taxonID	type	description	language	source
03D687E2FFB2905FEFBBAB3FFDC93580.taxon	type_taxon	Type species: Alitocoris schraderi Sailer, by original designation.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFB2905FEFBBAB3FFDC93580.taxon	diagnosis	Diagnosis: Oval body, general colour brown to black with yellow spots in some species. Punctures black, deep, and confluent, forming wrinkles on the dorsum of head, pronotum, and scutellum. Antennae fivesegmented, with sexual dimorphism.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFB2905FEFBBAB3FFDC93580.taxon	description	Head wider than long, declivous; punctures confluent, forming wrinkles at least from base of tylus to vertex. Juga longer than tylus by a distance not greater than the diameter of ocellus; apices of juga obtuse, not touching each other. Lateral margins of juga sinuous, deflected. Vertex flat to slightly tumid. Anteocular processes reduced to an obtuse tubercle, not projected. Posterior margin of head impunctate. Antennal tubercles partially visible from above, with lateral spine. Antennae five-segmented, second segment the shortest. Ocelli posterior to hind margin of eyes, interocellar distance about 1.5 times the width of eye. Area between ocelli and eyes impunctate. Ventrally with 1 + 1 longitudinal carina posterior to the eyes, concolorous with the surface, delimiting the base of eyes and gula. Gula slightly depressed, width equivalent to the diameter of eye; punctures evenly distributed. Bucculae elevated, evanescent and divergent posteriorly, not attaining posterior margin of head, with triangular anterior tooth in some species. Rostrum attaining or surpassing the posterior margin of urosternite IV; first segment reaching prosternum, second segment is the longest attaining mesocoxae; insertion of first segment of labium on head posterior to anterior limit of eyes. Pronotum wider than long, anterior half declivous; punctures confluent, forming wrinkles posterior to cicatrices. Cicatrices flat to slightly tumid. Anterior margin deeply concave in the middle third. Anterolateral angles with small acute projection, exceeding lateral limit of eyes for twice the diameter of ocellus or less. Anterolateral margins sinuous to subrectilinear; posterolateral margins slightly convex; posterior margin subrectilinear. Scutellum longer than wide, reaching connexival segment V; punctures confluent, wrinkles more dense on anterior half. Foveae at basal angles black, shallow, and punctured. Corium longer than scutellum, reaching connexival segment VI; punctures not confluent; 1 + 1 yellowish spot at apex of radial vein, equivalent to the diameter of ocellus or less. Membrane attaining or little surpassing apex of abdomen, bearing nine to 13 veins not forming cells. Pro-, meso-, and metapleura with punctures evenly distributed. Prosternum sulcate, meso- and metasternum with median carina along entire length. Mesopleural evaporatorium narrow, occupying a thin area along the posterior margin of mesopleura; metapleural evaporatorium black, occupying half the width of metapleura, lateral margin subrectilinear. Peritreme spout reaching at least half the width of evaporatorium. Tibiae shallowly sulcate dorsally; third segment of metatarsi of males and females depressed dorsally. Connexivum exposed from segments III to VII, narrow lateral strip impunctate, sometimes with light central spot. Abdomen with punctures more dense on lateral thirds, inconspicuous on central disc; longitudinal furrow inconspicuous on median third of sternites III to V. Trichobothria lateral to the line tangent to spiracles. Spiracles black; anteromedian margin elevated or flat, concolorous with abdomen or yellowish. Male. Antennae: segment III the widest, about twice the width of segment IV; also wider than segment III in females. Articular membrane along the posterior margin of urotergite VII expanded medially. Pygophore subrectangular. Posterolateral angles acuminate, apices obtuse. Dorsal rim concave with obtuse or spiniform projections. Inferior layer of ventral rim with 1 + 1 projection lateral to segment X and directed dorsoanteriorly over genital cup (Fig. 4 A, C). Superior layer of ventral rim bearing lighter, lobular areas beneath the projection of inferior layer (Fig. 7 B). Inferior layer of ventral rim continued laterally in low carina directed to posterolateral angles of pygophore. Median third of ventral rim concave (Fig. 4 B, D). Segment X subrectangular, exceeding the posterior limit of inferior layer of ventral rim, flattened dorsoventrally at apex, basal half membranous; fringe of setae at apex; anal opening apical; ventral genital opening ina longitudinal slit (Fig. 4 A, C). Parameres ventral to segment X, with apical setae; head of paramere projecting over ventral rim (Figs 4 B, D, 5 E – G). Phallus. Phallotheca globose, narrowing at apex, and with a pair of ample processes. Vesica bent; ductus seminis distalis sinuous, apex in gutter (Fig. 5 A – D). Female. Antennae segments uniformly wide. Genitalia. Genital plates convex, velvety. Gonocoxites 8 subrectangular, disc elevated; posterior margins sinuous, projecting over the base of laterotergites 9 or subrectilinear with posterolateral angles as small tooth over the base of laterotergites 8; callus present or not; disc tumid, bent or not toward head near the posterior margin, forming a flap; sutural margins juxtaposed. Laterotergites 8 triangular, wider than long; posterior margins subrectilinear; spiracles placed posterior to or lateral to lateral angle of gonocoxites 8. Gonocoxites 9 partially exposed, triangular. Laterotergites 9 digitiform, reaching the band connecting laterotergites 8; mesial margins juxtaposed. Segment X hidden by laterotergites 9 (Fig. 6 A – D). Pars comunis with conical microsculptures. Gonapophyses 8 and 9 with folds bearing conical microsculptures (Figs 6 E, F, 7 A). Thickenings of gonapophyses 9 present. Thickening of vaginal intima subrectangular (Fig. 6 E, F). Chitinellipsen ring-like. Median duct of KEY TO THE SPECIES OF ALITOCORIS	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFB2905FEFBBAB3FFDC93580.taxon	discussion	Comments: Alitocoris is in a polytomy in clade A (Fig. 1), so its relationships with the other genera in this clade are uncertain. Alitocoris species are recognized by the colour pattern, deep and confluent punctures, five antennal segments, and especially the morphology of female genitalia owing to the lack of knowledge about males of the genus. Alitocoris is distributed in Central America (Guatemala, Honduras, Costa Rica, and Panamá).	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFB09053ED1DA951FB08328E.taxon	description	(FIGS 3 A, 4 A, B, 5 A – G, 6 A, E, 7 A, B)	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFB09053ED1DA951FB08328E.taxon	materials_examined	Holotype: GUATEMALA, 1 male NMNH Type Cat. no. 59342 (Sailer, 1950). Allotype: GUATEMALA, 1 female (Sailer, 1950). Paratypes: COSTA RICA, Cartago: Turrialba, 1 m and 1 f; San Jose: San Isidro Del General, 1 m and 2 f (Sailer, 1950).. Material examined: HONDURAS, Santa Barbara: (13 km south-east of El Mochito), 1 f, 22. vii. 1977, OBrien (DARC). COSTA RICA, (Hamburg Farm), 1 f, 31. iii, C. P. Dodge (AMNH); Heredia: (Estación Biológica La Selva), 1 f and 1 m, 10 – 12. vi. 1986, B. C. Ratcliffe & party (DBTC); (Estación Biológica La Selva 2 km south of Puerto Viejo), 1 f, 15 – 22. iii. 1987: J. Negrón (DARC); (Estación Biológica La Selva 150 ma. s. l.), 1 f, vii. - viii. 1998, FIT, Nico Franz (JEE); (Finca La Selve Verde, 12 km south of Puerto Viejo, 125.4 ma. s. l.), 1 f, 23 – 26. ix. 1986, J. E. Eger, [10.4667; - 84.0167] (JEE); Guanacaste: (Estación Pitilla, 9 km south of Sta. Cecilia, 700 m a. s. l.), 1 f (Costa Rica INBio CRI 001 008706), vii. 1988, GNP Biodiversity Survey [11.1000; - 85.4167], 10 ° 59 ′ N, 85 ° 25 ′ W (INBio); 1 m (Costa Rica Inbio CRI 000154808) and 1 f (Costa Rica Inbio CRI 000154862), ii. 1990, P. Ríos, C. Morgada & R. Blanco, [11.1000; - 85.4167], 330200, 380200 (INBio); (Est. Pitilla, 9 km south of Sta Cecilia, P. N. Guanacaste, 700 m a. s. l.), 1 f (Costa Rica Inbio CRI 000808667), v. 1992, P. Ríos, [11.1000; - 85.4167], L-N 330200, 380200 (INBio); 1 f (Costa Rica CRI 000 819068), 10 – 17. vi. 1992, C. Moraga, [11.1000; - 854167], L-N 330200, 380200 (INBio); 1 f and 1 m (Costa Rica Inbio CRI 001818072), i. 1994, P. Ríos, [11.1000; - 854167], L-N 330200 _ 380200 (INBio); 1 f (Costa Rica Inbio CRI 002 048807), vii. 1994, P. Ríos, [11.1000; - 85.4167], L-N 330200 _ 380200 # 3140 (INBio); (Est. Biol. Cacao, A. C. Guanacaste, 1100 ma. s. l.), 1 m (Costa Rica Inbio CRI 002113078), xii. 1993 – ix. 1994, Fam. Guadamuz Chavarría, [10.9333; - 85.4500], L-N 323300 _ 375700 # 3318 (INBio); 1 f (Costa Rica Inbio CRI 002113079), xii. 1993 – i. 1994, Fam. Guadamuz Chavarría, [10.9333; - 85.4500], L-N 323300 _ 375700 # 3318, (INBio); (Estación Cacao, south-west side Volcan Cacao, 1000 – 1400 ma. s. l.), 1 m (Costa Rica Inbio CRI 000147453), 1 m (Costa Rica Inbio CRI 000147468) and 1 f (Costa Rica Inbio CRI 000 147462), xi. – xii. 1989, URCG, R. Blanco & C. Chavez, [10.9333; - 85.4500], 323300, 375700 (INBio); Chilamate (Selva Verde Lodge), 1 m, 11 – 13. viii. 1995, J. E. Eger, [10.4500; - 84.0667], Costa Rica collecting permit no. 00113412 (JEE); (Est. Magasasay, P. N. Braulio Carrillo, 200 m a. s. l.), 1 m (Costa Rica Inbio CRI 000298374), i. 1991, M. Barrelier, [10.1599; - 839744], L-N 264600, 531100, (INBio); Cartago: Turrialba, 1 m, 20. v. 1948, F. Schrader 445 [9.9000; - 83.6833] (AMNH); 1 f, 23. vi. 1948, F. Schrader 597, [9.9000; - 83.6833] (AMNH); Limon: (Sector Cerro Cocori, Fca de E. Rojas, 150 m a. s. l.), 1 f (Costa Rica Inbio CRI 000794714), 26. iii. – 24. iv. 1992, F. A. Quesada, L-N 286000, 567500 (INBio); Cerro Tortuguero (P. N. Tortuguero, 100 ma. s. l.), 1 f (Costa Rica Inbio CRI 000088898), iv. 1989, R. Aguilar & J. Solano, [10.5333; - 83.5167], 285000, 588000 (INBio); Amubri (70 m, Talamanca), 1 f (Costa Rica Inbio CRI 000 965969), 12 – 30. ix. 1992, G. Gallardo, [9.5167; - 82.9667] L-S 385500, 578050 (INBio). PANAMA, Bocas del Toro: Corriente Grande (100 m a. s. l.), 1 m, 25. iv. 1980, Henk Wolda, [9.5167; - 82.8667], 9 ° 17 ′ N, 82 ° 32 ′ W (DBTC).	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFB09053ED1DA951FB08328E.taxon	diagnosis	Diagnosis: Dark brown to black, punctures deep and concolorous. Legs not contrasting with body colour.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFB09053ED1DA951FB08328E.taxon	description	Head black, punctures not confluent at apical third of juga. Vertex slightly tumid. Antennae dark brown to black, except distal half of fourth segment and entire fifth segment light brown; dorsal pale blot on first segment, sometimes inconspicuous; third segment sometimes light brown. Proportional length of antennal segments: I <II <III> IV <V; fifth the longest. Bucculae lacking anterior tooth. Labium yellowish, reaching to posterior margin of urosternite V; proportinal length of labial segments: I <II> III> IV.. Pronotum bearing median transverse discontinuous line, lighter anteriorly, sometimes inconspicuous; punctures shallower between anterior margin of pronotum and cicatrices. Cicatrices flat. Anterolateral angles parallel to posterior margin of eyes, exceeding lateral limit of eyes for at most the diameter of ocellus. Anterolateral margins slightly sinuous. Scutellum reaching the level of middle of fifth segment of connexivum; punctures confluent on disc, forming wrinkles; 1 + 1 minute yellowish spots close to foveae. Corium with few pale spots; apex little surpassing anterior margin of sixth segment of connexivum. Yellowish spot at apex of radial vein equivalent to the diameter of ocellus. Membrane with nine veins, reaching or little surpassing apex of abdomen. Pro-, meso-, and metapleura black. Peritreme exceeding half the width of evaporatorium. Coxae and trochanters yellowish with coppery areas; femora and tibiae dark brown; tarsi yellowish. Connexivum black. Abdomen velvety, sparsely pubescent; shallow punctures inconspicuous on disc; tubercle of trichobothria brown. Anteromedian margin of spiracles yellowish, not elevated. Male. Measurements (N = 9): head length 1.76 ± 0.07 (1.64 – 1.83); width 2.42 ± 0.06 (2.33 – 2.52); pronotum length 2.32 ± 0.11 (2.08 – 2.46); width 5.97 ± 0.19 (5.73 – 6.30); scutellum length 4.03 ± 0.14 (3.78 – 4.22); width 3.59 ± 0.13 (3.34 – 3.78); length of antennal segments: I 0.73 ± 0.03 (0.69 – 0.76); II 0.68 ± 0.04 (0.63 – 0.76); III 1.63 ± 0.07 (1.51 – 1.70); IV 1.38 ± 0.06 (1.26 – 1.45); V 1.72 ± 0.13 (1.51 – 1.89); length of labial segments: I 1.23 ± 0.05 (1.13 – 1.26); II 2.18 ± 0.06 (2.08 – 2.27); III 1.76 ± 0.12 (1.58 – 1.89); IV 1.55 ± 0.12 (1.26 – 1.64); total length 10.61 ± 0.15 (10.40 – 10.80); width of abdomen 6.32 ± 0.14 (6.10 – 6.50).. Genitalia. Dorsal rim widely excavated with 1 + 1 spine projections directed towards the base of segment X. Inferior layer of ventral rim projecting lateral to segment Xover genital cup, perpendicular to the frontal plane of pygophore; projections spatulate, in dorsal view with an elevation midway between apex of projection and posterolateral angle of pygophore. Superior layer of ventral rim bent beneath the projection of inferior layer, forming a translucent, less sclerotized lobe, and looking almost membranous (Figs 4 A, 7 B). Median third of ventral rim concave, emarginate (Fig. 4 B). Lateral margins of segment X sinuous; 1 + 1 lateral tuft of short setae; apex covered with fringe of long setae and narrowed from the base (Fig. 4 A). Parameres sinuous, slender, and lacking projections; head spatulate with obtuse apex, slightly curved ventrally; fringe of setae on lateral and apex of head; base flattened, with dense, short setae (Fig. 5 E – G). Parameres placed ventral to segment X, projecting over and little surpassing the ventral rim of pygophore (Fig. 4 B, D). Phallus. Phallotheca globose, lateral projection wide; ventral and dorsal apical processes short, little projected. Vesica long, pseudojointed and bent backward, surpassing the base of phalotheca; dorsal expansion at basal half of vesica (Fig. 5 A – D). Female. Measurements (N = 15): head length 2.00 ± 0.07 (1.89 – 2.14); width 2.58 ± 0.07 (2.46 – 2.65); pronotum length 2.76 ± 0.11 (2.52 – 2.96); width 6.66 ± 0.26 (6.3 – 7.18); scutellum length 4.65 ± 0.2 (4.35 – 5.04); width 3.97 ± 0.17 (3.65 – 4.28); length of antennal segments: I 0.78 ± 0.04 (0.69 – 0.82); II 0.81 ± 0.07 (0.69 – 0.95); III 1.61 ± 0.09 (1.45 – 1.76); IV 1.49 ± 0.06 (1.39 – 1.58); V 1.82 ± 0.13 (1.58 – 1.95); length of labial segments: I 1.33 ± 0.07 (1.26 – 1.45); II 2.27 ± 0.11 (2.08 – 2.46); III 1.91 ± 0.16 (1.39 – 2.08); IV 1.70 ± 0.09 (1.58 – 1.83); total length 12.23 ± 0.45 (11.7 – 12.9); width of abdomen 7.11 ± 0.25 (6.7 – 7.5). Genitalia. Gonocoxites 8 wider than long; posterior margins sinuous, concave medially, projected over the base of laterotergites 9; disc with median callus. Spiracles of laterotergites 8 on low callus, placed lateral to lateral angle of gonocoxites 8 (Fig. 6 A). Gonocoxites 9 with longitudinal suture; anterior margin subrectilinear (Fig. 6 A, E). Laterotergites 9 not surpassing the band connecting laterotergites 8; median margins on a higher plane than lateral margins; lateral basal angle flat; anterior margin slightly sinuous (Fig. 6 A, E). Gonapophyses 9 with conical microsculptures occupying lateral portion of posterior folds (Figs 6 E, 7 A). Thickening of gonapophyses 9 in wide ‘ V’. Chitinellipsen lateral to thickening of vaginal intima. Ductus receptaculi anteriorly to vesicular area wider than median duct of vesicular area. Length of vesicular area twice the combined length of pars intermedialis and capsula seminalis; base of median duct clavate; inner duct uniformly cylindrical. Tubular portion of ductus receptaculi posterior to vesicular area short, not attaining half length of vesicular area. Pars intermedialis barrelshaped; at least four times longer than capsula seminalis; annular ridges convergent. Capsula seminalis hemispherical; projections long, slender, and twisted, sometimes bifid at apex, placed at base of capsula seminalis (Fig. 6 E).	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFB09053ED1DA951FB08328E.taxon	discussion	Comments: Alitocoris schraderi is in a clade with Ali. grandis sp. nov. and Ali. lateralis sp. nov. (Fig. 1), and can be distinguished by the rostrum reaching the posterior margin of urosternite V, and antennal segment Vbeing the longest. Gonocoxites 8 of Ali. schraderi bears a more tumid callus, and lateral angle more projected over laterotergites 9. The material examined in this study extends the distribution of the species to Honduras and Panama.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBC9052EF4AA861FF2730AC.taxon	materials_examined	Holotype: f, PANAMA, San Blas: (Nusagandi, Ina Trail, 250 – 350 m a. s. l.), 26. vii. 1995, C. W. & L. B. O’Brien, (JEE). Deposited in: AMNH, USA.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBC9052EF4AA861FF2730AC.taxon	etymology	Etymology: The name refers to the size of the species. Latin: grandis, big.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBC9052EF4AA861FF2730AC.taxon	diagnosis	Diagnosis: Black with concolorous punctures. Posterior margin of head and anterior margin of pronotum with yellowish spots. Body surface densely pubescent.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBC9052EF4AA861FF2730AC.taxon	description	Head with small, sparse, brown spots on juga and vertex; yellowish spot on posterior margin, smaller than ocellus; punctures not confluent on apical third of juga. Vertex slightly tumid. Antennal segments I – III black, IV – V brown. Setae uniformly distributed on antennae. Proportional length of antennal segments: I> II <III> IV <V; third the longest. Bucculae lacking anterior tooth. Rostrum black, pubescent, reaching anterior margin of urosternite V; proportional length of labial segments: I <II> III <IV. Pronotum with yellowish spot at middle of anterior margin. Cicatrices flat. Punctures rough, more sparse on disc. Anterolateral angles projected anterolaterally exceeding lateral limit of eyes by the ocellus width. Anterolateral margins slightly sinuous. Scutellum attaining posterior margin of connexival segment V; punctures not confluent on postfrenal lobe. Corium reaching middle of connexival segment VI; subtle, yellowish spots interspersed with punctures. Spot at apex of radial vein as wide as ocellum. Membrane with 12 veins, reaching apex of abdomen. Pleurae with sparse, yellowish spots. Peritreme surpassing half the width of evaporatorium. Coxae and trochanters yellowish with coppery areas; femora and tibiae black; tarsi coppery. Connexivum with yellowish spots at midpoints of segments IV- VII. Abdomen with deep punctures; yellowish spots, punctures, and setae more dense on lateral thirds and inconspicuous on disc; trichobothrial tubercles brown. Anteromesial margin of spiracles flat, yellowish. Male. Unknown. Female. Measurements (N = 1): head length 2.39; width 2.96; pronotum length 3.21; width 7.37; scutellum length 5.17; width 4.41; length of antennal segments: I 0.95; II 0.69; III 2.39; IV 1.70; V 2.02; length of labial segments: I 1.39; II 2.71; III 1.89; IV 1.51; total length 13.80; width of abdomen 8.30. Genitalia. Genital plates pubescent. Gonocoxites 8 wider than long; posterior margins sinuous, slightly concave medially, projected over the base of laterotergites 9; mid callus on disc. Spiracles of laterotergites 8 on ample callus delimited by few punctures, placed lateral to lateral angle of gonocoxites 8. Gonocoxites 9 scarcely exposed, triangular, with longitudinal suture. Laterotergites 9 not surpassing the band connecting laterotergites 8; median margins on a higher plane than lateral margins; lateral basal angle flat (Fig. 6 B).	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBC9052EF4AA861FF2730AC.taxon	discussion	Comments: Alitocoris grandis sp. nov. is in polytomy with Ali. lateralis sp. nov. and Ali. schraderi (Fig. 1). It can be distinguished by the larger size, the presence of a yellowish spot at the middle of the posterior margin of the head, the presence of pubescence on the body. The concavity of the posterior margin of gonocoxites 8 is shallower than in Ali. schraderi and Ali. lateralis sp. nov.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBD9055ECB0ABB0FDB8340B.taxon	materials_examined	Holotype: f, COSTA RICA, Guanacaste: (Fila Orosilito, Est. Pitilla, 9 km south of Santa Cecilia, 800 – 1000 m a. s. l.), 7. iv. 1995, M. Moraga, de Luz L-N 328650 378600 # 4722, [11.1000; - 85.4167], Costa Rica INBio CRI 002 1999 01 (INBio). Deposited in: INBio, Costa Rica.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBD9055ECB0ABB0FDB8340B.taxon	etymology	Etymology: The name refers to the lateral position of the callus on gonocoxites 8. Latin: lateralis, beside.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBD9055ECB0ABB0FDB8340B.taxon	diagnosis	Diagnosis: Dark brown to black; punctures black. Legs lighter coloured, contrasting with body.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBD9055ECB0ABB0FDB8340B.taxon	description	Head black; punctures not confluent on apical third of juga. Vertex slightly tumid. Antennal segments I – III and proximal third of IV brown, Vand distal 2 / 3 of IV yellowish; intersegmental areas translucent; segment Iwith lighter dorsal spot. Proportional length of antennal segments: I> II <III> IV <V; third the longest. Bucculae with anterior triangular tooth projected ventrally. Rostrum light brown, reaching posterior margin of urosternite VI; proportional length of labium segments: I <II <III <IV. Pronotum black, punctures confluent, forming wrinkles between posterior margin of cicatrices and posterior margin of pronotum. Cicatrices flat. Anterolateral angles projected laterally, exceeding lateral limit of eyes by almost three times the ocellus width. Anterolateral margins subrectilinear. Scutellum black, extending to middle of connexival segment V; punctures confluent, forming wrinkles throughout the scutellum. Corium dark brown, reaching middle of connexival segment VI; punctures more dense on exocorium. Spot at apex of radial vein slightly wider than ocellus. Membrane with nine veins, reaching apex of abdomen. Pleurae black. Peritreme surpassing half the width of evaporatorium. Coxae, trochanters, femora, and tibiae light brown; femora and tibiae with darker elliptical spots. Connexivum black. Abdomen black, velvety; punctures shallow, inconspicuous on disc; trichobothria tubercles black. Anteromesial margin of spiracles slightly elevated, concolorous. Male. Unknown. Female. Measurements (N = 1): head length 2.01; width 2.50; pronotum length 2.55; width 6.08; scutellum length 4.51; width 3.92; length of antennal segments: I 0.81; II 0.37; III 1.89; IV 1.36; V 2.17; length of labium segments: I 1.45; II 2.55; III 2.25; IV 2.20; total length 12.15; width of abdomen 6.76. Genitalia. Gonocoxites 8 little longer than wide; posterior margins sinuous, concave medially, projected over the base of laterotergites 9; callus near the lateral margin of gonocoxites, in line with laterotergites 8. Spiracles of laterotergites 8 little elevated, placed lateral to lateral angle of gonocoxites 8. Gonocoxites 9 triangular, lacking longitudinal suture. Laterotergites 9 not surpassing the band connecting laterotergite 8; median margins on a higher plane than lateral margins; lateral basal angle depressed (Fig. 6 C).	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBD9055ECB0ABB0FDB8340B.taxon	discussion	Comments: Alitocoris lateralis sp. nov. is in the polytomy that includes Ali. schraderi and Ali. grandis sp. nov. (Fig. 1). It can be distinguished from the other species by the anterior tooth of bucculae, a longer rostrum reaching posterior margin of urosternite VI, lighter legs with brown spots, and the concolorous anteromesial margin of spiracles. The more concave posterior margins and lateral callus of gonocoxites 8, gonocoxites 9 lacking a suture, and laterotergites 9 with the lateral basal angles depressed are also distinctive features of Ali. lateralis sp. nov.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBA9054ECD4AFE5FD5E30F8.taxon	materials_examined	Holotype: m, PANAMA, Chiriqui: (IHRE vivero 11 km north of Los Planes), 27 May 1995, B. Ratcliffe & M. Jameson, [7.8000; - 81.0333], collected at light (DBTC). Deposited in: AMNH, USA. Paratypes: PANAMA, Chiriqui: 1 f (Reserva Forestal La Fortuna, El Vivero), 20. vii. 1995, C. W. O’Brien, [8.6708; - 822299], UV trap (JEE) – Deposited in: AMNH; Fortuna, 1 f 20. v. 1978, O’Brien & Marshall, 82 ° 15 ′ W, 8 ° 44 ′ N (DARC) – Deposited in: NMNH; 1 f 17. iv. 1977, Henk Wolda, 1050 m a. s. l., 8 ° 42 ′ N, 82 ° 15 ′ W (DARC) – Deposited in: NMNH; 1 f 9. x. 1976, Henk Wolda, 1050 ma. s. l., 8 ° 44 ′ N, 82 ° 15 ′ W (DARC) – Deposited in: DARC.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBA9054ECD4AFE5FD5E30F8.taxon	etymology	Etymology: The name refers to the irregular, yellowish macula on pronotum. Latin: ornatus, ornamentation.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBA9054ECD4AFE5FD5E30F8.taxon	diagnosis	Diagnosis: Dark brown with black punctures and light brown spots; transverse, irregular, light brown macula on pronotum, across humeri. Ventral thorax and abdomen black; legs lighter coloured, contrasting with body.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBA9054ECD4AFE5FD5E30F8.taxon	description	Head black; punctures confluent, forming dorsal wrinkles. Vertex tumid. Antennal segments brown, dorsum of segment Iand sometimes apex of segments IV – V light brown; proportional length of antennal segments: I> II <III> IV <V; third and fifth the longest, subequal. Margins and apex of juga ventrally yellowish. Bucculae with small anterior tooth. Rostrum dark yellowish, surpassing anterior margin of urosternite VII. Proportional length of labium segments: I <II> III <IV. Pronotum bearing transverse, irregular, light brown macula across humeri; punctures sparse on macula, and forming shallow wrinkles posterior to cicatrices. Cicatrices slightly tumid, bearing a light brown middle spot. Anterolateral angles projected laterally, exceeding lateral limit of eyes by twice the ocellus width. Anterolateral margins slightly sinuous. Scutellum attaining middle of connexival segment V; punctures inconspicuous at apex; sometimes with irregular, small, light brown spots between punctures. Corium reaching middle of connexival segment VI, with irregular, small, light brown spots; punctures more dense on exocorium. Spot at apex of radial vein equals ocellus width. Membrane with ten to 13 veins, little surpassing apex of abdomen. Pleurae black with sparse, light brown spots; lateral margins of pronotum ventrally with light brown spots. Peritreme almost reaching lateral margin of evaporatorium. Legs dark yellowish, femora and dorsum of tibia with elliptical brown spots. Connexivum with middle yellowish spot continued ventrally. Abdomen velvety, sparsely pubescent, with shallow punctures; trichobothrial tubercles black; few yellowish spots between spiracles and lateral margin of abdomen. Anteromesial margin of spiracles slightly elevated, concolorous. Male. Measurements (N = 1): head length 2.06; width 2.60; pronotum length 2.65; width 6.57; scutellum length 4.46; width 3.38; length of antennal segments: I 0.78; II 0.31; III 1.86; IV 1.40; V 1.95; length of labial segments: I 1.60; II 2.80; III 2.30; IV 2.45; total length 12.15; width of abdomen 6.81. Genitalia. Dorsal rim of pygophore concave; 1 + 1 obtuse projections near base of segment X, and 1 + 1 spine-like projections halfway between base of segment Xand posterolateral angles of pygophore. Inferior layer of ventral rim projecting lateral to segment Xover genital cup, perpendicular to the frontal plane of pygophore; projections spatulate, apex obtuse and light brown. Superior layer of ventral rim beneath the projection of inferior layer forming striations and lobes, appears membranous (Fig. 4 C). Median third of ventral rim concave. Ventral wall of pygophore with low median carina (Fig. 4 D). Lateral margins of segment X sinuous; apex covered with short setae (Fig. 4 C). Parameres placed ventral to segment X, projecting over the ventral rim of pygophore; head of parameres spatulate with dense, long setae (Fig. 4 D). Female. Measurements (N = 4): head length 2.05 ± 0.12 (1.95 – 2.21); width 2.74 ± 0.04 (2.71 – 2.77); pronotum length 3.04 ± 0.29 (2.65 – 3.34); width 7.1 ± 0.16 (6.93 – 7.31); scutellum length 5.15 ± 0.17 (4.98 – 5.36); width 4.24 ± 0.23 (3.91 – 4.41); length of antennal segments: I 0.8 ± 0.06 (0.76 – 0.88); II 0.55 ± 0.03 (0.5 – 0.57); III 1.83 ± 0.07 (1.76 – 1.89); IV 1.43 ± 0.03 (1.39 – 1.45); V 2.05 ± 0.08 (1.95 – 2.14); length of labial segments: I 1.65 ± 0.08 (1.58 – 1.76); II 2.79 ± 0.14 (2.65 – 2.96); III 2.46 ± 0.09 (2.33 – 2.52); IV 2.55 ± 0.11 (2.46 – 2.71); total length 13.23 ± 0.47 (12.9 – 13.9); width of abdomen 6.96 ± 0.34 (6.67 – 7.46). Genitalia. Gonocoxites 8 wider than long, disc uniformly convex, tumid posteriorly forming a flap; few setae on posterior third; transverse wrinkles more dense on lateral third; posterior margin subrectilinear, posterolateral angles as small tooth over the base of laterotergites 8; sutural angles forming obtuse projection over gonocoxites 9; basal two thirds of sutural margins slightly elevated (Fig. 6 D). Laterotergites 8 sparsely punctate at base; spiracles on low callus, posterior to lateral angle of gonocoxites 8 (Fig. 6 D). Gonocoxites 9 triangular with longitudinal suture; anterior margin emarginate (Fig. 6 D, F). Laterotergites 9 not surpassing the band connecting laterotergites 8; median margins on a higher plane than lateral margins, the latter narrow and deflected from basal angle almost to the apex; lateral basal angle depressed; anterior margin subrectilinear (Fig. 6 D, F). Gonapophyses 8 slightly sclerotized at base; lateral folds with conical microsculptures. Gonapophyses 9 with folds bearing conical microsculptures along lateral third of gonocoxites 9. Thickening of gonapophyses 9 in wide ‘ V’. Chitinellipsen lateral to thickening of vaginal intima. Ductus receptaculi anterior to vesicular area wider than median duct of vesicular area. Length of vesicular area three times the combined length of pars intermedialis and capsula seminalis; base of median duct clavate; inner duct uniformly cylindrical. Tubular portion of ductus receptaculi posterior to vesicular area short, less than half length of vesicular area. Pars intermedialis long, at least four times longer than capsula seminalis; annular ridges convergent. Capsula seminalis hemispherical; projections long, slender, and twisted, sometimes bifid at apex, positioned at apex of capsula seminalis (Fig. 6 F).	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
03D687E2FFBA9054ECD4AFE5FD5E30F8.taxon	discussion	Comments: Alitocoris ornatus sp. nov. is included in clade Ain a polytomy with the clade of Ali. schraderi (Fig. 1). It is included in Alitocoris until there is a better understanding of its phylogenetic position, and also based on general morphology, in particular because it shares with Ali. schraderi the apomorphic character states of the male and internal female genitalia, and with Ali. lateralis sp. nov. the apomorphic character states of laterotergites 9. The presence of a pronotal macula, the gonocoxites 8 sutural angles projecting over gonocoxites 9, and the lateral angles of gonocoxites 8 little projected over laterotergites 8 are unique characters of Ali. ornatus sp. nov.	en	Garbelotto, Thereza De A., Campos, Luiz A., Grazia, Jocelia (2013): Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society 168: 452-472, DOI: 10.1111/zoj.12032
