identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D28781FFF5FFDB5AA663BCFC26FD33.text	03D28781FFF5FFDB5AA663BCFC26FD33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala eucoma Bates 1888	<div><p>Anomala eucoma Bates, 1888</p><p>(Fig. 2)</p><p>Material examined: Lectotype (here designated): 1ɞ "Syn-type / Type / Las Mercedes, 3000 ft. Champion. / Anomala eucoma Bates (handwritten) / B.C.A., Col., II(2) Anomala " (BMNH). Paralectotypes: 1ɞ "Las Mercedes, 3000 ft. Champion. / Anomala eucoma Bates (handwritten) / H.W. Bates Biol. Cent. Amer." (MNHN). ɞ "Las Mercedes, 3000 ft. Champion. / eucoma Bates (handwritten) / H.W. Bates Biol. Cent. Amer." (MNHN). 1ɞ "SYN- TYPE / Costa Rica. V.P. (handwritten) / Locality doubtful (handwritten) / B. C. A., Col. II (2) Anomala " (BMNH); 1ɞ "SYN-TYPE / Costa Rica / Salle Coll. / 1195 / Anomala eucoma Bates (handwritten) / Isonychus sp. apud Sallé (handwritten) / B. C. A., Col. II (2) Anomala " (BMNH). 1ɞ "Pantaleon, 1700 ft. Champion / Anomala eucoma (handwritten) / B. C. A., Col. II (2) Anomala ". 1Ƥ "San Isidro, 1600 ft. Champion / Anomala eucoma (handwritten) / B. C. A., Col. II (2) Anomala ".</p><p>Additional material: 1ɞ " MUSEO DE INSECTOS UNIVERSIDAD DE COSTA RICA, COSTA RICA, PROV. (printed) San José U.C.R. 12–XI–64. G. Fuentes (handwritten) / Anomala ? id: Sullivan 86" (MUCR); 1ɞ " MUSEO DE INSECTOS UNIVERSIDAD DE COSTA RICA, COSTA RICA, PROV. (printed) San José Univ. de Costa Rica San Pedro, San José. Alt. 1200 m. 10–oct 1968. P.L. Kazan (handwritten) / Anomala sp. Id: D. Corde 1994" (MUCR); 1ɞ "Amubri, A.C. Amistad, Prov. Limon, COSTA RICA. 70 m. 4–30 Abr 1994, G. Gallardo, L S 385500_578100 # 2822 / COSTA RICA INBIOCRI001785409"; 1Ƥ "Amubri, Prov. Limón, COSTA RICA. 70m. 1–22 JUN 1995. G. Gallardo, L S 385000 578100 #5333 / COSTA RICA INBIOCRI002235510"; 1ɞ "Amubri, Talamanca., A.C. Amistad, Prov. Limón, COSTA RICA. 70 m. 6–28 Jun 1994, G. Gallardo, L N 385000_578100 #3006 / COSTA RICA INBIOCRI001875336"; 1Ƥ "Amubri, A.C. Amistad, Prov. Limon, COSTA RICA. 70 m. 2–31 May 1994, G. Gallardo, L N 385000_578100 # 2928 / CEUA 00105270"; 1ɞ "Amubri, Prov. Limón, COSTA RICA. 70m. 1–22 Oct 1994. G. Gallardo, L S 385500 578000 #3266 / COSTA RICA INBIOCRI002008852"; 1ɞ "Amubri, Prov. Limón, COSTA RICA. 70m. 3–9 SET 1994. G.M. Gallardo, L S 385500 578000 #3201 / COSTA RICA INBIOCRI002018611"; 1ɞ "Amubri, A.C. Amistad, Prov. Limon, COSTA RICA. 70 m. 4–21 Dec 1993, G. Gallardo, L S 385500_578100 # 2480 / CEUA 00105271"; 1ɞ "Est. Sirena, P. N. Corcovado, 0 – 100m, Prov. Punt., COSTA RICA. N. Obando, Jun 1990, L- S 270500 _508300 / COSTA RICA INBIOCRI000644524"; 1ɞ "Est Sirena, Corcovado N. P., Puntarenas Prov. COSTA RICA, 0–100m, Jan. 1990, G. Fonseca, L_S_ 270500 _508300 / COSTA RICA INBIOCRI000198374"; 1ɞ "Sirena, Corcovado N. P. Puntarenas Province COSTA RICA. 0 – 100m. G. Fonseca, Dic 1989, L- S 270500 _508300 / COSTA RICA INBIOCRI000203726"; 1ɞ "Est. Sirena, P. N. Corcovado, 0–100m, Prov. Punt., COSTA RICA F. Quesada, Jun 1990, L- S 270500 _508300 / COSTA RICA INBIOCRI000376353"; 1ɞ "Est. Sirena, P. N. Corcovado, 0 –100m, Prov. Punt., COSTA RICA, G. Fonseca, Oct 1989, L- S 270500 _508300 / COSTA RICA INBIOCRI000446957"; 1ɞ "Est. Sirena, 0–100m, P. N. Corcovado, Prov. Puntarenas, Costa Rica, G. Rodriguez, Abr 1992, L- S 270500 _508300 / CEUA 00105265"; 1Ƥ "Est. Sirena, 0–100m, P. N. Corcovado, Prov. Punt., COSTA RICA, G. Fonseca, Ene 1992, L- S 270500 _508300 / COSTA RICA INBIOCRI000506778"; 1ɞ "Est. Sirena, P. N. Corcovado, 0–100m, Prov. Punt., COSTA RICA. N. Obando, Jun 1990, L- S 270500 _508300 / COSTA RICA INBIOCRI000644521"; 1Ƥ "Est. Sirena, 0–100m, P. N. Corcovado, Prov. Puntarenas, Costa Rica, Jun 1992, G. Fonseca, L -S 270500 _508300 / CEUA 00105264"; 1ɞ "Est. Sirena, 0–100m, P. N. Corcovado, Prov. Puntarenas, Costa Rica, Jun 1992, G. Fonseca, L -S 270500 _508300 / COSTA RICA INBIOCRI000714400"; 1Ƥ " COSTA RICA. Prov. Puntarenas. P.N. Corcovado, Est. Sirena. 0– 100m. 21 FEB 2004. E. Holzer. C. Libre. L_S_ 270850 _509176 #92946 / INB0004129881 INBIOCRI COSTA RICA "; 1Ƥ " COSTA RICA. Prov. Puntarenas. P.N. Corcovado, Est. Sirena. 0– 100m. 21 FEB 2004. E. Holzer. C. Libre. L_S_ 270850 _509176 #92946 / INB0004129887 INBIOCRI COSTA RICA "; 1ɞ " COSTA RICA. Prov. Puntarenas. P.N. Corcovado, Est. Sirena. 0– 100m. 21 FEB 2004. E. Holzer. C. Libre. L_S_ 270850 _509176 #92946 / INB0004129886 INBIOCRI COSTA RICA "; 1ɞ " COSTA RICA. Prov. Puntarenas. P.N. Corcovado, Est. Sirena. 0– 100m. 21 FEB 2004. E. Holzer. C. Libre. L_S_ 270850 _509176 #92946 / INB0004129883 INBIOCRI COSTA RICA "; 1 ɞ "Est. Palo Verde, 10m, Ref. Nac. Fauna Silv. R. L. Rodriguez, Prov. Guan., COSTA RICA, D. Acevedo, Jun 1991, L- N 259000_388400 / COSTA RICA INBIOCRI000652257"; 1ɞ "Est. Palo Verde, 10m, Ref. Nac. Fauna Silv. R. L. Rodriguez, Prov. Guan., COSTA RICA, D. Acevedo, Jun 1991, L- N 259000_388400 / COSTA RICA INBIOCRI000652258"; 1ɞ "Est. Las Pailas, 800m, P. N. Rincon de la Vieja, Prov. Guanacaste, Costa Rica, 1 a 22 jul 1992, D. Garcia, L- N 306300_388600 / COSTA RICA INBIOCRI000713719"; 1ɞ "Fca. Jenny, 30 km N de Liberia, P. N. Guanacaste, Prov. Guanacaste, Costa Rica, 20 ago a 12 set 1992, E. Araya, L N 316200_364400 / COSTA RICA INBIOCRI000849675"; 1ɞ "Est. Las Pailas, 800 m, P.N. Rincon de la Vieja, Prov. Guan. COSTA RICA. 15 May–11 Jun 1993, K. E. Taylor, L- N 306300_388600 / CEUA 00105267"; 1ɞ "Finca Jenny, 30 km N de Liberia, P.N. Guanacaste, Prov. Guana, COSTA RICA. 300m. Mar 1991. R. Espinoza, L N 316200_364400 # 1678 / COSTA RICA INBIOCRI001693034"; 1ɞ "Fca. Jenny, 300m, 31 Km N. Liberia, Guanacaste Prov. COSTA RICA, Nov 1988 GNP Biodiversity Survey W85 34 27 ", N10 51 55 " / COSTA RICA INBIOCRI002517326"; 1ɞ "Estac. Maritza, 600 m, W side Volcan Orosi Guanac. Pr. COSTA RICA. June 1988. Janzen &amp; Hallwachs W85 29`37". N10 57`39" / COSTA RICA INBIOCRI002517343"; 1ɞ "Estac. Maritza, 600 m, W side Volcan Orosi Guanac. Pr. COSTA RICA. June 1988. Janzen &amp; Hallwachs W85 29`37". N10 57`39" / COSTA RICA INBIOCRI002517348"; 1Ƥ "Tierras Morenas, 700m, Prov. Guan., COSTA RICA. May 1993. G. Rodriguez. L-N-283950, 424500 / COSTA RICA INBIOCRI001180822"; 1Ƥ "Tierras Morenas, 700m, Prov. Guan., COSTA RICA. May 1993. G. Rodriguez. L-N-283950, 424500 / COSTA RICA INBIOCRI001180834"; 1ɞ "Est. Las Pailas, P.N. Rincón de la Vieja, A.C. Guanacaste, Prov. Guana. COSTA RICA. 800 m. 8–26 May 1994. K.E. Taylor, L N 306300_388600 #2912 / COSTA RICA INBIOCRI001861399"; 1ɞ "Est. Las Pailas, P.N. Rincón de la Vieja, A.C. Guanacaste, Prov. Guana. COSTA RICA. 800 m. 8–26 May 1994. K.E. Taylor, L N 306300_388600 #2912 / COSTA RICA INBIOCRI001861400"; 1Ƥ "Est. Las Pailas, P.N. Rincón de la Vieja, A.C. Guanacaste, Prov. Guana. COSTA RICA. 800 m. 8–26 May 1994. K.E. Taylor, L N 306300_388600 #2912 / COSTA RICA INBIOCRI001861401"; 1Ƥ "Est. Las Pailas, P.N. Rincón de la Vieja, A.C. Guanacaste, Prov. Guana. COSTA RICA. 800 m. 7–26 May 1994. D. García, L N 306300_388600 #2910 / COSTA RICA INBIOCRI001879319"; 1Ƥ "Est. Las Pailas, P.N. Rincón de la Vieja, Prov. Guanacaste, Costa Rica. 800 m. 19 Jun– 1 Jul 1993. D. G. García, L N 306300_388600 #2189 / COSTA RICA INBIOCRI001967801"; 1Ƥ "Est. Las Pailas, P.N. Rincón de la Vieja, Prov. Guana. COSTA RICA. 800 m. 16–24 Ago 1993. D. García, L N 306300_388600 #2268 / CEUA 00105266".</p><p>Description. Male. Body shape oval. Length 12.15 mm. Width 6.90 mm. Head, pronotum, scutellum, and pygidium dark brown; legs and underside reddish brown; elytra dark brown with two slightly lighter horizontal bands (along anterior margin and on elytral disc). Surface covered with fine, blond setae.</p><p>Clypeus trapezoidal, densely punctate-reticulate; ratio width/length 1.91. Anterior angles rounded. Frons densely punctate, flat, with apparent boundary with vertex (due to an abrupt change in inclination of the two surfaces) (Fig. 6 a). Ocular canthum long and thin, with rounded apex. Interocular ratio (interocular width/width of eye): 3.15.</p><p>Pronotum trapezoidal, width 1.57 times length. Lateral margins forming an obtuse angle at 1/3 of pronotum length. Anterior angles right and sharp, posterior angles obtuse and rounded. Basal margin thin and complete, sinuate. Entire surface with deep punctures.</p><p>Scutellum sub-pentagonal in shape, with rounded sides and acute apex, ratio width/length 1.26; sparsely punctate.</p><p>Elytra with defined costae and interstices with sparse, large punctures, somewhat obliterated by the numerous and smaller setae-bearing punctures. Lateral margin convex, larger at base, disappearing on apex. Marginal membrane almost complete.</p><p>Pygidium finely granulate, with longer setae at hind margins. Triangular in shape. Slightly convex in lateral view. Ratio width/length 1.58.</p><p>Metasternal disc sulcated, with setigerous punctures on the entire surface. Space between the mesocoxae narrow, slightly convex, width 0.25 mm. Mesometasternal suture well defined, at base of mesocoxae.</p><p>Abdominal sternites with 5–8 irregular transverse rows of setae, concentrated on bottom on sternites 2–4. Surface with elongate punctures that coalesce at sides. Male last sternite strigate, with apical margin well defined and rounded.</p><p>Protibia with 2 teeth (Fig. 7), apical tooth long and curved; second tooth at the same level of internal apex of protibia, triangular in shape, acute. Metatibia stout, slightly narrower subapically. Ratio length/width 2.93. First carina somewhat obliterated by elongate and dense punctures, which become rugose below second carina.</p><p>Protarsal claws: external claw strongly curved; ratio length/height: 2.01; internal claw bifurcate, with upper branch slightly shorter and half the width of the lower one. Inferior margin slightly sinuate.</p><p>Aedeagus (Figs 11–13): parameres short and wide, with inferior margin slightly sinuate. Endophallus (Fig. 26) with one long sacculus with sclerotized setae in a wide patch on apex; a ring of short but thick setae is present subapically. The rest of surface is covered by microsetation. A smaller diverticle is present at base. Gonopore opening ventrally at basal third.</p><p>Female (Costa Rica): Similar to males. Apical tooth of protibia longer and wider than in male. Inferior margin of internal protarsal claw straight. Metatibia wider than in males. Last abdominal sternite with curved margin.</p><p>Variation: Body length 8.44–12.62 mm, body width 4.87–6.93 mm. Clypeus w/l: 1.75–2.05. Interocular ratio (interocular width/width of eye): 2.80–3.73. Antenna: ratio funiculus/club 0.66–0.75. Pronotum w/l: 1.54–1.67. Scutellum w/l: 1.21–1.48. Pygidium w/l: 1.53–1.78. Width between mesocoxae: 0.15–0.27 mm. Metatibia w/l: 2.35–3.08. External claw l/h: 1.32–2.01. Colour on head, pronotum, scutellum, and pygidium from greenish to reddish to bronze brown.</p><p>Three different populations are present in Costa Rica, with defined geographic ranges (Fig. 33), each presenting subtle differences with the others and the specimens from Las Mercedes, Guatemala. The two paralectotypes from Costa Rica and the specimens from San Pedro (Universidad de Costa Rica, near San José, Central Valley) and Amubri (the Talamanca mountain range) are similar in size (body length 11.45–12.62 mm, width 5.94–6.93 mm; Guatemalan specimens: body length 11.74–12.53 mm, width 6.71–6.90 mm) but are paler in colour; the frons gradually curving towards vertex (Fig. 6 b), clypeus trapezoidal with lateral sides straight. This is in contrast to the Las Mercedes specimens, where they are curved, with basal half perpendicular to the base (see Fig. 6). Other differences are: less curved external protarsal claw, metatibia punctate above first carina, which is well defined, slender parameres (Figs. 14) and shorter and narrower endophallus (Fig. 27).</p><p>The specimens from the Guanacaste province are smaller in size (body length 9.31–9.96 mm, width 5.05–5.93 mm), with frons gradually curving towards vertex, metatibia punctate above first carina, which is well defined, narrower apical angle of parameres (Fig. 15), thinner setation on endophallus (Fig. 28). The paralectotypes from Pantaleon and San Isidro, Guatemala, are more similar to these specimens in size and genitalia than to the ones from Las Mercedes.</p><p>The specimens from the Osa peninsula (P.N. Corcovado) are also smaller in size (body length 8.44–9.90 mm, width 4.87–5.63 mm), with frons gradually curving towards vertex, metatibia punctate above first carina, which is well defined, slender parameres with sinuate ventral margin (Fig. 16), endophallus without patches of sclerotized setae (Fig. 29).</p><p>Diagnosis. Anomala eucoma is separated from similar species by the combination of the following characters: two dark bands on elytra, parameres pentagonal in shape, with dorsal and ventral margin not strongly sinuate, endophallus with one long sacculus. It can easily be separated from A. amphicoma, about the same size and dorsally setose, by the latter having the following characteristics: a metallic green pronotum in most specimens, covered by velvety dense setae; the pronotum is largest at 1/3 of height, with base narrower than elytra; elytra uniformly dark in colour, shorter parameres with sinuate ventral margin, endophallus with two long sacculi. It is differentiated from A. flavacoma by size ( A. flavacoma has an intermediate size between the larger and smaller specimens of A. eucoma) and its darker colour, proportionally narrower scutellum, endophallus with one narrow and long sacculus and lack of thick setae, in contrast to the two short sacculi of A. flavacoma . It can be distinguished from A. pseudoeucoma by the longer and more curved apical tooth in male protibia; more convex intermesocoxal space, shorter parameres, endophallus with one long sacculus and lack of thick setae, instead of the two sacculi with patches of thick setae of A. pseudoeucoma . Anomala megaparamera is similar in size and colour to the Osa and Guanacaste populations, but A. megaparamera has a proportionally wider pygidium, aedeagus with proportionally larger parameres, endophallus with several inflations and diverticles and copiously covered with setae.</p><p>Distribution. Guatemala: Las Mercedes; Costa Rica: Guanacaste and the Talamanca mountain ranges; Central Valley; Parque Nacional Corcovado, in the Osa peninsula (Fig. 33).</p></div>	https://treatment.plazi.org/id/03D28781FFF5FFDB5AA663BCFC26FD33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Micó, Estefania;Galante, Eduardo	Filippini, Valentina, Micó, Estefania, Galante, Eduardo (2013): Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3670 (2): 255-273, DOI: 10.11646/zootaxa.3670.2.9
03D28781FFFEFFD95AA660E7FB1EFD33.text	03D28781FFFEFFD95AA660E7FB1EFD33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala flavacoma Filippini, Mico	<div><p>Anomala flavacoma Filippini, Micó, &amp; Galante, new species</p><p>Fig. 3</p><p>Material examined: Holotype: ɞ "Limon, Costa Rica Reserva Hitoy–Cerere Rio Cerere 100 msnm 24 marzo 1987 Angel Solis / 56 / Costa Rica INBIOCRI002517822" (INBIO).</p><p>Paratypes (11): 1ɞ "Est. Hitoy Cerere, 100m, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limon, Costa Rica, Nov 1992 G. Carballo L- N / Costa Rica INBIOCRI000926676"; 1ɞ "Est. Hitoy Cerere, 100 m, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limon, COSTA RICA. Abr 1993. G. Carballo. L- N 184200_643300 / Costa Rica INBIOCRI000975444"; 1Ƥ "Est. Hitoy Cerere, 100m, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limon, Costa Rica, Nov 1992 G. Carballo L- N / Costa Rica INBIOCRI000926686"; 1Ƥ "Est. Hitoy Cerere, 100m, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limon, Costa Rica, Nov 1992 G. Carballo L- N / Costa Rica INBIOCRI000926680"; 1ɞ "Cuatro esquinas, P.N. Tortuguero, Prov. Limon COSTA RICA. 0m. 26 Aug– 6 Set 1989. J. Solano 280000, 590500 / Costa Rica INBIOCRI000020620"; 1ɞ "Albergue Heliconias, Bijagua (Costa Rica). 750m 30–1–2006. Luz casa (18:45–22:00). Leg.: Micó, García, Galante. / CEUA 00003306"; 1ɞ "Volcán Tenorio. Alajuela. Costa Rica. 1160m. 7/4/2008. Leg. J.A. Azofeifa / CEUA 00003320"; 1ɞ "Estación Cabro Muco, Repr. ICE – Z.P. Miravalles, Guanacaste. Costa Rica. 1000m 12/06/2010. L. Filippini, Moraga. / CEUA 00105076"; 1Ƥ "Estación Cabro Muco, Repr. ICE – Z.P. Miravalles, Guanacaste. Costa Rica. 1000m 12/06/2010. L. Filippini, Moraga. / CEUA 00105077"; 1Ƥ "Dos de Tilarán (San Ramón), Prov. Guana. COSTA RICA. 1100m. MAY 1995. G. Rodriguez. L_N_262600_437500 #5891 / INBIOCRI002337626"; 1Ƥ "Dos de Tilarán (San Ramón), Prov. Guana. COSTA RICA. 1100m. MAY 1995. G. Rodriguez. L_N_262600_437500 #5891/ INBIOCRI002337602".</p><p>Description. Male. Body shape oval. Length 11.21 mm. Width 6.30 mm. Head, pronotum, scutellum, and pygidium dark brown; legs and underside from reddish brown to dark orange; elytra light brown with suture, humerus, and basal half of lateral margins dark brown. The dark stripe on suture broadens mid elytra, forming a small expansion. Surface all covered with fine, blonde setae.</p><p>Clypeus trapezoidal, densely punctate-reticulate; ratio width/length 1.88. Anterior angles rounded. Frons densely punctate, flat; vertex with apparent boundary. Ocular canthum long, thin, and rounded at apex. Interocular ratio (interocular width/width of eye): 3.20. Male antenna: ratio funiculus/club 0.76.</p><p>Pronotum trapezoidal, width 1.71 times length. Lateral margins forming an obtuse angle at 1/3 of pronotum length. Anterior angles right and sharp, posterior angles obtuse and sharp. Basal margin thin and complete, sinuate. Entire surface with deep punctures.</p><p>Scutellum subpentagonal to subtriangular in shape, with rounded sides and acute apex. Ratio width/length 1.38. Sparsely punctate.</p><p>Elytra with striae defined by dark punctures. Subsutural interstice with 2–3 rows of sparse punctures, 2nd–4th interstices with one irregular secondary striae. Surface covered with shallow setigerous punctures. Lateral margin convex, larger at base, disappearing on apex. Marginal membrane almost complete.</p><p>Pygidium finely granulate, with longer setae at hind margins. Triangular in shape. Slightly convex in lateral view. Ratio width/length 1.69.</p><p>Metasternal disc slightly sulcated, densely punctate. Space between the mesocoxae narrow, slightly convex; width 0.22 mm. Mesometasternal suture well defined, at base of mesocoxae.</p><p>Abdominal sternites completely covered with dense setae, arranged in about 7–10 irregular rows, with strigate surface. Male last sternite strigate-imbricate, with apical margin well defined and rounded.</p><p>Protibia with 2 teeth (Fig 8); apical tooth long and curved; second tooth at the same level of internal apex of protibia, triangular in shape, obtuse. Metatibia stout, slightly narrower subapically. Ratio length/width 2.95. First carina well developed. Surface with elongate punctures above second carina and rugose below. Protarsal claws: external claw strongly curved; ratio length/height: 1.37; internal claw bifurcate, with upper branch slightly shorter and half the width of the lower one. Inferior margin sinuate.</p><p>Aedeagus (Figs 17–19): parameres pointed and narrow. Endophallus (Fig. 30) wide, with 2 short sacculi. The wider, ventral one has a strip of thick spines, with defined margins that extend frontally from side to side in a horseshoe shape. The dorsal sacculus is covered with fine setation, slightly thicker than the microsetation that commonly covers the entire surface of endophallus. Median lobe sclerotized.</p><p>Female. Similar to male. Scutellum with blunt apex. Internal protarsal claw narrower than in male, inferior margin straight. Protibia (Fig. 8) with apical tooth wider and longer than in male; second tooth above internal apex. Metatibia wider than in males. Last abdominal sternite with curved margin.</p><p>Variation. Dark pigmentation of elytra margin can extend to the tip of elytra; the expansions of sutural dark stripe mid length of elytra can be inconspicuous or form a complete band across the elytra (as in specimens from P.N. Tortuguero, Est. Cabro Muco, Heliconias). Body length 10.86–11.21 mm, body width 6.04–6.30 mm. Clypeus w/l: 1.88–2.04. Interocular ratio (interocular width/width of eye): 3.15–3.32. Pronotum w/l: 1.56–1.71. Scutellum w/l: 1.33–1.49. Pygidium w/l: 1.66–1.77. Width between mesocoxae: 0.19–0.22 mm. Metatibia w/l: 2.68–2.95. External claw l/h: 1.37–1.58.</p><p>Diagnosis. Anomala flavacoma can be differentiated from similar species using the combination of the following characters: medium size (10–12 mm) elytra light brown, with a dark central spot to a complete medium transversal band; metatibia with oblong punctures above second carina, abdominal sternites with dense setae, endophallus with two short and swollen sacculi.</p><p>Slightly smaller and more slender than A. eucoma and A. pseudoeucoma, it can be distinguished by the much lighter colour of elytra, pronotum, and underside; slender apex of parameres and endophallus that is shorter and wider. The specimens that develop a horizontal dark band on elytra may look similar to the lighter specimens of A. eucoma, but in A. flavacoma the band is much darker than in A. eucoma, and there is just one. Moreover, the puncture on the elytra is shallow and not coalescent in A. flavacoma, with clear differentiation between setigerous punctures and striae punctures, the latter being larger and dark coloured; whereas in A. eucoma both types of punctures are deep, and coalescent. A. megaparamera is much smaller and darker in colour, with a proportionally wider pygidium, wider parameres and proportionally longer and thinner endophallus.</p><p>Distribution. Caribbean lowlands of Costa Rica (Fig. 34).</p><p>Etymology. From Latin "flava", blonde, and "coma", hair, for its pale colour and setation.</p></div>	https://treatment.plazi.org/id/03D28781FFFEFFD95AA660E7FB1EFD33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Micó, Estefania;Galante, Eduardo	Filippini, Valentina, Micó, Estefania, Galante, Eduardo (2013): Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3670 (2): 255-273, DOI: 10.11646/zootaxa.3670.2.9
03D28781FFFCFFDF5AA660FAFEDBFE43.text	03D28781FFFCFFDF5AA660FAFEDBFE43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala megaparamera Filippini, Mico	<div><p>Anomala megaparamera Filippini, Micó, &amp; Galante, new species</p><p>Fig. 4</p><p>Material examined. Holotype: ɞ "Est. Cuatro Esquinas, P. N. Tortuguero, 0 m, Prov. Limon, COSTA RICA, R. Delgado, Oct 1990, L- N 280000_590500 / COSTA RICA INBIOCRI000285104" (INBIO).</p><p>Paratypes (6): 1ɞ "Est. Cuatro Esquinas, 0m, P. N. Tortuguero, Prov. Limon, COSTA RICA. E. Quesada, Jun 1990, L- N 280000_590500 / CEUA 00105268"; 1ɞ "Cerro Tortuguero, 0–120m P. N. Tortuguero, Prov. Limon, COSTA RICA, R. Delgado, Feb 1992, L- N 285000_588000 / COSTA RICA INBIOCRI000552532"; 1ɞ "Est. Cuatro Esquinas, P. N. Tortuguero, 0m, Prov. Limon, COSTA RICA, E. Quesada, Jul 1990, L- N 280000_590500 / COSTA RICA INBIOCRI000670946"; 1ɞ "Cuatro Esquinas, P. N. Tortuguero, Prov. Limon, COSTA RICA. 0m. 26 Aug–6 Set 1989, J. Solano, L N 280000_590500 / COSTA RICA INBIOCRI000020633"; 1Ƥ "Cuatro Esquinas, P. N. Tortuguero, Prov. Limon, COSTA RICA. 0m, Set 1989. J. Solano, L N 280000_590500 / COSTA RICA INBIOCRI000086333"; 1Ƥ "Est. Cuatro Esquinas, P. N. Tortuguero, 0m, Prov. Limon, COSTA RICA, U. Chavarria, Jul 1990, L- N 280000_590500 / CEUA 00105269".</p><p>Description. Male. Body shape oval. Length 9.03 mm. Width 5.10 mm. Head, pronotum, and scutellum dark reddish brown. Pygidium, legs, and underside reddish brown; elytra light brown with two dark reddish brown horizontal bands, one median and one covering the bottom third. Surface covered with fine, blonde setae.</p><p>Clypeus trapezoidal, densely and deeply punctate, with rather coarse punctures; ratio width/length 2.21. Anterior angles rounded. Frons deeply punctate, with a shallow central depression and apparent boundary with vertex. Ocular canthum long, thin, and rounded at apex. Interocular ratio (interocular width/width of eye): 2.93. Antenna: ratio funiculus/club 0.62.</p><p>Pronotum trapezoidal, width 1.67 times length. Lateral margins form an obtuse and protruding angle at 1/3 of pronotum length, with width of pronotum at this level being slightly greater than at base. Anterior angles right and sharp, posterior angles obtuse and rounded. Basal margin thin and complete, sinuate. Whole surface with deep, large, and confluent punctures with sulcus not reaching hind margin. Surface with shallow wrinkles due to confluence of punctures.</p><p>Scutellum subpentagonal in shape, with rounded sides and blunt apex. Ratio width/length 1.30. Coarsely punctate.</p><p>Elytra with irregular striae. Subsutural interstice with 2 irregular rows of often confluent punctures, following interstices with irregular secondary striae; costae 2–6 not conspicuous. Setigerous punctures coincide with striae punctures, consequently the setae are not very dense. Lateral margin convex, larger at base, disappearing on apex. Marginal membrane almost complete.</p><p>Pygidium finely granulate, with longer setae at hind margins. Triangular in shape. Slightly convex in lateral view. Metasternal disc slightly sulcated, covered with sparse setae on the entire surface. Space between the mesocoxae narrow, slightly convex; width 0.21 mm. Mesometasternal suture well defined, at base of mesocoxae.</p><p>Abdominal sternites with 2–3 transverse rows of setae in the middle, and denser setae at sides. Punctures large and oval, sometimes fusing in diagonal streaks. Last sternite strigate-imbricate, with apical margin well defined and slightly sinuate.</p><p>Protibia with 2 teeth (Fig.9): apical tooth long and almost straight; second tooth at the same level of internal apex of protibia, triangular in shape, obtuse. Metatibia stout, narrower subapically. First carina well developed. Surface rugose.</p><p>Protarsal claws: external claw strongly curved; internal claw bifurcate, with upper branch slightly shorter and half as wide as the lower one. Inferior margin sinuate.</p><p>Aedeagus (Figs. 20–22) with very wide parameres, and proportionally shorter tectum. Endophallus (Fig. 31) composed of a long ventral sacculus covered with sclerotized setae and a median posterior inflation; on the opposite side, 2 frontal inflations and 2 dorsal diverticles; gonopore situated dorsally between one inflation and diverticles (ejaculatory duct partially everted in illustration).</p><p>Female: similar to males, but the antennal club is shorter than in males. Pronotum usually wider than in males. Apical tooth of protibia (Fig.9) wider and longer than in males; second tooth above internal apex. Upper branch of the internal protarsal claw half the width of the lower one, inferior margin straight. Metatibia stouter than in males. Last abdominal sternite with curved margin.</p><p>Variation. Second tooth of protibia at the same level or slightly superior in relation to the internal apex of protibia. Body length 9.02–10.14 mm, body width 5.10–6.18 mm. Clypeus w/l: 1.92–2.21. Interocular ratio (interocular width/width of eye): 2.93–3.52. Antenna: ratio funiculus/club 0.62–0.73. Pronotum w/l: 1.63–1.68. Scutellum w/l: 1.30–1.59. Pygidium w/l: 2.10–2.20. Width between mesocoxae: 0.21–0.25 mm. Metatibia w/l: 2.96–3.07. External claw l/h: 1.43.</p><p>Diagnosis. Anomala megaparamera is separated from similar species by the combination of the following characters: small size (8–10 mm); presence of depression in frons and irregular wrinkles on pronotum; pronotum wider at 1/3 of length than at base; metatibia surface rugose; parameres nearly as wide as tectum length, and similarly long; endophallus with additional inflations and diverticles.</p><p>Similar to the Osa and Guanacaste populations of A. eucoma, much smaller than the other species described here, it can be distinguished by the larger and coarser punctation on head, elytra, and sternites; the presence of depressions in frons and pronotum; much wider and longer parameres, and more complex and setose endophallus.</p><p>Distribution. Costa Rica, Parque Nacional Tortuguero (Fig. 35).</p><p>Etymology. From Greek mega "big" and paramere, referring to the large proportions of parameres in relation to size of aedeagus.</p></div>	https://treatment.plazi.org/id/03D28781FFFCFFDF5AA660FAFEDBFE43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Micó, Estefania;Galante, Eduardo	Filippini, Valentina, Micó, Estefania, Galante, Eduardo (2013): Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3670 (2): 255-273, DOI: 10.11646/zootaxa.3670.2.9
03D28781FFFAFFDC5AA6606AFB68F823.text	03D28781FFFAFFDC5AA6606AFB68F823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala pseudoeucoma Filippini, Mico	<div><p>Anomala pseudoeucoma Filippini, Micó, &amp; Galante, new species</p><p>Fig. 5</p><p>Material examined. Holotype: ɞ "Est. Hitoy Cerere, 100m, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limón, COSTA RICA, R. Guzman. 12 – 28 Abr 1992. L_N_184600_643400 #1137 / COSTA RICA INBIOCRI000393407" (INBIO).</p><p>Paratypes (20): 1ɞ "Alajuela, Costa Rica, San Ramon, Rio S. Lorencito, 800m 3 octubre 1986 Col: A. Solis Blanco. / COSTA RICA INBIOCRI002517836"; 1Ƥ "Alajuela, Costa Rica. Reserva de San Ramón. Río San Lorencito, 850m. 1 abril 1987 Angel Solís / COSTA RICA INBIOCRI002517838"; 1Ƥ "Alajuela, Costa Rica. Reserva de San Ramón. Río San Lorencito, 850m. 1 abril 1987 Angel Solís / COSTA RICA INBIOCRI002517839"; 1Ƥ "Alajuela, Costa Rica. Reserva de San Ramón. Río San Lorencito, 850m. 1 abril 1987 Angel Solís / COSTA RICA INBIOCRI002517840"; 2ɞ "Albergue Heliconias, Prov. Alajuela, Costa Rica. 11/11/ 2007 Leg. D. Gutiérrez" CEUA 00105078 and CEUA 00105079; 1Ƥ "Albergue Heliconias, Prov. Alajuela, Costa Rica. 31/01/2008 Leg. D. Gutiérrez / CEUA 00105081"; 1ɞ "Albergue Heliconias, Prov. Alajuela, Costa Rica. 14/ 09/2007 Leg. D. Gutiérrez / CEUA 00105082"; 1ɞ "Cerro Tortuguero, 0–100m, P. N. Tortuguero, Prov. Limon, COSTA RICA, R. Delgado, 26 Ene–14 Feb 1992, L- N 285000_588000 / COSTA RICA INBIOCRI000334543"; 1ɞ "Cerro Tortuguero, 0–120m, P. N. Tortuguero, Prov. Limon, COSTA RICA, R. Delgado, Set 1991, L- N 285000_588000 / COSTA RICA INBIOCRI000479636"; 1ɞ "Cerro Tortuguero, P. N. Tortuguero, 0 – 100m, Prov. Limon, COSTA RICA. J. Solano, May 1990. L- N 285000_588000 / COSTA RICA INBIOCRI000242771"; 1Ƥ "Cerro Tortuguero, P. N. Tortuguero, Prov. Limon, COSTA RICA. 100 m. Nov 1989. J. Solano, L N 285000_588000 / COSTA RICA INBIOCRI000055767"; 1ɞ "Cerro Tortuguero, P. N. Tortuguero, Prov. Limon, COSTA RICA. 100 m. Nov 1989. J. Solano, L N 285000_588000 / COSTA RICA INBIOCRI000141229"; 1ɞ "Est. La Casona, 1520 m, Res. Biol. Monteverde, Prov. Punt. COSTA RICA, N. Obando, Oct 1991, L- N 253250_4497001 / COSTA RICA INBIOCRI000632624"; 1ɞ "Est. La Casona, 1520 m, Res. Biol. Monteverde, Prov. Punt. COSTA RICA, N. Obando, Oct 1991, L- N 253250_449700 / COSTA RICA INBIOCRI000632689"; 1ɞ "Fca. Cafrosa, Est. Las Mellizas, P. N. Amistad, Puntarenas Pr. COSTA RICA, 1300 m. 20 Aug – 4 Set 1989, M. Ramirez &amp; G. Mora, L S 316100_596100 / COSTA RICA INBIOCRI000093803"; 1Ƥ " Isla Bonita, Prov. Alajuela, Costa Rica. 12/07/2007 Leg. M. Moraga / CEUA 00105080"; 1ɞ "Limon, Costa Rica Reserva Hitoy– Cerere Rio Cerere 100 msnm 24 marzo 1987 Angel Solis / COSTA RICA INBIOCRI002517319"; 1ɞ "San Luis, Monteverde, Prov. Punta., COSTA RICA, 1040 m. nov 1993. Z. Fuentes L- N 250850 _449250 #2443 / COSTA RICA INBIOCRI001937988"; 1ɞ "R.B. Hitoy Cerere, Valle La Estrella, Prov. Limon, COSTA RICA. 100– 200m. 9–14 May 1994, G. Carballo, L N 643400_184600 # 2856 / COSTA RICA INBIOCRI001799052".</p><p>Description. Male. Body shape oval. Length 12.16 mm. Width 6.57 mm. Head, pronotum, scutellum, and pygidium dark reddish brown; legs and underside reddish brown; elytra dark brown with a light horizontal band along the anterior margin. The entire surface is covered with fine, blonde setae.</p><p>Clypeus trapezoidal, with shallow and large reticulate punctures; ratio width/length 2.04. Anterior angles rounded. Frons densely punctate, flat, gradually curving towards vertex (no boundary clearly visible between the two parts). Ocular canthum long, thin, and rounded at apex. Interocular ratio (interocular width/width of eye): 3.03. Antenna: ratio funiculus/club 0.69.</p><p>Pronotum trapezoidal, width 1.58 times length. Lateral margins forming an obtuse angle at 1/3 of pronotum length. Anterior angles right and sharp, posterior angles obtuse and rounded. Basal margin thin and complete but partially obliterated in the middle, sinuate. Entire surface with deep punctures.</p><p>Scutellum subpentagonal to subtriangular in shape, with rounded sides, surface finely punctate. Ratio width/ length 1.47.</p><p>Elytra with striae defined by shallow grooves, surface with punctures coalescent. On light bands, dark spots show the striae-defining punctures and sparse punctures in the interstices. Lateral margin convex, larger at base, disappearing on apex. Marginal membrane almost complete.</p><p>Pygidium finely granulate, with all setae equal in size. Triangular in shape. Slightly convex in lateral view. Ratio width/length 1.94.</p><p>Metasternal disc slightly sulcated, densely setose. Space between the mesocoxae narrow, slightly convex, width 0.17 mm. Mesometasternal suture well defined at base of mesocoxae.</p><p>Abdominal sternites with numerous (5–10) irregular transverse rows of setae, and elongate punctures that fuse at sides. Last sternite strigate, with apical margin well defined and slightly sinuate.</p><p>Protibia with 2 teeth (Fig.10), apical tooth long and slightly curved; second tooth at the same level of internal apex of protibia, triangular in shape, obtuse. Metatibia stout, slightly narrower subapically. Ratio length/width 3.97. First carina poorly defined by isolated spines. Surface punctate above second carina and rugose below.</p><p>Protarsal claws: external claw strongly curved; ratio length/height: 1.82; internal claw bifurcate, with upper branch slightly shorter and more than half the width of the lower one. Inferior margin sinuate.</p><p>Aedeagus (Figs. 23–25): parameres with blunt apex, protruding from the strongly sinuate ventral margin. Endophallus (Fig. 32) composed by a long ventral sacculus with a ridge of thick spines on dorsal margin, and a swollen shorter dorsal sacculus with a wide patch of sclerotized setae on left side. Gonopore ventral, between the two sacculi.</p><p>Female: similar to male, but the antennal club is slightly shorter than in males. Apical tooth of protibia (Fig. 10) wider and longer than in males; second tooth above internal apex. Internal protarsal claw narrower than in males, inferior margin straight. Metatibia stouter than in males. Last abdominal sternite with curved margin.</p><p>Variation: Underside from reddish brown to dark brown; elytra dark brown to bluish black; in some specimens with homogeneous colour (without the basal lighter band), and rarely, with a barely distinguishable median band. In some specimens, particularly the darker ones, the denser and thicker setae give the pronotum a velvety blond appearance. Second tooth of protibia at the same level or slightly above internal apex of protibia. On endophallus, the patch of setae on dorsal sacculus extends further frontally in some specimens.</p><p>Body length 11.34–12.63 mm, body width 6.57–7.20 mm. Clypeus w/l: 1.81–2.10. Interocular ratio (interocular width/width of eye): 2.94–3.45. Antenna: ratio funiculus/club 0.69–0.71. Pronotum w/l: 1.58–1.65. Scutellum w/l: 1.26–1.47. Pygidium w/l: 1.65–1.94. Width between mesocoxae: 0.16–0.24 mm. Metatibia w/l: 2.79–3.97. External claw l/h: 1.37–1.82.</p><p>Diagnosis. Anomala pseudoeucoma can be distinguished from the other species described here by its dark and uniform brown colour, median size, frons nearly continuous with apex, first carina of metatibia obliterated, surface of metatibia punctate above second carina, parameres with strongly sinuate inferior margin, and endophallus with two patches of sclerotized spines and setae that can be distinguished even in the inverted endophallus.</p><p>This species is very similar to A. eucoma, especially in the lighter specimens; A. pseudoeucoma can be distinguished by the following characters: shorter and less curved apical tooth in male protibia; large internal claw in males, with inferior border more strongly sinuate; narrower intermesocoxal space, and less convex than in A. eucoma . As in the case of A. eucoma, the brown and trapezoidal pronotum easily distinguishes this species from A. amphicoma . Anomala flavacoma is smaller and much lighter in colour, with marked sides of frons with vertex, metatibia with oblong punctures above second carina. Anomala megaparamera is much smaller in size, with larger parameres and endophallus with diverticles and setae less sclerotized and more diffuse.</p><p>Distribution. Widely distributed, with localities recorded in Guanacaste and the Tilarán mountain ranges to the north, the Talamanca Mountain Range near the border with Panama to the south, and Caribbean lowlands. Altitudinal range varies from 0 m to above 1500 m (Fig. 36).</p><p>Etymology. From Greek "pseudo-", false, and " eucoma ", for its similarity with A. eucoma .</p></div>	https://treatment.plazi.org/id/03D28781FFFAFFDC5AA6606AFB68F823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Micó, Estefania;Galante, Eduardo	Filippini, Valentina, Micó, Estefania, Galante, Eduardo (2013): Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3670 (2): 255-273, DOI: 10.11646/zootaxa.3670.2.9
03D28781FFF8FFC25AA6673FFA34FDF0.text	03D28781FFF8FFC25AA6673FFA34FDF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala	<div><p>Key to dorsally setose Anomala species of the Neotropical Region</p><p>Only the species with setae on the elytral surface and two protibial teeth are included. Of the additional species included in the key, only A. amphicoma probably belongs to the same group of the species described here. A. semitonsa, A. balzapambae, and especially A. hispidula and A. tindakua are likely distant phylogenetically.</p><p>1. Rounded shape, with widest point mid length of elytra; two transversal bands on elytra: a median transversal wavy band, hind band usually not reaching the posterior margin of elytra....................................................... 2</p><p>-. Elongate shape, with widest point at last third of elytra; elytra with uniform colour, or with 1 or more, continuous, transversal bands, the hind band covering the apical third of elytra; when bands are defined by spots, they are arranged in 3 rows...... 3</p><p>2. Median band on elytra continuous, hind band reaching sides, male genitalia illustrated in Fig. 37. Panama, Ecuador ........................................................................................ A. semitonsa Bates, 1888</p><p>-. Median band on elytra composed of isolated spots, hind band usually not reaching sides, male genitalia illustrated in Fig. 38. Costa Rica, Ecuador ............................................................ A. balzapambae Ohaus, 1897</p><p>3. Pronotum metallic green, larger at 1/3 of height than at base, elytra uniform dark brown, total length about 10 mm, male genitalia illustrated in Fig. 39. Panama ................................................... A. amphicoma Bates, 1888</p><p>-. Pronotum dark brown, largest at base, if a green luster is present then elytra have transversal light bands................ 4</p><p>4. Pronotum almost rectangular, head large, almost as long as pronotum and 2/3 the width of the pronotum. Elytra light brown dappled with darker spots, concentrated in three transversal bands (post humeral, median, and posterior half of elytra), or expanding until covering almost all surface................................................................ 5</p><p>-. Pronotum trapezoidal, head small, about half wide as the pronotum. Elytra with uniform colour or maximum two transversal bands, never a posthumeral band........................................................................ 6</p><p>5. Elytra with setae on the entire surface, pronotum of uniform colour or with slightly lighter border in light coloured specimens, pattern on elytra not defined, aedeagus illustrated in Fig. 40. Mexico, Guatemala ................ A. hispidula Bates, 1888</p><p>-. Elytra with setae only on posterior third, border of pronotum always of lighter colour, pattern on elytra well defined, aedeagus in Morón &amp; Nogueira (2002). Mexico ...................................... .. A. tindakua Morón &amp; Nogueira, 2002</p><p>6. Elytra uniformly light brown or with one transversal dark band. Size approximately 11 mm. Costa Rica ...................................................................................... A. flavacoma Filippini, Micó, &amp; Galante</p><p>-. Elytra uniformly dark brown or with two bands, apical third of elytra always dark.................................. 7</p><p>7. Pronotum with irregular surface due to small depressions on sides of median sulcus. Parameres wide, maximum length 3/4 of tectum length, with blunt and wide apex. Length less than 10.1 mm. Costa Rica ..................................................................................................... A. megaparamera Filippini, Micó, &amp; Galante</p><p>-. Pronotum with uniform surface, sometimes a median sulcus is present. Parameres slender, with defined narrow apex; length of parameres not reaching 3/4 of tectum length. Length 8.4–12.7 mm .............................................. 8</p><p>8. Elytra uniformly dark or with a lighter posthumeral band (rarely a second light band on disc); male protibia upper tooth short (less than 1/4 of total length) and almost straight; aedeagus with wide apex and strongly sinuate ventral margin. Costa Rica ................................................................... A. pseudoeucoma Filippini, Micó, &amp; Galante</p><p>-. Two darker transversal bands are visible on elytra; male protibia upper tooth long (more than 1/4 of total length) and oblique, parameres with short rounded apex and slightly sinuate ventral margin. Guatemala, Costa Rica ...... A. eucoma Bates, 1888</p></div>	https://treatment.plazi.org/id/03D28781FFF8FFC25AA6673FFA34FDF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Micó, Estefania;Galante, Eduardo	Filippini, Valentina, Micó, Estefania, Galante, Eduardo (2013): Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3670 (2): 255-273, DOI: 10.11646/zootaxa.3670.2.9
