taxonID	type	description	language	source
03D2230FB373F61489D4DF3EB70CE654.taxon	description	urn: lsid: zoobank. org: act: E 2 A 3 B 029 - 11 D 7 - 4 F 43 - A 091 - 62641 A 6 A 197 A Type-species: Sigalopella rufifemoralis sp. n. (here designated).	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB373F61489D4DF3EB70CE654.taxon	diagnosis	Diagnosis. Predominantly glossy black-bodied flies with long antennae, setal pile longer in male. Body length: 6.5 – 8.0 mm (male), 7.0 – 9.5 mm (female). Head relatively broad, wider than thorax; male eyes contiguous dorsomedially; male with single row of postocular macrosetae dorsally; face distinctively rounded to bulbous laterally, glabrous except for silver pubescence along eye margin, parafacial setae present, more extensive and elongate in male; antenna longer than head; scape narrow elongate, longer than flagellum; flagellum tapered cylindrical; extensive black setal pile along all antennal segments, especially scape and pedicel; palpus incrassate; prosternum without setal pile; metanepisternum without postspiracular setal patch; 1 – 3 dorsocentral macrosetae; one pair of scutellar macrosetae; posterior lateral surface of midcoxa lacking setal pile; velutum patches absent on femora and gonocoxites; legs with distinct short black setal pile, denser on tibia and tarsi; hind leg relatively longer than fore and mid legs; single anteroventral seta present apically on hind femur; wing mostly uniform dark infuscate, paler posteriorly, cell r 1 hyaline basally; costal vein circumambient around wing margin, wing cell m 3 open, vein R 1 with setae absent; abdominal tergite 2 with setae uniform in size medially; male genitalia with inner gonocoxal process well developed; gonocoxites with medial atrium lacking; hypandrium small; aedeagus with distiphallus narrow apically, dorsal apodeme of parameral sheath broadly triangular, ventral apodeme narrowly forked; female genitalia with acanthophorite setae as two sets (A 1 & A 2), A 1 usually enlarged; three spermathecae present, ducts joining to spermathecal sac duct; spermathecal sac present and simple in shape (not lobed). Included species. Sigalopella armeniaca sp. n., S. ammophila n. sp., S. nigrofemoralis sp. n. and S. rufifemoralis sp. n.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB373F61489D4DF3EB70CE654.taxon	etymology	Etymology. Sigalopella is derived from the Greek sigaloma – smooth, polished, and pellos – dusky, dark, for the dark polished appearance of members of this genus; gender is feminine.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB373F61489D4DF3EB70CE654.taxon	discussion	Comments. Sigalopella gen. n. is known only from central Chile (Fig. 14). Pachyrrhiza, Entesia and Mela- nothereva form a monophyletic clade of South American Agapophytinae called the Entesia genus-group (sensu Winterton et al., 2016), which also includes this new genus. We consider Sigalopella gen. n. as the likely sister to Pachyrrhiza. Amongst the South American Agapophytinae, these two genera share characteristics such as a protruding rounded parafacial with parafacial setae present, and the male post-ocular setae elongate and arranged in a single row dorsally. Sigalopella gen. n. can be easily recognised by the overall black coloration, narrow elongate antennae, absence of setal pile on the prosternum and incrassate palpi. Pachyrrhiza has a notably short and bulbous antennal scape (cf. elongate and narrow in Sigalopella gen. n.) and an abundance of macrosetae on the head and scutum. Most Pachyrrhiza have mottled or hyaline wings, never uniformly infuscate like in Sigalopella gen. n. Species of Sigalopella gen. n. are easily separated from each other using external characters such as colour and vestiture, as well as head and antennal shape; male and female genitalia is remarkably uniform amongst all members of the genus and is not useful for differentiating species in either sex.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB370F61289D4D8E2B7AAE50C.taxon	description	urn: lsid: zoobank. org: act: E 0330000 - D 674 - 484 B-A 696 - 9 FCE 64794 C 42	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB370F61289D4D8E2B7AAE50C.taxon	materials_examined	Type material — Holotype female, CHILE: Coquimbo Region: Choapa Province, 7 km N. Los Vilos, 13 – 20. xi. 1997, 215 m [- 31.851, - 70.495], Malaise in coastal dunes, M. E. Irwin, E. I. Schlinger (MEI 106826, MEUC). Paratypes. CHILE: Coquimbo Region: 2 females, same data except Fundo Agua Amarilla, Malaise in stable dunes, M. E. Irwin, 11 – 28. xii. 2003, 28. xii. 2003 – 8. i. 2004 [- 31.849, - 71.493] (MEI 147503 – 4, CSCA). Common name. Sand-loving sigalopella.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB370F61289D4D8E2B7AAE50C.taxon	diagnosis	Diagnosis. Face broadly rounded but not as protruding or bulbous anteriorly as in other species in the genus; scape noticeably thicker than base of flagellum; abdomen uniformly black; femora dark yellow, apex of hind femur with dark band; basal part of wing with yellowish suffusion of primary veins.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB370F61289D4D8E2B7AAE50C.taxon	description	Description. Body length ca. 7.0 mm. Head: Polished black, frons smooth and flat, polished with sparse dark setae laterally above antennae, extending ventrally onto parafacial; parafacial rounded, narrow silver pubescent stripe laterally along eye margin, extending along gena and more expansive on occiput; gena polished black with sparse black setae, overlain with silver pubescence; occiput black overlain with silver pubescence, occipital setae black, sparse dorsolaterally; maxillary palpus brown with extensive black setal pile; antennal scape slightly longer than head, thicker than pedicel and flagellum, numerous strong, erect black setae along entire scape, many with a length equal to width of pedicel (female), scape dark yellow basally and along ventral surface to approximately mid point; flagellum no more than four times length of pedicel. Thorax: Scutum and scutellum glossy black with short dark setal pile admixed with sparse glaucous pubescence, no vittae evident; pleuron black with silver pubescence, denser on lower half, relatively few fine, pale setae largely confined to anepisternum and katatergite; chaetotaxy: notopleural: 3, supra-alar: 1, post-alar 1, dorsocentral: 1, scutellar: 1; coxae polished black with sparse glaucous pubescence, macrosetae black, numerous on forecoxa; femora dark yellow, hind femur with dark brown band apically; tibia and tarsi dark yellow with blackish suffusion, darker distally on each segment and tarsomere; haltere cream-white coloured; wing dark infuscate, basal part of wing with yellowish suffusion on veins. Abdomen: glossy black with sparse, scattered, black setae, shorter dorsally, more elongate anteroventrally; segments 2 – 4 with posterior margin dark yellow and inter-segmental membrane white. Female terminalia: as per genus description.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB370F61289D4D8E2B7AAE50C.taxon	etymology	Etymology. Derived from the Greek: ammo — sand, and phil — loving. A feminine adjective.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB370F61289D4D8E2B7AAE50C.taxon	discussion	Comments. Only females are known for this species. Sigalopella ammophila sp. n. can be distinguished from all other species of the genus by the thickness of the scape and the face only being broadly rounded in the female rather than having any distinctive bulbous calli laterally below the antennal insertion. Although few in number, collection records indicate that it is essentially a coastal species (Fig. 15).	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB376F61089D4DAA5B641E680.taxon	description	(Figs 3 – 5, 14) urn: lsid: zoobank. org: act: A 56238 A 6 - E 565 - 4 C 16 - BC 9 E-AA 174342 CA 04	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB376F61089D4DAA5B641E680.taxon	materials_examined	Type material — Holotype female, CHILE: Valparaíso Region: Aconcagua, Rio Blanco [- 32.932, - 70.281], 1958, L. Peña (MEI 121486, CNC). Paratypes: CHILE: Valparaíso Region: 1 male, 4 females, Aconcagua, Piscicultura, Rio Blanco [- 32.932, - 70.281], 17. xi – 20. xi. 1958, L. Peña (MEI 1121302, MEI 121488, CSCA; MEI 121485, MEI 121487, MEI 121489, CNC). Common name. Bicoloured sigalopella.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB376F61089D4DAA5B641E680.taxon	diagnosis	Diagnosis. Parafacial with domed callus; scape longer than head, barely thicker than base of flagellum; abdomen black with segments 2 – 4 yellow laterally; femora yellow; wing uniformly dark infuscate.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB376F61089D4DAA5B641E680.taxon	description	Description. Body length ca. 8.5 mm. Head: Polished black; frons smooth and slightly rounded with slight transverse furrow immediately below ocellar tubercle (female), sparse dark setae laterally around antennal base, extending ventrally onto parafacial along eye margin; parafacial domed laterally, narrow silver pubescent stripe laterally along eye margin, extending along gena; gena with sparse black setae; occiput black, overlain with silver pubescence laterally on postocular ridge, occipital setae black, sparse dorsolaterally; maxillary palpus black with extensive black setal pile; antennal scape longer than head, barely thicker than pedicel and flagellum, yellow at base only, extensive erect, black setae, shorter in female, much longer in male. Thorax: Scutum and scutellum glossy black with dark setal pile admixed with sparse glaucous pubescence, faint dorsocentral vittae present anteriorly; pleuron black with silver pubescence, denser on lower half; dark setae on katatergite; chaetotaxy: notopleural: 4, supra-alar: 1, post-alar 1, dorsocentral: 1, scutellar: 1; coxae polished black with sparse glaucous pubescence; coxae with black macrosetae, more abundant on forecoxa that other coxae, especially in male; femora yellow, hind femur dark brown dorsoapically; fore- and midtibia yellow, black distally; hind tibia black; tarsi black, some tarsomeres yellow basally; haltere cream-white to yellow; wing dark infuscate, paler posteriorly. Abdomen: glossy black with sparse, scattered, black setae, shorter dorsally, more elongate anteroventrally; segments 2 – 3 dark yellow laterally. Male and female terminalia: as per genus description.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB376F61089D4DAA5B641E680.taxon	etymology	Etymology. From the Latin: armeniacus — apricot coloured, yellow with a perceptible mixing of red; refers to the colour of the distinctive abdominal colour of this species. A feminine adjective.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB376F61089D4DAA5B641E680.taxon	discussion	Comments. Sigalopella armeniaca sp. n. can be distinguished from all other species of the genus by the striking yellow-orange areas laterally on abdominal segments 2 and 3. In common with S. rufifemoralis sp. n. and S. nigrifemoralis sp. n., this species possesses inflated bulbous parafacial calli laterally as well as a narrower and more elongate antennal scape. In common with S. rufifemoralis sp. n. and S. ammophila sp. n., all femora are yellow. This species occurs inland in central Chile.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	description	(Figs 6 – 8, 14) urn: lsid: zoobank. org: act: 7 CAA 30 B 2 - A 149 - 4 AA 0 - 8 F 69 - BF 3 D 5693 A 688	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	materials_examined	Type material — Holotype male, CHILE: Santiago Region: Milipilla [Melipilla], El Bollenar, 2 km N Pichi Alhue [- 33.573, - 71.022], 100 m, 23. xi – 13. xii. 1998, Malaise trap nr. wash, G. Baria (MEI 121460, MEUC). Paratypes: CHILE: Santiago Region: 9 males, 12 females, same data as holotype, various dates (MEI 121461 – 7, 121476 – 82, CSCA; 121469 – 75, MEUC). Maule Region: 1 female, Curicó, Rio Teno, 1400 – 1600 m, 24. i – 26. i. 1968, L. E. Peña (MEI 121484, MEUC); 1 female, Curicó, Estero la Jaula, [- 35.083, - 70.8], i. 1964, L. E. Peña, Nothofagus (MEI 121483, CNC). Common name. Black-thighed sigalopella.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	diagnosis	Diagnosis. Face with domed callus laterally; scape longer than head, barely thicker than base of flagellum; abdomen uniformly black except for pale intersegmental membranes 2 – 4; femora black; wing uniformly dark infuscate.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	description	Description. Body length ca. 6.5 – 7.5 mm. Head: Polished black; frons smooth and slightly rounded with irregular transverse furrows (female), medially divided above antennal bases in male, sparse dark setae laterally around antennal base, extending ventrally onto parafacial; parafacial domed laterally, narrow silver pubescent stripe laterally along eye margin, extending along gena; gena with sparse black setae; occiput black, overlain with silver pubescence along postocular ridge, occipital setae black, sparse dorsolaterally; maxillary palpus black with extensive black setal pile; antennal scape longer than head, barely thicker than pedicel and flagellum, yellow at base only, extensive erect, black setae, shorter in female, much longer in male. Thorax: Scutum and scutellum glossy black with dark setal pile admixed with sparse glaucous pubescence, faint dorsocentral vittae sometimes barely evident anteriorly in female; pleuron black with silver pubescence, denser on lower half; dark setae on katatergite; chaetotaxy: notopleural: 4, supra-alar: 1, post-alar 1, dorsocentral: 1, scutellar: 1; coxae polished black with glaucous pubescence, macrosetae black, more abundant on fore coxa, especially in male; femora black; fore- and mid tibia yellow; hind tibia yellow with extensive black suffusion; tarsi black, most tarsomeres yellow basally; haltere cream-white to yellow; wing dark infuscate, paler posteriorly. Abdomen: glossy black with sparse, scattered, black setae, shorter dorsally and more elongate anteroventrally; intersegmental membranes 2 – 4 white, more pronounced laterally. Male and female terminalia: as per genus description.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	etymology	Etymology. Derived from the Latin: niger — black, and femur — thigh, referring to the colouration of the legs. A noun in apposition.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	discussion	Comments. The uniformly black femora easily differentiate this species from others in the genus. This species is found in central Chile. Sigalopella rufifemoralis sp. n. (Figs 9 – 14) urn: lsid: zoobank. org: act: F 68 A 3 A 1 E- 8 D 3 C- 4826 - AC 34 - 672 DAF 8325 FD	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	materials_examined	Type material — Holotype male, CHILE: Valparaiso Region: Quillota Prov., Parque Nacional Campanas, Palmas de Ocoa [- 32.932, - 71.078], 215 m, 2. i. 2000, malaise trap, M. E. Irwin, E. I. Schlinger, in hillside draw (MEI 121494, MEUC). Paratypes: CHILE: Valparaiso Region: 20 males, 7 females, [various dates], same data as holotype (MEI 121403 – 9, 121414 – 9, CSCA; MEI 121490 – 500, 1214501 – 503, MEUC). Coquimbo Region: 2 males, El Naranjo, Tilama, [- 32.083, - 71.167], iii. 1968, J. Molina (MEI 121504, 121410, MEUC); 3 females, Santiago Region: Metropolitana de Santiago, La Rinconada de Maipú, Quebrada de la Plata [- 33.517, - 70.783], 510 m, xii. 1966 [various dates], malaise trap, M. E. Irwin (MEI 127066, 121507, CSCA; MEI 127067, CASC). Common name. Yellow-thighed sigalopella.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	diagnosis	Diagnosis. Face with domed calli laterally; scape longer than head, barely thicker than base of flagellum; abdomen uniformly black except for pale intersegmental membranes 2 – 4; femora yellow; wing uniformly dark infuscate. This species is found in central Chile.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	description	Description. Body length ca. 6.0 – 7.5 mm. Head: Polished black; frons smooth and slightly rounded with a transverse furrow dorsally (female), medially divided above antennal bases in male, sparse dark setae laterally around antennal base, extending ventrally onto parafacial; parafacial domed laterally, narrow silver pubescent stripe laterally along eye margin, extending along gena; gena with sparse black setae; occiput black, overlain with silver pubescence along postocular ridge, occipital setae black, sparse dorso-laterally; maxillary palpus black with extensive black setal pile; antennal scape longer than head, barely thicker than pedicel and flagellum, yellow ventrally at base only, extensive erect, black setae, shorter in female, much longer in male. Thorax: Scutum and scutellum glossy black with dark setal pile admixed with sparse glaucous pubescence, faint dorsocentral vittae sometimes barely evident anteriorly in female; pleuron black with silver pubescence; dark setae on katatergite; chaetotaxy: notopleural: 4, supra-alar: 1, post-alar 1, dorsocentral: 1, scutellar: 1; coxae polished black with glaucous pubescence, macrosetae black, more abundant on fore coxa, especially in male; femora bright yellow, sometimes with dark spot dorsoapically, male with dark setae more abundant and longer in places; tibiae black, fore and mid tibiae suffused with yellow in female; tarsi black, mid basitarsus yellow basally; haltere cream-white to yellow; wing dark infuscate, paler posteriorly. Abdomen: glossy black with sparse, scattered, black setae, shorter dorsally and more elongate anteroventrally; intersegmental membranes 2 – 4 sometimes white, more pronounced laterally. Male and female terminalia: as per genus description.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	etymology	Etymology. Derived from the Latin: rufus – reddish, and femoralis – pertaining to the thigh, referring to the colouration of the legs.	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
03D2230FB374F61B89D4D811B7F6E0EB.taxon	discussion	Comments. The domed parafacial calli, narrow scape, black abdomen and yellow femora differentiate this species from others in the genus. This species is found in central Chile (Fig. 16).	en	Irwin, Michael E., Winterton, Shaun L. (2020): A new stiletto fly genus from South America (Diptera: Therevidae: Agapophytinae). Zootaxa 4751 (2): 276-290, DOI: 10.11646/zootaxa.4751.2.4
