taxonID	type	description	language	source
F6FE9D0E175B48464A0DACB3691F05CB.taxon	description	Description. FEMALE. Total body length, measured from tip of rostrum to posterior margin of caudal rami, 474 µm (holotype), 468 µm (paratypes mean value; n = 3). Body depigmented and eyeless. Habitus slender, no clear demarcation between prosome and urosome. Integument with surface pits, moderately sclerotized. Cephalosome subquadrate, with a dorsal rounded protuberance, hardly observable, referable to the dorsal integumental window (Fig. 10 A). Couples of setule rows located on surface of cephalic shield. Rostrum elongate, subrectangular in shape, clearly articulated to the cephalosome; two dorsal sensilla laterally inserted on distal third, and one pore apically. Cephalosome and both thoracic and abdominal somites with cuticular ornamentation represented by reduced number of paired sensilla (Fig. 10 A). First pedigerous somite free. Hyaline frills of cephalosome, somites bearing P 1 - P 4 and urosome dorsally smooth. P 5 - bearing somite with large paired pores laterodorsally. Genital double-somite with three lateral spinule rows and three pairs of setule rows inserted dorsally. Female genital field located at the middle of genital double-somite. Genital apparatus simplified, copulatory pore located at half of the genital double-somite. Seminal receptacles laterally located and condensed close to the lamellar sixth legs. Urosomites with smooth hyaline frill ventrally, except third urosomite (Fig. 10 B). Last two urosomites with spinular fringe on proximal third; anal somite with distal continuous spinule row. Anal somite with paired sensilla on dorsal surface (Fig. 11 A), and two short spinule rows close to the anal operculum. Anal operculum rounded, protruding beyond insertion line of caudal rami and armed with strong spinules on free distal margin (Fig. 11 A). Caudal rami rectangular with strongly expanded inner margin, slightly divergent, distinctly longer than wide (length / width ratio: about 1.5), with incomplete setal pattern (6 setae) (Figs 10 C, 11 A); anterolateral accessory seta (I) absent, anterolateral seta (II) smooth, inserted at proximal third of caudal ramus; posterolateral seta (III) inserted on distal third of ramus, transformed in a short and stout spiniform seta, with tuft of spinules apically. Outer terminal seta (IV) very short, thin, and naked, without articulation at base (Fig. 10 C), distinctly shorter than caudal ramus; inner terminal seta (V) not transformed, very long, without articulation at base; terminal accessory seta (VI) as long as outer terminal seta, thin and naked; dorsal seta (VII) inserted at half of caudal ramus, about as long as caudal ramus or slightly shorter. A continuous spinule row along inner margin of caudal ramus and three spinule rows inserted close to the anterolateral seta (Figs 10 B, 11 A), at the basis of the posterolateral seta and at distal margin of ramus ventrally, respectively. Two pores are located dorsally on each caudal ramus, and one pore ventrally. Antennule (Fig. 11 B): short, 8 - segmented. Segment 1 with ventral spinule row. Both segments 1 and 2 bearing long and flaccid tube-pores. Armature formula: 1 - [1], 2 - [8], 3 - [5], 4 - [1 + (1 + ae)], 5 - [1], 6 - [3], 7 - [4], 8 - [6 + (1 + ae)]. Aesthetasc on segment 4 very large, reaching about the last antennulary segment. Antenna (Fig. 11 C): coxa unarmed; basis with 1 transverse spinule row on surface; exopod 1 - segmented, well-defined at base, with spinule row on surface, bearing 3 lateral unipinnate and 2 apical bipinnate setae; free endopod 2 - segmented; both segments robust, of about the same length; segment 1 with inner spinule row; segment 2 with one inner and one surface spinule rows; armature consisting of 2 inner spines and 1 thin seta, 1 apical unipinnate spine, 4 geniculate setae, and 1 apical and 1 surface slender setae; two rows of spinules at outer corner and in subapical position on free distal margin, respectively. Mandible (Fig. 12 A): coxal gnathobase elongate, cutting edge with 3 large and coarse teeth, 5 smaller fringed teeth; naked seta at dorsal corner. Mandibular palp biramous, basis with inner strong spinule row, exopod with 1 apical and 1 inner bipinnate setae; endopod with 1 inner bipinnate, and 1 spiniform and 2 bipinnate apical setae. Ornamentation as in Fig. 12 A. Maxillule (Fig. 12 B): well developed arthrite incorporated into praecoxa, with 7 strong curved spines inserted on free distal margin, and 2 anterior surface setae. Proximal surface bipinnate seta inserted on tubercle absent (vs. present in Phyllognathopus viguieri, see Fig. 4 D). Coxal epipodite represented by 2 setae; coxo-endite with 2 plumose setae. Exopod and endopod incorporated into basis, bearing 7 plumose setae. Maxilla (Fig. 12 C): syncoxa with 3 endites. Proximal endite free, with 6 setae; medial and distal endites incorporated to syncoxa, each with 3 plumose setae, inserted as in Fig. 12 C. Allobasis drawn out into a strong claw apparently smooth, accompanied by 1 robust and 2 thin setae; endopod 3 - segmented; segment 1 with 1 robust curved seta; segment 2 with 2 robust curved setae; segment 3 with 2 robust curved and 2 slender setae. Maxilliped (Fig. 12 D): phyllopodial, lamelliform, 1 - segmented, and slender than in Phyllognathopus viguieri. Trace of ancestral 2 - segmented condition marked by the presence of outer incision, probably representing original segmentation boundary between segments 1 and 2. Armature consisting of 10 elements, of which 5 bipinnate setae in apical position, two of which with independent insertion, 1 unipinnate seta inserted along inner margin together with 4 strong unipinnate spines. No trace of incision along inner margin. P 1 - P 3 with 3 - segmented exopods and endopods. P 4 with 2 - segmented exopod and endopod. Intercoxal sclerites: boundary between intercoxa and basis not well defined at posterior surface in P 2 - P 4. P 1 - P 3 praecoxa well developed, with 1 outer spinule row. P 4 praecoxa absent. P 1 (Fig. 12 E): praecoxa and coxa with outer spinule row on anterior surface. Basis with 1 outer spiniform seta and 1 inner spine; with spinule rows along outer margin, between exopod and endopod and at the insertion of inner spine, respectively. Exopod slightly longer than endopod: exp- 1 and - 2 with 1 outer unipinnate spine; exp- 3 with 2 outer unipinnate spines and 2 setae, respectively inserted apically and subapically. Endopod: enp- 1 unarmed, about as long as enp- 2 and enp- 3. Enp- 2 cylindrical, with 1 inner short seta. Enp- 3 with 1 inner bipinnate seta, 1 long unipinnate curved seta and 1 spiniform curved seta in apical position. Ornamentation as in Fig. 12 E. P 2 (Fig. 13 A): praecoxa and coxa as in P 1; basis with 1 outer spiniform seta, with spinule rows along outer margin, and between exopod and endopod. Exopod distinctly longer than endopod; exopodal segments of about the same length; exp- 1 and - 2 with 1 outer bipinnate spine; exp- 3 with 2 outer unipinnate spines, 1 apical unipinnate seta and 1 subapical long bipinnate seta. Endopod: enp- 1 and- 2 unarmed; enp- 3 with 1 spine and 1 long bipinnate seta in apical position, 1 subapical short bipinnate seta. Ornamentation as in Fig. 13 A. P 3 (Fig. 13 B): ornamentation of praecoxa and coxa as in P 1 - P 2. Basis with long outer seta and spinule rows along outer margin and at the insertion of the endopod. Exopod distinctly longer than endopod. Exp- 1 and - 2 with 1 outer bipinnate spine; exp- 3 with 2 outer unipinnate spines, and 2 apical long bipinnate setae. Endopod: enp- 1 and - 2 unarmed; enp- 3 with 1 unipinnate spine and 1 long bipinnate seta in apical position, 1 subapical bipinnate seta. Ornamentation as in Fig. 13 B. P 4 (Fig. 13 C): small sized, if compared to P 1 - P 3; praecoxa absent, coxa and basis without ornamentation; basis with outer long and naked seta; exopod as long as endopod. Exp- 1 with 1 outer long unipinnate spine; exp- 2 with 1 outer long spine and 2 apical setae. Endopod: enp- 1 unarmed; enp- 2 with 2 apical plumose setae. Ornamentation as in Fig. 13 C. P 5 (Fig. 13 D): free, with clear articulation to P 5 - bearing somite; right and left legs distinct; baseoendopod and exopod coalescent, incision marked original segmentation still present; basipodal outer seta present, exopodal armature consisting of 1 outer spine, 2 apical short setae, of about the same length, and 1 apical spine; baseoendopod armed with 2 robust bipinnate setae, the outer the longest. P 6 (Fig. 13 E): rudimentary, consisting of small paired chitinous lamellar plates not coalescent along medial margin, partially covering the genital field. Armature consisting of 1 long and slender bipinnate seta on each side. Male unknown.	en	Galassi, Diana M. P., Laurentiis, Paola De, Fiasca, Barbara (2011): Systematics of the Phyllognathopodidae (Copepoda, Harpacticoida): re-examination of Phyllognathopus viguieri (Maupas, 1892) and Parbatocamptus jochenmartensi Dumont and Maas, 1988, proposal of a new genus for hyllognathopus bassoti Rouch, 1972, and description of a new species of Phyllognathopus. ZooKeys 104: 1-65, DOI: http://dx.doi.org/10.3897/zookeys.104.763, URL: http://dx.doi.org/10.3897/zookeys.104.763
F6FE9D0E175B48464A0DACB3691F05CB.taxon	etymology	Etymology. The specific name derives from the Latin adjective inexspectatus which means " unexpected ", alluding to the surprising geographical location of the species, being the taxonomically related Phyllognathopus distributed in the Southern Hemisphere, and to the ecological finding of this species, which was collected from a large karstic aquifer in Central Italy, whereas all the other members of the genus are epigean.	en	Galassi, Diana M. P., Laurentiis, Paola De, Fiasca, Barbara (2011): Systematics of the Phyllognathopodidae (Copepoda, Harpacticoida): re-examination of Phyllognathopus viguieri (Maupas, 1892) and Parbatocamptus jochenmartensi Dumont and Maas, 1988, proposal of a new genus for hyllognathopus bassoti Rouch, 1972, and description of a new species of Phyllognathopus. ZooKeys 104: 1-65, DOI: http://dx.doi.org/10.3897/zookeys.104.763, URL: http://dx.doi.org/10.3897/zookeys.104.763
394F29EF91DFB2645CC8A01944AE5889.taxon	diagnosis	Diagnosis. Phyllognathopodidae. Habitus slightly dorsoventrally flattened with no clear demarcation between prosome and urosome. Integumental dorsal window on cephalosome not confirmed. Integument without surface pits, moderately sclerotized. Ceph alosome rounded; rostrum elongate, clearly articulated to cephalosome. Cephalosome and both thoracic and abdominal somites with cuticular ornamentation represented by dorsal sensilla. First pedigerous somite free. Hyaline frills of cephalosome, somites bearing P 1 - P 4 plain both dorsally and ventrally. P 5 - bearing somite with large paired pores laterodorsally. Sexual dimorphism in antennule, P 5, P 6, urosomal segmentation and ornamentation, and morphology of anal operculum. Female first and second abdominal somites fused forming the genital double-somite. Female urosomal segments with plain hyaline frills ventrally. Female genital apparatus simplified; copulatory pore located at the end of the proximal third of the genital double-somite. Seminal receptacles laterally located and condensed close to the lamellar sixth legs. Male urosome with different arrangement of hyaline frill ornamentations: urosome consisting of 6 segments, second urosomite with indented hyaline frill, third and fourth urosomites with deep ventral sockets; socket on third urosomite plicate, with smooth free distal margin, and 2 setules laterally inserted close to the socket opening; socket on fourth urosomite with free distal margin ornamented by strong and long spinules, covering the opening; fifth urosomite with indented hyaline frill. Anal somite with paired sensilla on dorsal side. Anal operculum protruding free distal margin of anal somite and extruded in strong spinular processes. Sexual dimorphism in the number of spinular processes of anal operculum (3 in females vs. 4 in males; and, in general, anal operculum in male more armed than in female). Caudal rami sub-quadrate, with incomplete setal pattern (6 setae). Dorsal seta inserted on distal third of caudal ramus. Antennule: 8 - segmented in female, basically 9 - segmented in male; geniculation between segments 7 and 8; penultimate and last segments, each with suture line marking original segmentation between former segments 8 and 9, and 10 and 11, respectively. Long tube-pores on segments 1 and 2 in both sexes. Antenna: armature of the second endopodal segment as in Phyllognathopus and Parbatocamptus, consisting of 10 elements. Exopod 1 - segmented, with 3 lateral and 2 apical setae. Mandible: mandibular palp biramous, basis with inner spinule row, exopod with 1 apical and 1 inner setae; endopod with 1 inner, 1 subapical and 2 apical setae. Armature of maxillule and maxilla as in Phyllognathopus. Maxilliped: phyllopodial, lamelliform, 1 - segmented. Clear trace of ancestral 2 - segmented condition marked by the presence of outer and inner incisions as in Parbatocamptus. Armature consisting of 11 elements: 1 strong spine inserted at inner corner of former segment 1; 4 spines and 1 spiniform short seta inserted along inner margin, 5 bipinnate setae in apical position, armature topology basically referable to that of Parbatocamptus. P 1 - P 3 with 3 - segmented exopods and endopods. P 4 with 2 - segmented exopod and endopod. P 1 - P 3 praecoxa well developed. P 1 exopod and endopod of about the same length; P 2 - P 3 endopods shorter than exopods, reaching about tip of exp- 2. P 4 small - sized, praecoxa missing. Female P 5: free, with clear articulation to P 5 - bearing somite; right and left legs distinct; baseoendopod and exopod coalescent, deep incision marking original segmentation between them; endopodal lobe well developed, elongate, longer than exopodal lobe, rectangular in shape, bearing 1 long pinnate seta, subdistally inserted, close to outer margin and a spinule row apically inserted. Exopodal lobe wide, fully incorporated into baseoendopod; exopodal armature consisting of 4 elements, the outermost bipinnate seta inserted in subdistal position, and three apical elements: 2 spinulose and 1 short setae; basipodal outer seta present. Female P 6 rudimentary, each leg defined by a small cuticular lateral plate bearing a short, naked seta with rounded tip. Male P 5: free, with clear articulation to P 5 - bearing somite; right and left legs separate, intercoxal sclerite rudimentary, but still discernible. Basis of each leg expanded, endopod strongly trasformed, consisting of a sclerotized and strong protrusion articulated to basis. Endopodal seta bipinnate, inserted on posterior surface of the endopod, close to its articulation to basis. Exopod distinct, clearly articulated to basis, wide and short, rectangular in shape, representing most part of the free distal margin of each leg; exopodal armature consisting of 6 elements, the innermost spiniform seta curved inward. Male P 6: right and left legs distinct but closely adjacent to each other along their medial margin, and symmetrical; each leg consisting of a well developed lamellar plate, with spinule row on the anterior surface; armature consisting of 2 inner spines of different length and 1 outer seta.	en	Galassi, Diana M. P., Laurentiis, Paola De, Fiasca, Barbara (2011): Systematics of the Phyllognathopodidae (Copepoda, Harpacticoida): re-examination of Phyllognathopus viguieri (Maupas, 1892) and Parbatocamptus jochenmartensi Dumont and Maas, 1988, proposal of a new genus for hyllognathopus bassoti Rouch, 1972, and description of a new species of Phyllognathopus. ZooKeys 104: 1-65, DOI: http://dx.doi.org/10.3897/zookeys.104.763, URL: http://dx.doi.org/10.3897/zookeys.104.763
394F29EF91DFB2645CC8A01944AE5889.taxon	etymology	Etymology. The genus name is derived from the type genus Phyllognathopus and the Latinised Greek prefix νέοσ which means " new ", referring to the new position of Phyllognathopus bassoti in the systematics of the family Phyllognathopodidae.	en	Galassi, Diana M. P., Laurentiis, Paola De, Fiasca, Barbara (2011): Systematics of the Phyllognathopodidae (Copepoda, Harpacticoida): re-examination of Phyllognathopus viguieri (Maupas, 1892) and Parbatocamptus jochenmartensi Dumont and Maas, 1988, proposal of a new genus for hyllognathopus bassoti Rouch, 1972, and description of a new species of Phyllognathopus. ZooKeys 104: 1-65, DOI: http://dx.doi.org/10.3897/zookeys.104.763, URL: http://dx.doi.org/10.3897/zookeys.104.763
