identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F0CFE84E2D7CBE731D8A89A020D11F9C.text	F0CFE84E2D7CBE731D8A89A020D11F9C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phyllognathopus Mrazek 1893	<div><p>Genus Phyllognathopus Mrazek, 1893</p><p>Emended diagnosis.</p><p>Phyllognathopodidae . Habitus slender, with no clear demarcation between prosome and urosome. Integumental dorsal window on cephalosome not confirmed for all members of the genus. Integument with surface pits, moderately sclerotized. Cephalosome rounded; rostrum clearly articulated to cephalosome. First pedigerous somite free. P5-bearing somite with large paired pores laterodorsally. Anal operculum plain or ornamented by fine spinules or extruded in strong spinular processes. Sexual dimorphism in antennule, P5, P6, urosomal segmentation and ornamentation. Female first and second abdominal somites fused forming genital double-somite. Anal somite with paired sensilla on dorsal side. Male urosome consisting of 6 segments. Caudal rami sub-quadrate, or longer than wide, with incomplete setal pattern (6 setae). Dorsal seta inserted on distal third of caudal ramus. Setae III and V variable in morphology among species. Antennule: 8-segmented in female, basically 10-segmented in male, although a suture line marking original segmentation between former segments 10 and 11 may be still discernible in some species; geniculation between segments 7 and 8; segment 9 discrete. Long tube-pores on segments 1 and 2 in both sexes. Antenna: armature of the second endopodal segment consisting of 10 elements. Exopod 1-segmented, with 3 lateral and 2 apical setae. Mandible: mandibular palp biramous, basis unarmed; exopod with 1 apical and 1 inner setae; endopod with 1 inner, 1 subapical and 2 apical setae. Armature of maxillule and maxilla as in Phyllognathopus viguieri . Maxilliped: phyl lopodial, lamelliform, 1-segmented. Trace of ancestral 2-segmented condition marked by the presence of outer incision; armature consisting of 10 elements.</p><p>P1-P3 with praecoxa and 3-segmented exopods and endopods. P4 small-sized, praecoxa missing, with 3- or 2- or 1-segmented exopod and 2- or 1-segmented endopod. Female P5 free, with clear articulation to P5-bearing somite; right and left legs distinct; intercoxal sclerite absent; baseoendopod and exopod coalescent, feeble incision marking original segmentation between them; endopodal lobe not pronounced, bearing 2 apical setae. Exopodal lobe fully incorporated into baseoendopod, not pronounced; basipodal outer seta present. Female P6 present, right and left legs represented by small chitinous lamellar plates, each leg bearing 1 normal seta or a stout spine with rounded tip. Male P5 free, with clear articulation to P5-bearing somite; right and left legs coalescent, intercoxal sclerite absent. Exopod discrete, but sometimes incorporated to basis. Endopod 1-segmented, normally conformed, cylindrical, bearing 1 leaf-like seta, alternatively transformed in a curved and stout element bearing 1 bipinnate seta, inserted on posterior surface of the endopod, close to its articulation to basis. Male P6 present; symmetrical, right and left legs coalescent along their medial margin, forming a continuous lamellar plate; each leg bearing 2 inner spines of different length and 1 outer seta.</p></div>	https://treatment.plazi.org/id/F0CFE84E2D7CBE731D8A89A020D11F9C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Galassi, Diana M. P.;Laurentiis, Paola De;Fiasca, Barbara	Galassi, Diana M. P., Laurentiis, Paola De, Fiasca, Barbara (2011): Systematics of the Phyllognathopodidae (Copepoda, Harpacticoida): re-examination of Phyllognathopus viguieri (Maupas, 1892) and Parbatocamptus jochenmartensi Dumont and Maas, 1988, proposal of a new genus for hyllognathopus bassoti Rouch, 1972, and description of a new species of Phyllognathopus. ZooKeys 104: 1-65, DOI: http://dx.doi.org/10.3897/zookeys.104.763, URL: http://dx.doi.org/10.3897/zookeys.104.763
DA88F11657E2BCCDA9A2F4CA76DBB06E.text	DA88F11657E2BCCDA9A2F4CA76DBB06E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phyllognathopus viguieri (Maupas 1892) Maupas 1892	<div><p>Phyllognathopus viguieri (Maupas, 1892) Figs 19</p><p>Material examined.</p><p>11 ♀♀ and 2 ♂♂,completely dissected and mounted in polyvinyl lactophenol, S. Anna D’Alfaedo, Progno di Valpantena (Verona, Italy), hyporheic habitat, 25.06.2002, E. Gattone coll.; 1 ♀, karstic spring in the hydrogeological basin of Rio Biondo, Progno di Valpantena (Verona, Italy), karstic habitat, 7. 07. 2003, B. Fiasca coll.; 3 ♀♀, Lake Bracciano (Latium, Italy), interstitial habitat, 27.05.02, V. Cottarelli coll.; 10 ♀♀ and 5 ♂♂, Avisio floodplain (Trento, northern Italy), hyporheic habitat, 30.05.2006, T. Di Lorenzo coll.; 3 ♀♀ and 1 ♂, Oignin stream (French Jura Mountains), hyporheic habitat, 30.07.2002, M.-J. Dole-Olivier coll.; 2 ♀♀ and 1 ♂, Ariège floodplain (France), P. Dumas coll.; 1 ♀, Lac Léman (France), slide code MNHN - Cp922, Paris; 2 ♂♂, S. Pierre, (France), slide code MNHN - Cp456, Paris; 3 ♂♂ and 2 ♀♀, Ruhr floodplain (Germany), T. Glatzel coll.; 6 ♀♀ and 1 ♂, R. Krishna, India, Y. Ranga Reddy coll.; 1 ♀, in pitcher of Nepenthes mirabilis (Hong Kong), B. Coker det., slide code 1982.329, Natural History Museum, London; 2 ♀♀ deposited at the Smithsonian Institution, Washington D.C., code USMN 251806, 204501; 1 ♂, code USMN 204500; 1 ♂ (juvenile), code USMN 204501.</p><p>Phyllognathopus cf. viguieri A. 2 ♂♂, 2 ♀♀, Madagascar, vial code, MNHN - Cp910, Paris, B. Dussart coll..</p><p>Phyllognathopus cf. viguieri B. 1 ♀, slide code 66/52, freshwater well, Mindoro Island, Philippines, 17.8.1992, V. Cottarelli coll.</p><p>Phyllognathopus viguieri ?. 10 ♀♀ and 5 ♂♂, Andhra Loyola College Campus, Vijaya-Woda, Andhra Pradesh, India, Y. Ranga Reddy coll..</p><p>Phyllognathopus viguieri menzeli . 1♀, vial code USNM 150192, labelled: Pacific Ocean, Mariana Islands, Guam, 19 November 1970, Watkins R.L. coll. (remaining material in the vial: 17 ♀♀).</p><p>Phyllognathopus viguieri menzeli 2. ♂♂, vial code USNM 150193, labelled: Pacific Ocean, Mariana Islands, Guam, 1 April 1971, Belk and Watkins R.L. coll. (remaining material in the vial: 8 specimens, of which several copepodids).</p><p>Supplementary description.</p><p>FEMALE. Body length, measured from tip of rostrum to posterior margin of caudal rami, from 400 to 600 µm (mean = 439 µm; n = 27). Habitus slender, no clear demarcation between prosome and urosome. Integument with surface pits, moderately sclerotized as in Fig. 1A. Cephalosome sub-quadrate, with a dorsal rounded protuberance, hardly observable, plausibly referable to a dorsal integumental window. Setule pattern as in Fig. 1A. Rostrum elongate, subrectangular in shape, clearly articulated to the cephalosome; two dorsal sensilla laterally inserted on its distal third, and one pore apically. Cephalosome and both thoracic and abdominal somites with cuticular ornamentation apparently represented by reduced number of paired sensilla (Fig. 1A). First pedigerous somite free. Hyaline frills of cephalosome, somites bearing P1-P4 and urosome dorsally smooth. Urosomites with smooth hyaline frill ventrally, except third urosomite (Figs 1B, 2A). Last three urosomites with spinular fringe on proximal third ventrally; anal somite with distal continuous spinule row. Anal somite with paired sensilla on dorsal side only (Fig. 1C), and two short spinule rows close to the anal operculum. Anal operculum rounded, only slightly protruding beyond insertion line of caudal rami (Figs 1C, 2B). P5-bearing somite with large paired pores laterodorsally and paired spinule rows laterally inserted on distal third of somite (Fig. 1A). Genital double-somite with three lateral spinule rows. Female genital field located between first and second third of genital double-somite. Genital apparatus apparently simplified; copulatory pore located halfway of genital double-somite (Figs 1B, 3A). Seminal receptacles located laterally and condensed close to the lamellar P6 (Fig. 3B).</p><p>Caudal rami rectangular, parallel, distinctly longer than wide (length/width ratio about 1.7), with incomplete setal pattern (6 setae) (Fig. 1 B–C). Anterolateral accessory seta (I) absent, anterolateral seta (II) smooth, inserted at half of caudal ramus; posterolateral seta (III) inserted on distal third of ramus, transformed in a large and stout spiniform element. Outer terminal seta (IV) very short, thin, and naked, shorter than caudal ramus, inner terminal seta (V) unipinnate and relatively short, without articulation at base, with very enlarged proximal part tapering in a subtle tip; terminal accessory seta (VI) slightly shorter than outer terminal seta, thin and naked; dorsal seta (VII) inserted on distal third of caudal ramus, about as long as caudal ramus. Three spinule rows inserted dorsolaterally and two spinule rows inserted at distal margin of caudal ramus ventrally. Two pores located close to the insertion of setae II and IV, ventrally.</p><p>Antennule (Fig. 4A): short, 8-segmented. Segment 1 with 1 spinule row. Both segments 1 and 2 bearing long and flaccid tube-pores. Armature formula: 1-[1], 2-[8], 3-[5], 4-[1 + (1+ae)], 5-[1], 6-[3], 7-[4], 8-[6 + (1+ae)]. Aesthetasc on segment 4 large, reaching about the proximal part of the penultimate antennulary segm ent .</p><p>Antenna (Fig. 4B): coxa unarmed; basis with 1 transverse spinule row on surface, a spinule row inserted on inner margin; exopod 1-segmented, well-defined at base, with surface spinule row, bearing 3 lateral and 2 apical setae; free endopod 2-segmented; both segments robust, of about the same length; segment 1 with inner spinule row; segment 2 with two inner spinule rows; armature consisting of 2 inner spines and 1 seta, 1 unipinnate apical spine, 4 geniculate setae, 1 apical slender seta and 1 subapical slender seta; a row of spinules at outer corner.</p><p>Labrum (Fig. 1D): trapezoidal, with two spinule rows on free distal margin. Paired rows of hair-like elements on medioventral surface.</p><p>Mandible (Fig. 4C): coxal gnathobase elongate, cutting edge with 2 large and coarse teeth, three smaller teeth and row of fringed teeth; naked seta at dorsal corner. Mandibular palp biramous, basis with inner spinule row, exopod with 1 apical and 1 inner bipinnate setae; endopod with 1 inner, 1 subapical and 2 apical bipinnate setae.</p><p>Maxillule (Fig. 4D): well developed arthrite incorporated into praecoxa, with 7 strong curved spines inserted on free distal margin and 1 short seta inserted on a sort of surface peduncle and 2 anterior surface setae. Coxal epipodite represented by 2 setae; coxo-endite with 2 plumose setae. Exopod and endopod incorporated into basis, with a total of 7 plumose setae.</p><p>Maxilla (Fig. 4E): syncoxa with 3 endites. Proximal endite with 6 setae; medial and distal endites, each with 3 plumose setae. Allobasis drawn out into a strong claw, distally spinulose, accompanied by 2 robust and 1 thin setae; endopod 3-segmented; segment 1 with 1 robust curved seta; segment 2 with 2 robust curved setae; segment 3 with 2 robust curved and 2 slender setae.</p><p>Maxilliped (Fig. 4F): phyllopodial, lamelliform, and 1-segmented. Trace of ancestral 2-segmented condition marked by the presence of outer incision, representing original segmentation boundary between former segments 1 and 2. Armature consisting of 10 elements, of which 5 bipinnate setae in apical position, 1 unipinnate seta inserted along inner margin together with 4 strong unipinnate stout spines. No trace of incision along inner margin.</p><p>P1-P3 with 3-segmented exopods and endopods. P4 with 3-segmented exopod and 2-segmented endopod. Intercoxal sclerites: boundary between intercoxa and basis not well defined at posterior surface of P2-P4 (Fig. 5 B–D). P1-P3 praecoxa well developed, with outer spinule row. P4 praecoxa absent.</p><p>P1 (Fig. 5A): praecoxa and coxa with outer spinule row on anterior surface; one posterior row of thin spinules inserted close to coxo-basis boundary. Basis with 1 outer spiniform seta and 1 inner spine, with spinule rows along outer margin, between exopod and endopod and at the insertion of inner spine, respectively. Exopod about as long as endopod; exp-1 and -2 with 1 outer unipinnate spine; exp-3 with 2 unipinnate spines in apical position, and 1 apical and 1 subapical inner setae. Endopod: enp-1 unarmed, about as long as enp-2 and enp-3, wider than enp-2 and enp-3. Enp-2 cylindrical, with short inner seta inserted at the middle of segment. Enp-3 with 1 inner spine, 1 apical seta and 1 curved apical spine. Ornamentation as in Fig. 5A.</p><p>P2 (Fig. 5B): ornamentation of praecoxa and coxa as in P1. Basis with 1 outer spine, with spinule rows along outer margin, and between exopod and endopod. Exopod slightly longer than endopod; exopodal segments of about the same length; exp-1 and -2 with 1 outer unipinnate spine; exp-3 with 2 outer unipinnate spines, 1 apical unipinnate seta and 1 subapical long inner seta. Endopod: enp-1 unarmed; enp-2 wit h 1 naked inner seta; enp-3 with 1 spine and 1 long bipinnate seta in apical position, and 1 short bipinnate subapical seta. Ornamentation as in Fig. 5B.</p><p>P3 (Fig. 5C): ornamentation of praecoxa and coxa as in P2. Basis with short outer seta and spinule rows along outer margin and at the insertion of the endopod. Exopod distinctly longer than endopod. Exp-1 and -2 with 1 unipinnate outer spine; exp-3 with 2 unipinnate outer spines, 1 bipinnate apical seta and 1 long bipinnate subapical seta. Endopod: enp-1 and -2 unarmed; enp-3 with 1 spine and 2 bipinnate setae in apical position. Ornamentation as in Fig. 5C.</p><p>P4 (Fig. 5D): reduced in size, praecoxa absent, coxa and basis without ornamentation; basis with long outer naked seta; exopod and endopod about as long as half of remaining legs; the exopod only slightly longer than endopod. Exp-1 with 1 unipinnate outer spine; exp-2 unarmed; exp-3 with 1 bipinnate outer spine and 2 apical setae of different length. Endopod: enp-1 unarmed; enp-2 with 3 apical setae. Ornamentation as in Fig. 5D.</p><p>P5 (Figs 3A, 5E): free, with clear articulation to P5-bearing somite; right and left legs distinct; baseoendopod and exopod coalescent, incision marked original segmentation between them still present; basipodal outer seta present, exopodal armature consisting of 3 bipinnate setae and 1 stout spine: all elements in apical position; baseoendopod armed with 2 robust bipinnate setae, the outermost the longest.</p><p>P6 (Figs 3B, 5F): rudimentary, consisting of paired small chitinous lamellar plates not coalescent along medial margin, partially covering seminal receptacles. Armature consisting of 1 short naked spine with rounded tip on each leg.</p><p>Male.</p><p>Body length, measured from tip of rostrum to posterior margin of caudal rami, from 370 to 541 µm, with mean of 424 µm based on 8 individuals. Sexual dimorphism in antennule, abdominal segmentation, P5, P6 and caudal setae morphology. Habitus, cephalosome (Figs 6A, 7 A–B), sensilla and pore patterns as in female. Integument with surface pits. Urosome as in Figs 6B, 7C. Caudal rami with 6 setae (Fig. 6B). Anterolateral seta (II) as in female, posterolateral seta (III) setiform, not transformed (length seta/length caudal ramus: about 2) and bipinnate. Outer terminal seta (IV) as in female, inner terminal seta (V) not transformed, plumose and long, not articulated at base; terminal accessory seta (VI) and dorsal seta (VII) as in female (Fig. 6). Ornamentation and pore patterns as in female. Anal operculum as in female.</p><p>Antennule (Figs 6C, 7 D–F): elongate, basically 10-segmented, last segment still showing a surface suture line only on anterior surface, indicating an incipient 11-segmented condition. Segment 1 with 1 ventral spinule row and 1 tube-pore (Fig. 8A). Segment 2 with tube-pore. Segment 4 represented by small U-shaped sclerite. Segment 6 the largest, sclerotized. Segment 8 elongate and transformed, moderately sclerotized, segment 9 distinct, not incorporated into segment 8, segment 10 derived by incomplete fusion between former segments 10 and 11 (Fig. 7 D–F). Armature formula: 1-[1], 2-[9], 3-[8], 4-[2], 5-[7+(1 + ae)], 6-[2], 7-[2], 8-[0], 9-[1], 10-[10 + (1 + ae)]. Aesthetasc on segment 5 very large. Segment 8 with medial pointed protrusions as in Fig. 6C.</p><p>P 1-P4 as in female; for morphological details of P1-P4 see Fig. 8B. P5 (Figs 6D, 9A): free, with clear articulation to P5-bearing somite; right and left legs coalescent; exopod clearly discernible but incorporated to basis: no trace of articulation between them observable, bearing 2 inner, 2 apical and 2 outer bipinnate setae; endopod discrete, distinctly 1-segmented, bearing 1 large leaf-like transformed seta and a spinule row along its free outer margin. Basipodal outer seta slender and naked, one pore near its insertion.</p><p>P6 (Figs 6E, 9B): right and left legs coalescent, forming a single linear lamellar plate, with no trace of incision between right and left P6; armature consisting of 2 spines and 1 outer seta.</p></div>	https://treatment.plazi.org/id/DA88F11657E2BCCDA9A2F4CA76DBB06E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Galassi, Diana M. P.;Laurentiis, Paola De;Fiasca, Barbara	Galassi, Diana M. P., Laurentiis, Paola De, Fiasca, Barbara (2011): Systematics of the Phyllognathopodidae (Copepoda, Harpacticoida): re-examination of Phyllognathopus viguieri (Maupas, 1892) and Parbatocamptus jochenmartensi Dumont and Maas, 1988, proposal of a new genus for hyllognathopus bassoti Rouch, 1972, and description of a new species of Phyllognathopus. ZooKeys 104: 1-65, DOI: http://dx.doi.org/10.3897/zookeys.104.763, URL: http://dx.doi.org/10.3897/zookeys.104.763
F6FE9D0E175B48464A0DACB3691F05CB.text	F6FE9D0E175B48464A0DACB3691F05CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phyllognathopus inexspectatus Galassi & De Laurentiis	<div><p>Phyllognathopus inexspectatus Galassi &amp; De Laurentiis sp. n. Figs 1013</p><p>Phyllognathopus sp. (in Di Lorenzo et al. 2005). (Synonymy)</p><p>Type material.</p><p>♀ holotype completely dissected and mounted in glycerine, deposited at the Natural History Museum, London (UK); January 2003, D. Cipriani coll.; 3 ♀♀ paratypes completely dissected and mounted in lactophenol; May 2003; January 2004. D. Cipriani coll..</p><p>Type locality.</p><p>Mazzoccolo karstic spring (Latium, central Italy), coordinates: 41°15'17"N, 13°27'08"E; Western Aurunci Mountains; 20 m a.s.l.; water temperature 13.5 ± 0.3 °C; pH 7.5 ± 0.1; O2 9.1 ± 0.9 mg/L (n = 11).</p><p>Description.</p><p>FEMALE. Total body length, measured from tip of rostrum to posterior margin of caudal rami, 474 µm (holotype), 468 µm (paratypes mean value; n = 3). Body depigmented and eyeless. Habitus slender, no clear demarcation between prosome and urosome. Integument with surface pits, moderately sclerotized. Cephalosome subquadrate, with a dorsal rounded protuberance, hardly observable, referable to the dorsal integumental window (Fig. 10A). Couples of setule rows located on surface of cephalic shield. Rostrum elongate, subrectangular in shape, clearly articulated to the cephalosome; two dorsal sensilla laterally inserted on distal third, and one pore apically. Cephalosome and both thoracic and abdominal somites with cuticular ornamentation represented by reduced number of paired sensilla (Fig. 10A). First pedigerous somite free. Hyaline frills of cephalosome, somites bearing P1-P4 and urosome dorsally smooth. P5-bearing somite with large paired pores laterodorsally. Genital double-somite with three lateral spinule rows and three pairs of setule rows inserted dorsally. Female genital field located at the middle of genital double-somite. Genital apparatus simplified, copulatory pore located at half of the genital double-somite. Seminal receptacles laterally located and condensed close to the lamellar sixth legs.</p><p>Urosomites with smooth hyaline frill ventrally, except third urosomite (Fig. 10B). Last two urosomites with spinular fringe on proximal third; anal somite with distal continuous spinule row.</p><p>Anal somite with paired sensilla on dorsal surface (Fig. 11A), and two short spinule rows close to the anal operculum. Anal operculum rounded, protruding beyond insertion line of caudal rami and armed with strong spinules on free distal margin (Fig. 11A). Caudal rami rectangular with strongly expanded inner margin, slightly divergent, distinctly longer than wide (length/width ratio: about 1.5), with incomplete setal pattern (6 setae) (Figs 10C, 11A); anterolateral accessory seta (I) absent, anterolateral seta (II) smooth, inserted at proximal third of caudal ramus; posterolateral seta (III) inserted on distal third of ramus, transformed in a short and stout spiniform seta, with tuft of spinules apically. Outer terminal seta (IV) very short, thin, and naked, without articulation at base (Fig. 10C), distinctly shorter than caudal ramus; inner terminal seta (V) not transformed, very long, without articulation at base; terminal accessory seta (VI) as long as outer terminal seta, thin and naked; dorsal seta (VII) inserted at half of caudal ramus, about as long as caudal ramus or slightly shorter. A continuous spinule row along inner margin of caudal ramus and three spinule rows inserted close to the anterolateral seta (Figs 10B, 11A), at the basis of the posterolateral seta and at distal margin of ramus ventrally, respectively. Two pores are located dorsally on each caudal ramus, and one pore ventrally.</p><p>Antennule (Fig. 11B): short, 8-segmented. Segment 1 with ventral spinule row. Both segments 1 and 2 bearing long and flaccid tube-pores. Armature formula: 1-[1], 2-[8], 3-[5], 4-[1 + (1 + ae)], 5-[1], 6-[3], 7-[4], 8-[6 +(1+ ae)]. Aesthetasc on segment 4 very large, reaching about the last antennulary segment.</p><p>Antenna (Fig. 11C): coxa unarmed; basis with 1 transverse spinule row on surface; exopod 1-segmented, well-defined at base, with spinule row on surface, bearing 3 lateral unipinnate and 2 apical bipinnate setae; free endopod 2-segmented; both segments robust, of about the same length; segment 1 with inner spinule row; segment 2 with one inner and one surface spinule rows; armature consisting of 2 inner spines and 1 thin seta, 1 apical unipinnate spine, 4 geniculate setae, and 1 apical and 1 surface slender setae; two rows of spinules at outer corner and in subapical position on free distal margin, respectively.</p><p>Mandible (Fig. 12A): coxal gnathobase elongate, cutting edge with 3 large and coarse teeth, 5 smaller fringed teeth; naked seta at dorsal corner. Mandibular palp biramous, basis with inner strong spinule row, exopod with 1 apical and 1 inner bipinnate setae; endopod with 1 inner bipinnate, and 1 spiniform and 2 bipinnate apical setae. Ornamentation as in Fig. 12A.</p><p>Maxillule (Fig. 12B): well developed arthrite incorporated into praecoxa, with 7 strong curved spines inserted on free distal margin, and 2 anterior surface setae. Proximal surface bipinnate seta inserted on tubercle absent (vs. present in Phyllognathopus viguieri, see Fig. 4D). Coxal epipodite represented by 2 setae; coxo-endite with 2 plumose setae. Exopod and endopod incorporated into basis, bearing 7 plumose setae.</p><p>Maxilla (Fig. 12C): syncoxa with 3 endites. Proximal endite free, with 6 setae; medial and distal endites incorporated to syncoxa, each with 3 plumose setae, inserted as in Fig. 12C. Allobasis drawn out into a strong claw apparently smooth, accompanied by 1 robust and 2 thin setae; endopod 3-segmented; segment 1 with 1 robust curved seta; segment 2 with 2 robust curved setae; segment 3 with 2 robust curved and 2 slender setae.</p><p>Maxilliped (Fig. 12D): phyllopodial, lamelliform, 1-segmented, and slender than in Phyllognathopus viguieri . Trace of ancestral 2-segmented condition marked by the presence of outer incision, probably representing original segmentation boundary between segments 1 and 2. Armature consisting of 10 elements, of which 5 bipinnate setae in apical position, two of which with independent insertion, 1 unipinnate seta inserted along inner margin together with 4 strong unipinnate spines. No trace of incision along inner margin.</p><p>P1-P3 with 3-segmented exopods and endopods. P4 with 2-segmented exopod and endopod. Intercoxal sclerites: boundary between intercoxa and basis not well defined at posterior surface in P2-P4. P1-P3 praecoxa well developed, with 1 outer spinule row. P4 praecoxa absent.</p><p>P1 (Fig. 12E): praecoxa and coxa with outer spinule row on anterior surface. Basis with 1 outer spiniform seta and 1 inner spine; with spinule rows along outer margin, between exopod and endopod and at the insertion of inner spine, respectively. Exopod slightly longer than endopod: exp-1 and -2 with 1 outer unipinnate spine; exp-3 with 2 outer unipinnate spines and 2 setae, respectively inserted apically and subapically. Endopod: enp-1 unarmed, about as long as enp-2 and enp-3. Enp-2 cylindrical, with 1 inner short seta. Enp-3 with 1 inner bipinnate seta, 1 long unipinnate curved seta and 1 spiniform curved seta in apical position. Ornamentation as in Fig. 12E.</p><p>P2 (Fig. 13A): praecoxa and coxa as in P1; basis with 1 outer spiniform seta, with spinule rows along outer margin, and between exopod and endopod. Exopod distinctly longer than endopod; exopodal segments of about the same length; exp-1 and -2 with 1 outer bipinnate spine; exp-3 with 2 outer unipinnate spines, 1 apical unipinnate seta and 1 subapical long bipinnate seta. Endopod: enp-1 and-2 unarmed; enp-3 with 1 spine and 1 long bipinnate seta in apical position, 1 subapical short bipinnate seta. Ornamentation as in Fig. 13A.</p><p>P3 (Fig. 13B): ornamentation of praecoxa and coxa as in P1-P2. Basis with long outer seta and spinule rows along outer margin and at the insertion of the endopod. Exopod distinctly longer than endopod. Exp-1 and -2 with 1 outer bipinnate spine; exp-3 with 2 outer unipinnate spines, and 2 apical long bipinnate setae. Endopod: enp-1 and -2 unarmed; enp-3 with 1 unipinnate spine and 1 long bipinnate seta in apical position, 1 subapical bipinnate seta. Ornamentation as in Fig. 13B.</p><p>P4 (Fig. 13C): small sized, if compared to P1-P3; praecoxa absent, coxa and basis without ornamentation; basis with outer long and naked seta; exopod as long as endopod. Exp-1 with 1 outer long unipinnate spine; exp-2 with 1 outer long spine and 2 apical setae. Endopod: enp-1 unarmed; enp-2 with 2 apical plumose setae. Ornamentation as in Fig. 13C.</p><p>P5 (Fig. 13D): free, with clear articulation to P5-bearing somite; right and left legs distinct; baseoendopod and exopod coalescent, incision marked original segmentation still present; basipodal outer seta present, exopodal armature consisting of 1 outer spine, 2 apical short setae, of about the same length, and 1 apical spine; baseoendopod armed with 2 robust bipinnate setae, the outer the longest.</p><p>P6 (Fig. 13E): rudimentary, consisting of small paired chitinous lamellar plates not coalescent along medial margin, partially covering the genital field. Armature consisting of 1 long and slender bipinnate seta on each side.</p><p>Male unknown.</p><p>Etymology.</p><p>The specific name derives from the Latin adjective inexspectatus which means “unexpected”, alluding to the surprising geographical location of the species, being the taxonomically related Phyllognathopus distributed in the Southern Hemisphere, and to the ecological finding of this species, which was collected from a large karstic aquifer in Central Italy, whereas all the other members of the genus are epigean.</p><p>Ecology.</p><p>At present knowledge the species is to be considered a stygobiotic species, collected from a karstic aquifer of the Western Aurunci Mountains (Latium) (Di Lorenzo et al. 2005). Although this aquifer is intensively fissured and karstified, with diffuse landforms of sinkholes and a discharge which is strictly linked to rainy events, stygoxene species were only sporadically present and represented by few individuals, due to the absence of a surface hydrological network, a landscape feature which is typical for coastal Mediterranean areas.</p></div>	https://treatment.plazi.org/id/F6FE9D0E175B48464A0DACB3691F05CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Galassi, Diana M. P.;Laurentiis, Paola De;Fiasca, Barbara	Galassi, Diana M. P., Laurentiis, Paola De, Fiasca, Barbara (2011): Systematics of the Phyllognathopodidae (Copepoda, Harpacticoida): re-examination of Phyllognathopus viguieri (Maupas, 1892) and Parbatocamptus jochenmartensi Dumont and Maas, 1988, proposal of a new genus for hyllognathopus bassoti Rouch, 1972, and description of a new species of Phyllognathopus. ZooKeys 104: 1-65, DOI: http://dx.doi.org/10.3897/zookeys.104.763, URL: http://dx.doi.org/10.3897/zookeys.104.763
76948213A95A9CEC09A9E69191E6E0F8.text	76948213A95A9CEC09A9E69191E6E0F8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parbatocamptus jochenmartensi Dumont and Maas 1988	<div><p>Parbatocamptus jochenmartensi Dumont and Maas, 1988 Figs 14-16</p><p>Material examined.</p><p>♂ holotype, completely dissected and mounted on slide labeled SMF 14657, deposited at the Senckenberg Museum (Germany).</p><p>Emended diagnosis of the genus Parbatocamptus Dumont and Maas, 1988.</p><p>Phyllognathopodidae . Body flattened. First pedigerous somite free. Hyaline frills of abdominal somites ventrally smooth. Anal operculum prominent, subdistally crenulated. Caudal ramus subquadrate in ventral view, tapering on free distal margin in dorsal view, with 6 setae; anterolateral accessory seta (I) absent; dorsal seta inserted on spinulose protuberance closely located to free distal margin. Male antennule 10-segmented, with geniculation between segments 7 and 8; segment 9 distinct, as usual in the family; tube-pores on segments 1 and 2; aesthetascs on segments 5 and 10. Antenna: exopod 1-segmented, well-defined at base, bearing 3 lateral and 2 apical unipinnate setae; free endopod 2-segmented; both segments robust, of about the same length. Mandibular palp biramous; basis with inner long bipinnate seta; exopod with 2 setae; endopod with 2 apical geniculate setae; 1 inner and 1 subapical bipinnate setae. Maxillary syncoxa with 3 endites. Proximal endite quadrilobate, with 6 apical setae; medial and distal endites, each with 3 setae. Allobasis drawn out into a strong claw, accompanied by 1 curved and 2 normal setae; endopod 3-segmented; segment 1 with 1 robust curved seta; segment 2 with 2 robust curved setae; segment 3 with 2 robust curved and 2 slender setae. Maxilliped phyllopodial, lamelliform, 1-segmented. Clear trace of ancestral 2-segmented condition well discernible and marked by outer and inner incisions; armature consisting of 11 elements: 1 strong spine inserted on inner corner of former proximal segment; 4 spines and 6 setae along free distal margin of former second segment, two of which inserted on independent little knob. P1-P2 with 3-segmented exopods and endopods. P3-P4 with 3-segmented exopods and 2-segmented endopods. P1-P4 praecoxa present. P1 exp-1 long, about as long as exp-2 and -3 together. P1-P4 endopods distinctly shorter than exopods, not overreaching distal margin of exp-2. P2 enp-2 transformed: outer margin produced into a comb-like structure; P2 enp-3 with 1 apical strong curved spine and 1 apical seta. P5 with 2-segmented exopod; endopod incorporated to basis, forming a baseoendopod; suture line still observable on posterior surface; rudimentary intercoxal sclerite still discernible. P6 symmetrical, consisting of a well developed, deeply incised lamella, marking original division between left and right legs. Armature consisting of 1 outer seta and 2 inner short spines of different length.</p><p>Female unknown.</p><p>Supplementary description of the holotype.</p><p>The description deals with major morphological details, omitted or overlooked in the original description, and with improvements of observational errors.</p><p>Integumental pitting not detectable on the dissected holotype; integument well sclerotized.</p><p>Urosomites 3-5 with smooth hyaline frills ventrally (Fig. 14A); third and fourth urosomites with spinular fringe closely located to hyaline frill; fifth urosomite with surface spinule rows ventrally, anal somite with distal continuous spinule row. Other ornamentation as in Fig. 14A. Urosomites 3-5 with crenulated hyaline frills dorsally. Indented surface rows are observable on third, fourth and fifth urosomites dorsally (Fig. 14B). Anal somite with row of fine spinules on proximal third; spinule row at the insertion of each caudal ramus; paired dorsal sensilla. Anal operculum slightly protruding the insertion line of caudal rami, subdistally crenulated (Fig. 14B).</p><p>Caudal rami ventrally subquadrate, parallel, longer than wide (length/width ratio: about 1.4), with incomplete setal pattern (6 setae) (Fig. 14 A–B). Anterolateral accessory seta (I) absent, anterolateral seta (II) unipinnate, inserted at second third of caudal ramus; posterolateral seta (III) inserted in subdistal position, transformed in spiniform bifid seta. Outer terminal seta (IV) well developed, unipinnate, with articulation at base; inner terminal seta (V) bipinnate and long; terminal accessory seta (VI) thin and naked, about as long as posterolateral seta; dorsal seta (VII) inserted on a spinulose knob, close to free distal margin of caudal ramus, distinctly longer than caudal ramu s . A spinule row on proximal third of each caudal ramus dorsally; three spinule rows inserted ventrolaterally and one spinule row at distal margin of caudal ramus, ventrally.</p><p>Antennule (Fig. 14C): 10-segmented. Segment 1 with spinule row and tube-pore. Segment 2 with tube-pore. Segment 4 represented by small U-shaped sclerite. Segment 5 large, sclerotized (Fig. 14D). Segment 8 very large and transformed, moderately sclerotized, segment 9 short, discrete, segment 10 derived by complete fusion of former segments 10 and 11. Armature formula: 1-[1], 2-[9], 3-[8], 4-[2], 5-[5?+(1 + ae)], 6-[2], 7-[2], 8-[0], 9-[1], 10-[10 + (1 + ae)].</p><p>Antenna (Fig. 15A): coxa unarmed; basis with 1 transverse spinule row on surface, a spinule row inserted on apical inner margin; exopod 1-segmented, well-defined at base, bearing 3 lateral and 2 apical unipinnate setae; free endopod 2-segmented; both segments robust, of about the same length; segment 1 naked; segment 2 with one inner spinule row, armature consisting of 2 inner spines and 1 seta, 1 apical unipinnate spine, 4 geniculate setae, 1 apical slender seta and 1 subapical tiny seta; a row of spinules at outer corner.</p><p>Mandible (Fig. 15B): coxal gnathobase elongate, cutting edge with 2 large and coarse teeth, three smaller teeth and row of tiny teeth; naked seta at dorsal corner. Mandibular palp biramous, basis with inner long bipinnate seta and spinule row, exopod with 2 bipinnate setae; endopod with 2 apical geniculate setae, 1 inner and 1 subapical bipinnate setae.</p><p>Maxillule not observable.</p><p>Maxilla (Fig. 15C): syncoxa with 3 endites fully incorporated to syncoxa. Proximal endite quadrilobate, with 6 setae; the first two distal lobes each bearing 2 setae; the proximal ones, each with 1 plumose seta; medial and distal endites, each with 3 bipinnate setae. Allobasis drawn out into a strong claw, distally spinulose, accompanied by 1 robust and curved seta and 2 naked setae, respectively; endopod 3-segmented; segment 1 with 1 robust curved seta; segment 2 with 2 robust curved setae; segment 3 with 2 curved and 2 slender setae.</p><p>Maxilliped (Fig. 15D): phyllopodial, lamelliform, 1-segmented. Clear trace of ancestral 2-segmented condition marked by the presence of both outer and inner incisions. Armature consisting of 11 elements: 1 strong spine inserted at inner corner of former segment 1; 4 strong unipinnate spines along inner margin, 1 bipinnate seta inserted along inner margin and 5 bipinnate setae in apical position.</p><p>P1-P2 with 3-segmented exopods and endopods. P3-P4 with 3-segmented exopods and 2-segmented endopods. P1-P4 praecoxa well developed. P1 (Fig. 15E): praecoxa and coxa with outer spinule row on anterior surface; one posterior row of thin spinules inserted on coxo-basis boundary. Basis with 1 outer spiniform seta and 1 inner spine, with spinule rows along outer margin, between exopod and endopod and at the insertion of inner spine, respectively. Endopod distinctly shorter than exopod, reaching about distal third of exp-2; exp-1 long, about as long as exp-2 and -3 together; exp-1 and -2 with 1 outer spine; exp-3 with 2 outer curved, unipinnate spines, and 1 apical and 1 subapical geniculate setae. Endopod: enp-1 unarmed, about as long as enp-2 and enp-3, wider than enp-2 and enp-3. Enp-2 cylindrical, unarmed. Enp-3 with 1 inner long seta, and 2 apical geniculate setae of different length. Ornamentation as in Fig. 15E.</p><p>P 2 (Fig. 16A): praecoxa unornamented, coxa as in P1 and P2. Basis with 1 outer spiniform seta, with spinule rows along outer margin. Exopod distinctly longer than endopod; endopod reaching about the proximal half of exopodal segment 2; exopodal segments of about the same length; exp-1 and -2 with 1 outer spine; exp-3 with 2 outer unipinnate spines, 1 apical fringed seta and 1 subapical long and slender seta. Endopod: enp-1 unarmed; enp 2 transformed, with outer strong comb-like process; enp-3 with 1 transformed spine and 1 long slender and naked seta in apical position. Ornamentation as in Fig. 16A.</p><p>P3 (Fig. 16B): praecoxa unornamented; ornamentation of coxa as in P1 and P2. Basis with long outer plumose seta and spinule rows along outer margin and at the insertion of the endopod. Exopod distinctly longer than endopod; endopod reaching about half of exp-2. Exp-1 and -2 with 1 outer spine; exp-3 with 2 outer fringed spines, 1 apical unipinnate seta and 1 subapical long and naked seta. Endopod: enp-1 unarmed; enp-2 with 2 apical spinulose setae and 1 subapical thin and naked seta. Ornamentation as in Fig. 16B.</p><p>P4 (Fig. 16C): slightly smaller than the other swimming legs, praecoxa present, unornamented; coxa with outer spinule row, one posterior row of thin spinules inserted on coxo-basis boundary; basis with outer plumose seta, and spinule rows along outer margin and at the insertion of endopod; exopod distinctly longer than endopod. Exp-1 and -2 with 1 outer bipinnate spine; exp-3 with 2 outer unipinnate spines and 2 apical bipinnate setae of different length. Endopod: enp-1 unarmed; enp-2 with 1 spine and 1 seta in apical position, and 1 subapical short unipinnate seta. Ornamentation as in Fig. 16C.</p><p>P5 (Fig. 16D): free, with clear articulation to P5-bearing somite; right and left legs distinct, trace of intercoxal sclerite present but hardly observable, with coxo-basis protrusions; exopod discrete, 2-segmented, segment 1 with 1 outer spine and 1 inner seta distally fringed; segment 2 with 4 elements: 1 outer slender and naked seta, 1 apical long spine, 1 medial short spine and 1 short seta distally crested; endopod incorporated to basis forming a baseoendopod, trace of original segmentation still recognizable on posterior surface (Fig. 16D); rudimentary endopod 1-segmented, bearing 1 strong spiniform element, crested on its distal margin. Basipodal outer seta slender and naked.</p><p>P6 (Fig. 16E): well developed, symmetrical, right and left legs distinct, deep medial incision marking boundary between legs; armature consisting of 1 outer long, bipinnate seta and two inner short spines, the innermost the shortest.</p></div>	https://treatment.plazi.org/id/76948213A95A9CEC09A9E69191E6E0F8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Galassi, Diana M. P.;Laurentiis, Paola De;Fiasca, Barbara	Galassi, Diana M. P., Laurentiis, Paola De, Fiasca, Barbara (2011): Systematics of the Phyllognathopodidae (Copepoda, Harpacticoida): re-examination of Phyllognathopus viguieri (Maupas, 1892) and Parbatocamptus jochenmartensi Dumont and Maas, 1988, proposal of a new genus for hyllognathopus bassoti Rouch, 1972, and description of a new species of Phyllognathopus. ZooKeys 104: 1-65, DOI: http://dx.doi.org/10.3897/zookeys.104.763, URL: http://dx.doi.org/10.3897/zookeys.104.763
394F29EF91DFB2645CC8A01944AE5889.text	394F29EF91DFB2645CC8A01944AE5889.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neophyllognathopus Galassi & De Laurentiis	<div><p>Neophyllognathopus Galassi &amp; De Laurentiis gen. n.</p><p>Diagnosis.</p><p>Phyllognathopodidae . Habitus slightly dorsoventrally flattened with no clear demarcation between prosome and urosome. Integumental dorsal window on cephalosome not confirmed. Integument without surface pits, moderately sclerotized. Ceph alosome rounded; rostrum elongate, clearly articulated to cephalosome. Cephalosome and both thoracic and abdominal somites with cuticular ornamentation represented by dorsal sensilla. First pedigerous somite free. Hyaline frills of cephalosome, somites bearing P1-P4 plain both dorsally and ventrally. P5-bearing somite with large paired pores laterodorsally. Sexual dimorphism in antennule, P5, P6, urosomal segmentation and ornamentation, and morphology of anal operculum. Female first and second abdominal somites fused forming the genital double-somite. Female urosomal segments with plain hyaline frills ventrally. Female genital apparatus simplified; copulatory pore located at the end of the proximal third of the genital double-somite. Seminal receptacles laterally located and condensed close to the lamellar sixth legs. Male urosome with different arrangement of hyaline frill ornamentations: urosome consisting of 6 segments, second urosomite with indented hyaline frill, third and fourth urosomites with deep ventral sockets; socket on third urosomite plicate, with smooth free distal margin, and 2 setules laterally inserted close to the socket opening; socket on fourth urosomite with free distal margin ornamented by strong and long spinules, covering the opening; fifth urosomite with indented hyaline frill. Anal somite with paired sensilla on dorsal side. Anal operculum protruding free distal margin of anal somite and extruded in strong spinular processes. Sexual dimorphism in the number of spinular processes of anal operculum (3 in females vs. 4 in males; and, in general, anal operculum in male more armed than in female). Caudal rami sub-quadrate, with incomplete setal pattern (6 setae). Dorsal seta inserted on distal third of caudal ramus. Antennule: 8-segmented in female, basically 9-segmented in male; geniculation between segments 7 and 8; penultimate and last segments, each with suture line marking original segmentation between former segments 8 and 9, and 10 and 11, respectively. Long tube-pores on segments 1 and 2 in both sexes. Antenna: armature of the second endopodal segment as in Phyllognathopus and Parbatocamptus, consisting of 10 elements. Exopod 1-segmented, with 3 lateral and 2 apical setae. Mandible: mandibular palp biramous, basis with inner spinule row, exopod with 1 apical and 1 inner setae; endopod with 1 inner, 1 subapical and 2 apical setae. Armature of maxillule and maxilla as in Phyllognathopus . Maxilliped: phyllopodial, lamelliform, 1-segmented. Clear trace of ancestral 2-segmented condition marked by the presence of outer and inner incisions as in Parbatocamptus . Armature consisting of 11 elements: 1 strong spine inserted at inner corner of former segment 1; 4 spines and 1 spiniform short seta inserted along inner margin, 5 bipinnate setae in apical position, armature topology basically referable to that of Parbatocamptus .</p><p>P1-P3 with 3-segmented exopods and endopods. P4 with 2-segmented exopod and endopod. P1-P3 praecoxa well developed. P1 exopod and endopod of about the same length; P2-P3 endopods shorter than exopods, reaching about tip of exp-2. P4 small - sized, praecoxa missing. Female P5: free, with clear articulation to P5-bearing somite; right and left legs distinct; baseoendopod and exopod coalescent, deep incision marking original segmentation between them; endopodal lobe well developed, elongate, longer than exopodal lobe, rectangular in shape, bearing 1 long pinnate seta, subdistally inserted, close to outer margin and a spinule row apically inserted. Exopodal lobe wide, fully incorporated into baseoendopod; exopodal armature consisting of 4 elements, the outermost bipinnate seta inserted in subdistal position, and three apical elements: 2 spinulose and 1 short setae; basipodal outer seta present. Female P6 rudimentary, each leg defined by a small cuticular lateral plate bearing a short, naked seta with rounded tip. Male P5: free, with clear articulation to P5-bearing somite; right and left legs separate, intercoxal sclerite rudimentary, but still discernible. Basis of each leg expanded, endopod strongly trasformed, consisting of a sclerotized and strong protrusion articulated to basis. Endopodal seta bipinnate, inserted on posterior surface of the endopod, close to its articulation to basis. Exopod distinct, clearly articulated to basis, wide and short, rectangular in shape, representing most part of the free distal margin of each leg; exopodal armature consisting of 6 elements, the innermost spiniform seta curved inward. Male P6: right and left legs distinct but closely adjacent to each other along their medial margin, and symmetrical; each leg consisting of a well developed lamellar plate, with spinule row on the anterior surface; armature consisting of 2 inner spines of different length and 1 outer seta.</p><p>Type species by monotypy.</p><p>Phyllognathopus bassoti Rouch, 1972 = Neophyllognathopus bassoti (Rouch, 1972), comb. n.</p><p>Etymology.</p><p>The genus name is derived from the type genus Phyllognathopus and the Latinised Greek prefix νέοσ which means “new”, referring to the new position of Phyllognathopus bassoti in the systematics of the family Phyllognathopodidae .</p></div>	https://treatment.plazi.org/id/394F29EF91DFB2645CC8A01944AE5889	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Galassi, Diana M. P.;Laurentiis, Paola De;Fiasca, Barbara	Galassi, Diana M. P., Laurentiis, Paola De, Fiasca, Barbara (2011): Systematics of the Phyllognathopodidae (Copepoda, Harpacticoida): re-examination of Phyllognathopus viguieri (Maupas, 1892) and Parbatocamptus jochenmartensi Dumont and Maas, 1988, proposal of a new genus for hyllognathopus bassoti Rouch, 1972, and description of a new species of Phyllognathopus. ZooKeys 104: 1-65, DOI: http://dx.doi.org/10.3897/zookeys.104.763, URL: http://dx.doi.org/10.3897/zookeys.104.763
2DDFE2EE7722D9A2EA94FC14B77A4DBC.text	2DDFE2EE7722D9A2EA94FC14B77A4DBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neophyllognathopus bassoti (Rouch 1972) Rouch 1972	<div><p>Neophyllognathopus bassoti (Rouch, 1972) comb. n. Figs 1725</p><p>Neotype designation.</p><p>Female neotype completely dissected and mounted in polyvinyl lactophenol, deposited at the Natural History Museum, London (reg. No. NHM.2008. neotype). Other material: 5 ♀♀ and 3 ♂♂ mounted on slides, 5 ♀♀ and 5 ♂♂ processed for SEM; India, 7 January 1999, Y. Ranga Reddy coll.; 1 ♀, slide code 66/49, 1 ♀, slide code 66/53, 1 ♂, slide code 66/55, Santa Fe, Bantayan island, Pooc, Philippines, V. Cottarelli coll. (see Bruno and Cottarelli 1999, for locality details).</p><p>Neophyllognathopus bassoti is proposed herein as new combination for Phyllognathopus bassoti assigned by Rouch (1972) in the original description to the genus Phyllognathopus . According to ICZN (2000), a neotype may be designated when no name-bearing type specimen (i.e. holotype, lectotype, syntype or prior neotype) is believed to be extant and an author considers that a name-bearing type is necessary to define the nominal taxon objectively (Article 75.1). Article 75.3 asks also for qualifying conditions for the establishment of a neotype; among them: …. a statement that it is designated with the express purpose of clarifying the taxonomic status or the type locality of a nominal taxon (Article 75.3.1), and the author’s reasons for believing the name–bearing type specimen(s) … to be lost or destroyed, and the steps that have been taken to trace it or them (Article 73.3.4).</p><p>The specimens on which Rouch (1972) based the original description of Phyllognathopus bassoti no longer exist, and most part of the Rouch’s collection has been lost (Rouch, in litt.). Consequently, the ICZN (2000) recommendation 75A cannot be met, because no extant paratypes or paralectotypes, nor topotypic specimens are available, in order to select among them a neotype. The need to clarify the taxonomic status of this species, which is ranked herein to a new genus, imposed to follow another formal procedure, which, if not completely fulfills the ICZN rules (Article 75.3.6), is accepted by the Code (Article 76.3): the place of origin of the neotype becomes the type locality of the nominal species-group taxon, despite any previously published statement of the type locality. Following these arguments, a consistent population from India has been selected to establish the new genus, together with additional material from Indonesia. It is relevant to observe that other researchers have given consensus (Bruno and Cottarelli 1999, Karanovic and Ranga Reddy 2004) on the attribution of both populations to the species Phyllognathopus bassoti described by Rouch (1972) from the Lake Wisdom (New Guinea).</p><p>Neotype locality.</p><p>India, Andhra Pradesh, town of Guntur, Brindavan Gardens, domestic water reservoir filled by a freshwater bore well; coordinates: approx. 16°18'N, 80°29'E (see Karanovic and Ranga Reddy 2004, for more details).</p><p>Description based on the designed neotype.</p><p>FEMALE NEOTYPE. Body length, measured from tip of rostrum to posterior margin of caudal rami, 348 µm . Habitus slig htly dorsoventrally flattened (Fig. 17A), with no clear demarcation between prosome and urosome. Body depigmented and eyeless. Integumental dorsal window on cephalosome not confirmed. First pedigerous somite free. Integument without surface pits, moderately sclerotized. Cephalosome rounded; rostrum elongate, clearly articulated to cephalosome. Hyaline frills of cephalosome, somites bearing P1-P4 and urosome plain both dorsally and ventrally (Fig. 17 A, B). Cephalosome and both thoracic and abdominal somites (except fourth urosomite) with cuticular ornamentation represented by dorsal sensilla. P5-bearing somite with lateral paired and large pores (Fig. 18A). Female genital field located between first and second third of genital double-somite. Genital apparatus simplified; copulatory pore located at half of genital double-somite. Seminal receptacles laterally located and condensed close to the lamellar sixth legs. Three spinular processes on free distal margin of anal operculum (Figs 17 A–C). Caudal rami sub-quadrate, with incomplete setal pattern (6 setae). Dorsal seta inserted close to free distal margin of caudal ramus (Fig. 17C).</p><p>Antennule (Fig. 17D): consisting of 8 segments, segments 1 and 2 with long tube-pores. Armature formula: 1-[1], 2-[8], 3-[5], 4-[1 + (1 + ae)], 5-[1], 6-[3], 7-[4], 8-[6 + (1 + ae)]. Aesthetasc on segment 4 very large and long, well overreaching the last antennulary segment.</p><p>Antenna (Fig. 17E): exopod and armature of the second endopodal segment as in Phyllognathopus and Parbatocamptus . Exopod 1-segmented, with 3 lateral and 2 apical setae.</p><p>Mandible (Fig. 19A): mandibular palp biramous, basis with inner spinule row, exopod with 1 apical and 1 inner setae; endopod with 1 inner, 1 subapical and 2 apical setae. Armature of maxillule (Fig. 19B) and maxilla (Fig. 19C) as in Phyllognathopus .</p><p>Maxilliped (Fig. 19D): phyllopodial, lamelliform, 1-segmented. Clear trace of ancestral 2-segmented condition marked by the presence of both outer and inner incisions. Armature consisting of 11 elements: 1 strong spine inserted at inner corner of former segment 1; 4 spines and 1 spiniform short seta inserted along inner margin, 5 bipinnate setae in apical position, armature topology basically referable to that of Parbatocamptus .</p><p>P1-P3 with 3-segmented exopods and endopods. P4 with 2-segmented exopod and endopod. P1-P3 praecoxa well developed. P4 praecoxa absent (Fig. 19E). P1 exopod of about the same length of endopod. P2-P3 exopods longer than endopods, endopod not overreaching exp-2, fitting the original description (Rouch 1972) and the subsequent ones (Bruno and Cottarelli 1999, Karanovic and Ranga Reddy 2004).</p><p>P5 (Fig. 19F, G): free, with clear articulation to P5-bearing somite; right and left legs separate; baseoendopod and exopod coalescent, deep incision marking original segmentation between them; endopodal lobe well developed, elongate, rectangular in shape, longer than exopod, bearing 1 long pinnate seta, subdistally inserted, close to outer margin, and 2 spinule rows, the proximal one composed by tiny elements, the distal one of long spinules; exopodal lobe well discernible, with armature consisting of 4 (rarely 5 elements, observed in only one female) elements, the outermost seta inserted in subdistal outer position, the remaining ones in apical position; the outer apical seta slender and bipinnate, the remaining two spiniform. Basipodal outer seta present.</p><p>P6 (Figs 19H, 20): rudimentary, consisting of small paired chitinous lamellar plates not coalescent along medial margin, partially covering seminal receptacles. Armature consisting of 1 short smooth spine with rounded tip on each side.</p><p>Male.</p><p>No marked sexual dimorphism in body size. Body length, measured from tip of rostrum to posterior margin of caudal rami, 335 µm . Rostrum and ornamentation of cephalosome as in female (Fig. 21A). Male urosome consisting of 6 segments (Fig. 21A), third and fourth urosomites with deep ventral sockets (Figs 21B, 22A); socket on third urosomite plicate, with smooth free distal margins, and 2 setules laterally inserted close to the socket opening (Fig. 22B); socket on fourth urosomite with ornamented anterior margin, armed by strong spinules covering the opening (Fig. 22C). Anal somite with paired sensilla on dorsal side. Anal operculum protruding free distal margin of anal somite and extruded in 4 strong spinular processes, rarely 5 (in general anal operculum in males more armed than in females) (Fig. 18B). Antennule (Fig. 21C): basically 9-segmented, geniculation between segments 7 and 8; penultimate and last segments, each with suture line marking original segmentation between former segments 8 and 9, and 10 and 11, respectively. Long tube-pores on segments 1 and 2. Armature formula: 1-[1], 2-[8], 3-[8], 4-[1], 5-[7+(1 + ae)], 6-[2], 7-[1], 8-[1], 9-[10 + (1 + ae)].</p><p>P5 (Figs 21D, 23A): free, with clear articulation to P5-bearing somite; right and left legs separate, intercoxal sclerite rudimentary but still discernible (Figs 21D, 23B). Basis of each leg well developed, representing most part of each leg; endopod rudimentary, consisting of a sclerotized and strong process articulated to basis. Row of surface spinules inserted near articulation between endopod and basis. Endopodal seta bipinnate, inserted on posterior surface of the endopod, close to its articulation to basis. Exopod distinct, clear articulated to the basis, wide and short, with unusual topology, being placed at the inner free distal margin of basis; exopodal armature consisting of 6 elements, all of which in apical position. Inner spinulose seta short and distinctly curved inward, the remaining setae of about the same length, 2 of which (the second and the fourth, beginning from the inner margin of the exopod) are respectively bipinnate and unipinnate; the remaining 3 smooth and slender, frequently closely adherent to each other and not easily discernible as distinct (Fig. 24). Male P6 (Figs 21E, 25 A–B): right and left legs distinct, closely adherent along inner margin, and symmetrical, each leg consisting of a well developed lamellar plate, with some spinule rows on the anterior surface. A membranous lamell a is observable between right and left P6 (rudimentary intercoxa?) (Fig. 25B); armature consisting of 2 inner spines of different length and 1 outer naked seta.</p></div>	https://treatment.plazi.org/id/2DDFE2EE7722D9A2EA94FC14B77A4DBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Galassi, Diana M. P.;Laurentiis, Paola De;Fiasca, Barbara	Galassi, Diana M. P., Laurentiis, Paola De, Fiasca, Barbara (2011): Systematics of the Phyllognathopodidae (Copepoda, Harpacticoida): re-examination of Phyllognathopus viguieri (Maupas, 1892) and Parbatocamptus jochenmartensi Dumont and Maas, 1988, proposal of a new genus for hyllognathopus bassoti Rouch, 1972, and description of a new species of Phyllognathopus. ZooKeys 104: 1-65, DOI: http://dx.doi.org/10.3897/zookeys.104.763, URL: http://dx.doi.org/10.3897/zookeys.104.763
