taxonID	type	description	language	source
8737C9C36BF7AF054586D5391737C63F.taxon	diagnosis	Diagnosis. The genus Doryodes is easily recognized because of the elongated, apically pointed wings and the elongated abdomen. Males have broadly bipectinate antennae with pectinations 3 - 5 x as long as the width of the antennal shaft; females have filiform antennae. The frons is bare ventrally, covered with rough scaling dorsally. The eyes are rounded, without hairs or lashes. Ocelli are present. The labial palpus usually is directed forward, occasionally with the terminal segment angled ventrally; the basal and apical segments are about half as long as the middle segment. Forewing length varies from 13 - 21 mm, females, on average, are longer winged than males. The forewing in the male usually is pale buffy brown; some species have longitudinal streaks of gray, brown, or yellowish orange. There is a blackish-brown stripe along the middle of the wing, almost straight from the wing base to three-quarters of the distance to the apex, at which point it curves upward toward the apex and tends to narrow and fade out between the curve and the apex. The dark stripe is bordered dorsally by a narrow white line extending from the wing base to the point where the dark stripe curves upward; the dark stripe is bordered ventrally by a narrow white line on the outer half or third of the wing to, or almost to, the wing apex. Usually there are one or two dark dots representing the reniform spot, orbicular spot, or both. The forewing of the female usually is paler and more acutely-pointed apically than that of the male. The dark longitudinal stripe along the middle of the forewing is narrower in most females than in males. The hindwing is white to buff, and the color can vary with seasonal generations. In particular, many individuals captured from November to March are much darker than those found in spring and summer. The winter forms are often misidentified because superficially they differ more from the " summer " forms than most species differ from each other. The legs have spiniform setae on the tibia; the basitarsus has two or three ventral rows of spiniform setae. The abdomen is without tufts or abdominal brushes. In the male genitalia the tegumen, and especially the vinculum, are long and have a simple articulation with each other and with the valves; the vinculum ends ventrally as a deep V-shaped saccus. The uncus is shorter than the tegumen, bulbous at the base, then slightly tapered to the apex, with a spine-like tip. The juxta is strongly fused to the base of the valves, so it is difficult to spread the valves laterally without distorting or tearing the middle part of the genitalia. The valve is elongate and tripartite with the medial part lightly sclerotized apically. The base of the valve has a long lens-shaped sacculus extending about half way to the valve apex. The sacculus is extended as a sclerotized tube along (and fused with) the ventral margin of the valve, except toward the apex of the valve where the saccular extension is free from the valve and tapered to a pointed or blunt apex. A medial sclerotized ridge extends posteroventrally across the ventral surface of the valve from the base of the tegumen to fuse with the saccular extension on the lower edge of the valve (transverse ridge most prominent in Doryodes tenuistriga); the costal margin of the valve also is heavily sclerotized, more so toward the apex of the valve where it ends in a pointed or bluntly-rounded process, often free from the medial part of the valve at the apex. In one species (Doryodes tenuistriga) the sclerotized costal and ventral margins of the valves are broadly rounded at their apices and end well before the valve apex. The middle part of the valve is lightly sclerotized, especially the almost membranous, rounded, apical part of the valve. The aedeagus is elongated and cylindrical, 7 - 10 x as long as its mesial width in most species (4 - 5 x as long in Doryodes tenuistriga and Doryodes okaloosa). The vesica is about as long as the aedeagus in most species, although abruptly curved ventrally or laterally near its middle. The vesica has numerous diverticula, typically five (numbered on Figs 33, 34, 36, 37, 38, 39), each usually with a cornutus that may be broad and shaped like a shark fin, or slightly to markedly serrated into several to many basally-fused spines; the basal part of the vesica, just beyond the end of the aedeagus is often slightly to markedly swollen and has one to five sclerotized plates, often armed with one to many spinules. The narrow ductus seminalis arises near the middle of the vesica ventrally and typically clogs the narrow opening rather than everting during dissection. The recognition of most species requires examination of the relative position, size, and shape of the diverticula and their associated cornuti. The female genitalia are elongated with an oblong corpus bursae, which occasionally has a small signum, and has a well-developed and variably-sclerotized ventro-lateral appendix bursae posteriorly on the right. The ductus bursae is straight and varies in length, with the sclerotized plate wider at each end than mesially, except in Doryodes reineckei and Doryodes tenuistriga. The ductus bursae extends posteriorly as a sclerotized quadrangular plate over the ostium bursae. The ductus seminalis arises at the base of the appendix bursae posteriorly; in most species it is abruptly tapered at its base and thread-like after that, however in Doryodes reineckei and Doryodes tenuistriga it is wider and only gradually tapered. The anal papillae are lightly sclerotized and apically rounded or tapered, covered laterally with hair-like setae. The anterior apophyses are rod-like and usually are about as long as the posterior apophyses.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
2B0C61999421D0806D18DE560AF3F493.taxon	materials_examined	Other material examined and distribution. We have examined material from North Carolina east of the Piedmont, from most of Florida except for the Keys and Panhandle and a single male from a power cut near Grand Bay National Wildlife Refuge, Jackson County, Mississippi. Specimens from Florida, Mississippi and North Carolina are closely similar in appearance, genitalia and barcodes.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
2B0C61999421D0806D18DE560AF3F493.taxon	diagnosis	Diagnosis. Superficially, adults usually can be identified by the narrower dark stripe on the forewing and their relatively small size (forewing length: ♂ 13.0 - 15.5 mm, ♀ 14.5 - 16.0 mm). Compared to Doryodes spadaria the medial longitudinal stripe on the forewing of Doryodes bistrialis is much narrower and the hindwings are whitish not buff through June emergences. Later in the year, Doryodes spadaria hindwings become more whitish, so wing length and the width of the longitudinal line must be relied upon to distinguish the species, or genital characters must be examined for positive identification. Females of Doryodes bistrialis do not overlap those of Doryodes spadaria in size, females of Doryodes spadaria having a forewing length of 18.0 - 21.0 mm. The male vesica is also diagnostic in Doryodes bistrialis in that there is a line of smaller cornuti extending along the trunk usually in three patches, whereas in Doryodes spadaria there are two side-by-side cornuti on the basal trunk of the vesica. In Doryodes bistrialis, diverticulum 1 is 2 - 3 x as long as wide and has a deeply-serrated rooster-comb-like cornutus at the apex; Doryodes spadaria has no diverticulum in this position and the cornuti in the vesica are triangular, sometimes minutely serrated on one side. Doryodes bistrialis can be distinguished from Doryodes fusselli by the shape of diverticulum 1, which is rounded in Doryodes fusselli and about as long as wide; also the apex of the vesica in Doryodes bistrialis is symmetrical with a triangular cornutus on a pouch on each side (diverticula 4 and 5). Doryodes bistrialis can be distinguished from Doryodes reineckei and Doryodes latistriga by the wing pattern and by size and from Doryodes broui by the characters of the vesica. Along the Gulf Coast Doryodes bistrialis could be confused with Doryodes broui and Doryodes tenuistriga, but characters of the vesica (Doryodes broui, Fig. 40 and Doryodes tenuistriga, Fig. 42, versus Doryodes bistrialis, Fig. 33) readily distinguish these three species and Doryodes bistrialis occurs farther inland. The female genitalia of Doryodes bistrialis are elongated with a more compact, less differentiated, appendix bursae than in Doryodes spadaria.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
D8C1ED1D1FF2DE8F0D03E5B35F32BF1A.taxon	etymology	Etymology. This species is named after Ft. De Soto Park, Florida.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
D8C1ED1D1FF2DE8F0D03E5B35F32BF1A.taxon	diagnosis	Diagnosis. This species is superficially indistinguishable from Doryodes spadaria, Doryodes fusselli, and Doryodes broui, although its range on the Gulf Coast of Florida is north and west of the range of Doryodes spadaria and Doryodes fusselli, and east of the range of Doryodes broui. Forewing length is 16 mm, on the small size for Doryodes spadaria, but within the normal range of the other similar species. The species can be definitively identified only by the male vesica, particularly by the long, narrow diverticulum 1, which is about 4 - 5 x as long as its basal width and with an apical, deeply serrated rooster-comb-like cornutus; diverticulum 2 is on the left side of the vesica, not posterior as in Doryodes bistrialis; diverticulum 4 is preapical on the right, without a cornutus, and diverticulum 5 forms a projecting lobe with a cone-shaped diverticulum on its left side.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
D8C1ED1D1FF2DE8F0D03E5B35F32BF1A.taxon	description	Description. External structural characters as described for genus. Forewing length 16 mm; forewing whitish buff with slightly darker-buff and pale-gray streaks; a prominent blackish-brown stripe along the middle of wing, curving upward and tapered at about 3 / 4 from base; stripe narrower than for Doryodes spadaria and Doryodes okaloosa, but wider than for Doryodes bistrialis; stripe bordered by narrow white line above extending to 3 / 4 from base, and with similar white line below stripe extending from above forewing tornus almost to wing apex. Hind wing white with very faint buffy tone. Male genitalia mainly as described for genus. Dorsal heavily-sclerotized margin of valve extending beyond middle membranous part, then tapered abruptly into a sharp spine; ventral sclerotized margin of valve not evenly tapered, but widens slightly preapically then tapered to blunt point free from inner membranous part of valve. Aedeagus cylindrical, 8 - 9 x as long as mesial width. Vesica with swollen area distal to end of aedeagus, 0.4 - 0.5 x as long as aedeagus and about 2 x as long as wide, with two spinule-covered sclerotized plates, these partially or completely divided into as many as four plates each with less spinules; diverticulum 1 finger-like, 4.5 - 5.5 x as long as mesial width, with apical serrated cornutus; diverticulum 2 rounded, on left side at base of diverticulum 1, with shark-fin-like cornutus; diverticulum 3 quadrate, on right side and without a cornutus; diverticulum 4 preapical, on right side without a cornutus; diverticulum 5 forming a bulbous apical lobe with a rounded spine-tipped cornutus on left side.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
DC29BD655602B484D4B34902E96F63B2.taxon	etymology	Etymology. The species name is in honor of the Okaloosa, a tribe of the Creek Nation and longtime inhabitants of the area.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
DC29BD655602B484D4B34902E96F63B2.taxon	diagnosis	Diagnosis. This species probably occurs with Doryodes desoto and Doryodes reineckei in the salt marshes and tidal creeks throughout the coastal panhandle area of Florida. The species is slightly larger than Doryodes bistrialis, which may occur nearby but inland. The washed out appearance of Doryodes reineickei immediately distinguishes it from Doryodes okaloosa. Doryodes desoto is very similar and at present dissection of the male genitalia is the only reliable way to distinguish these two species, although based on the single specimen of Doryodes okaloosa, it appears that Doryodes okaloosa is broader winged than Doryodes desoto, but not as broad winged as Doryodes reineckei. The abundance of large cornuti in the vesica, as well as the spatulate lower process of the valve and the short, broad aedeagus distinguishes Doryodes okaloosa from all other Doryodes species.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
DC29BD655602B484D4B34902E96F63B2.taxon	description	Description. Forewing length 16.5 mm; forewing stripe dark brown, narrower than for Doryodes spadaria but wider than for Doryodes desoto; forewing wider and browner than in Doryodes spadaria and Doryodes desoto, but not as wide as in Doryodes reineckei, and longitudinal stripe sharply defined, unlike that of Doryodes reineckei. Antennae missing but presumed to be bipectinate as in other species in genus. Hind wing white with buff tinge. Male genitalia mainly as described for genus. Dorsal heavily-sclerotized margin of valve wider and less well defined than in other species except for tapered spine-like apex; apex not rounded as in Doryodes tenuistriga, but wider than in other species; ventral sclerotized margin of valve slightly tapered to a broadly spatulate apex, much as in Doryodes tenuistriga. Aedeagus cylindrical, about 5 x as long as mesial width. Vesica with swollen area distal to end of aedeagus about 0.5 x as long as aedeagus and about 2 x as long as wide, with four large spine-covered sclerotized plates; diverticulum 1 slightly longer than wide with large deeply-serrated cornutus near apex; three preapical diverticula, each with large shark-fin-like cornutus; arrangement of large cornuti not seen in any other species of Doryodes.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
6FCA54E40E64DAB2DA53FBD01B3EB9D1.taxon	materials_examined	Other material examined and distribution. Canada: Quebec, Nova Scotia, Prince Edward Island. USA: Maine, New York, New Jersey, Maryland, North Carolina, South Carolina, Georgia, Florida.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
6FCA54E40E64DAB2DA53FBD01B3EB9D1.taxon	diagnosis	Diagnosis. External structural characters as described for genus. Doryodes spadaria is the most widespread and common species in the genus, and except for Doryodes fusselli in coastal North Carolina, all specimens of Doryodes from salt marshes along the Atlantic Coast of Canada and the United States that we have seen are Doryodes spadaria. Adults are relatively larger (forewing length: ♂ 13 - 20 mm, most commonly 16 - 18 mm; ♀ 18 - 21 mm, most commonly 19 mm) than those of Doryodes bistrialis (forewing length: ♂, 13.5 - 15.5 mm, ♀ 14.5 - 16.0 mm), the species most likely to be confused with it in southeastern United States outside of North Carolina where the smaller Doryodes fusselli also occurs in salt marshes. The forewing ground color in males varies from whitish buff to yellow buff with gray streaks; the longitudinal dark stripe is dark brown and conspicuously wider than in Doryodes bistrialis; the ground color in females averages paler than that of males and the wings and dark longitudinal stripe are narrower. In the male genitalia, most structural characters are as described for the genus; the sclerotized costal margin of the valve is more heavily sclerotized apically and extends farther beyond the central membranous part of the valve than in most other species; the sclerotized ventral margin of the valve ends in a blunt point before the end of the central part of the valve. Two examples of the genitalia are illustrated, one to show the typical orientation of the vesica in comparison with other species (Fig. 37), and a ventral orientation of the vesica (Fig. 38) to illustrate the shape and positions of the two preapical diverticula and their cornuti. The vesica is slightly shorter than the aedeagus; the basal, swollen part of the vesica is armed with two thorn-like cornuti on sclerotized plates; the left preapical diverticulum (# 5) is tapered to a large triangular cornutus, whereas the right preapical diverticulum (# 4) has a bulge in the middle and a smaller cornutus. In the female genitalia the corpus bursae is elongated with the part anterior to the opening of the ductus bursae swollen laterally and with longitudinal " ribbons " of sclerotization. The appendix bursae is lightly sclerotized and slightly bilobed posteriorly. The ductus bursae is almost as long as the corpus bursae, and is heavily sclerotized dorsally and ventrally with the plates slightly narrower mesially and expanded laterally and more heavily sclerotized at their junction with the corpus bursae; the ventral plate at the end of the ductus bursae is extended posteriorly as a quadrangular plate that projects over the opening to the ductus with the posterior margin of the plate concave. The anterior and posterior apophyses are similar in length (posterior slightly longer), about half the length of the ductus. The anal papillae are lightly sclerotized, produced ventrally anteriorly, rounded posteriorly, with the surface setose.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
FD38239826C78431C3CD5097F59BDCB9.taxon	etymology	Etymology. The species name is in honor of John Fussell from Morehead City, North Carolina, who has worked tirelessly for decades to describe and protect the unique flora and fauna of the North Carolina coastal plain, particularly the Croatan National Forest. All of our lives are richer for his efforts.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
FD38239826C78431C3CD5097F59BDCB9.taxon	diagnosis	Diagnosis. This species occurs with Doryodes spadaria in the salt marshes and tidal creeks throughout coastal North Carolina Adults are slightly larger than Doryodes bistrialis, but noticeably smaller (especially females) than Doryodes spadaria. The medial chocolate stripe on the forewing is broader than in Doryodes bistrialis, but narrower than that of Doryodes spadaria. Spring males are larger than those of the summer and fall generations, so they are more easily confused with Doryodes spadaria. The hind wing is white, without the buff coloring of Doryodes spadaria; in late summer some males of Doryodes spadaria can have white hind wings, but size ranges for the two species do not overlap in this generation. Similar changes in size with season are seen for many other species (Sullivan and Miller 2007). The vesica differs in having a diffuse line of cornuti along the basal trunk and by the symmetrical pair of cornuti-tipped diverticula at the apex. Doryodes spadaria is described in full, so only differences from it are given in the description.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
FD38239826C78431C3CD5097F59BDCB9.taxon	description	Description. Smaller than Doryodes spadaria; spring males (forewing length: 16 - 17 mm [16 - 20 mm for Doryodes spadaria]), slightly larger than summer males (forewing length: 14 - 15 mm); some small individuals (forewing length: 12 - 13 mm) late in season; females of Doryodes fusselli (forewing length: 16 mm) smaller than those of Doryodes spadaria (forewing length: 18 - 21 mm) but larger than those of Doryodes bistrialis. Hindwing pearly white. Male genitalia with valves similar to those of Doryodes spadaria but slightly smaller (valve length: 4.45 mm versus 4.75 mm for Doryodes spadaria). Vesica basal trunk with single band of cornuti, usually separated into two elongated sections; diverticulum 1 rounded with toothed, rooster-comb-like cornutus on dorsal side; diverticulum 2 rounded, about 1.0 - 1.5 x as long as wide, with triangular cornutus on ventral surface near apex; terminus of vesica with two similar cornuti-tipped diverticula projecting distally [in Doryodes spadaria two terminal diverticula are more elongated, different in shape from each other, and project laterally]. Female genitalia differ from those of Doryodes spadaria in more even width of ductus bursae, and appendix bursae more elongated with posterior margin even, not bilobed as in Doryodes spadaria.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
1FDD2C1F36D77EB228530156D5BA3964.taxon	etymology	Etymology. The name of this species refers to the width of the longitudinal dark stripe on the forewing in both sexes.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
1FDD2C1F36D77EB228530156D5BA3964.taxon	materials_examined	Other material examined and distribution. We have examined material from southern Alabama, Mississippi, and Louisiana. Specimens in Bold database exhibit considerable heterogeneity but all sequences are within 0.8 % of each other.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
1FDD2C1F36D77EB228530156D5BA3964.taxon	diagnosis	Diagnosis. This species occurs with Doryodes broui, Doryodes reineckei, and Doryodes tenuistriga. It is the largest species and with the most distinctive forewing pattern. It should not be confused with Doryodes tenuistriga, which has a narrow forewing stripe, or with Doryodes broui, which is smaller (forewing length: 13.0 - 15.5 mm) and also has a narrower forewing stripe. It can be distinguished from Doryodes reineckei by the breadth and distinctness of the longitudinal stripe; in females of Doryodes reineckei the forewing stripe is absent or pale gray, hardly contrasting with the ground color. The most distinguishing character of the male genitalia is the elongated terminal diverticulum of the vesica that projects ventrally. Females are similar to males but have longer, more pointed forewings and whiter hindwings. The female genitalia are the most robust of any Doryodes species. The corpus bursae has a bulge on the right side that is more pointed than in the other species, and the appendix bursae is quadrate with the posterior margin almost straight and heavily sclerotized.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
1FDD2C1F36D77EB228530156D5BA3964.taxon	description	Description. Forewing ground color in spring and summer specimens yellowish white to buff with gray streaking, hindwing white to whitish buff; forewing in fall and winter specimens darker with more brown shading, hindwing variably suffused with brown, especially along wing margin. Forewing length 14.5 - 18.0 mm (males), 17.5 - 20.0 mm (females), similar in length to those of Doryodes spadaria, but wings somewhat broader. Longitudinal stripe broader than in any other species in genus. Male genitalia with cornuti in variable patches on basal trunk of vesica, larger patch dorsal and most basal, two smaller patches of cornuti lateral and more distal (but one or both can be absent). Vesica with diverticulum 1 absent, its position represented by rounded curve of vesica posterior to swollen part of vesica after aedeagus; diverticulum 2 a rounded bulge on left side of vesica with large, slightly-serrated shark-fin-like cornutus; diverticula 3 a short quadrate pouch on right side almost opposite position of diverticulum 2; diverticulum 4 cone shaped, dorsolaterally on right, with bulbous spine-tipped cornutus at base near junction with elongated ventral lobed representing diverticulum 5. Female genitalia similar to those of the other species but more robust, and appendix bursae quadrate with posterior margin almost straight and heavily sclerotized.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
A8ABAFC70328E5FDFE9BE9364900D2E5.taxon	etymology	Etymology. We name this species after Vernon A. Brou, Jr. in recognition of his impressive and tireless efforts in collecting and researching the Lepidoptera of Louisiana.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
A8ABAFC70328E5FDFE9BE9364900D2E5.taxon	diagnosis	Diagnosis. This species occurs with Doryodes latistriga, Doryodes reineckei, and Doryodes tenuistriga in coastal salt marsh habitats from Alabama to southern Texas. Doryodes broui is superficially indistinguishable from Doryodes spadaria, but differs from it in male genitalia, barcodes, and occurs far to the west of the known range of Doryodes spadaria. It can be distinguished from Doryodes reineckei in having a sharply-defined longitudinal dark stripe on the forewing, and from Doryodes latistriga by the narrower forewing stripe in Doryodes broui. It is most likely to be confused with Doryodes tenuistriga, which typically is larger, but because of variation in both species, some specimens must be dissected or barcoded, for positive identification. The male genitalia of Doryodes broui are most similar to those of Doryodes latistriga, but the diverticula and cornuti are smaller, especially the terminal diverticulum (# 5), which is short and rounded, not elongated as in Doryodes latistriga.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
A8ABAFC70328E5FDFE9BE9364900D2E5.taxon	description	Description. Forewing length: 13.0 - 15.5 mm (males), 13.5 - 17.0 mm (females), Forewing buffy brown to whitish gray with faint buffy streaks, darker forms in colder months; longitudinal stripe dark brown, similar in width to that of Doryodes spadaria, narrower than for Doryodes latistriga, wider than for Doryodes tenuistriga. Male genitalia. Aedeagus 8 x as long as mesial width; vesica with dorsolateral toothed triangular cornutus on left side of basal part of vesica distal to end of aedeagus; a sclerotized plate in anterior 90 ° bend in vesica at position of ductus seminalis; posterior curve in vesica extended posteriorly into rounded diverticulum 1 with toothed preapical cornutus; diverticulum 2 on left side reduced to low bulge with large conical cornutus in middle; preapical posterior diverticulum (# 4) tapered with conical cornutus at distal base and also at base of short rounded apical diverticulum 5. The female genitalia are similar to those of Doryodes latistriga, but the appendix bursae is rounded posteriorly and only lightly sclerotized.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
7195AB76472F1DDEE2992F5C777A31A6.taxon	etymology	Etymology. The species is named for John P. Reinecke, a retired USDA entomologist who worked in Mississippi and developed insect organ culture techniques and detailed the anatomy of the hindgut of larval Lepidoptera.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
7195AB76472F1DDEE2992F5C777A31A6.taxon	diagnosis	Diagnosis. This species occurs with Doryodes latistriga, Doryodes broui, and Doryodes tenuistriga in coastal salt marsh habitats from Alabama to Texas. Doryodes bistrialis may occur nearby on the Gulf Coast, but inhabits inland longleaf pine savannas rather than salt marshes. Doryodes reineckei is immediately distinguished from other species of Doryodes in having a broader forewing in the male, the wing pattern appears faded and smudged, and the ventral margin of the longitudinal stripe appears to blend into the forewing ground color below it, not sharply defined as in other species. The male genitalia of Doryodes reineckei are smaller than those of Doryodes latistriga, but the characters of the vesica are definitive. In Doryodes reineckei the three patches of cornuti on the basal trunk of the vesica are rotated to the left, so the basal one is more lateral than dorsal; diverticulum 1 of the vesica projects dorsally and has a rooster-comb-like cornutus, whereas in Doryodes latistriga and Doryodes broui there is a tapered diverticulum (or rounded lobe) projecting posteriorly with a small spined cornutus on the side, and diverticulum 2 has no cornutus in Doryodes reineckei, whereas in the other two species there is a large cornutus at the base of the diverticulum. Females are immediately distinguishable by their whitish-gray forewing color with the longitudinal stripe is absent, or very faint, compared with the normal pattern seen in other species of Doryodes. Females have acutely pointed forewings.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
7195AB76472F1DDEE2992F5C777A31A6.taxon	description	Description. Forewing length 15.0 - 17.0 mm (males), 17.0 - 20.0 mm (females). Male noticeably broader winged than other species of Doryodes. Forewing pale brown to dark gray brown, darker forms in colder months; longitudinal dark stripe paler and less sharply defined than in other species, lower margin of stripe blending into darker ground color below it; wing margin more rounded than in other species. Male genitalia with valve similar in size to that of Doryodes spadaria but aedeagus much shorter (2.29 versus 2.97 mm), about 7.5 x as long as mesial width; basal trunk of the vesica with three patches of cornuti, two ventrolaterally on right side of aedeagus, largest one basal, dorsolaterally on left with multiple spinules on a heavily-sclerotized oval plate. Vesica above basal trunk T-shaped, elongated posteriorly-directed diverticulum 1 with a preapical rooster-comb-like cornutus on anterior surface, a small rounded diverticulum 2 on posterior right side opposite end of aedeagus, a sclerotized plate at 90 ° ventral angle in vesica next to ductus seminalis, and a ventral rounded apical diverticulum 5 with a conical cornutus on left side. Female genitalia disproportionally small, especially sclerotized plate in ventral wall of ductus bursae only 2.5 x as long as posterior width, plate tapered anteriorly, only 0.55 - 0.65 x as long as ductus bursae; ductus seminalis broad at base and gradually tapered, so appearing much wider than other species except Doryodes tenuistriga, and, like Doryodes tenuistriga, corpus bursae more rounded than in other species; appendix bursae a rounded lightly-sclerotized lobe with slightly bilobed posterior margin.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
9C157D5FB9BA43151C50EF0C7B51C349.taxon	materials_examined	Other material examined and distribution. USA: Louisiana, Texas.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
9C157D5FB9BA43151C50EF0C7B51C349.taxon	diagnosis	Diagnosis. Doryodes tenuistriga adults usually can be distinguished from other species in the genus by the narrow forewing stripes. The forewing length varies from 16.5 - 18.0 mm (males) and 15.5 - 18.5 mm (females). Some specimens of Doryodes broui can be similar, but the male and female genitalia are diagnostic. In the male genitalia the sclerotized areas on the dorsal and ventral margins of the valve end in broadly rounded processes well before the mainly membranous apex of the valve; in other species the process on the dorsal margin of the valve is pointed and extends to the apex of the valve, or beyond it, and the ventral process extends to, or almost to, the valve apex and is bluntly pointed or narrowly rounded. The aedeagus is short and wide, only about 5 x as long as wide. In the vesica the dorsal and ventral diverticula are similarly elongated, giving the vesica a T-shape; the vesica immediately posterior to the aedeagus has a rounded ventral lobe with a massive spine-covered sclerotized apical plate that is unique in the genus. In the female genitalia the ventral sclerotized plate on the ductus bursae is short and wide, about as long as wide and extending about 1 / 2 length of ductus; corpus bursae rounded, with protruding sclerotized lobe on posterior left opposite appendix bursae; appendix bursae rounded, lightly sclerotized.	en	Lafontaine, J. Donald, Sullivan, J. Bolling (2015): A revision of the genus Doryodes Guenee, 1857, with descriptions of six new species (Lepidoptera, Erebidae, Catocalinae, Euclidiini). ZooKeys 527: 3-30, DOI: http://dx.doi.org/10.3897/zookeys.527.6087, URL: http://dx.doi.org/10.3897/zookeys.527.6087
