taxonID	type	description	language	source
03D487ADFFA9FF9CFF2AFC61EA3B176A.taxon	description	Only 16 species of Eustrophopsis, including E. laurae sp. nov., have their distributional pattern known, with the remaining species only being recorded for the general territory of Brazil (Lopes-Andrade et al. 2024). Eustrophopsis was first described by Champion, 1889, later becoming a senior synonym for Eustrophinus Seidlitz, 1898 (Nikitsky 1998; Pollock 2012). We present the bibliography which includes the studied species, synonyms, and their known distribution in Brazil and other countries.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFA9FF9CFF2AFB25E9E21685.taxon	distribution	Distr.: recorded for “ Amazones ” in the original description, it is unclear if it refers to the Brazilian state of Amazonas or the Amazon Rainforest.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFA9FF9CFF2AFA08EC0215B5.taxon	synonymic_list	Syn. Eustrophinus bimaculata Pic, 1912: 93.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFA9FF9CFF2AFA08EC0215B5.taxon	distribution	Distr.: Brazil — states of Espírito Santo, Rio de Janeiro, Santa Catarina, and São Paulo.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFA9FF9CFF2AF918EAC61484.taxon	distribution	Distr.: Brazil — states of Amazonas and Goiás.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFA8FF9DFF2AFF7DE972124E.taxon	distribution	Distr.: Brazil.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFA8FF9DFF2AFE40EA1B117E.taxon	distribution	Distr.: Brazil — state of Rio de Janeiro.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFA8FF9DFF2AFD50EA1B106E.taxon	distribution	Distr.: Brazil — state of Rio de Janeiro.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFA8FF9DFF2AFB30EAA1166C.taxon	distribution	Distr.: Brazil — states of Amazonas and Rio de Janeiro.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFA8FF9DFF2AFA27EAD1157B.taxon	distribution	Distr.: Brazil — states of Amazonas and Pará.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFA8FF9DFF2AF955E972146B.taxon	distribution	Distr.: Brazil.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAFFF9BFF2AF8D8EA051256.taxon	diagnosis	Diagnosis (adapted from Nikitsky (1998) and Pollock (2012 )): Eyes large, covering most of the head, deeply emarginated around antennal insertions, nearly contiguous dorsally. Antennomeres I – IV distinctly narrow, antennomeres V – X distinctly widened and flattened, with annular antennal sensilla around the entire circumference of antennomeres. Sexually dimorphic antennomeres, males with distinctive sensilla on antennomeres V – X. Prosternal process separating procoxa. Prothoracic episterna with distinct, transverse suture. Front legs separated by the prothoracic process. Outer faces of meso- and metatibiae with numerous oblique, comb-like ridges. Tarsal formula 5 - 5 - 4, with the first tarsomere being the longest, the other subequal in length; metatarsomere I subequal in length to the other four metatarsomeres.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAFFF9AFF2AFF7DEC961264.taxon	synonymic_list	Syn.: Eustrophus bombinus Champion, 1889: 76.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAFFF9AFF2AFF7DEC961264.taxon	distribution	Distr.: Brazil — states of Rio de Janeiro and São Paulo; Guatemala, México, Nicarágua, Panamá.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAFFF9AFF2AFE2FEA931194.taxon	distribution	Distr.: Brazil — state of Rio de Janeiro; Guatemala.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAFFF9AFF2AFD3FEAD51084.taxon	distribution	Distr.: Brazil — state of Rio de Janeiro; Peru.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAFFF9AFF2AFC0FE92217D6.taxon	synonymic_list	Syn.: Eustrophinus marginatus Seidlitz, 1898: 440	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAFFF9AFF2AFC0FE92217D6.taxon	distribution	Distr.: Brazil — Peru.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAFFF9AFF2AFAF8EA1916C5.taxon	distribution	Distr.: Brazil — state of Espírito Santo.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAFFF9AFF2AF9C8EA1915F5.taxon	distribution	Distr.: Brazil — state of Espírito Santo.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAEFF98FF2AFE6CEA24110E.taxon	description	(Figs 2, 4 A)	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAEFF98FF2AFE6CEA24110E.taxon	diagnosis	Diagnosis: Body length of nearly half a centimeter. Overall body brown to testaceous, covered by many fine setae. Eyes nearly contiguous dorsally; antennomeres I – IV and the tip of antennomere XI yellow, antennomeres V – X dark brown; clypeus inflated. Apex of the prosternal process bifurcated. The most noticeable feature is the two black rounded maculae in the upper portion of the elytra. Length: 5.71 – 6.30 mm (based on three individuals). Male aedeagus median lobe (Fig. 4 A): slender, overall shape oval, with apical segment slightly longer than basal segment, with distal apex spear-like, Studied specimens: 2 ♂, 1 ♀ BRAZIL, GO, Goianápolis, Parque Altamiro de Moura Pacheco-PEAMP; Malaise Mata — Trilha do Tamanduá - 16,553250 - 49,131511 — 880 m; 29. vi. 2022 - 02. vii. 2022; Cols. Oliveira, S. S. & Lopes, W. R. 1 ♂ and 1 ♀ housed in MZUSP, and 1 ♂ housed in ZUFG.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAEFF98FF2AFE6CEA24110E.taxon	discussion	Remarks: No photos or illustrations have previously been made for this species. This species appears very similar to E. bimaculatus (Pic, 1912) due to the two black elytral maculae. The main subtle difference is the position of the black maculae, the maculae in E. bimaculatus is placed in the median portion of the elytron, while that of E. bilunulata is placed in the upper portion of the elytron, which is pointed out in the original description. Another difference is the geographical occurrence, E. bilunulata was first described from Jataí, Goiás, within the Cerrado (Pic, 1912), while E. bimaculataus from Itatiaia, Rio de Janeiro, and there are also records for Espírito Santo and Santa Catarina, all within the Atlantic Forest area (Pic, 1912). Other more recent records are also known for E. bimaculatus in the same biome (iNaturalist 2024). Whether these are just geographical variations of the same species, or two truly distinct taxa remains to be seen, as a proper dissection of both male and female, as well as a complete type revision, would be required. Furthermore, E. bilunulata has a record for the Amazon Rainforest, in Tefé, Amazonas (Champion, 1916), but given the distance and dissimilarity of the environment, we are not sure if this is truly a record for E. bilunulata or another similar species.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFAEFF98FF2AFE6CEA24110E.taxon	distribution	Distribution: Brazil, Goiás, Goianópolis, Parque Altamiro de Moura Pacheco (this work), and Jataí (Pic, 1912). Brazil, Amazonas, Tefé (Champion, 1916).	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFADFF96FF2AFD81ECE1102E.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: pub: 81 A 58 D 39 - 1 C 56 - 421 F-B 988 - FCF 8088 A 94 FC (Figs 2, 3, 4 B)	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFADFF96FF2AFD81ECE1102E.taxon	diagnosis	Diagnosis: Body length of nearly one centimeter. Overall body honey colored, edges of pronotum and elytra with slightly lighter tone in dorsal view, pronotum with well-defined lighter edge and elytropleura also yellow. Body completely covered by many fine setae. Eyes large, nearly contiguous dorsally. Prosternal truncated, nearly reaching the apex of the procoxae. Length: 8.93 – 9.87 mm (based on three individuals). Head: testaceous. Eyes large, covering most of the head (ocular index 3,902), deeply emarginated around antennal insertions, nearly contiguous dorsally; antennomeres I – IV distinctly narrow, antennomeres V – X distinctly widened and ventrally flattened; with annular antennal sensilla around entire circumference of antennomeres; clypeus inflated, uniform punctuation; labrum rounded; region of the gena light orange. Prothorax: pronotum with testaceous disc and lighter margins; prosternum black to dark brown, outer margins of the pronotum light orange posterior margin of the pronotum bisinuated, median portion deeply sinuated, rounded; prothoracic episterna with distinct, transverse suture; prosternal process truncated, nearly reaching the apex of the procoxae. Prothoracic episterna each with distinct, transverse suture. Mesothorax and Metathorax: black to dark brown; mesosternum short and keeled; metasternum with well-marked median suture. Elytra: testaceous with lighter margins; elytropleura light orange, wider at the humeral region and tapering towards the apex; elytra with rows of punctures. Legs: all legs similar; metathoracic legs longer than meso- or prothoracic legs; outer faces of meso- and metatibiae with numerous oblique, comb-like ridges; tarsal formula 5 - 5 - 4, all tarsi cylindrical, with first tarsomere being the longest, the other subequal in length; metatarsomere I subequal in length to the other four metatarsomeres; all claws simple. Abdomen: ventrite I dark brown to black, median portion testaceous; ventrites II – IV curved, testaceous with inner margin dark brown, ventrite V testaceous. Female ovipositor (Fig. 4 B): membranous, gonocoxites parallels, inner portions with spike-like projection, with apex bearing many fine setae; apical portion of vulva tapered, sclerotized portion parallel, spike-like; stylus with rounded apex slightly wider than base, bearing fine setae at apex.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFADFF96FF2AFD81ECE1102E.taxon	materials_examined	Holotype material: ♀, BRAZIL, GO, Goianápolis, Parque Altamiro de Moura Pacheco-PEAMP; Malaise Mata — Trilha do Tamanduá - 16,553250 - 49,131511 — 880 m; 18. i. 2023 - 16. ii. 2023; Cols. Oliveira, S. S. & Lopes, W. R. (MZUSP). Paratype material: ♀, BRAZIL, GO, Goianápolis, Parque Altamiro de Moura Pacheco-PEAMP; Malaise Mata — Trilha do Tamanduá - 16,553250 - 49,131511 — 880 m; 12. xii. 2022 - 18. i. 2023; Cols. Oliveira, S. S. & Lopes, W. R; ♀ (MZUSP). ♀, Malaise Mata — Trilha da Onça - 16,534879 - 49,113812 — 790 m; 16. ii. 2023 - 21. iii. 2023; Cols. Oliveira, S. S. e equipe cols. (ZUFG).	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFADFF96FF2AFD81ECE1102E.taxon	etymology	Etymology: the specific epithet is a patronym honoring Dr. Laura Rocha Prado, for her important contributions to the taxonomy and systematics of Coleoptera.	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
03D487ADFFADFF96FF2AFD81ECE1102E.taxon	distribution	Distribution: Brazil, Goiás, Goianópolis, Parque Altamiro de Moura Pacheco (this work).	en	Begha, Bruno Piotrovski, Souza, Isabella Magalhães De, Oliveira, Sarah Siqueira (2024): Polypore fungus beetles (Coleoptera: Tetratomidae) in a remnant of semi-deciduous forest of the Brazilian Cerrado: checklist and the description of a new species of Eustrophopsis Champion, 1889. Zootaxa 5501 (4): 542-552, DOI: 10.11646/zootaxa.5501.4.4, URL: http://dx.doi.org/10.11646/zootaxa.5501.4.4
