taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03D25556FFA53131FC2CFA6BFD48DCCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351568/files/figure.png	https://doi.org/10.5281/zenodo.5351568	Fig. 1. Arachnothelphusa sarang, new species. A, male (12.1 × 9.9 mm) (ZRC 2020.0351), Batu Rusa cave, Bukit Sarang, Bintulu, Sarawak, Malaysia; B, female (12.9 × 10.4 mm) (ZRC 2020.0351), Batu Kelelut, Bukit Sarang, Bintulu, Sarawak, Malaysia, specimen; C–E, paratype male (18.7 × 15.3 mm) (ZRC 2020.0099), Bukit Sarang, Bintulu, Sarawak, Malaysia; F, paratype female (15.8 × 11.8 mm) (ZRC 2020.0099), Bukit Sarang, Bintulu, Sarawak, Malaysia. A, B, photographed in situ; C, F, overall dorsal habitus; D, ventral view of cephalothorax; E, frontal view of cephalothorax and chelae. Photographs: Tan Heok Hui.	Fig. 1. Arachnothelphusa sarang, new species. A, male (12.1 × 9.9 mm) (ZRC 2020.0351), Batu Rusa cave, Bukit Sarang, Bintulu, Sarawak, Malaysia; B, female (12.9 × 10.4 mm) (ZRC 2020.0351), Batu Kelelut, Bukit Sarang, Bintulu, Sarawak, Malaysia, specimen; C–E, paratype male (18.7 × 15.3 mm) (ZRC 2020.0099), Bukit Sarang, Bintulu, Sarawak, Malaysia; F, paratype female (15.8 × 11.8 mm) (ZRC 2020.0099), Bukit Sarang, Bintulu, Sarawak, Malaysia. A, B, photographed in situ; C, F, overall dorsal habitus; D, ventral view of cephalothorax; E, frontal view of cephalothorax and chelae. Photographs: Tan Heok Hui.	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
03D25556FFA53131FC2CFA6BFD48DCCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351570/files/figure.png	https://doi.org/10.5281/zenodo.5351570	Fig. 2. Arachnothelphusa sarang, new species. A–E, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), Bukit Sarang, Bintulu, Sarawak, Malaysia; F, G, paratype female (19.8 × 15.8 mm) (ZRC 2020.0100), Bukit Sarang, Bintulu, Sarawak, Malaysia. A, F, dorsal view of habitus; B, G, ventral view of carapace showing pleon; C, frontal view of cephalothorax; D, outer view of left chela; E, outer view of right fourth ambulatory leg. Scales: A, B, C, F, G = 10 mm; D, E = 5 mm.	Fig. 2. Arachnothelphusa sarang, new species. A–E, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), Bukit Sarang, Bintulu, Sarawak, Malaysia; F, G, paratype female (19.8 × 15.8 mm) (ZRC 2020.0100), Bukit Sarang, Bintulu, Sarawak, Malaysia. A, F, dorsal view of habitus; B, G, ventral view of carapace showing pleon; C, frontal view of cephalothorax; D, outer view of left chela; E, outer view of right fourth ambulatory leg. Scales: A, B, C, F, G = 10 mm; D, E = 5 mm.	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
03D25556FFA53131FC2CFA6BFD48DCCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351572/files/figure.png	https://doi.org/10.5281/zenodo.5351572	Fig. 3. Arachnothelphusa sarang, new species, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), limestone cave, Bukit Sarang, Bintulu, Sarawak, Malaysia. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, ventral view of left G1; D, ventral view of distal part of left G1; E, dorsal view of left G2. Scales = 0.5 mm.	Fig. 3. Arachnothelphusa sarang, new species, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), limestone cave, Bukit Sarang, Bintulu, Sarawak, Malaysia. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, ventral view of left G1; D, ventral view of distal part of left G1; E, dorsal view of left G2. Scales = 0.5 mm.	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
03D25556FFA53131FC2CFA6BFD48DCCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351574/files/figure.png	https://doi.org/10.5281/zenodo.5351574	Fig. 4. Carapace morphology of Arachnothelphusa species. A, A. sarang, new species, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), limestone cave, Bukit Sarang, Bintulu, Sarawak; B, A. merarapensis Grinang, Pui & Ng, 2015, holotype male (22.5 × 16.8 mm) (ZRC 2016.0297), Merarap Hot Spring, Lawas, Sarawak; C, A. terrapes Ng, 1991, male (30.8 × 20.5 mm) (ZRC 2017.1205), Danum Valley, Lahad Datu, Sabah; D, A. kadamaiana (Borradaile, 1900), male (20.1 × 14.9 mm) (SMF 4282), Kadamian River, Sabah; E, A. melanippe (De Man, 1899), paratype female (21.4 × 16.7 mm) (RMNH D1303b), Mt. Liang Koebeng, Kalimantan, Indonesia. E after Ng (1991).	Fig. 4. Carapace morphology of Arachnothelphusa species. A, A. sarang, new species, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), limestone cave, Bukit Sarang, Bintulu, Sarawak; B, A. merarapensis Grinang, Pui & Ng, 2015, holotype male (22.5 × 16.8 mm) (ZRC 2016.0297), Merarap Hot Spring, Lawas, Sarawak; C, A. terrapes Ng, 1991, male (30.8 × 20.5 mm) (ZRC 2017.1205), Danum Valley, Lahad Datu, Sabah; D, A. kadamaiana (Borradaile, 1900), male (20.1 × 14.9 mm) (SMF 4282), Kadamian River, Sabah; E, A. melanippe (De Man, 1899), paratype female (21.4 × 16.7 mm) (RMNH D1303b), Mt. Liang Koebeng, Kalimantan, Indonesia. E after Ng (1991).	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
03D25556FFA53131FC2CFA6BFD48DCCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351576/files/figure.png	https://doi.org/10.5281/zenodo.5351576	Fig. 5. Gonopod morphology of Arachnothelphusa species. A–E, A. merarapensis Grinang, Pui & Ng, 2015, holotype, male (22.5 × 16.8 mm) (ZRC 2016.0297), Merarap Hot Spring, Lawas, Sarawak; F–J, A. terrapes Ng, 1991, holotype, male (17.6 × 13.3 mm) (ZRC 1992.7918), Danum Valley, Lahad Datu, Sabah; K–N, A. kadamaiana (Borradaile, 1900), male (20.1 × 14.9 mm) (SMF 4282), Kadamian River, Sabah; O–S, A. melanippe (De Man, 1899), lectotype, male (18.9 × 14.4 mm) (RMNH D1303a), Mt. Liang Koebeng, Kalimantan, Borneo. C, F, K, O, ventral view of left G1; D, G, L, P, ventral view of distal part of left G1; A, H, Q, dorsal view of left G1; B, I, M, R, ventral view of distal part of left G1; E, J, N, S, left G2. A–E, K–N after Grinang et al. (2015); F–J, O–S after Ng (1991).	Fig. 5. Gonopod morphology of Arachnothelphusa species. A–E, A. merarapensis Grinang, Pui & Ng, 2015, holotype, male (22.5 × 16.8 mm) (ZRC 2016.0297), Merarap Hot Spring, Lawas, Sarawak; F–J, A. terrapes Ng, 1991, holotype, male (17.6 × 13.3 mm) (ZRC 1992.7918), Danum Valley, Lahad Datu, Sabah; K–N, A. kadamaiana (Borradaile, 1900), male (20.1 × 14.9 mm) (SMF 4282), Kadamian River, Sabah; O–S, A. melanippe (De Man, 1899), lectotype, male (18.9 × 14.4 mm) (RMNH D1303a), Mt. Liang Koebeng, Kalimantan, Borneo. C, F, K, O, ventral view of left G1; D, G, L, P, ventral view of distal part of left G1; A, H, Q, dorsal view of left G1; B, I, M, R, ventral view of distal part of left G1; E, J, N, S, left G2. A–E, K–N after Grinang et al. (2015); F–J, O–S after Ng (1991).	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
03D25556FFA53131FC2CFA6BFD48DCCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351578/files/figure.png	https://doi.org/10.5281/zenodo.5351578	Fig. 6. Live colours of Arachnothelphusa rhadamanthysi from Gomantong limestone cave in Sabah, specimen not collected. A, male, from outside of the cave; B, inside of cave. Photographs: Keith Christenson.	Fig. 6. Live colours of Arachnothelphusa rhadamanthysi from Gomantong limestone cave in Sabah, specimen not collected. A, male, from outside of the cave; B, inside of cave. Photographs: Keith Christenson.	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
03D25556FFA03131FCA9FDFFFA0BDCC2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351568/files/figure.png	https://doi.org/10.5281/zenodo.5351568	Fig. 1. Arachnothelphusa sarang, new species. A, male (12.1 × 9.9 mm) (ZRC 2020.0351), Batu Rusa cave, Bukit Sarang, Bintulu, Sarawak, Malaysia; B, female (12.9 × 10.4 mm) (ZRC 2020.0351), Batu Kelelut, Bukit Sarang, Bintulu, Sarawak, Malaysia, specimen; C–E, paratype male (18.7 × 15.3 mm) (ZRC 2020.0099), Bukit Sarang, Bintulu, Sarawak, Malaysia; F, paratype female (15.8 × 11.8 mm) (ZRC 2020.0099), Bukit Sarang, Bintulu, Sarawak, Malaysia. A, B, photographed in situ; C, F, overall dorsal habitus; D, ventral view of cephalothorax; E, frontal view of cephalothorax and chelae. Photographs: Tan Heok Hui.	Fig. 1. Arachnothelphusa sarang, new species. A, male (12.1 × 9.9 mm) (ZRC 2020.0351), Batu Rusa cave, Bukit Sarang, Bintulu, Sarawak, Malaysia; B, female (12.9 × 10.4 mm) (ZRC 2020.0351), Batu Kelelut, Bukit Sarang, Bintulu, Sarawak, Malaysia, specimen; C–E, paratype male (18.7 × 15.3 mm) (ZRC 2020.0099), Bukit Sarang, Bintulu, Sarawak, Malaysia; F, paratype female (15.8 × 11.8 mm) (ZRC 2020.0099), Bukit Sarang, Bintulu, Sarawak, Malaysia. A, B, photographed in situ; C, F, overall dorsal habitus; D, ventral view of cephalothorax; E, frontal view of cephalothorax and chelae. Photographs: Tan Heok Hui.	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
03D25556FFA03131FCA9FDFFFA0BDCC2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351570/files/figure.png	https://doi.org/10.5281/zenodo.5351570	Fig. 2. Arachnothelphusa sarang, new species. A–E, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), Bukit Sarang, Bintulu, Sarawak, Malaysia; F, G, paratype female (19.8 × 15.8 mm) (ZRC 2020.0100), Bukit Sarang, Bintulu, Sarawak, Malaysia. A, F, dorsal view of habitus; B, G, ventral view of carapace showing pleon; C, frontal view of cephalothorax; D, outer view of left chela; E, outer view of right fourth ambulatory leg. Scales: A, B, C, F, G = 10 mm; D, E = 5 mm.	Fig. 2. Arachnothelphusa sarang, new species. A–E, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), Bukit Sarang, Bintulu, Sarawak, Malaysia; F, G, paratype female (19.8 × 15.8 mm) (ZRC 2020.0100), Bukit Sarang, Bintulu, Sarawak, Malaysia. A, F, dorsal view of habitus; B, G, ventral view of carapace showing pleon; C, frontal view of cephalothorax; D, outer view of left chela; E, outer view of right fourth ambulatory leg. Scales: A, B, C, F, G = 10 mm; D, E = 5 mm.	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
03D25556FFA03131FCA9FDFFFA0BDCC2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351574/files/figure.png	https://doi.org/10.5281/zenodo.5351574	Fig. 4. Carapace morphology of Arachnothelphusa species. A, A. sarang, new species, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), limestone cave, Bukit Sarang, Bintulu, Sarawak; B, A. merarapensis Grinang, Pui & Ng, 2015, holotype male (22.5 × 16.8 mm) (ZRC 2016.0297), Merarap Hot Spring, Lawas, Sarawak; C, A. terrapes Ng, 1991, male (30.8 × 20.5 mm) (ZRC 2017.1205), Danum Valley, Lahad Datu, Sabah; D, A. kadamaiana (Borradaile, 1900), male (20.1 × 14.9 mm) (SMF 4282), Kadamian River, Sabah; E, A. melanippe (De Man, 1899), paratype female (21.4 × 16.7 mm) (RMNH D1303b), Mt. Liang Koebeng, Kalimantan, Indonesia. E after Ng (1991).	Fig. 4. Carapace morphology of Arachnothelphusa species. A, A. sarang, new species, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), limestone cave, Bukit Sarang, Bintulu, Sarawak; B, A. merarapensis Grinang, Pui & Ng, 2015, holotype male (22.5 × 16.8 mm) (ZRC 2016.0297), Merarap Hot Spring, Lawas, Sarawak; C, A. terrapes Ng, 1991, male (30.8 × 20.5 mm) (ZRC 2017.1205), Danum Valley, Lahad Datu, Sabah; D, A. kadamaiana (Borradaile, 1900), male (20.1 × 14.9 mm) (SMF 4282), Kadamian River, Sabah; E, A. melanippe (De Man, 1899), paratype female (21.4 × 16.7 mm) (RMNH D1303b), Mt. Liang Koebeng, Kalimantan, Indonesia. E after Ng (1991).	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
03D25556FFA03131FCA9FDFFFA0BDCC2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351572/files/figure.png	https://doi.org/10.5281/zenodo.5351572	Fig. 3. Arachnothelphusa sarang, new species, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), limestone cave, Bukit Sarang, Bintulu, Sarawak, Malaysia. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, ventral view of left G1; D, ventral view of distal part of left G1; E, dorsal view of left G2. Scales = 0.5 mm.	Fig. 3. Arachnothelphusa sarang, new species, holotype male (20.4 × 14.7 mm) (ZRC 2020.0098), limestone cave, Bukit Sarang, Bintulu, Sarawak, Malaysia. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, ventral view of left G1; D, ventral view of distal part of left G1; E, dorsal view of left G2. Scales = 0.5 mm.	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
03D25556FFA03131FCA9FDFFFA0BDCC2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5351576/files/figure.png	https://doi.org/10.5281/zenodo.5351576	Fig. 5. Gonopod morphology of Arachnothelphusa species. A–E, A. merarapensis Grinang, Pui & Ng, 2015, holotype, male (22.5 × 16.8 mm) (ZRC 2016.0297), Merarap Hot Spring, Lawas, Sarawak; F–J, A. terrapes Ng, 1991, holotype, male (17.6 × 13.3 mm) (ZRC 1992.7918), Danum Valley, Lahad Datu, Sabah; K–N, A. kadamaiana (Borradaile, 1900), male (20.1 × 14.9 mm) (SMF 4282), Kadamian River, Sabah; O–S, A. melanippe (De Man, 1899), lectotype, male (18.9 × 14.4 mm) (RMNH D1303a), Mt. Liang Koebeng, Kalimantan, Borneo. C, F, K, O, ventral view of left G1; D, G, L, P, ventral view of distal part of left G1; A, H, Q, dorsal view of left G1; B, I, M, R, ventral view of distal part of left G1; E, J, N, S, left G2. A–E, K–N after Grinang et al. (2015); F–J, O–S after Ng (1991).	Fig. 5. Gonopod morphology of Arachnothelphusa species. A–E, A. merarapensis Grinang, Pui & Ng, 2015, holotype, male (22.5 × 16.8 mm) (ZRC 2016.0297), Merarap Hot Spring, Lawas, Sarawak; F–J, A. terrapes Ng, 1991, holotype, male (17.6 × 13.3 mm) (ZRC 1992.7918), Danum Valley, Lahad Datu, Sabah; K–N, A. kadamaiana (Borradaile, 1900), male (20.1 × 14.9 mm) (SMF 4282), Kadamian River, Sabah; O–S, A. melanippe (De Man, 1899), lectotype, male (18.9 × 14.4 mm) (RMNH D1303a), Mt. Liang Koebeng, Kalimantan, Borneo. C, F, K, O, ventral view of left G1; D, G, L, P, ventral view of distal part of left G1; A, H, Q, dorsal view of left G1; B, I, M, R, ventral view of distal part of left G1; E, J, N, S, left G2. A–E, K–N after Grinang et al. (2015); F–J, O–S after Ng (1991).	2021-01-12	Grinang, Jongkar;Ng, Peter K. L.		Zenodo	biologists	Grinang, Jongkar;Ng, Peter K. L.			
