taxonID	type	description	language	source
03D77E1287136B1CFF0EFC7D656EFEDD.taxon	materials_examined	Type species: Carabus emarginatus Gmelin, 1790 (= Carabus dentatus P. Rossi, 1790) = Desera Dejean, 1825 (nec Hope, 1831) Type species: Drypta longicollis MacLeay, 1835	en	Sciaky, Riccardo, Anichtchenko, Alexander (2020): Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530, DOI: 10.11646/zootaxa.4731.4.5
03D77E1287136B1CFF0EFC7D656EFEDD.taxon	description	Liang et al. (2004) have correctly pointed out the reasons for which Desera must be regarded as a synonym of Drypta and its long-lasting confusion with Dendrocellus, due to the misinterpretation of Drypta longicollis MacLeay.	en	Sciaky, Riccardo, Anichtchenko, Alexander (2020): Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530, DOI: 10.11646/zootaxa.4731.4.5
03D77E1287106B1CFF0EFEF2631FFB19.taxon	materials_examined	Type species: Dendrocellus discolor Schmidt-Goebel, 1846 (= Desera nepalensis Hope, 1831) = Desera Hope, 1831 (nec Dejean, 1825) Type species: Desera nepalensis Hope, 1831 Bousquet (2002) and Liang et al. (2004) have correctly reconstructed the complex nomenclatorial history of this genus, reaching the conclusion that its valid generic name is Dendrocellus Schmidt-Goebel, 1846, instead of Desera Dejean, 1825, as it was frequently considered previously. We only observe that back in 1949 Jeannel already wrote: “ Les Dendrocellus Schm. - Goeb. (type: discolor Schm. - Goeb., = nepalensis Hope) sont généralement classés à tort sous le nom de Desera. … Malgré leurs ongles tarsaux pectinés, ils sont très voisins des Drypta s. str. ” (Jeannel, 1949). So, it seems the french author had already reached the same conclusions. This genus, recently revised by Liang & Kavanaugh (2007), contains 22 species occuring in Africa, Asia and Australia. Systematically it is extremely close to Drypta, differing only in its tarsal claws pectinate instead of smooth. The genitalic characters of both sexes are the same, the external resemblance among members of the two genera is sometimes puzzling and a few species of Dendrocellus show very slight tarsal pectination, sometimes leaving a doubt on their generic pertinence; only a comparative study of all the characters of the species belonging to the two genera will allow to decide whether they can be really maintained as separate genera. Already in 1968 Darlington observed: “ Desera differs from Drypta only in having pectinate tarsal claws. A modern revision of the species is needed to show whether both genera are really monophyletic and distinct ” (Darlington, 1968, p. 218). The unique combination of characters distinguishing this genus from the others of the tribe is: pronotal bead absent or very rudimental (fig. 10); punctuation on head and pronotum dense, regular, the punctures usually well distinct from each other; pronotum very feebly constricted towards base; elytral microsculpture well developed; elytral pubescence dense, usually arranged in two-three more or less regular rows; scutellar pore constantly single; intervals flat or slightly convex; tarsal claws slender, more or less pectinate on inner side; two to five evident setae on outer side of stylomere (fig. 8).	en	Sciaky, Riccardo, Anichtchenko, Alexander (2020): Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530, DOI: 10.11646/zootaxa.4731.4.5
03D77E1287106B1CFF0EFAB661E4F81C.taxon	materials_examined	Type species: Drypta mandibularis Castelnau, 1897 This genus, very synthetically described by Jeannel (1949), is scarcely known, so that none of the few authors who have mentioned it has noticed its peculiar characters and its complete independence from the “ true ” Drypta, rather treating it as a simple subgenus of Drypta (Lorenz, 2005), or implicitly considering it a synonym of Drypta (Baehr, 2003, 2017). In our opinion it is a well distinguished genus, probably related to Nesiodrypta and forming with it and with Megadrypta, here described, a group of Gondwanian origin, The completely beaded sides of pronotum seem to us a more primitive character compared to the unbeaded sides of Drypta and Dendrocellus, indicating a more ancient group from which probably the other two genera have later evolved. It contains 8 species from South-East Asia (India, Myanmar, Indonesia, China, Laos), still inadequately known; a revision would be useful. The unique combination of characters distinguishing this genus is: pronotal bead well marked, with margin serrate (fig. 11); punctuation on head sparse, often leaving the frons smooth, on pronotum dense and irregular, with the punctures often confluent; pronotum evidently constricted towards base; elytral microsculpture almost absent, intervals smooth; elytral pubescence only aligned along the striae; single scutellar pore; intervals rather convex; tarsal claws thick, strongly curved, smooth on inner side; no setae, or a rudimental one, on outer side of stylomere (fig. 6).	en	Sciaky, Riccardo, Anichtchenko, Alexander (2020): Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530, DOI: 10.11646/zootaxa.4731.4.5
03D77E1287116B1DFF0EFAC26453F83B.taxon	materials_examined	Type species Megadrypta mirabilis sp. n. by monotypy. Etymology. Prefix “ Mega - “ refers to the unusual large size of the only known species, and Drypta to the nominotypical genus of the tribe. Diagnosis. This new genus can be easily distinguished from all the other known genera by the combination of: pronotal lateral bead well marked (fig. 12), tarsal claws smooth, completely reduced humera and hind wings, smooth stylomere (fig. 4) and presence of 7 to 8 periscutellar pores on elytra; pronotum very elongate, constricted near anterior margin and near base, with elevated median part along median line, antennae rather short, with antennomere 1 nearly as long as the sum of antennomeres 2 – 7 together. Systematic notes. The systematic position of this genus is in our opinion near the genera Nesiodrypta and Prionodrypta. These three genera, in fact, share two important characters, namely the lateral margins of pronotum completely beaded and the stylomere much longer and more curved, with at most one small setae on each side, beyond the claws completely smooth. The occurrence of these two characters seem to us to suggest a true relationship among these three genera. Jeannel alone (1949) had noticed the relationship between Nesiodrypta and Prionodrypta, not followed by any subsequent author. The discovery of Megadrypta is certainly unexpected, but this genus seems to fit into the group, possibly of Gondwanian origin, including the other two above-mentioned genera.	en	Sciaky, Riccardo, Anichtchenko, Alexander (2020): Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530, DOI: 10.11646/zootaxa.4731.4.5
03D77E1287116B1DFF0EFC606399FAC7.taxon	materials_examined	Type sp. Drypta costigera Chaudoir, 1860 This monobasic genus is the only known member of tribe Dryptini from the Neotropical region. The single species, known only from Brazil (“ Ega ”, today Tefé, in the state of Amazonas), is easily distinguishable by its unique combination of characters, i. e. elytra with strong, smooth costae, intervals wide and transversely striate; apex of elytra with external angle spiniform; claws smooth.	en	Sciaky, Riccardo, Anichtchenko, Alexander (2020): Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530, DOI: 10.11646/zootaxa.4731.4.5
03D77E1287116B1DFF0EFF0D60FCFC60.taxon	materials_examined	Type species: Drypta perrieri Fairmaire, 1834	en	Sciaky, Riccardo, Anichtchenko, Alexander (2020): Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530, DOI: 10.11646/zootaxa.4731.4.5
03D77E1287116B1DFF0EFF0D60FCFC60.taxon	description	This genus contains 18 species primarily occuring in Madagascar (15 species), but marginally also in continental Africa (3 species), with two species in East Africa (Kenya and Tanzania) and one in Gabon, recently described (Facchini & Czeppel, 2011). The unique combination of characters distinguishing this genus is: pronotal bead well marked, with sides not serrate (fig. 13); punctuation on head sparse, often leaving the frons smooth, on pronotum often limited to the lateral parts, the disc covered with transversal wrinkles; elytral microsculpture well developed; elytral pubescence limited to some small and sparse punctures, only in few cases some more regular punctures are aligned along the striae (except N. setigera, that shows a punctuation on head, pronotum and elytra almost as dense as in Prionodrypta; it is possible that future studies will show the necessity to isolate this species in a separate genus); scutellar pore single; intervals rather convex; tarsal claws thick, strongly curved, smooth on inner side; no setae, or a rudimental one, on outer side of stylomere (fig. 5).	en	Sciaky, Riccardo, Anichtchenko, Alexander (2020): Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530, DOI: 10.11646/zootaxa.4731.4.5
03D77E1287166B19FF0EFF756042FE6D.taxon	materials_examined	Holotype: male, China, Guangdong, Ruyuan, Naling Nature Reserve, 24.92579 º, 113.01638 º / 1021 m, 2007.6.18 – 21 Huang Hao Coll. in IZCS. We have added a red label: “ Holotypus Megadrypta mirabilis gen. & sp. n., des. Sciaky & Anichtchenko, 2019 ” Paratypes: 2 males (IZCS), China, Guangdong, Ruyuan, Naling Nature Reserve, 24.93450 º, 113.01588 º / 1266 m, 2008.7.20 D, Liang Hongbin Coll.; 1 female (IZCS, body teneral), China, Guangdong, Ruyuan, Naling, Xiaohuangshan, 24.89541, 113.02075 / 1318 m, 2008.7.18 D, Tang Guo, vegetation, Institute of Zoology; 1 female (IZCS), China, Guangdong, Ruyuan, Naling, Xiaohuangshan, 24.89541, 113.02075 / 1318 m, 2008.7.17 D, Tang Guo, vegetation, Institute of Zoology; 1 female (IZCS), China, Guangdong, Ruyuan, Naling, Xiaohuangshan, 24.89541, 113.02075 / 1318 m, 2008.7.18 D, Yang G. Y., vegetation, Institute of Zoology; 1 female (cRS), China — Guangdong, Nanling Park, 11. VII. 2011; 1 male (IZCS, left elytron missing) China, Guangxi Prov., Ziyuan, Mao-er Shan, Hongjunting / 2003.6.19, Wang Shaoneng coll.; 1 male (IZCS), China Guangxi Prov., Cewanglao Shan Nature Reserve / 2007.8.5 day, Liu Chunxiang coll. All paratypes have been labelled with a red label: “ Paratypus Megadrypta mirabilis gen. & sp. n., des. Sciaky & Anichtchenko, 2019 ” Description. Length: 14,5 – 16,0 mm (15,0 in the holotype). Head, pronotum and elytra black with very weak bluish luster; ventral side black; femora black, tibiae and tarsi dark brown; tarsal claws smooth; mouth parts and antennae dark brown, apical half of scape blackish brown (fig. 1). Ventral (fig. 2) and dorsal (fig. 1) side of body and femora covered with yellow pubescence of almost equal length. Head densely and coarsely punctate on vertex; labrum bisinuate, with medial lobe slightly protruded anteriorly; clypeus smooth; genae developed; eyes large, moderately convex, 1.88 times as long as genae. All palpi with last segment evidently dilated, securiform in the male (fig. 1), slender, feebly dilated at tip in the female (fig. 3). Microsculpture irregularly transverse. Pronotum (fig. 12) relatively long, widest in the middle, 1.62 times as long as wide; apical angles obtuse; lateral sides cariniform, not emarginated, widely sinuate before anterior and basal angles; basal angles obtuse; pronotal lateral bead well marked. Median line distinct. Sides of pronotum abruptly flattened, median part of pronotum elevated. Disc irregularly coarsely punctured, punctures sometimes confluent. Microsculpture irregularly transverse. Elytra wide, 1.5 times as long as wide; widest in apical third; with 7 to 8 periscutellar pores; 4 to 5 discal pores hardly distinguishable from punctures in interval 3; shoulders completely reduced, strongly dilated to apical third, then narrowed to apex; all striae deep and closely punctate, punctures separated by less of one diameter distance from each other; all intervals slightly convex, uniformly and fairly closely punctate; outer apical angles widely rounded, sutural angle subacute. Microsculpture irregularly isodiametric. Hind wings completely reduced. Pro-, meso- and metasterna, epipleurae and ventral sternites similarly punctate (fig. 2). Punctures large, separated by 1 – 2 diameters distance from each other.). Protarsi moderately dilated in the male (fig. 1), simple in the female; protarsomeres 2 and 3 strongly asymmetrical in males, symmetrical in females; tarsal claws smooth. Median lobe of aedeagus short and robust, 2,5 – 2,8 long (2,8 in the holotype). In lateral view inferior margin straight at base, in apical fourth slightly bent upwards. In dorsal view apex short and widely rounded (figs. 14 – 16). Female with stylomere smooth and strongly arched, without spines on either margin (fig. 4). Distribution. This species is known only from three localities in two different provinces of Southern China. The type locality, that is a national nature reserve established in 1994 and located in northern Guangdong Province, in the centralsouth part of the Nanling Mountains. It is a forest ecological protected area that for subtropical evergreen broadleaf forest and rare flora and fauna. The size of the Nanling National Nature Reserve is 584 km ² and it is a typical representative of the evergreen broadleaved forest in southeast China. It is regarded as being the largest national forest in southeastern China. Therefore the habitat of Megadrypta mirabilis should be considered protected and the species not in danger. The two other localities are in Guangxi, some hundreds kilometers from the type locality, but the specimens here collected do not show differences both in the external and in the aedeagal characters from those of the type locality. Etymology. From Latin “ mirabilis ” meaning “ worth of admiration ”. Ecology. Part of the type-series was collected it by sweeping vegetation and flowers by two of its collectors. Since this species has reduced hind wing, we can suppose that it was most probably living on vegetation. This is, at our knowledge, the first time that a Dryptini has these living habits, since more frequently they are found in marshes or near streams.	en	Sciaky, Riccardo, Anichtchenko, Alexander (2020): Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini). Zootaxa 4731 (4): 522-530, DOI: 10.11646/zootaxa.4731.4.5
