointed and strongly curved............................ Scapheremaeus palaciosi Ríos, 1996 (CDMX: Contreras; Edo. de Mex.: Popocatépetl volcano).</p><p>- Notogaster with nine or more pairs of setae................................................................ 4</p><p>4. Notogaster with nine pairs of setae, genitoanal chaetotaxy 6-1-2-2, round lenticule.................................................................... Scapheremaeus dirzoius Palacios-Vargas and Páez, 2018 (Veracruz: Los Tuxtlas).</p><p>- Notogaster with ten to 13 pairs of setae, distinct genito-anal chaetotaxy.......................................... 5</p><p>5. Notogaster irregular with 14 pairs of setae, bidactylous, humeral process pointed........................................ Scapheremaeus baloghius Ríos and Palacios-Vargas, 1998 (Jalisco: Chamela; Chiapas: Chajul; Veracruz: Los Tuxtlas).</p><p>- Notogaster with ten pairs of setae........................................................................ 6</p><p>6. Epimeral chaetotaxy 3-1-3-3, bidactylous, triangular humeral process present; three pairs of setae in the centrodorsal region........................................ Scapheremaeus schatzi Ríos and Palacios-Vargas, 1998 (Jalisco, Chamela).</p><p>- Epimeral chaetotaxy 3-1-2-2............................................................................ 7</p><p>7. Notogaster with a well-developed diagonal extensions of centrodorsal region; setae of the centrodorsal region spiniform, c2, la, hi-3 and p1-p3 bacilliform shape; sensillum (Bothridial seta) with a striate capitulum........................................................................................ Scapheremaeus subiasi sp. nov. (El Zapote, Morelos).</p><p>- Notogaster without diagonal extensions; total body length less than two times its width............................. 8</p><p>8. Centrodorsal region of notogaster with a longitudinal ridge.................................................... 9</p><p>- Centrodorsal region of notogaster without a longitudinal ridge................................................ 11</p><p>9. Notogaster setae rod-shape with normal alveoli............................................................ 10</p><p>- Notogaster setae flame-shaped with relatively large alveoli, lichen-shaped sensillum (Bothridial seta), three longitudinal carinae.... Scapheremaeus flamiferus Palacios-Vargas and Ríos, 1998 (Nicaragua: León; Mex: Q. Roo, Guerrero, Veracruz).</p><p>10. Sensillum (Bothridial seta) black, barbed and elongated, an incomplete crest between the lenticule and the centrodorsal plate.................. Scapheremaeus grahamius Ríos and Palacios-Vargas, 1998 (CDMX: Reserva Pedregal de San Ángel).</p><p>- Sensillum (Bothridial seta) moruliform and elliptical. A lateral and posterior crest to the lenticule that borders the central area present. Posterior margin of the notogaster truncated and a well-developed concavity........................................................... Scapheremaeus tillandsiophilus Ríos and Palacios-Vargas, 1998 (Morelos: Chichinautzin).</p><p>11. Centrodorsal region of notogaster with two lateral ridges joining the lm and lp setae; anterior region of notogaster semi rectangular. Rostral setae minute (3 μm).................................................................................................. Scapheremaeus eugenius Ríos y Palacios-Vargas, 1998 (Jalisco: Sierra de Manantlán).</p><p>- No lateral ridges and average rostral setae about 10 μm or more............................................... 12</p><p>12. Notogaster with ridges between lenticule and central zone................................................... 16</p><p>- Notogaster without ridges between the lenticule and the central zone........................................... 13</p><p>13. Centrodorsal region of the notogaster almost circular. Prodorsum with very long lamellae, inhabitant rock surfaces........................................................ Scapheremaeus chaac Ríos and Palacios-Vargas, 1998 (Yucatán).</p><p>- Centrodorsal region of notogaster elliptic. Prodorsum without lamellae.......................................... 14</p><p>14. All setae of notogaster and ad1 and ad2 fungiform, humeral processes spiniform...................................................................... Scapheremaeus fungisetosus Ríos and Palacios-Vargas, 1998 (Jalisco: Chamela).</p><p>- Centrodorsal region of the notogaster with or without longitudinal crest, setae no fungiform......................... 15</p><p>15. Humeral processes absent. Lamellar setae setiform, lichen-shaped sensillum (Bothridial seta), with a small notch near seta c2 .................................. Scapheremaeus mahunkaius Ríos and Palacios-Vargas, 1998 (Jalisco, Chamela).</p><p>- Humeral process present. Lamellar setae bacilliform, sensillum (bothridial seta) head black; no notch near setae c2 .......................................... Scapheremaeus simplex Ríos and Palacios-Vargas, 1998 (Morelos: Chichinautzin).</p><p>16. Body very flattened (body length/thickness = 3.4). Lamellar and notogastral setae minute (4μm)........................................................... Scapheremaeus nogueraius Ríos and Palacios-Vargas, 1998 (Jalisco: Chamela).</p><p>- Body normal, not flattened (length/thickness = 1.6–2.3). Lamellar setae medium to long........................... 17</p><p>17. Sensillum (Bothridial seta) lichen-shaped. Notogaster broad (length/width ratio = 1.5), capitulum sensilla hyaline................................ Scapheremaeus volcanicus Ríos and Palacios-Vargas, 1998 (Edo. de Mex.: volcán Popocatépetl)</p><p>- Sensillum (Bothridial seta) has a different shape. Notogaster not as wide........................................ 18</p><p>18 Sensilum (Bothridial seta) with a morula-like appearance, body relatively elongated (length/width = 1.8). Humeral process absent......................... Scapheremaeus morulisensillatus Ríos and Palacios-Vargas, 1998 (CDMX: Contreras)</p><p>- Sensilla not morula-shaped............................................................................. 19</p><p>19 Notogastral setae rod-shaped, long (15 μm).Posterior margin of notogaster with irregular edges. Posterior margin of circumdorsal region strongly projected and triangular........................................................ Scapheremaeus magdalenae Ríos and Palacios-Vargas, 1998 (Baja California: Cedros Island, Baja California Sur: Sierra La Laguna).</p><p>- Body setae not rod-shaped, distinct, shorter. Posterior margin of notogaster rounded............................... 20</p><p>20 Prodorsum with short lamellar cusps, translamella incomplete. Cerotegument strongly developed. Notogaster with two circular central areas. Posterior notogastral setae fungiform............................................................ Scapheremaeus tonatiuh Palacios-Vargas, Rìos and Vázquez, 1998 (Jalisco: Chamela, Q. Roo: Sian Ka’an; Chiapas: Chajul).</p><p>- Prodorsum without lamellar cusps and without translamella. Cerotegument inconspicuous. Notogaster with a single central area, with tiny setae slightly clavate and large alveolus..................................................................................... Scapheremaeus hectorperezius Ríos and Palacios-Vargas, 1998 (Veracruz, Los Tuxtlas).</p></div>	https://treatment.plazi.org/id/03C887F4FFA5FFE3D7B6FE71B2E90C4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Palacios-Vargas, José G.;Jaimes Murillo, Kevin A.;Ojeda, Margarita	Palacios-Vargas, José G., Jaimes Murillo, Kevin A., Ojeda, Margarita (2024): New species of Scapheremaeus (Acari: Oribatida: Cymbaeremaeidae) from an epiphytic environment in Morelos, Mexico. Zootaxa 5556 (1): 97-105, DOI: 10.11646/zootaxa.5556.1.9, URL: https://doi.org/10.11646/zootaxa.5556.1.9
