identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D61C0EFF95F96F350461EA181A6940.text	03D61C0EFF95F96F350461EA181A6940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichopagurus	<div><p>Trichopagurus de Saint Laurent, 1968</p><p>Trichopagurus de Saint Laurent, 1968: 927; 1970: 210; McLaughlin, 2003a: 128.</p><p>Type species. Catapaguroides ? trichophthalmus Forest, 1954 . Gender: Masculine.</p><p>Emended diagnosis. Gills biserial, 11 pairs, including 1 pleurobranch above third pereopod (sixth thoracic somite) and 2 arthrobranchs on each third maxilliped to fourth pereopods; most lamellae distinctly narrowed distally. Rostrum distinct, triangular. Ocular acicles simple. Third maxilliped with 1 accessory tooth. Chelipeds strongly unequal, with weak degree of sexual dimorphism. Fourth pereopod semichelate, propodal rasp consisting of 1 row of corneous scales; no preungual process. Coxa of male right fifth pereopod with moderately long sexual tube directed toward exterior, distal part of tube semitransparent, coiled; left with very short tube. Female with single left gonopore; coxae of fifth pereopods slightly to somewhat asymmetrical, left largest. Anterior lobe of sixth thoracic somite transversely oblong. Eighth thoracic sternite composed of 2 distinct, rounded lobes. Male with unpaired left third to fifth pleopods; female with unpaired left second to fifth pleopods, no paired first pleopods modified as gonopods. Uropods markedly asymmetrical. Telson with faint or distinct transverse indentations, terminal margins oblique; posterior lobes each with fringe of long setae on lateral margin.</p><p>Remarks. As noted before, the gills of Trichopagurus trichophthalmus have been reported as ‘intermediaire’ (de Saint Laurent 1968, 1970) or quadriserial (McLaughlin 2003a). De Saint Laurent (1970) illustrated a shallowly divided gill lamella. However, our examination of the holotype and additional specimens has shown that the gill lamellae of T. trichophthalmus are actually entire, lacking distal division. Most gill lamellae, however, are abruptly narrowed distally, and the partially overlapping distal parts of a few gill lamellae might give the incorrect impression that each lamella is divided distally.</p><p>It is worth comparing Trichopagurus with Anapagrides de Saint Laurent­Dechancé, 1966 and Catapagurus A. Milne­Edwards, 1880 (rediagnosed by McLaughlin 2004), as all are characterized by 11 pairs of biserial gills, strongly unequal chelipeds, single row of scales in the propodal rasp on the fourth pereopod, and some development of a right sexual tube of male. Anapagrides primarily differs from Trichopagurus in the structure of the right sexual tube. In Anapagrides, the right sexual tube is short (about 0.5–1.2 times the length of the coxa of the fifth pereopod) and is posteriorly directed. Catapagurus is easily distinguished from Trichopagurus by the broadly rounded rostral lobe, the possession of a ventral row of very small spinules on the ambulatory dactyli, instead of rather strong corneous spines, and the presence of paired female gonopores. In addition, Catapagurus species occasionally have elongate ocular acicles and usually a well developed preungual process on the dactylus of the fourth pereopod.</p></div>	https://treatment.plazi.org/id/03D61C0EFF95F96F350461EA181A6940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Komai, Tomoyuki;Osawa, Masayuki	Komai, Tomoyuki, Osawa, Masayuki (2005): Decapoda: Anomura: Paguridae) from the Ryukyu and Yap Islands, and redescription of T. trichophthalmus (Forest). Zootaxa 801: 1-20, DOI: 10.5281/zenodo.170561
03D61C0EFF96F967350461B819076EF0.text	03D61C0EFF96F967350461B819076EF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichopagurus macrochela	<div><p>Trichopagurus macrochela, n. sp. (Figs. 1–3)</p><p>Trichopagurus trichophthalmus . — Baba, 1982: 68 (part).</p><p>Type material. HOLOTYPE: male SL 1.8 mm, Hiyajo, Kume­jima Island, Ryukyu Islands, 10 m, 12 June 1995, SCUBA diving, coll. K. Nomura, CBM­ZC 7853.</p><p>PARATYPES: 2 males SL 1.6, 2.0 mm, 1 ovigerous female SL 2.5 mm, Ahra Beach, Kume­jima Island, Ryukyu Islands, subtidal, 15 June 1995, hand, coll. T. Komai, CBM­ ZC 7854; 1 male SL 1.5 mm, 1 female SL 1.3 mm, Urabishi, Kuroshima Island, Ryukyu Islands, 3–4 m, 10 July 1998, skin diving, coll. K. Nomura and T. Komai, CBM­ZC 7855.</p><p>Other material examined. 1 ovigerous female SL 1.7mm, Off Inuf, Yap Islands, lagoon, 29 June 1980, coll. K. Baba, NSMT­Cr 14559.</p><p>Description. Gill lamellae narrowed distally with concave distolateral margins (Fig. 1 A).</p><p>Shield (Fig. 1 B, C) approximately as long as broad; anterior margin between rostrum and lateral projections concave; anterolateral margins sloping; posterior margin truncate; dorsal surface weakly convex, with 1 median and 2–5 dorsolateral tufts of setae. Rostrum triangular or roundly subacute, distinctly overreaching lateral projections, terminating bluntly or subacutely. Lateral projections obsolete, with prominent marginal or submarginal spine. Carapace lateral lobes (Fig. 1 C) narrow, moderately calcified. Posterior carapace (Fig. 1 C) generally membranous except for weakly calcified anterior part of posteromedian plate; posteromedian plate with 1 pair of tufts of setae; cardiac sulci obsolescent; sulci cardiobranchialis appearing as opaque lines on either side of posteromedian plate, extending posterolaterally; branchial region with few short setae.</p><p>Ocular peduncles (including cornea) (Fig. 1 B) stout, about 0.7 times of shield length; not inflated basally; cornea slightly dilated; dorsal surface with tuft of long setae near base of cornea. Ocular acicles subtriangular or subovate, separated basally by width of one acicle, with small submarginal spine terminally; dorsal surface concave.</p><p>Antennular peduncles (Fig. 1 B), when fully extended, overreaching ocular peduncles (including cornea) by 0.5–0.7 length of ultimate segment. Ultimate segment approximately twice length of penultimate segment, somewhat widened distally in lateral view, with tuft of long setae at dorsodistal lateral angle. Penultimate segment short, without setae. Basal segment with distolateral margin distinctly produced as short process, statocyst lobe weakly inflated, unarmed laterally.</p><p>Antennal peduncles (Fig. 1 B) overreaching ocular peduncles by 0.2–0.3 length of fifth segment, with supernumerary segmentation. Fifth and fourth segments with few setae distally. Third segment with spinule at ventromesial distal margin. Second segment with dorsolateral distal angle produced, not reaching midlength of fourth segment, terminating in simple, acute spine; dorsomesial distal angle with small spine; mesial and lateral faces with few setae. First segment with small spine at dorsolateral distal angle; ventromesial distal margin strongly produced anteriorly, with 1 minute spinule laterally. Antennal acicle reaching or slightly overreaching distal margin of cornea, weakly arcuate, distal part abruptly tapering and terminating in small spine; dorsomesial margin with sparse setae. Antennal flagellum (missing in holotype) 4.0–5.0 length of shield; each article with 3–5 very short to moderately long setae on distal margin.</p><p>Mandible without distinctive features. Maxillule (Fig. 2 A) with subquadrate coxal endite; endopod with outer lobe obsolete, inner lobe slightly produced, with 1 apical seta. Maxilla (Fig. 2 B) with broad proximal lobe of coxal endite; endopod overreaching anterior margin of moderately broad scaphognathite. First maxilliped (Fig. 2 C) with moderately broad exopod; endopod distinctly overreaching anterior margin of basial endite. Second maxilliped (Fig. 2 D) with basis­ischium fusion incomplete; dactylus and carpus stout. Third maxilliped (Fig. 2 E) moderately stout; carpus unarmed on dorsodistal margin; merus with strong dorsodistal spine and small ventromesial spine; ischium (Fig. 2 F) with welldeveloped crista dentata composed of very small corneous teeth and 1 accessory tooth; basis­ischium fusion incomplete; basis apparently unarmed on ventromesial margin; exopod slightly falling short of carpus.</p><p>Chelipeds strongly unequal in both sexes. Right cheliped (Fig. 3 A–C) very large, moderately stout. Chela elongate, subovate in dorsal view, 2.0–2.1 times as long as broad (greatest width at about midlength of palm). Dactyl about 0.7 length of palm, weakly curved ventrally; dorsal surface minutely granular proximally, dorsomesial margin not delimited; all surfaces with scattered tufts of short to moderately short setae; cutting edge with 2 or 3 large calcareous teeth and very small corneous teeth interspersed, terminating in small calcareous claw. Palm equal in length to carpus; dorsomesial margin not delimited, dorsolateral margin slightly delimited by multiple rows of small granules; dorsal surface weakly convex, nearly smooth medially and granular laterally and mesially, and with few short to moderately short setae; lateral and mesial surfaces minutely granular; ventral surface convex, smooth or microscopically granular, with scattered tufts of long setae extending onto fixed finger. Fixed finger with scattered tufts of short setae on dorsal surface; cutting edge with 1 large tooth proximally and row of small calcareous claw, terminating in small calcareous claw. Carpus flattened dorsoventrally, approximately equal to merus in length; dorsomesial margin clearly delimited by row of moderately large spines and some distal tubercles, dorsodistal margin unarmed, dorsolateral margin weakly delimited by row of low tubercles or protuberances; dorsal surface with some small, low tubercles laterally and scattered tufts of moderately long setae; lateral surface granular, ventrolateral margin with row of small, low protuberances; mesial surface shallowly concave, microscopically granular; distal part of ventral surface concave, bordered by distinct tuberculate ridge bearing tufts of long setae, posterior part convex, with few small tubercles and tufts of long setae. Merus with subdistal transverse ridge extending onto mesial face and few tufts of setae on dorsal surface, dorsodistal margin unarmed; lateral and mesial faces smooth, ventrolateral margin with row of small spines becoming smaller and blunt proximally and tufts of long setae, ventromesial margin with large, blunt tooth proximally and row of long stiff setae; ventral surface with tufts of long setae distally. Ischium with short spiniform setae and moderately short setae on surfaces, ventromesial margin nearly smooth. Coxa unarmed.</p><p>Left cheliped (Fig. 3 D–F) slender. Chela about 3.5 times as long as broad (greatest breadth at about base of fingers). Dactylus slightly longer than palm; dorsomesial margin not delimited, surfaces nearly smooth, with tufts of short to long setae; cutting edge with row of small, slender corneous teeth, terminating in small corneous claw. Palm with dorsal surface weakly elevated in midline, bearing several very small tubercles and scattered short setae; dorsolateral margin of palm and proximal half of fixed finger weakly delimited by row of small, low protuberances, dorsomesial margin not delimited; lateral face with scattered low tubercles; mesial face with low protuberances and long setae; ventral surface convex, with few low protuberances and long setae. Cutting edge of fixed finger with row of small, blunt corneous teeth, terminating in small corneous claw. Carpus equal to or slightly shorter than merus; dorsolateral and dorsomesial margins each with row of small, low protuberances accompanied by spiniform setae; lateral and mesial faces with scattered low protuberances, and short to long setae; ventral surface slightly convex, with low protuberances and scattered long setae. Merus with row of low transverse ridges and few tufts of short setae on dorsal surface, dorsodistal margin unarmed; lateral face with scattered very small, low tubercles and short stiff setae, ventrolateral margin with 2 or 3 small spines distally and row of long stiff setae; mesial face with short, low transverse ridges bearing long stiff setae adjacent to ventromesial margin, ventromesial margin also with row of long stiff setae; ventral surface without distinct spines or tubercles, but with short oblique rows of long stiff setae. Ischium with short to long setae on dorsal and ventral surfaces and cluster of short spiniform setae at ventrolateral angle. Coxa unarmed.</p><p>Ambulatory legs (Fig. 2 G, H) elongate, overreaching tip of extended left cheliped, but not reaching tip of right cheliped. Dactyli slender, in dorsal view straight, in lateral view almost straight with slightly curved claw, 1.1–1.3 times longer than propodi (third pair slightly longer than second); dorsal margins with row of tufts of short to moderately short setae; lateral faces with few short setae; mesial faces (Fig. 2 I, J) also with few short setae, unarmed on second, but armed with corneous spinules dorsally on third; ventral margins each with 8–10 long corneous spines. Propodi 1.75–2.00 times length of carpi; dorsal surfaces each with widely­spaced tufts of moderately long stiff setae; lateral and mesial faces with few setae; ventral surface with 3 or 4 widely­spaced, long corneous spines, including 2 or 3 spines on ventrodistal margin. Carpi short, 0.50–0.70 length of meri; dorsal surfaces unarmed but with moderately short to long stiff setae; lateral faces each with row of tufts of setae on midline; ventral surfaces with few short to long, stiff setae. Meri with row of short transverse ridges and stiff setae on dorsal surfaces; lateral and mesial faces mostly naked; ventral surfaces smooth or with row of low protuberances and tufts of moderately long setae, ventrolateral distal margins unarmed. Ischia with short to long setae on dorsal and ventral margins.</p><p>Fourth pereopods (Fig. 1 D) semichelate. Dactyli each with row of minute corneous teeth on distal half of ventral margin, terminating in tiny corneous claw; no preungual process. Propodal rasp composed of single row of small corneous scales on distal 0.7 of ventral margin. Carpi with tufts of long setae on dorsal surface, dorsodistal margin unarmed. Meri with tuft of long setae on dorsodistal margin.</p><p>Fifth pereopods (Fig. 1 E) chelate. Male coxae (Fig. 1 H) slightly asymmetrical, both with short stiff setae on anterior and posterior margins. Female coxae (Fig. 1 I) asymmetrical, left larger, with oblique transverse row of numerous long setae on ventral surface.</p><p>Third thoracic sternite with anterior margin slightly produced medially, unarmed. Anterior lobe of sixth thoracic sternite (Fig. 1 F, G) transversely oblong or subrectangular, with row of short setae on anterior margin. Eighth thoracic sternite (Fig. 1 H, I) composed of 2 rounded, contiguous lobes.</p><p>Male with moderately long (3–4 times of coxal length) right sexual tube directed toward exterior and upward along lateral side of body, distal part transparent and coiled (Fig. 1 E, H). Right sexual tube (Fig. 1 H) very short. Female with only left gonopore.</p><p>Abdomen twisted. Male with 3 unpaired, very unequally biramous left pleopods (third to fifth pleopods). Female with 4 unpaired left pleopods, second to fourth subequally biramous; fifth pleopod very small, uniramous. Uropods markedly asymmetrical; protopods unarmed.</p><p>Telson (Fig. 1 J) with only slight lateral indentations; posterior lobes triangular or rounded, terminal margins oblique, each with row of 4–7 very small spinules, lateral margins with fringe of long setae (Fig. 1 K).</p><p>Color in life. Carapace, antennae, chelipeds and ambulatory legs generally pale reddish brown, without conspicuous markings; setae at base of cornea accentuated with dark reddish brown.</p><p>Ecology. Found to inhabit small crevices or holes on dead coral. Various gastropod shells were used as housing.</p><p>Distribution. Presently known only from Ryukyu and Yap Islands; subtidal to 10 m.</p><p>Etymology. From the Latin macro meaning large and chela meaning cheliped and referring to the very large right cheliped.</p><p>Remarks. This new species is characterized by the very large right cheliped with a dorsoventrally flattened carpus and markedly asymmetrical coxae of the female fifth pereopods, of which the left is the largest and is provided with numerous long setae on the ventral surface. Furthermore, the following notable differences are found between T. macrochela and T. trichophthalmus . The dorsomesial margin of the carpus of the right cheliped is distinctly delimited by a row of spines in T. macrochela, while in T. trichophthalmus, the carpus is subcylindrical or slightly flattened, without distinct delineation of the dorsomesial margin. The surfaces of the right cheliped are granular or tuberculate in A. macrochela, rather than smooth and polished in T. trichophthalmus . The chelipeds and ambulatory legs are distinctly less setose in T. macrochela than in T. trichophthalmus . In addition, males of T. macrochela possess unpaired, unequally biramous pleopods, but those of T. trichophthalmus have uniramous pleopods.</p><p>Baba (1982) recorded T. trichophthalmus from the Palau and Yap Islands based on four specimens. Two of the four specimens are now deposited in the collection of the National Science Museum, Tokyo, although other two were not located. Reexamination of these two specimens revealed that the male (NSMT­Cr 14558) from Palau is referable to true T. trichophthalmus, but an ovigerous female (NSMT­Cr 14559) from the Yap Islands represents T. macrochela .</p></div>	https://treatment.plazi.org/id/03D61C0EFF96F967350461B819076EF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Komai, Tomoyuki;Osawa, Masayuki	Komai, Tomoyuki, Osawa, Masayuki (2005): Decapoda: Anomura: Paguridae) from the Ryukyu and Yap Islands, and redescription of T. trichophthalmus (Forest). Zootaxa 801: 1-20, DOI: 10.5281/zenodo.170561
03D61C0EFF9EF97C350465201FBA6988.text	03D61C0EFF9EF97C350465201FBA6988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichopagurus trichophthalmus (Forest 1954) Forest 1954	<div><p>Trichopagurus trichophthalmus (Forest, 1954) (Figs. 4–6)</p><p>Catapaguroides ? trichophthalmus Forest, 1954: 74, figs. 20–24; 1956: 1069.</p><p>Trichopagurus trichophthalmus .— de Saint Laurent, 1968: 927; 1970: 212, figs. 1–16; Baba, 1982: 68 (part); Poupin, 1996: 19; Hogarth et al., 1998: 168; Asakura 2004: 460, fig. 3.</p><p>Type material. HOLOTYPE: male SL 1.7 mm, Tahiti, French Polynesia, 1953, coll. Ranson, MNHN­Pg 602.</p><p>Other material examined. Marshall Islands: 1 male SL 1.4 mm, Bikini, 16 August 1947, coll. USNM, MNHN­Pg 603; 1 female SL 1.6 mm, same locality, April 1946, coll. USNM, MNHN­Pg 604; 1 male SL 2.2 mm, similar locality, 17 August 1947, coll. USNM, MNHN­Pg 606. Palau Islands: 1 male SL 1.1mm, Northeast coast of Kayangel, 8 July 1980, coll. K. Baba, NSMT­Cr 14558. Ryukyu Islands: 1 ovigerous female SL 1.7 mm, Near Sesoko Station of the University of the Ryukyus, Sesoko Island, 1 m, 16 July 2003, coll. M. Osawa and Y. Fujita, NSMT­Cr 15987; 1 male SL 1.7 mm, Ahra, Kumejima Island, outer reef flat, intertidal, 13 June 1995, coll. M. Osawa, NSMT­Cr 15233; 1 female SL 1.6 mm, same locality, moat, intertidal, 14 June 1995, coll. M. Osawa, NSMT­ Cr 15234; 1 male SL 1.7 mm, Shimajiri, Kume­jima Island, 1 m, 30 June 2000, coll. M. Osawa, NSMT­Cr 15988; 1 female SL 1.6 mm, Hizushihama, Aka­shima Island, moat, subtidal, 23 April 1993, coll. M. Osawa, NSMT­Cr 15989; 1 male SL 1.8 mm, in front of the Yaeyama Marine Park Research Station, Kuro­shima Island, moat, subtidal, 7 September 1992, coll. M. Osawa, NSMT­Cr 15990; 1 male SL 1.7 mm, same locality, moat, subtidal, 9 September 1992, coll. M. Osawa, NSMT­Cr 15991. Maldives: 1 female SL 1.5 mm, South coast of Vadoo, 10 m, 19 April 1996, SCUBA diving, coll. K. Nomura, CBM­ ZC 7802; 1 male SL 1.7 mm, Helmut Reef, South Male Atoll, 3 m, 30 August 2003, on SCUBA, coll. K. Nomura, CBM­ZC 7803.</p><p>Redescription. Eleven pairs of biserial phyllobranchiae including 1 pleurobranch above third pereopod (sixth thoracic somite) and 2 arthrobranchs on each third maxilliped to fourth pereopods; gill lamellae (Fig. 4 A) narrowed distally with concave distolateral margins.</p><p>Shield (Fig. 4 B, C) slightly longer or as long as broad; anterior margin between rostrum and lateral projections concave; anterolateral margins sloping; posterior margin truncate; dorsal surface weakly convex, with 1 or 2 median and 5–7 lateral tufts of setae; paragastric grooves indistinct. Rostrum narrowly rounded, distinctly overreaching lateral projections, abruptly tapering and terminating in spinule. Lateral projections low, broadly triangular, with prominent submarginal spine. Carapace lateral lobes narrow, moderately calcified. Posterior carapace (Fig. 4 C) with few short setae, generally membranous except for weakly calcified anterior parts of posteromedian and posterolateral plates; cardiac sulci short, indistinct; posterolateral plates each with 1 or 2 tufts of setae on median part; sulci cardiobranchialis appearing as opaque lines, extending posterolaterally.</p><p>Ocular peduncles (including cornea) (Fig. 4 B) stout, 0.5–0.6 times of shield length; not inflated basally; cornea slightly dilated; dorsal surfaces with tufts of long setae along distal half of dorsomesial margin and near dorsolateral base of cornea. Ocular acicles subovate, separated basally by width of one acicle, with small submarginal spine terminally; dorsal surface concave; mesial margins with short and moderately long setae.</p><p>Antennular peduncles (Fig. 4 B), when fully extended, overreaching ocular peduncles (including cornea) by full length of ultimate segment. Ultimate segment approximately 1.5 times longer than penultimate segment, widened distally in lateral view, with tuft of long setae at dorsodistal margin; dorsomesial margin with few setae medially. Penultimate segment apparently naked. Basal segment with distolateral margin distinctly produced, terminating acutely; statocyst lobe weakly inflated, with minute spinule laterally.</p><p>Antennal peduncles (Fig. 4 B) overreaching ocular peduncles by 0.2–0.5 length of fifth segment, with supernumerary segmentation. Fifth and fourth segments with tufts of short and long setae. Third segment with small spine on ventromesial distal margin and few tufts of setae. Second segment with dorsolateral distal angle produced, reaching midlength of fourth segment, terminating in simple, acute spine, with tuft of long setae; dorsomesial distal angle with small spine; mesial margin with short setae. First segment with minute spine at dorsolateral distal angle; ventromesial distal margin strongly produced anteriorly, with 1 minute spinule laterally. Antennal acicle overreaching distal margin of cornea by 0.3 length, weakly arcuate, distal part abruptly tapering and terminating in spinule; dorsomesial margin with tufts of long setae. Antennal flagellum approximately 4.0 times as long as shield, with articles each bearing 4 or 5 short and long setae on distal margin.</p><p>Maxillule (Fig. 4 D) with subquadrate coxal endite; endopod with outer lobe moderately produced, inner lobe slightly produced, with 1 apical seta. Third maxilliped (Fig. 4 E) moderately stout; carpus unarmed on dorsodistal margin; merus with dorsodistal spine; ischium (Fig. 4 F) with well­developed crista dentata composed of small corneous teeth and 1 accessory tooth; basis­ischium fusion incomplete; basis with 1 spinules on ventromesial margin; exopod reaching half length of carpus.</p><p>Chelipeds strongly unequal in both sexes. Right cheliped (Figs. 5 A, B, 6A, B) large, occasionally strongly elongate; rotation of propodal­carpal articulation 30–45º counterclockwise from perpendicular. Chela subovate in dorsal view, 2.5 times as long as broad (greatest width at about midlength of palm). Dactylus 0.5–0.6 length of palm in male, 0.8 times in female, weakly curved ventrally; articulation with palm strongly oblique; dorsal surface convex, dorsomesial and ventromesial margins not delimited; all surfaces with tufts of short to moderately short setae; cutting edge with 2 large and some smaller calcareous teeth in proximal half and small, calcareous or corneous teeth in distal half, terminating in small calcareous claw. Palm 0.7–0.8 times length of carpus in males, subequal in females; dorsomesial and dorsolateral margin weakly delimited; dorsal surface convex, apparently smooth, with numerous tufts of short to moderately long setae; lateral and mesial surfaces also smooth, with scattered tufts of setae; ventral surface convex, smooth, with scattered tufts of short and long setae extending onto fixed finger. Fixed finger with scattered tufts of moderately long setae on dorsal surface; cutting edge with 1 or 2 large, blunt teeth in proximal half and small rounded calcareous teeth in distal half, terminating in small calcareous claw. Carpus subcylindrical or slightly flattened dorsoventrally, longer than merus; dorsal surface flattish or slightly convex, with numerous tufts of short to long setae, dorsodistal margin unarmed; dorsomesial margin not delimited or slightly delineated, unarmed in holotype, but armed with 1–3 small spines on proximal half in other specimens, dorsolateral margin not delimited or weakly delineated, unarmed; mesial and lateral surfaces also with scattered tufts of short to long setae; ventromesial and ventrolateral margins unarmed; ventral surface smooth, concave in proximal and distal parts and convex in median part, with scattered tufts of short and long setae. Merus nearly smooth on dorsal, mesial and lateral surfaces; dorsodistal margin unarmed but with moderately long setae; ventrolateral margin with 1–3 small spines on distal half and tufts of long setae, ventromesial margin with 2 small spines on distal half and tufts of short and long setae; ventral surface flat, mostly smooth. Ischium with short to moderately long setae on surfaces, ventromesial margin with small, low protuberances. Coxa unarmed but with short setae on ventrodistal margin.</p><p>Left cheliped (Figs. 5 C, D, 6C, D) slender; rotation of propodal­carpal articulation 20– 45º clockwise from perpendicular. Chela 3.5–3.6 times as long as broad (greatest width at about midlength of palm). Dactylus 0.9–1.0 length of palm; articulation with palm strongly oblique; surfaces smooth, with tufts of short to long setae; dorsal surface slightly convex, dorsomesial margin not delimited; cutting edge with row of small, slender corneous teeth, terminating in small corneous claw. Palm 0.5–0.6 times longer than carpus, surfaces smooth, with scattered tufts of short to long setae; dorsal surface slightly convex; dorsomesial and dorsolateral margins slightly delimited proximally; ventral surface convex. Cutting edge of fixed finger with row of small, blunt calcareous teeth, terminating in small corneous claw. Carpus elongate, as long as or slightly longer than merus; dorsomesial margin delimited, with 2–4 small spines and short, faint transverse ridges; dorsolateral margin delimited by short, faint transverse ridges, unarmed or armed with 1 or 2 small spines; dorsal surface flattish, with scattered tufts of long or moderately long setae; mesial and lateral surfaces with short, indistinct, low transverse ridges each bearing tuft of short to long setae; ventral surface slightly convex, scattered long or moderately long setae. Merus with row of low, very short transverse ridges and few tufts of moderately long setae on dorsal surface, dorsodistal margin unarmed but with long or moderately long setae; lateral surface with row of few tufts of short to moderately long setae on midline and row of longer setae ventrally, ventrolateral margin with 1–4 small spines on distal half and small low protuberances; mesial surface with tufts of short to moderately long setae ventrally, ventromesial margin with 1 or 2 small spines on distal half; ventral surface nearly smooth. Ischium with short to moderately long setae on dorsal and ventral surfaces, ventromesial margin with small, low protuberances. Coxa unarmed but with short setae on ventrodistal margin.</p><p>Ambulatory legs (Fig. 6 E, G) generally similar from right to left, right slightly longer than left, slender, elongate, overreaching anterior tip of extended right cheliped by approximately half length of dactylus. Dactyli (Fig. 6 F) slender, in dorsal view slightly curved, in lateral view almost straight with curved slender claw, approximately equal in length to propodi; dorsal margins with row of tufts of short to moderately long setae; lateral and mesial faces unarmed but with few short to moderately long setae; ventral margins each with 7–9 (second) and 8–10 (third) slender corneous spines, decreasing in size proximally. Propodi 1.2–1.4 times length of carpi, somewhat tapering distally in lateral and dorsal views; dorsal surfaces with numerous tufts of short to long setae; lateral faces with weak or very weak, rounded crest on midline and row of tufts of short to long setae each along dorsal and ventral margins; mesial faces also each with short to long setae along dorsal and ventral margins; ventral surfaces each with 4­6 widely­spaced, slender corneous spines, including 1 or 2 longer spines on ventrodistal margin. Carpi moderately long, 0.7–0.8 length of meri; dorsal surfaces unarmed but with moderately long or long setae; lateral faces each with row of tufts of setae on midline; mesial faces nearly naked; ventral surfaces with few short to long setae. Meri with row of short transverse ridges bearing short to long setae on dorsal surfaces; lateral faces with tufts of setae on proximoventral and distal parts, more numerous in third; mesial faces with few short setae; ventral surfaces smooth or with row of low protuberances, and with long to moderately long setae, ventrolateral distal margins unarmed. Ischium with short to long setae on dorsal and ventral margins; lateral and mesial faces naked. Female with only left gonopore.</p><p>Fourth pereopods (Fig. 6 H) semichelate. Dactyli each with row of minute corneous teeth on distal half of ventral margin, terminating in tiny corneous claw; no preungual process. Propodal rasp composed of single row of small corneous scales on distal 0.7 of ventral margin. Carpi with tufts of long setae on dorsal surface, dorsodistal margin unarmed. Meri with tuft of long setae on dorsodistal margin.</p><p>Fifth pereopods chelate. Coxae of male and female slightly asymmetrical (Fig. 4 H, I), with short and long setae on anterior and posterior margins in males but only on anterior margin in females. Male with moderately long (about 4 times longer than coxa of fifth pereopod measured on ventral surface) right sexual tube (Fig. 4 H, J) directed toward exterior and upward along lateral side of body, distal part transparent and slightly to somewhat coiled. Right sexual tube (Fig. 4 H) very short, papilla­like.</p><p>Third thoracic sternite with anterior margin slightly produced medially, with pair of spinules. Anterior lobe of sixth thoracic sternite (Fig. 4 G) transversely oblong, with row of long setae on anterior margin. Eighth thoracic sternite (Fig. 4 H, I) composed of 2 rounded, contiguous lobes, with few setae on anterior surface.</p><p>Abdomen twisted. Male with 3 left unpaired, uniramous pleopods (third to fifth pleopods); fifth pleopod shorter than others. Female with 4 unpaired left pleopods, second to fourth pleopods uniramous or biramous with endopods about half­length of exopods; fifth shorter than others, uniramous. Uropods markedly asymmetrical; protopods unarmed. Telson (Fig. 4 K) with distinct lateral indentations; dorsal surface with short setae; posterior lobes rounded, terminal margins oblique, each with row of 3–7 small spines, posterolateral margins each with fringe of long setae.</p><p>Color in life. Carapace, antennae, chelipeds and ambulatory legs yellowish orange, with no conspicuous markings.</p><p>Ecology. Found living among branches of dead corals; intertidal to 10 m. Shells carried by crabs were small compared with the body size, and thus the cephalothorax and thoracic appendages were not completely retracted in shell. The animals were observed to move very rapidly like some Pagurixus species when disturbed.</p><p>Distribution. Previously known from Tahiti, Bikini, Palau, Ponape, the Sulu Archipelago of the Philippines, and Sanguinisiapo Archipelago of Indonesia and Japan in the West Pacific Ocean (de Saint Laurent 1970, Baba 1982, Asakura 2004), and the Maldives in the Indian Ocean (Hogarth et al. 1998).</p><p>Remarks. The present specimens are very similar, although they came from widely distant locations in the Indian and Pacific oceans. Two females from the Ryukyu Islands (SL 1.6 mm, NSMT­Cr 15234; SL 1.6 mm, NSMT­Cr 15989) have uniramous pleopods (the former possesses second to fourth pleopods, but the latter lacks the fourth pleopod), although other females all have biramous second to fourth pleopods. This seems to be due to immaturity since these two specimens each have a left gonopore, which is distinctly smaller than that of other females.</p><p>De Saint Laurent (1970) tentatively referred two small specimens collected from Indonesia and the Philippines by the Siboga Expedition to T. trichophthalmus because of slightly longer ocular peduncles, slightly more slender chelipeds, and less setose ocular peduncles and chelipeds. Although the Siboga specimens were not available for study, examination of the present material suggests that these differences are size­related.</p><p>Asakura (2004) reported examining the holotype of T. trichophthalmus, with the registration number MNHN­Pg 3403. In fact, the holotype of T. trichophthalmus, from Tahiti, French Polynesia, bears the registration number MNHN–Pg 602. Régis Cleva of MNHN kindly informed us that the lot, MNHN­Pg 3403, contains a non­type specimen of T. trichophthalmus collected from Tuléar, Madagascar (personal communication).</p></div>	https://treatment.plazi.org/id/03D61C0EFF9EF97C350465201FBA6988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Komai, Tomoyuki;Osawa, Masayuki	Komai, Tomoyuki, Osawa, Masayuki (2005): Decapoda: Anomura: Paguridae) from the Ryukyu and Yap Islands, and redescription of T. trichophthalmus (Forest). Zootaxa 801: 1-20, DOI: 10.5281/zenodo.170561
