taxonID	type	description	language	source
03D574268E5AFFB4FF1AFB28D837FD7F.taxon	materials_examined	Material examined Saudi Arabia: 1 ♂, 3 ♀♀, Khamis Mushayt, Wadi Bishah, 18 ° 20 ' 01 '' N, 42 ° 42 ' 13 '' E, 1990 m, 27. IV. 2011, leg. Sharaf (cAss); 1 ex., Wadi Aljora near Abadan, 17.29 ° N, 43.07 ° E, 465 m, 12. XI. 2012, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5AFFB4FF1AFB28D837FD7F.taxon	discussion	Comment: This species, the type species of the subgeneric name Phanerosphena PACE 1991, was previously known only from two localities in Yemen (PACE 1991) and originally described as Atheta (Phanerosphena) rougemonti. An examination of the above material, which represents the first record from Saudi Arabia, revealed that A. rougemonti is a close relative of Acrotona pellucida (FAUVEL 1878) and A. oxypodoides (BRUNDIN 1952). In his revisionary work, BRUNDIN (1952) attributed them to Acrotona THOMSON 1859, which in those days was regarded as a subgenus of Atheta THOMSON 1858, but today is treated as a distinct genus by most authors. Subsequently, A. pellucida and A. oxypodoides were assigned to the genus Paradilacra BERNHAUER 1909 (see SMETANA 2004) and then reinstated in Acrotona by ASSING (2010). Acrotona rougemonti, A. pellucida, and A. oxypodoides are distinguished from other Acrotona species particularly by a different morphology of the median lobe of the aedeagus, of the parameres, and by longer and more slender legs, suggesting that both taxa form separate lineages. On the other hand, a close relationship is indicated by similar external characters and a similar morphology of the spermatheca. In consequence, until the systematic affiliations have been thoroughly studied based on a phylogenetic approach, Phanerosphena is proposed to represent a subgenus of Acrotona. Other species that have been attributed to Phanerosphena are: Atheta perlaeta PACE 1986 (= A. tronqueti PACE 1988) (Himalaya, Oriental region), A. retroarmata PACE 1998 (Hong Kong), A. alboguttata BERNHAUER 1915 (Afrotropical region), A. tridentata (KRAATZ, 1859) (Oriental region), A. coprophila CAMERON 1950 (Oriental region), and A. spinosa SCHEERPELTZ 1962 (China, Thailand). However, at least A. retroarmata is unlikely to be correctly assigned to Phanerosphena, as can be inferred from the illustrations provided by PACE (1998).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E59FFB4FF1AFBFBDBF7FB7B.taxon	materials_examined	Material examined: Saudi Arabia: 2 exs., Tamnia Dam (Abha), 18.02 ° N, 42.76 ° E, 2320 m, 14. XI. 2012, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E59FFB4FF1AFBFBDBF7FB7B.taxon	discussion	Comment Atheta coriaria is a common Cosmopolitan species, but was previously unknown from Saudia Arabia (SMETANA 2004).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E59FFB4FF1AFD38DB2FFC30.taxon	materials_examined	Material examined: SaudiArabia: 2 ♂♂, Wadi Aljora near Abadan, 17.29 ° N, 43.07 ° E, 465 m, 12. XI. 2012, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E59FFB4FF1AFD38DB2FFC30.taxon	discussion	Comment Brachida africana was originally described from Tanzania and subsequently reported also from Kenya (PACE 1994). The aedeagus of the above males is practically identical to the drawing of the aedagus of one of the type specimens provided by PACE (1994). The observed slight differences are most likely explained by intraspecific variation. The above specimens represent the first records from Saudi Arabia and the Palaearctic region sensu SMETANA (2004).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E59FFB4FF1AFB33DD5DFA7D.taxon	materials_examined	Material examined: Saudi Arabia: 5 ♀♀, Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 740 m, 11. V. 2011, leg. Sharaf (cAss); 2 ♀♀, same data, but 19 ° 55 ' 47 '' N, 41 ° 26 ' 34 '' E, 23. IX. 2011, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E59FFB4FF1AFB33DD5DFA7D.taxon	discussion	Comment: The above specimens, all them unfortunately females, evidently represent an undescribed species of Peltodonia BERNHAUER 1936 closely related to P. bodemeyeri BERNHAUER 1936 from Turkey. Males would be desirable for a description.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E59FFB5FF1AFA3EDBADFE8A.taxon	materials_examined	Material examined: Saudi Arabia: 1 ♀, Khamis Mushayt, W Ben Hashbal, 18 ° 35 ' 41 '' N, 42 ° 39 ' 01 '' E, 1890 m, 26. IV. 2011, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E59FFB5FF1AFA3EDBADFE8A.taxon	discussion	Comment: Species of Piochardia HEYDEN 1870 are associated with ants of the genus Cataglyphis FÖRSTER. The previously known distribution of P. oberthuri was confined to North Africa from Algeria in the west to Libya in the east (ASSING 1999). The above specimen considerably expands the distribution towards the east. It was found in association with Cataglyphis holgerseni COLLINGWOOD & AGOSTI 1996 (det. Sharaf). The nest was built in sandy dry soil under a large rock.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E58FFB5FF1AFD6FDA34FCC9.taxon	materials_examined	Material examined: Saudi Arabia: 26 exs., Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 740 m, 15. V. 2011, leg. Sharaf (MNHUB, cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E58FFB5FF1AFD6FDA34FCC9.taxon	discussion	Comment: This species is widespread and common in North Africa and the Middle East. It had been recorded from Saudi Arabia by GUSAROV (1997).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E58FFB5FF1AFE6CDA18FD8C.taxon	materials_examined	Material examined: SaudiArabia: 1 ♂, Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 56 ' N, 41 ° 27 ' E, 740 m, 23. IX. 2011, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E58FFB5FF1AFE6CDA18FD8C.taxon	discussion	Comment: This widespread species has been recorded from the Oriental, the Afrotropical, and the Neotropical regions. In the West Palaearctic it was previously only once reported from Israel. For more information, a redescription, and illustrations of the external and male sexual characters see ASSING & FELDMANN (2012).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E58FFB5FF1AFB7DDA44FAB3.taxon	materials_examined	Material examined: Saudi Arabia: 3 exs., Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 740 m, 15. V. 2011, leg. Sharaf (cAss); 1 ex., same data, but 11. V. 2011 (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E58FFB5FF1AFB7DDA44FAB3.taxon	discussion	Comment: The vast distribution of this common species ranges from the Afrotropical region across the Palaearctic and Oriental regions to Australia. According to SMETANA (2004), it was previously unknown from Saudi Arabia.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E58FFB5FF1AFA7BD8B0F9D1.taxon	materials_examined	Material examined: Saudi Arabia: 1 ♀, Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 740 m, 11. V. 2011, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E58FFB5FF1AFA7BD8B0F9D1.taxon	discussion	Comment: According to SMETANA (2004), P. indicus is distributed from the Afrotropical to the Oriental region. It had been recorded from Saudi Arabia by COIFFAIT (1982) and GUSAROV (1997).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E58FFB5FF1AFCA2DBA3FBBA.taxon	materials_examined	Material examined: Saudi Arabia: 1 ♀, Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 740 m, 15. V. 2011, leg. Sharaf (cAss); 1 ♀, same data, but 11. V. 2011 (cAss); 1 ♀, same data, but 19 ° 55 ' 47 '' N, 41 ° 26 ' 34 '' E, 23. IX. 2011 (cAss); 1 ♀, Al Bahah, Shada Mt., 19 ° 50 ' 20 '' N, 41 ° 18 ' 42 '' E, 1620 m, 22. IX. 2011, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E58FFB5FF1AFCA2DBA3FBBA.taxon	discussion	Comment Schatzmayrina oxyclypea oxyclypea had previously been reported from the Afrotropical region, North Africa, and Yemen; for data and a distribution map see PUTHZ (2007). Two other subspecies are distributed in the Oriental region. The above specimens represent the first records from Saudi Arabia.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5FFFB2FF1AFF32DABDFE7A.taxon	materials_examined	Material examined: Saudi Arabia: 1 ♂, 1 ♀, Al Urdiyah, Wadi Gonouna, 19 ° 25 ' 46 ' N, 41 ° 36 ' 18 '' E, 350 m, 12. V. 2011, leg. Sharaf (cAss). Oman: 2 ♂♂, Dhofar, " DS ", 31. VIII. 1994, leg. Rihane (cAss); 1 ♂, Dhofar, Rouri, 13. X. 1004, leg. Rihane (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5FFFB2FF1AFF32DABDFE7A.taxon	discussion	Comment Astenus indicus was previously known from the Afrotropical region, Sicily, Egypt, the East Palaearctic, and the Australian region (SMETANA 2004). The above specimens from Saudi Arabia and Oman represent new country records.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5FFFB2FF1AFE3DDAAEFD57.taxon	materials_examined	Material examined: Saudi Arabia: 2 ♀♀, Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 740 m, 15. V. 2011, leg. Sharaf (cAss); 1 ♀, same data, but 11. V. 2011 (cAss); 1 ♀, Al Bahah, W Elzaraeb, 20 ° 04 ' 24 '' N, 41 ° 23 ' 12 '' E, 2090 m, 9. V. 2011, leg. Sharaf (cAss); 1 ♀, Al Urdiyah, Wadi Gonouna, 19 ° 25 ' 46 ' N, 41 ° 36 ' 18 '' E, 350 m, 12. V. 2011, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5FFFB2FF1AFE3DDAAEFD57.taxon	discussion	Comment: The distribution of A. nigromaculatus ranges from the Canary Islands and the Iberian Peninsula across North Africa to the Middle East (SMETANA 2004). The species had already been recorded from Saudi Arabia by COIFFAIT (1981, 1982).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5FFFB2FF1AFB15DA27F9C6.taxon	materials_examined	Material examined: Saudi Arabia: 2 exs., Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 740 m, 11. V. 2011, leg. Sharaf; 2 exs., same data, but 15. V. 2011.; 1 ex., same data, but 19 ° 55 ' 47 '' N, 41 ° 26 ' 34 '' E, 20. IX. 2011, leg. Sharaf; 1 ex., Al Bahah, W Turabah, Al Mandaq, 20 ° 12 ' 40 '' N, 41 ° 17 ' 18 '' E, 1790 m, 10. V. 2011, leg. Sharaf; 1 ex., Khamis Mushayt, Wadi Bishah, 18 ° 20 ' 01 '' N, 42 ° 42 ' 13 '' E, 1990 m, 27. IV. 2011, leg. Sharaf (material in MNHUB and cAss); 3 exs., Tamnia Dam (Abha), 18.02 ° N, 42.76 ° E, 2320 m, 14. XI. 2012, leg. Sharaf (cAss); 1 ex., Wadi Shahdan (Jizan), 17.45 ° N, 42.72 ° E, 200 m, 13. XI. 2012, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5FFFB2FF1AFB15DA27F9C6.taxon	discussion	Comment: The vast distribution of G. maritimus includes the Afrotropical, Palaearctic, and Oriental regions (SMETANA 2004). The species had already been reported from Saudi Arabia by COIFFAIT (1981) and GUSAROV (1997).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5FFFB2FF1AFD27D992FC48.taxon	materials_examined	Material examined: Saudi Arabia: 2 ♂♂, Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 740 m, 11. V. 2011, leg. Sharaf (cAss); 1 ♀, Wadi Aljora near Abadan, 17.29 ° N, 43.07 ° E, 465 m, 12. XI. 2012, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5FFFB2FF1AFD27D992FC48.taxon	discussion	Comment: The previously known distribution of H. galilaeus was confined to Israel and Lebanon (ASSING 2008 b). The above material represents the first records from Saudi Arabia.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5FFFB2FF1AFC22DBBBFB25.taxon	materials_examined	Material examined: Saudi Arabia: 1 ♂, Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 740 m, 11. V. 2011, leg. Sharaf (cAss); 1 ♀, Al Atawla (Baha-Taif Rd), Wadi Bawah, 20 ° 45 ' N, 41 ° 15 ' E, 1310 m, 8. XI. 2012, leg. Sharaf (cAss); 1 ♂, Wadi Shahdan (Jizan), 17.45 ° N, 42.72 ° E, 200 m, 13. XI. 2012, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5FFFB2FF1AFC22DBBBFB25.taxon	discussion	Comment: This species was originally described from Chad (Ennedi) and subsequently reported also from the Middle East (ASSING 2007, SMETANA 2004), including Saudi Arabia (COIFFAIT 1979, 1981, GUSAROV 1997).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5EFFB3FF1AFE55DD74FD78.taxon	materials_examined	Material examined: Saudi Arabia: 1 ♂, 2 ♀♀, Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 740 m, 11. V. 2011, leg. Sharaf (cAss); 1 ♂, Al Bahah, Shada Mt., 19 ° 50 ' 20 '' N, 41 ° 18 ' 42 '' E, 1620 m, 22. IX. 2011, leg. Sharaf.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5EFFB3FF1AFE55DD74FD78.taxon	discussion	Comment Philonthus bisignatus is widespread in the Afrotropical region. It had been unknown from the Palaearctic region sensu SMETANA (2004) until it was recently recorded from Yemen for the first time (ASSING 2012). The above specimens represent the first records from Saudi Arabia and the second record from the Palaearctic region.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5EFFB3FF1AFF32DA66FE62.taxon	materials_examined	Material examined Saudi Arabia: 1 ♂, Khamis Mushayt, Wadi Bishah, 18 ° 20 ' 01 '' N, 42 ° 42 ' 13 '' E, 1990 m, 27. IV. 2011, leg. Sharaf (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5EFFB3FF1AFF32DA66FE62.taxon	discussion	Comment Philonthus turbidus is widespread in the southern Palaearctic, the Afrotropical, Oriental, and Nearctic regions (SMETANA 2004). It had been reported from Saudi Arabia by COIFFAIT (1979, 1981) and GUSAROV (1997).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5EFFBEFF1AFCCADA40FDD5.taxon	description	(Figs 1 - 5, Map 1)	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5EFFBEFF1AFCCADA40FDD5.taxon	materials_examined	Type material: Holotype ♂: “ SAUDI ARABIA 740 m, Al Bahah, Al Mukhwah Dhi Ayn Arch. vill., 19 ° 55 ' 46 '' N, 41 ° 26 ' 30 '' E, 11. V. 2011, leg. Sharaf / Holotypus ♂ Pinophilinus spinosus sp. n., det. V. Assing 2012 ” (KSMA). Paratype ♀: “ Saudi Arabia, Al Bahah, Shada Mnt., 22. IX. 2011, 19 ° 50 ' 19.7 '' N, 41 ° 18 ' 42.2 '' E, 1620 m, Leg. M. R. Sharaf 0036 ” (cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5EFFBEFF1AFCCADA40FDD5.taxon	etymology	Etymology: The specific epithet (Latin, adjective) refers to the presence of a large spine in the internal sac of the aedeagus.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5EFFBEFF1AFCCADA40FDD5.taxon	description	Description: Body length 5.8 - 6.5 mm; length of forebody 2.2 - 2.4 mm. Habitus as in Fig. 1. Coloration: whole body pale-reddish; legs and antennae yellow. Head (Fig. 2) strongly transverse, approximately 1.4 times as wide as long; posterior margin concave in the middle; frons practically impunctate; punctation of vertex dense, somewhat sparser in posterior median portion, rather coarse, and defined; interstices glossy, without microsculpture, on average as broad as, or slightly narrower than diameter of punctures, somewhat broader in posterior median portion. Eyes weakly convex and rather large, distinctly longer than postocular region in dorsal view. Antenna approximately 1.4 mm long; antennomere III approximately twice as long as broad, IV-V approximately 1.5 times as long as broad, VI and VII weakly oblong, VIII approximately as long as broad, and IX and X weakly transverse. Pronotum (Fig. 2) indistinctly oblong, at anterior angles approximately as wide as head, and distinctly tapering posteriad in posterior half; posterior angles abruptly rounded, moderately marked; punctation similar to that of head; interstices without microsculpture, on average somewhat narrower than diameter of punctures. Elytra (Fig. 2) short and narrow, approximately 0.75 times as long and 0.81 - 0.82 times as broad as pronotum, slightly widened posteriad; humeral angles almost obsolete; punctures similar to those of pronotum, or slightly larger; interstices without microsculpture. Hind wings completely reduced. Metatarsomere I as long as the combined length of II-IV. Abdomen (Fig. 3) long and slender, broader than elytra; maximal width at segment VII; punctation of tergites III-V coarse and dense, that of tergites VII and VIII rather fine and much sparser, that of tergite VI intermediate; interstices without microsculpture; posterior margin of tergite VII without palisade fringe. ♂: sternite VIII oblong, distinctly tapering posteriad, and with V-shaped posterior excision (Fig. 4); aedeagus approximately 1.2 mm long, with slightly asymmetric and in the middle pointed ventral process (ventral view) and with large sclerotized spine in the internal sac (Fig. 5).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5EFFBEFF1AFCCADA40FDD5.taxon	discussion	Comparative notes: Two Pinophilinus species were previously known from the Arabian region, P. schatzmayri (KOCH 1934) from the south of the Sinai Peninsula and P. socotranus ASSING 2012 from Socotra Island, Yemen. Pinophilinus spinosus is distinguished from these species as follows: from P. schatzmayri (male unknown) by much smaller size (P. schatzmayri: length of forebody 3.4 mm) (compare Figs 1 and 6), much shorter antennae (P. schatzmayri: antennae 2.1 mm long; antennomere III approximately three times as long as broad, IV-VIII distinctly oblong), larger and less convex eyes, finer setation of the head and pronotum (P. schatzmayri: head and pronotum with long and dark lateral setae), the posteriorly more distinctly narrowed pronotum (P. schatzmayri: lateral margins in anterior two thirds of pronotum almost parallel), and the less coarse punctation of the abdomen; from P. socotranus by much paler coloration (P. socotranus: body dark-brown), denser punctation of the whole body, distinctly longer antennae with more slender antennomeres (P. socotranus: antenna approximately 1.2 mm long; antennomere III approximately 1.5 times as long as broad, IV-VII weakly oblong), much larger eyes (P. socotranus: shorter than postocular region in dorsal view), the more convex (cross-section) and posteriorly less strongly narrowed pronotum, the longer, more slender, and less convex elytra, the larger and deeper posterior excision of the male sternite VIII, and by the completely different morphology of the aedeagus. For illustrations of P. schatzmayri and P. socotranus see Figs 6 - 8 and ASSING (2012), respectively.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E5EFFBEFF1AFCCADA40FDD5.taxon	distribution	Distribution and natural history The type specimens were collected in two localities in Al Bahah, southwestern Saudi Arabia (Map 1), at altitudes of 740 and 1620 m in May and September. The reduced hind wings, the short elytra, and the absence of a palisade fringe at the posterior margin of tergite VII suggest that the distribution of P. spinosus is restricted. The holotype was collected from leaf litter unter an old Ficus tree, the paratype from soil under a large plastic bag.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E53FFBEFF1AFDA5D8FEFC37.taxon	description	(Figs 6 - 8, Map 1)	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E53FFBEFF1AFDA5D8FEFC37.taxon	materials_examined	Type material examined: Syntype ♀: " Sinai W. Isla, 28.2.1933, A. Schatzmayr / Syntypus ♀ Pinophilus schatzmayri Koch, rev. V. Assing 2012 / Pinophilinus schatzmayri (Koch), det. V. Assing 2012 " (NHMB).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E53FFBEFF1AFDA5D8FEFC37.taxon	discussion	Comment: The original description is based on two female syntypes from " Wadi Isla, 28.2.1933 (Süd-Sinai) " (KOCH 1934). One of the type specimens was located in the Frey collection, which is currently housed in the NHMB. This species is characterized particularly by its large size (total length 7.5 mm; length of forebody 3.4 mm), palereddish coloration, long and slender antennae (2.1 mm) with distinctly oblong antennomeres I-IX, and by the dense and coarse punctation of the whole body, particularly of the abdomen. The habitus, the forebody, and the abdomen of the examined syntype are illustrated in Figs 6 - 8.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E52FFBDFF1AFE57D98CFEDF.taxon	description	(Figs 9 - 12)	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E52FFBDFF1AFE57D98CFEDF.taxon	materials_examined	Type material: Holotype ♂: “ SAUDI ARABIA 740 m, Al Bahah, W Turabah, Al Mandaq, 20 ° 12 ' 40 '' N, 41 ° 17 ' 18 '' E, 1790 m, 10. V. 2011, leg. Sharaf / Holotypus Tachyporus saudicus sp. n., det. M. Schülke 2012 ” (KSMA). Paratypes: 3 ♂♂, 3 ♀♀ [1 ♂, 3 ♀♀ teneral]: same data as holotype (cSch, cAss).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E52FFBDFF1AFE57D98CFEDF.taxon	etymology	Etymology: The specific epithet (Latin, adjective) is derived from Saudi Arabia where the species was discovered.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E52FFBDFF1AFE57D98CFEDF.taxon	description	Description: Measurements (in mm) and indices: body length 2.7 - 3.4; length of forebody 1.7 - 1.9; head width 0.59 - 0.60; pronotal width 0.96 - 1.00; pronotal length 0.64 - 0.66; elytral sutural length (apex of scutellum to sutural angle) 0.77 - 0.79; elytral humeral length (anterior to posterior margin) 0.96 - 1.01; elytral width 1.03 - 1.10; aedeagus length 0.69 - 0.72. Ratios: head width / pronotal width 0.60 - 0.62; pronotal length / elytral length 0.77 - 0.83; elytral humeral length / elytral width 0.90 - 0.97. Habitus as in Fig. 9. Coloration: head black, pronotum and elytra yellowish-brown, disc of pronotum dark-brown, only one specimen with oblique brown elytral markings, abdomen dark-brown to black, posterior margins of segments yellowish-brown, legs and basal antennomeres yellow, antennomeres IV-XI and maxillary palpus brownish. Head transverse; posterior part concealed under pronotal anterior margin; punctation of frons and vertex fine and sparse, distance between punctures approximately 30 µm; without microsculpture. Eyes weakly convex and rather large, distinctly longer than postocular region in dorsal view. Antenna approximately 1.1 mm long; antennomeres I-III more than twice as long as broad, antennomeres IV-X of decreasing length, antennomere X as long as broad, antennomere XI slightly elongated, 1.7 times as long as broad. Antennomeres I-III only with sensory bristles, antennomeres IV-XI with dense pubescence. Pronotum distinctly transverse, 1.6 - 1.7 times as wide as head; anterior angles bent lateroventral, posterior angles broadly rounded; punctation similar to that of head, pubescence almost invisible, only 1 - 2 µm long; microsculpture absent. Anterior, posterior, and lateral margins each with four distinct bristle-bearing punctures, bristles approximately 100 µm long. Elytra moderately long, 1.2 - 1.3 times as long as pronotum, slightly widened posteriad, 0.90 - 0.97 times as long as wide; punctures as sparse as those of pronotum, but distinctly coarser with pubescence of approximately 50 µm length. Elytra covered with rows of sensory bristles of approximately 100 µm length forming distinct rows: humeral 0 - 1, sublateral 1 - 2, inner discal 2, middle discal 3, outer discal 2, sutural 3 - 4, apical 3 and lateral 5 - 6; surface of elytra without any microsculpture. Hind wings fully developed. Abdomen with punctation and pubescence of tergites similar to those of pronotum; tergites with indistinct microsculpture composed of transverse striae or meshes (2 meshes per 10 µm); posterior margin of tergite VII with distinct palisade fringe. ♂: protarsomeres I-IV distinctly dilated; sternite VIII with distinct posterior emargination (Fig. 11); aedeagus approximately 0.7 mm long, without large sclerotized structures in the internal sac (Fig. 10).	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E52FFBDFF1AFE57D98CFEDF.taxon	discussion	Comparative notes No records of Tachyporus species were previously known from the Arabian Peninsula. Only few species have been recorded from the adjacent regions, such as T. nitidulus (FABRICIUS 1781) from Egypt, Lebanon, Syria, Iran, Israel and Iraq, T. abner SAULCY 1865 from Israel, Jordan and Iran, T. caucasicus KOLENATI 1846 from Israel, Jordan, Syria and Iran, T. hypnorum (FABRICIUS 1775) from Lebanon and Iran, and T. pusillus GRAVENHORST 1806 from Egypt, Israel, Syria and Iran. The record of T. solutus ERICHSON 1839 from Syria (SMETANA 2004) remains doubtful, most likely it is based on a confusion with T. caucasicus. Tachyporus saudicus is highly similar in size, habitus, and elytral chaetotaxy to T. abner and T. pusillus, but distinguished as follows: from T. abner by the much paler coloration of the whole body, especially of the elytra (with characteristic black markings in T. abner), variability of the presence of humeral bristle-bearing punctures on the elytra, and the larger size of the aedeagus (T. abner: 0.58 - 0.63 mm). from T. pusillus by usually paler coloration (T. pusillus: body dark-brown), the variability of the presence of humeral bristle-bearing punctures on the elytra, the absence of elytral microsculpture, and the smaller aedeagus (T. pusillus: 0.79 - 0.85 mm). Other pale-coloured species with similar oblique elytral markings (T. himalayicus BERNHAUER 1920, T. montanus BERNHAUER, 1915, and other probably undescribed species) are distinguished by the reduced number of elytral sensory bristles, especially the reduced number of discal bristles.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
03D574268E52FFBDFF1AFE57D98CFEDF.taxon	distribution	Distribution and natural history The type specimens were collected in a pasture, probably from under stones, in Wadi Turabah, Al Bahah, southwestern Saudi Arabia, at an altitude of 1790 m in May. The type locality is illustrated in Fig. 12.	en	Assing, V., Schülke, M., M. R, A. S (2013): On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera). Linzer biologische Beiträge 45 (1): 141-154, DOI: 10.5281/zenodo.5341014
