taxonID	type	description	language	source
03D787DCFFA5FFDAC69C5C8BFA22FB0D.taxon	diagnosis	Diagnosis. Surstylus submedially strongly curved inward, apically with 1 small prensiseta (Figs 3 – 9 A or B). Only those characters that depart from the description of the coleoptrata species group are provided here for the sake of brevity. For more detailed description see Laštovka & Máca 1982; Hu & Toda 1994; Bächli et al. 2004.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFA5FFD5C69C5A4EFAA7FECE.taxon	description	(Fig. 3)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFA5FFD5C69C5A4EFAA7FECE.taxon	diagnosis	Diagnosis. Surstylus acute apically, with 2 minute projections (Fig. 3 A); aedeagus concave medially in lateral view (Fig. 3 C); gonopods nearly triangular in ventral view (Fig. 3 D).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFA5FFD5C69C5A4EFAA7FECE.taxon	description	Description. Male: Frons black. First flagellomere greish yellow. Clypeus and palpus yellow. Mesonotum brown to dark brown, with 1 thin, yellow longitudinal stripe medially, yellowish brown laterally. Katepisternum entirely white. Scutellum black. Femur of foreleg with 4 long erect setae near distal part of posteroventral surface. Abdominal tergites dark brown to black medially, brown to dark brown laterally. Male terminalia (Fig. 3): Epandrium slightly protruded ventrally, with ca. 30 setae on posterior portion per side. Surstylus with numerous setulae distally. Cercus lacking pubescence. Paramere with 2 setulae. Tenth sternite convex dorsolaterally. Measurements. BL = 2.90 – 3.02 mm in 5 ♂, ThL = 1.26 – 1.35 mm in ♂, WL = 2.66 – 3.02 mm in ♂, WW = 1.13 – 1.21 mm in ♂, arb = 3 – 4 / 4 – 6, avd = 0.61 – 0.67, adf = 1.21 – 1.42, flw = 1.74 – 2.00, FW / HW = 0.36 – 0.37, ch / o = 0.22 – 0.25, prorb = 0.29 – 0.30, rcorb = 0.65 – 0.83, vb = 0.58 – 0.72, dcl = 0.57 – 0.59, presctl = 0.65 – 0.68, sctl = 1.27 – 1.30, sterno = 1.29 – 1.30, orbito = 1.57 – 1.95, dcp = 0.28 – 0.30, sctlp = 1.59 – 1.64, C = 2.09 – 2.23, 4 c = 0.90 – 1.00, 4 v = 1.50 – 1.70, 5 x = 1.32 – 1.69, ac = 6.40 – 7.77, M = 0.41 – 0.47, C 3 F = 0.58 – 0.61. Specimens examined. China: 1 ♂ (SCAU, No. 110905), Gangshan Forest Park, Xinbin, Liaoning, 41 ° 37 ' 07 '' N, 125 ° 24 ' 03 '' E, altitude 539 m, 15. vii. 2018, tree trunk, Y Lin; 2 ♂ (SCAU, Nos 110906, 07), Green Stone Valley Forest Park, Benxi, Liaoning, 41 ° 04 ' 09 '' N, 124 ° 12 ' 12 '' E, altitude 570 m, 17. vii. 2018, tussock, L Gong; 2 ♂ (SCAU, Nos 110908, 09), Dayangcha, Kuandian, Liaoning, 124 ° 7 ' 01 '' E, 41 ° 1 ' 48 '' N, altitude 485 m, 23. vii. 2018, tree trunk, BX Li.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFA5FFD5C69C5A4EFAA7FECE.taxon	distribution	Distribution. China * (Liaoning), Russia (Far East, Europan), Japan (Hokkaido, Honshu), Finland.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAAFFD5C69C5B92FA28F862.taxon	description	(Figs 2 B, 5)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAAFFD5C69C5B92FA28F862.taxon	diagnosis	Diagnosis. Surstylus broadened and short (Fig. 5 A, B); aedeagus nearly round apically (Fig. 5 C); gonopods semicircular and broadened laterally in ventral view (Fig. 5 E). The minimum interspecific genetic distance to S. pililobosa is 8.3 %, which is smaller than to other species (Table 2).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAAFFD5C69C5B92FA28F862.taxon	description	Description. Male: Frons brown, with 1 black band below. First flagellomere yellow, black at apical margin. Clypeus and palpus yellow. Mesonotum brownish to brown, with 3 pairs of yellow longitudinal stripes sublaterally (Fig. 2 B). Katepisternum white (Fig. 2 B). Scutellum brown (Fig. 2 B). Femur of foreleg with 3 long erect setae near distal part of posteroventral surface. Abdominal tergites dark brown medially, brownish laterally (Fig. 2 B). Male terminalia (Fig. 5): Epandrium protruded ventrally, with ca. 28 setae on posterior portion per side. Surstylus with several setulae distally. Cercus lacking pubescence. Paramere with 2 setulae. Tenth sternite inverted triangular per side. Measurements. BL = 3.01 mm in holotype, ThL = 1.41 mm, WL = 2.95 mm, WW = 1.23 mm, arb = 6 / 4, avd = 0.63, adf = 1.20, flw = 1.76, FW / HW = 0.40, ch / o = 0.20, prorb = 0.73, rcorb = 0.70, vb = 0.51, dc 1 = 0.47, presct 1 = 0.54, sct 1 = 0.90, sterno = damaged, orbito = 1.76, dcp = 0.28, sct 1 p = 1.67, C = 1.88, 4 c = 1.11, 4 v = 1.67, 5 x = 1.29, ac = 9.33, M = 0.49, C 3 F = 0.63. Specimens examined. Holotype ♂ (SCAU, No. 110945), China: Pangquangou, Jiaocheng, Shanxi, 37 ° 04 ' 47 '' N, 111 ° 30 ' 20 '' E, altitude 2200 m, 30. vii. 2016, tree trunk, YL Wang.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAAFFD5C69C5B92FA28F862.taxon	etymology	Etymology. A combination of the Latin words: latus and tabula, reference to the surstylus broadened distally.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAAFFD5C69C5E10FCAAFB49.taxon	description	(Figs 2 A, 4)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAAFFD5C69C5E10FCAAFB49.taxon	diagnosis	Diagnosis. Mesonotum black medially, brown laterally (Fig. 2 A); aedeagus convex apicomedially (Fig. 4 C, D).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAAFFD5C69C5E10FCAAFB49.taxon	description	Description. Male: Frons black. First flagellomere orange on basal 2 / 3, black on distal 1 / 3. Clypeus and palpus yellow. Katepisternum entirely white. Scutellum black (Fig. 2 A). Femur of foreleg with 4 long erect setae near distal part of posteroventral surface. Abdominal tergites nearly black (Fig. 2 A). Male terminalia (Fig. 4 A – G): Epandrium slightly protruded ventrally, with ca. 30 setae on posterior portion per side. Surstylus slender, with several setulae distally. Cercus lacking pubescence. Paramere with 1 setula. Tenth sternite broadened, with 1 pair of processes submedially. Female terminalia (Fig. 4 H – J): Epiproct and cercus with setae, lacking pubescence. Hypoproct entirely pubescent, with dense setae along posterior margin. Eighth sternite glabrous on basal 3 / 4 region, with pubescence and a few of setae on distal 1 / 4 region. Measurements. BL = 3.27 mm in 1 ♂, 2.93 mm in 1 ♀, ThL = 1.42 mm in ♂, 1.54 mm in ♀, WL = 3.32 mm in ♂, 2.59 mm in ♀, WW = 1.21 mm in ♂, 1.23 mm in ♀, arb = 5 – 7 / 4 – 6, avd = 0.65 – 0.75, adf = 1.26 – 1.27, flw = 1.76 – 1.82, FW / HW = 0.30 – 0.33, ch / o = 0.23 – 0.25, prorb = 0.27 – 0.28, rcorb = 0.74 – 0.90, vb = 0.57 – 0.60, dcl = 0.57 – 0.62, presctl = 0.64 – 0.72, sctl = damaged, sterno = 0.82, orbito = 1.59 – 1.68, dcp = 0.32 – 0.33, sctlp = 1.73 – 1.74, C = 1.94 – 2.08, 4 c = 0.95 – 1.13, 4 v = 1.30 – 1.45, 5 x = 1.26 – 1.63, ac = 8.39 – 9.92, M = 0.42 – 0.50, C 3 F = 0.63 – 0.70. Specimens examined. China: 2 ♂, 1 ♀ (SCAU, Nos 110910, 11), Laobiangou, Benxi, Liaoning, 41 ° 04 ' 39 '' N, 124 ° 30 ' 03 '' E, altitude 575 m, 21. vii. 2018, tree trunk, L Gong, NN Wang.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAAFFD5C69C5E10FCAAFB49.taxon	distribution	Distribution. China * (Liaoning), Japan (Hokkaido).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFA9FFD1C69C5A34FD9BFECE.taxon	description	(Figs 2 C, 6)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFA9FFD1C69C5A34FD9BFECE.taxon	diagnosis	Diagnosis. This species resembles S. toyaensis Okada & Sidorenko, 1992 in some characters of the male terminalia, but can be distinguished by the mesonotum dark brown, with 1 thin, unconspicuous, yellow longitudinal stripe medially, yellowish brown laterally (Fig. 2 C); aedeagus slightly flat apically in ventral view (Fig. 6 C, D). The minimum interspecific genetic distance to S. toyaensis is 3.5 %, which is smaller than to other species (Table 2).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFA9FFD1C69C5A34FD9BFECE.taxon	description	Description. Male: Frons black. First flagellomere yellow on basal 2 / 3, black on distal 1 / 3. Clypeus and palpus yellow. Katepisternum entirely white. Scutellum black (Fig. 2 C). Femur of foreleg with 4 long erect setae near distal part of posteroventral surface. Abdominal tergites dark brown to black (Fig. 2 C). Male terminalia (Fig. 6): Epandrium slightly protruded ventrally, with numerous setae on posterior portion per side. Surstylus with numerous setulae basally. Cercus lacking pubescence. Tenth sternite broadened, with 1 pair of processes submedially. Paramere with 1 setula. Measurements. BL = 4.07 mm, ThL = 1.87 mm, WL = 3.20 mm, WW = 1.50 mm, arb = 8 / 6, avd = 1.00, adf = 1.11, flw = 1.89, FW / HW = 0.45, ch / o = 0.10, prorb = 1.11, rcorb = 0.84, vb = 0.35, dc 1 = 0.64, presct 1 = 0.64, sct 1 = damaged, sterno = damaged, orbito = 2.83, dcp = 0.18, sct 1 p = 1.90, C = 2.23, 4 c = 0.87, 4 v = 1.40, 5 x = 1.22, ac = 8.66, M = 0.37, C 3 F = 0.84. Specimens examined. Holotype ♂ (SCAU, No. 122193), China: Fengtongzhai, Baoxing, Sichuan, 30 ° 32 ' 55 '' N, 102 ° 53 ' 56 '' E, altitude 1500 m, 18. vii. 2017, tree trunk, L Gong.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFA9FFD1C69C5A34FD9BFECE.taxon	etymology	Etymology. Pertaining to the type locality, where Giant Panda was distributed in.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFA9FFD1C69C5A34FD9BFECE.taxon	distribution	Distribution. China (Sichuan).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAFFFD3C69C5ACCFE11FEB6.taxon	description	(Figs 2 D, 7)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAFFFD3C69C5ACCFE11FEB6.taxon	diagnosis	Diagnosis. This species resembles S. longifibula Takada, 1968 in some characters of the body colour patterns (Fig. 2 D), but can be distinguished by the male terminalia, in this species, the surstylus without minute projections, except acute apically (Fig. 7 A); aedeagus medially expanded in lateral view, strongly narrowed in ventrally (Fig. 7 B, C); gonopods nearly vaulted in ventral view (Fig. 7 D). The minimum interspecific genetic distance to S. longifibula is 4.3 %, which is smaller than to other species (Table 2).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAFFFD3C69C5ACCFE11FEB6.taxon	description	Description. Male: Frons black. First flagellomere greish yellow. Clypeus and palpus yellow. Mesonotum brown to dark brown, with 1 thin, unconspicuous, yellow longitudinal stripe medially, yellowish brown laterally. Katepisternum entirely white. Scutellum black. Femur of foreleg with 4 long erect setae near distal part of posteroventral surface. Abdominal tergites nearly black. Male terminalia (Fig. 7): Epandrium slightly protruded ventrally, with ca. 15 setae on posterior portion per side. Surstylus with numerous setulae distally. Cercus lacking pubescence. Paramere with 2 setulae. Tenth sternite flat dorsolaterally. Measurements. BL = 3.01 mm in holotype, ThL = 1.47 mm, WL = 2.95 mm, WW = 1.26 mm, arb = 6 / 4, avd = 0.62, adf = 1.36, flw = 1.87, FW / HW = 0.41, ch / o = 0.22, prorb = 1.30, rcorb = 0.73, vb = 0.52, dc 1 = damaged, presct 1 = damaged, sct 1 = damaged, sterno = damaged, orbito = 1.73, dcp = 0.35, sct 1 p = 1.68, C = 2.08, 4 c = 1.02, 4 v = 1.55, 5 x = 1.26, ac = 9.25, M = 0.45, C 3 F = 0.65. Specimens examined. Holotype ♂ (SCAU, No. 110944), China: Pangquangou, Jiaocheng, Shanxi, 37 ° 04 ' 47 '' N, 111 ° 30 ' 20 '' E, altitude 2200 m, 30. vii. 2016, tree trunk, L Zhu.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFAFFFD3C69C5ACCFE11FEB6.taxon	etymology	Etymology. A combination of the Latin words: pinguis and foliaceous, reference to the aedeagus medially expanded in lateral view.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFACFFD2C69C59A8FD8EFD56.taxon	description	(Figs 2 E, 8)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFACFFD2C69C59A8FD8EFD56.taxon	diagnosis	Diagnosis. This species resembles S. nigrithorax Strobl, 1898 in some characters of the colour patterns of body (Fig. 2 E) and semicircular gonopods (Fig. 8 E), but can be distinguished by the median flaps of hypandrium (Hu & Toda, 1994; fused gonopod in Bächli et al., 2004) concave, spoon-like (Fig. 8 E, F); aedeagus with numerous, serrated, minute processes medially (Fig. 8 C, D). However, the minimum interspecific genetic distance is 3.1 % to S. pililobosa (Table 2).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFACFFD2C69C59A8FD8EFD56.taxon	description	Description. Male: Frons brown. First flagellomere yellow on basal 1 / 4, brown on distal 3 / 4. Clypeus yellow, slightly brownish medially. Palpus yellow. Katepisternum entirely white. Scutellum brown (Fig. 2 E). Femur of foreleg with 5 long erect setae near distal part of posteroventral surface. Abdominal tergites entirely dark brown medially, yellowish brown laterally (Fig. 2 E). Male terminalia (Fig. 8): Epandrium protruded ventrally, with ca. 20 setae on posterior portion per side. Surstylus distally with numerous setulae on inner surface. Cercus lacking pubescence. Tenth sternite with 1 pair of processes submedially. Paramere lacking setula. Measurements. BL = 3.47 mm, ThL = 1.46 mm, WL = 2.90 mm, WW = 1.14 mm, arb = 6 / 5, avd = 0.77, adf = 1.31, flw = 1.82, FW / HW = 0.28, ch / o = 0.23, prorb = 0.30, rcorb = 0.79, vb = 0.66, dc 1 = 0.33, presct 1 = 0.35, sct 1 = damaged, sterno = 0.88, orbito = 1.45, dcp = 0.30, sct 1 p = 1.59, C = 2.00, 4 c = 1.14, 4 v = 1.36, 5 x = 1.41, ac = 8.42, M = 0.54, C 3 F = 0.63. Specimens examined. Holotype ♂ (SCAU, No. 110902), China: Guojiapuzi, Kuandian, Liaoning, 40 ° 46 ' 46 '' N, 124 ° 45 ' 07 '' E, altitude 342 m, 26. vii. 2018, tree trunk, NN Wang.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFACFFD2C69C59A8FD8EFD56.taxon	etymology	Etymology. From the Latin word: spatulate, referring to the median flaps of hypandrium spoon-like.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFACFFD2C69C59A8FD8EFD56.taxon	distribution	Distribution. China (Liaoning).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB2FFCCC69C5A32FD9FFE2D.taxon	description	(Figs 2 F, 9)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB2FFCCC69C5A32FD9FFE2D.taxon	diagnosis	Diagnosis. This species resembles S. malanostoma Chen & Chen, 2008 from Hubei, China in the colour patterns of body (Fig. 2 F) and some characters of male terminalia (Fig. 9), but can be distinguished by the aedeagus with 1 pair of fringe-like processes ventrally (Fig. 9 C, D); gonopods slightly protruded dorsomedially, with a few serrated, minute processes (Fig. 9 E, F). The minimum interspecific genetic distance to S. malanostoma is 3.4 %, which is smaller than to other species (Table 2).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB2FFCCC69C5A32FD9FFE2D.taxon	description	Description. Male: Frons black on upper half, with 1 yellow band medially, brown below. First flagellomere yellow on basal 2 / 3, black on distal 1 / 3. Clypeus black medially, yellow laterally. Palpus yellow, slightly brownish apically. Mesonotum brown medially, yellowish brown laterally (Fig. 2 F). Katepisternum entirely white. Scutellum brown (Fig. 2 F). Foreleg femur with 4 long suberect setae near distal part of posteroventral surface. Abdominal tergites entirely brown (Fig. 2 F). Male terminalia (Fig. 9): Epandrium with ca. 24 setae on dorsal to posterolateral portion per side. Cercus lacking pubescence. Surstylus distally with numerous setulae on inner surface. Paramere with ca. 2 setulae. Aedeagus with numerous, thin setae along apical margin and serration processes on dorsal portion. Gonopods with 1 small projection distodorsally in lateral view. Measurements. BL = 2.80 mm in holotype (4 ♂ paratypes: 2.47 – 2.67), ThL = 1.33 mm (1.11 – 1.58), WL = 2.20 mm (1.87 – 2.00), WW = 1.01 mm (0.87 – 1.00), arb = 7 / 4 (7 / 4 – 8 / 4), avd = 0.75 (0.50 – 0.67), adf = 1.60 (1.43 – 1.80), flw = 2.4 (1.57 – 2.20), FW / HW = 0.41 (0.40 – 0.44), ch / o = 0.10 (0.10 – 0.12), prorb = 0.94 (0.93 – 1.07), rcorb = 0.63 (0.71 – 0.79), vb = 0.30 (0.26 – 0.36), dcl = 0.48 (0.44 – 0.63), presctl = 0.40 (0.45 – 0.53), sctl = 1.60 (1.61 – 1.75), sterno = 0.79 (0.75 – 0.83), orbito = 1.82 (1.50 – 1.80), dcp = 0.23 (0.22 – 0.24), sctlp = 2.5 (2.67 – 3.3), C = 2.06 (2.00 – 2.07), 4 c = 0.89 (0.79 – 1.07), 4 v = 1.42 (1.21 – 1.79), 5 x = 1.16 (1.16 – 2.00), ac = 8.5 (7.5 – 8.5), M = 0.37 (0.41 – 0.43), C 3 F = 0.71 (0.66 – 0.84), A = 1.17 (1.03 – 1.17), H = 0.17 (0.17 – 0.22). Specimens examined. Holotype ♂ (SCAU, No. 121491), China: Yixiang, Pu'er, Yunnan, 22 ° 42 ' N, 101 ° 09 ' E, altitude 1400 m, 2. x. 2011, tree trunk, HW Chen. Paratypes: 3 ♂ (SCAU, Nos 121492 – 94), 13. v. 2012, same data as holotype; 1 ♂ (KIZ, No. 0090662), Guanlei, Mengla, Yunnan, 21 ° 37 ' 39 '' N, 101 ° 9 ' 55 '' E, altitude 670 m, 14. ix. 2012, tussock, JJ Gao; 1 ♂ (SCAU, No. 124705), Husa, Longchuan, Yunnan, 24 ° 27 ' 37 '' N, 97 ° 45 ' 9 '' E, altitude 1227 m, 7. ix. 2017, tree trunk, HW Chen.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB2FFCCC69C5A32FD9FFE2D.taxon	etymology	Etymology. From the Greek word: stachydifolius, referring to the aedeagus with 1 pair of fringe-like processes.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB2FFCCC69C5A32FD9FFE2D.taxon	distribution	Distribution. China (Yunnan).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB3FFCCC69C59C2FD36F83C.taxon	diagnosis	Diagnosis. Surstylus with several thin, long setae basally and 1 strong prensiseta apically (Figs 10 B, 11 B); gonopods with 1 pair of projections sublaterally (Figs 10 D, 11 D); aedeagus mostly ciliate and separated to leafiness from basally (Figs 10 C, 11 C); 10 th sternite mostly narrow, nearly arcuate, with 1 pair of projections posterolaterally (Figs 10 E, 11 E). Only characters departing from the diagnosis of the ornatipes species group (Cheng et al. 2009) are provided for brevity.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB0FFCEC69C5FE8FD9FFCEB.taxon	description	(Figs 2 G, 10)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB0FFCEC69C5FE8FD9FFCEB.taxon	diagnosis	Diagnosis. This species resembles S. unguiculata sp. nov. in the colour patterns of body (Fig. 2 G) and male terminalia (Fig. 10), but can be distinguished by the aedeagus with 4 short and thick processes per side (Fig. 10 C); sublateral processes of gonopods shorter and broadly separated than in S. unguiculata sp. nov. (Fig. 10 D). The minimum interspecific genetic distance to S. unguiculata is 3.3 %, which is smaller than to other species (Table 2); see description of S. unguiculata for more detail.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB0FFCEC69C5FE8FD9FFCEB.taxon	description	Description. Male: Pedicel yellow medially, black laterally; first flagellomere nearly black. Palpus yellow basally, brown on distal 1 / 3 in male. Mesonotum brown (Fig. 2 G). Katepisternum dark brown on upper part, whiteyellow on lower part (Fig. 2 G). Scutellum black (Fig. 2 G). Legs dark brown to black, yellow on basal part of fore femora and 2 nd to 5 th tarsomeres. Fore femur with 3 long setae on distal half of posteroventral surface. All abdominal tergites brown (Fig. 2 G); all sternites yellowish brown. Male terminalia (Fig. 10): Epandrium with ca. 20 setae near posterior portion per side. Surstylus broad in lateral view. Cercus lacking pubescence. Tenth sternite with 1 pair of asymmetric projections sublaterally. Paramere with ca. 2 sensilla. Aedeagus with pubescence. Gonopods with 1 suberect lobe-like process bearing several pubescence. Measurements. BL = 2.80 mm in holotype (1 ♂ paratype: 2.75), ThL = 1.13 mm (1.26), WL = 2.00 mm (2.65), WW = 1.00 mm (1.21), arb = 6 / 5 (5 / 4), avd = 0.60 (0.84), adf = 1.43 (1.02), flw = 1.57 (1.90), FW / HW = 0.31 (0.40), ch / o = 0.13 (0.11), prorb = 1.07 (0.90), rcorb = 0.60 (0.62), vb = 0.25 (0.27), dc 1 = damaged (damaged), presct 1 = 0.50 (damaged), sct 1 = 1.59 (damaged), sterno = damaged (0.87), orbito = 2.25 (1.91), dcp = 0.24 (0.25), sct 1 p = 1.08 (1.15), C = 2.00 (2.19), 4 c = 1.13 (1.42), 4 v = 1.57 (1.82), 5 x = 1.33 (1.13), ac = 9.00 (7.47), M = 0.50 (0.62), C 3 F = 0.71 (0.58). Specimens examined. Holotype ♂ (SCAU, No. 124706), China: Dashuigou, Lüchun, Yunnan, 22 ° 57 ' 19 '' N, 101 ° 57 ' 4 '' E, altitude 1310 m, 2. viii. 2016, tussock, HW Chen. Paratype: China: 1 ♂ (SCAU, No. 124709), Mengma, Menglian, Yunnan, 22 ° 12 ' 59 '' N, 99 ° 22 ' 51 '' E, altitude 1070 m, 15. iv. 2018, tree trunk, HW Chen.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB0FFCEC69C5FE8FD9FFCEB.taxon	etymology	Etymology. A combination of the Latin words: hirtus + foliaceus, referring to the aedeagus with several setae.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB0FFCEC69C5FE8FD9FFCEB.taxon	distribution	Distribution. China (Yunnan).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB1FFC8C69C5CF3FD3EFF20.taxon	description	(Figs 2 H, 11)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB1FFC8C69C5CF3FD3EFF20.taxon	diagnosis	Diagnosis. This species resembles S. linnanensis Cheng, Gao & Chen, 2009 from southern China in the colour patterns of body (Fig. 2 H) and male terminalia (Fig. 11), but can be distinguished from it by the aedeagus nearly bilobed, with 7 pair of slender processes and thick, long pubescence laterally (Fig. 11 C); gonopods with 1 suberect lobe-like process bearing several pubescence (Fig. 11 D). However, the minimum interspecific genetic distance is 3.3 % to S. hirtifoliacea (Table 2).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB1FFC8C69C5CF3FD3EFF20.taxon	description	Description. Male and Female: Pedicel brown medially, black laterally; first flagellomere nearly black. Palpus yellow basally, brown on distal 1 / 3 in male, brown on distal 2 / 3 in female. Mesonotum dark brown medially, brown laterally (Fig. 2 H). Katepisternum dark brown on upper part, white-yellow on lower part (Fig. 2 H). Scutellum black (Fig. 2 H). Legs dark brown to black, yellow on basal part of fore femora and 2 nd to 5 th tarsomeres. Fore femur with 3 long setae on distal half of posteroventral surface. All abdominal tergites brown (Fig. 2 H); all sternites yellowish brown. Male terminalia (Fig. 11 A – E): Epandrium with approximately 22 setae near posterior portion per side. Surstylus broad in lateral view. Cercus lacking pubescence. Tenth sternite with 1 pair of asymmetric projection sublaterally. Plate between hypandrium and aedeagus with 4 sensilla. Aedeagus composed of 1 bilobed medial process with pubescence. Female terminalia (Fig. 11 F – H): Epiproct and cercus with setae, lacking pubescence. Hypoproct pubescent medially, with dense setae along lateral and posterior margins. Eighth sternite glabrous, with a pair of small setae near posterior margin. Measurements. BL = 2.83 mm in holotype (mm in five ♂ and five ♀ paratypes: 2.37 – 2.83 in ♂, 2.87 – 3.27 in ♀), ThL = 1.17 mm (1.20 – 1.30 in ♂, 1.20 – 1.47 in ♀), WL = 1.83 mm (1.93 – 2.17 in ♂, 2.00 – 2.23 in ♀), WW = 1.00 mm (1.00 – 1.10 in ♂, 1.07 – 1.17 in ♀), arb = 7 / 5 (7 – 9 / 5 – 6), avd = 0.90 (0.60 – 0.92), adf = 1.67 (1.11 – 1.83), flw = 2.17 (1.57 – 2.50), FW / HW = 0.46 (0.34 – 0.44), ch / o = 0.10 (0.09 – 0.16), prorb = 0.94 (1.00 – 1.27), rcorb = 0.59 (0.60 – 0.91), vb = 0.29 (0.18 – 0.31), dc 1 = damaged (0.31 – 0.48), presctl = damaged (0.55 – 0.71), sct 1 = 2.00 (1.40 – 1.90), sterno = 1.00 (0.85 – 1.25), orbito = 2.25 (1.67 – 2.50), dcp = 0.29 (0.17 – 0.36), sct 1 p = 1.30 (1.09 – 1.42), C = 1.89 (1.71 – 2.06), 4 c = 1.20 (1.00 – 1.29), 4 v = 1.60 (1.50 – 1.81), 5 x = 1.50 (1.18 – 1.54), ac = 9.00 (7.00 – 10.05), M = 0.60 (0.43 – 0.67), C 3 F = 0.39 (0.50 – 0.74). Specimens examined. Holotype: ♂ (SCAU, No. 111477), China: Qimaba, Lüchun, Yunnan, 22 ° 49 ' 8 '' N, 102 ° 17 ' 25 '' E, altitude 1280 m, 3,4. viii. 2016, tree trunk, HW Chen. Paratypes: China: 3 ♂, 7 ♀ (SCAU, Nos 111478 – 7), same data as holotype, HW Chen, L Gong, YQ Liu; 1 ♀ (SCAU, No. 111488), Dashuigou, Lühun, Yunnan, 22 ° 57 ' 19 '' N, 101 ° 57 ' 4 '' E, altitude 1310 m, 2. viii. 2016, tree trunk, HW Chen; 1 ♂, 1 ♀ (SCAU, Nos 111540, 41), Jinshan Forest Park, Menglian, Yunnan, 22 ° 20 ' 6 '' N, 99 ° 34 ' 34 '' E, altitude 1049 m, 24. x. 2017, tree trunks, HW Chen, L Gong; 1 ♂, 1 ♀ (KIZ, Nos 0 0 90663, 64), Moli Forest Park, Ruili, Yunnan, 24 ° 7 ' 5 '' N, 97 ° 58 ' 51 '' E, altitude 920 m, 5. xi. 2017, tree trunks, HW Chen, L Gong; 1 ♀ (SCAU, No. 124146), Beibeng, Motuo, Xizang, 29 ° 19 ' N, 95 ° 20 ' E, altitude 800 m, 29. ix. 2010, tree trunk, L Wang.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB1FFC8C69C5CF3FD3EFF20.taxon	etymology	Etymology. From the Latin word: unguiculus, referring to the aedeagus with claw-shape processes. Distribution. China (Yunnan, Xizang).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB7FFC8C69C5DFBFD31F859.taxon	description	(Figs 2 I, 12)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB7FFC8C69C5DFBFD31F859.taxon	diagnosis	Diagnosis. This species resembles S. flavichypeata Chen & Chen in Lu et al. 2011 a in the yellow clypeus and palpus, and colour patterns of body (Fig. 2 I), but can be distinguished by having the face black on upper 2 / 3, yellow on lower 1 / 3; aedeagus with triangular opening in ventral view (Fig. 12 B). In S. flavichypeata: Face black, with yellow band medially; aedeagus with rhombic opening in ventral view (fig. 24 in Lu et al. 2011 a). The minimum interspecific genetic distance to P. flavichypeata is 7.1 %, which is smaller than to other species in this group (Table 2).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB7FFC8C69C5DFBFD31F859.taxon	description	Description. Male and female. Frons black, with orange yellow stripe medially and band submedially. First flagellomere medially grayish yellow on basal half, black on distal half. Mesonotum black, with thin, yellow longitudinal stripes sublaterally (Fig. 2 I). Katepisternum entirely white (Fig. 2 I). Scutellum black, yellow at tip (Fig. 2 I). Veins R 4 + 5 and M 1 lacking setae on basal section of dorsal surface. Legs white on basal half of all femora, brown on distal 1 / 3 of foreleg femur, black on distal half of femora and basal half of tibiae of mid and hindleg, yellow on the rest. Fore femur distally with 3 or 4 setae on posteroventral surface. Abdominal tergites 1 st to 5 th medially brown, laterally yellowish brown, nearly black on 6 th tergite in male; all tergites black in female. Sternites yellow to brownish yellow. Male terminalia (Fig. 12 A – F): Hypandrium anteromedially slightly elongated, about 1.2 times as long as width of epandrium, with a few paramedian setae posteroventrally. Gonopods slightly triangular. Aedeagus slightly expanded distally in ventral view, about 3 / 5 times as long as aedeagal apodeme. Female terminalia (Fig. 12 G – I): Epiproct, cercus and hypoproct with dense setae and pubescence. Eighth sternite glabrous on anterior 2 / 3 region, with dense setae and pubescence on posterior 1 / 3 region. Measurements. BL = 3.60 mm in holotype (mm in 2 ♂ and 3 ♀ paratypes: 3.67 – 4.13 in ♂, 4.00 – 4.13 in ♀), ThL = 1.73 mm (1.53 – 2.33 in ♂, 1.87 – 2.07 in ♀), WL = 3.00 mm (2.80 – 3.47 in ♂, 3.00 – 3.87 in ♀), WW = 1.53 mm (1.40 – 1.87 in ♂, 1.33 – 1.60 in ♀), arb = 4 / 2 (7 – 8 / 3 – 5), avd = 0.50 (0.33 – 0.89), adf = 2.40 (1.50 – 1.88), flw = 1.60 (1.40 – 2.40), FW / HW = 0.51 (0.40 – 0.50), ch / o = 0.17 (0.16 – 0.33), prorb = 0.93 (0.82 – 1.42), rcorb = 0.71 (0.70 – 1.00), orbito = 1.80 (2.20 – 3.00), vb = 0.75 (0.18 – 0.80), sctl = 1.60 (1.25 – 1.60), sterno = 0.67 (0.50 – 0.89), sctlp = 1.45 (1.27 – 1.83), C = 1.92 (1.25 – 2.00), 4 c = 1.50 (1.08 – 1.40), 4 v = 2.38 (1.10 – 2.00), 5 x = 1.50 (1.25 – 2.00), ac = 6.00 (6.50 – 12.00), M = 0.75 (0.50 – 0.80), C 3 F = 0.67 (0.67 – 0.79). Specimens examined. Holotype ♂ (SCAU, No. 111533), China: Mulun, Huanjiang, Guangxi, 25 ° 07 ' N, 108 ° 02 ' E, altitude 480 m, 16. ix. 2015, tree trunk, YQ Liu. Paratypes: China: 2 ♀ (SCAU, Nos 111534, 35), same data as holotype; 1 ♂ (KIZ, No. 0090669), Qimaba, Lühun, Yunnan, 22 ° 49 ' 8 '' N, 102 ° 17 ' 25 '' E, altitude 1280 m, 3. viii. 2016, tree trunk, HW Chen; 1 ♂ (KIZ, No. 0090670), Yakou, Huanglianshan, Lühun, Yunnan, 22 ° 53 ' 40 '' N, 102 ° 18 ' 17 '' E, altitude 1900 m, 31. x. 2016, tree trunk, HW Chen; 2 ♂, 1 ♀ (SCAU, Nos 111536 – 38), Wengang, Libo, Guizhou, 25 ° 13 ' N, 107 ° 56 ' E, altitude 754 m, 19. ix. 2015, tree trunks, YQ Liu, Y Zhang, L Zhu.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB7FFC8C69C5DFBFD31F859.taxon	etymology	Etymology. A combination of the Latin words: bi- + flavus, referring to the yellow clypeus and palpus.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB7FFC8C69C5DFBFD31F859.taxon	distribution	Distribution. China (Yunnan, Guizhou).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB7FFC8C69C5E39FA31FDE3.taxon	diagnosis	Diagnosis. Surstylus not strongly arcuate, mostly with 1 small acute prensiseta submedially on inner margin, lacking setae basally, with dense setae distally (Figs 12 A, 13 A); 10 th sternite erect and expanded distally (Figs 12 F, 13 F). Only characters that depart from the diagnosis of the undulata species group (Lu et al. 2011 a) are provided.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB4FFC7C69C587EFD9FFB08.taxon	description	(Figs 2 J, 13)	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB4FFC7C69C587EFD9FFB08.taxon	diagnosis	Diagnosis. This species resembles S. flavipalpata Chen & Chen in Lu et al. 2011 a in the yellow clypeus and palpus, and colour patterns of body (Fig. 2 J), but can be distinguished by the aedeagus lacking serrated, minute processes basally in lateral view, and with round opening in ventral view (Fig. 13 B, C). The minimum interspecific genetic distance to S. flavipalpata is 6.9 % (Table 2).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB4FFC7C69C587EFD9FFB08.taxon	description	Description. Male and female. Frons black, with orange yellow stripe medially and band submedially. Frontoorbital plate orange yellow. First flagellomere medially grayish yellow on basal 2 / 3, black on distal 1 / 3. Face black on upper 2 / 3, yellow on lower 1 / 3. Mesonotum medially brown and with yellow, thin longitudinal stripe, laterally dark brown, with yellow longitudinal stripes and a few patches (Fig. 2 J). Katepisternum entirely white (Fig. 2 J). Scutellum brownish yellow medially, dark brown to balck laterally (Fig. 2 J). Veins R 4 + 5 and M 1 lacking setae on basal section of dorsal surface. Legs white, brownish knee articulations of fore and hindlegs, and black on distal half of hindleg, distal half of femur and basal half of tibia of midleg. Fore femur distally with 6 – 8 setae on posteroventral surface. Abdominal 1 st to 5 th tergites dark brown to black medially, yellow laterally; 6 th tergite nearly black. Sternites nearly yellow. Male terminalia (Fig. 13 A – F): Distal part of 10 th sternite diamond-shaped, about 1.85 times as wide as long. Hypandrium anteromedially elongated, as long as width of epandrium and with a few paramedian setae posteroventrally. Gonopods slightly rounded apicoomedially. Aedeagus about 0.5 times as long as aedeagal apodeme. Female terminalia (Fig. 13 G – I): Epiproct, cercus and hypoproct with dense setae and pubescence. Eighth sternite mostly glabrous, with several setae and pubescence on posterolateral corners. Measurements. BL = 4.87 mm in holotype (mm in 5 ♂ and 5 ♀ paratypes: 3.20 – 4.67 in ♂, 4.13 – 4.53 in ♀), ThL = 2.53 mm (1.73 – 2.40 in ♂, 2.07 – 2.27 in ♀), WL = 3.47 mm (2.53 – 3.67 in ♂, 3.00 – 3.47 in ♀), WW = 1.73 mm (1.20 – 1.80 in ♂, 1.33 – 1.87 in ♀), arb = 10 / 6 (8 – 10 / 4 – 6), avd = 0.67 (0.67 – 0.83), adf = 1.80 (1.50 – 2.00), flw = 1.50 (1.30 – 2.13), FW / HW = 0.42 (0.36 – 0.48), ch / o = 0.21 (0.20 – 0.36), prorb = 0.65 (0.67 – 1.00), rcorb = 0.87 (0.60 – 1.10), orbito = 2.14 (1.50 – 2.33), vb = 0.40 (0.18 – 0.45), sctl = 1.69 (1.29 – 2.50), sterno = 1.00 (0.50 – 0.94), sctlp = 1.25 (1.67 – 2.50), C = 2.80 (1.59 – 2.92), 4 c = 0.83 (1.00 – 1.38), 4 v = 1.92 (1.77 – 2.50), 5 x = 1.75 (1.50 – 2.50), ac = 5.00 (5.50 – 9.00), M = 0.58 (0.58 – 1.00), C 3 F = 0.80 (0.67 – 0.83). Specimens examined. Holotype ♂ (SCAU, No. 111489), China: Hesong, Menghai, Yunnan, 21 ° 49 ′ N, 100 ° 06 ′ E, altitude 1800 m, 7. v. 2012, tree trunk, HW Chen. Paratypes: China: 6 ♂, 5 ♀ (SCAU, Nos 111490 – 500), same data as holotype; 1 ♂, 4 ♀ (SCAU, Nos 111501 – 05), altitude 1600 m, 12. v. 2012, same data as holotype; 1 ♂, 1 ♀ (SCAU, Nos 111506, 07), Mengyuan, Mengla, Yunnan, 21 ° 43 ' 56 '' N, 101 ° 23 ' 15 '' E altitude 876 m, 18. xii. 2017, tree trunks, HW Chen; 1 ♀ (SCAU, No. 111508), Wangtianshu Forest Park, Mengla, Yunnan, 21 ° 36 ' 9 '' N, 101 ° 35 ' 6 '' E, altitude 640 m, 16. iv. 2010, tree trunk, HW Chen; 4 ♀ (SCAU, Nos 111509 – 12), Menglai Grand Canyon, Cangyuan, Yunnan, 23 ° 13 ' 20 '' N, 99 ° 15 ' 4 '' E, altitude 1240 m, 30. x. 2017, tree trunks, HW Chen, L Gong, BX Li; 3 ♂, 1 ♀ (KIZ, Nos 0090665 – 68), Longtan Forest Park, Ximeng, Yunnan, 22 ° 38 ' 44 '' N, 99 ° 35 ' 39 '' E, altitude 1145 m, 27. x. 2017, tree trunks, HW Chen, L Gong, BX Li; 1 ♂ (SCAU, No. 111513), Moli Forest Park, Ruili, Yunnan, 24 ° 7 ' 5 '' N, 97 ° 58 ' 51 '' E, altitude 920 m, 5. xi. 2017, tree trunk, L Gong; 1 ♂, 8 ♀ (SCAU, Nos 111514 – 22), Yixiang, Pu'er, Yunnan, 22 ° 42 ' N, 101 ° 09 ' E, altitude 1400 m, 13. v. 2012, tree trunks, HW Chen; 1 ♀ (SCAU, No. 111539), Tongbiguan, Yingjiang, Yunnan, altitude 1310 m, 19. v. 2016, tree trunks, J Huang; 2 ♀ (SCAU, Nos 111523, 24), Husa, Longchuan, Yunnan, 24 ° 25 ' 19 '' N, 97 ° 46 ' 38 '' E, altitude 1227 m, 20,21. viii. 2016, tree trunks, HW Chen, L Gong; 2 ♂, 5 ♀ (SCAU, Nos 111525 – 1), Bangda, Mangshi, Yunnan, 24 ° 7 ' 55 '' N, 98 ° 13 ' 42 '' E, altitude 1780 m, 8. xi. 2017, tree trunks, HW Chen, L Gong, BX Li; 1 ♀ (SCAU, No. 111532), Botanic Garden, Ruili, Yunnan, 24 ° 4 ' 19 '' N, 97 ° 49 ' 8 '' E, altitude 1174 m, 22. v. 2016, tussock, J Huang; 1 ♂ (SCAU, No. 111542), Mengdong, Cangyuan, Yunnan, 23 ° 10 ' 8 '' N, 99 ° 13 ' 52 '' E, altitude 1320 m, tussock, 6. V. 2016, J Huang.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB4FFC7C69C587EFD9FFB08.taxon	etymology	Etymology. A combination of the Latin words: flavus and vittatus, referring to the mesonotum with yellow longitudinal stripes.	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
03D787DCFFB4FFC7C69C587EFD9FFB08.taxon	distribution	Distribution. China (Yunnan).	en	Li, Bingxue, Zhang, Yuan, Chen, Hongwei (2019): Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae). Zootaxa 4576 (1): 81-108, DOI: 10.11646/zootaxa.4576.1.4
