taxonID	type	description	language	source
03D39C15FFDD3F15FE60C6669EA35CEA.taxon	description	New records. CHINA: SHAANXI: 18 spec. (ASC, MSC): ‘ (S-Shaanxi) Qinling Shan mount. range W pass on road X’ian- Shagoujie, 45 km SSW X’ian 33 ° 52 ′ N 108 ° 46 ′ E 2675 m (N-slope / Abies, Betula, Larix) / subalp. meadows / along road) 25. VII. 2001 Wrase, 20 [= sample number] ’; 16 spec. (ASC, MSC): same data, ‘ N-slope / Abies, Betula, Larix, Rhododendron / subalpine meadows (sifted), C 01 - 20 [sample ID], leg. M. Schülke’; 12 spec. (ASC, MSC): same data, ‘ 26. VII. 2001, C 0 - 20 A [sample ID], leg. M. Schülke’.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDD3F15FE60C6669EA35CEA.taxon	discussion	Comment. These are additional specimens from the type locality. Quedius io is the dominant Quedius species in the habitat. Quedius (Microsaurus) duh Smetana, 2001 is another species living in the same habitat.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDD3F15FE63C7FB9D725B94.taxon	description	New records. CHINA: YUNNAN: 2 spec. (MSC): ‘ (Yunnan) Dali Bai Aut. Pref., Diancang Shan E pass, 43 km NW Dali, 2700 m (secondary pine forest, litter, moss sifted) 25 ° 59 ′ 50 ″ N / 100 ° 00 ′ 30 ″ E 23. VIII. 2009, D. W. Wrase, 02 [sammple no.] ’; 4 spec. (ASC, MSC): same data, ‘ litter, moss and mushrooms sifted, leg. M. Schülke, CH 09 - 02 [= sample ID] ’; 2 spec. (MSC): same data, ‘ 25 ° 59 ′ 33.5 ″ N 99 ° 52 ′ 12.5 ″ E 3104 m, pasture & shrubs, litter, moss and mushrooms sifted, 23. V. III. 2009, leg. M. Schülke, CH 09 - 01 [sample ID] ’.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDD3F15FE63C7FB9D725B94.taxon	discussion	Comment. This is the second record of this species from Diancang Shan. Quedius li is at present known only from the Diancang Shan.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDD3F10FE67C1319CEF5DA2.taxon	description	(Figs. 1 – 7)	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDD3F10FE67C1319CEF5DA2.taxon	materials_examined	Type locality. People’s Republic of China, Yunnan, Nujiang Lieu Pref., Gaoligong Shan, W ‘ Cloud Pass’ 24 km NW Liuku, 25 ° 59 ′ 02 ″ N 98 ° 39 ′ 56.5 ″ E, 2940 m a. s. l. Type material. HOLOTYPE: J (MSC, to be deposited in Naturhistorisches Museum in Berlin), ‘ CHINA: Yunnan Nujiang Lieu Pref., Gaoligong Shan, W “ Cloud Pass ” 24 km NW Liuku, 25 ° 59 ′ 02 ″ N 98 ° 39 ′ 56.5 ″ E, 2940 m, small cleft, wet moss & litter sifted, 3. IX. 2009, leg. M. Schülke [CH 09 - 24] ’. ALLOTYPE: ♀ (ASC), same data as holotype. PARATYPES: 8 JJ 2 ♀♀ (ASC, MSC), same data as holotype.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDD3F10FE67C1319CEF5DA2.taxon	description	Description. Head piceous-black, pronotum piceous, somewhat paler laterally, elytra brunneous to brunneopiceous; head and pronotum with faint metallic bronze lustre, abdomen iridescent, dark brunneous to brunneopiceous with apical margins of tergites more or less paler; both maxillary and labial palpi pale testaceous, antennae testaceous, vaguely darkened toward apex, legs uniformly testaceous. Head rounded, wider than long (ratio 1.22); eyes very large and convex, tempora very short, considerably shorter than length of eyes seen from above (ratio 0.15); no additional setiferous punctures between anterior frontal punctures; posterior frontal puncture touching posteriomedian margin of eye, one puncture between it and posterior margin of head; temporal puncture small, touching posterior margin of eye; surface of head with fine, dense microsculpture of transverse and oblique waves gradually changing into meshes on middle of clypeus. Antenna moderately long, segments 2 and 3 subequal in length, segments 4 – 7 longer than wide, gradually becoming shorter, segments 8 – 10 about as long as wide, segment 11 as long as two preceding segments combined. Pronotum as long as wide, to vaguely wider than long (ratio 1.08), widely rounded basally, slightly narrowed anteriad, evenly transversely convex; dorsal rows each with three punctures; sublateral rows variable, each usually with three punctures with posterior puncture situated slightly after level of large lateral puncture, or with only two punctures with posterior puncture situated slightly before level of large lateral puncture; surface of pronotum with microsculpture of transverse waves similar to those on head. Scutellum with 5 – 9 punctures, surface with microsculpture of rudimentary striae. Elytra short, at suture markedly shorter (ratio 0.71), at sides shorter (ratio 0.81) than pronotum at midline; punctation moderately dense, rather coarse, asperate; transverse interspaces between punctures about as large as diameters of punctures; pubescence golden-yellowish; surface between punctures with microscopic irregularities. Wings each reduced to non-functional, narrow stump. Abdomen with tergite 7 (fifth visible) without whitish apical seam of palisade setae; tergite 2 (in front of first fully visible tergite) with a few fine punctures; punctation of tergites markedly finer than that on elytra, dense, rather evenly covering each tergite, becoming in general sparser toward apex of abdomen; pubescence golden-yellowish; surface between punctures with excessively fine microsculpture of transverse striae. Male. First four segments of front tarsus dilated, subbilobed, each with tenent setae ventrally, segment two narrower than apex of tibia; segment 4 narrower than preceding segments. Sternite 8 with two or three long setae on each side, apical margin with moderately wide and deep, obtusely triangular medioapical emargination, small triangular area before emargination flatteded and smooth (Fig. 1). Genital segment with tergite 10 narrow, evenly narrowed toward arcuate apex, setose as in Fig. 2; sternite 9 with narrow basal portion, apical portion emarginate apically, with a pair of differentiated apical and subapical setae, otherwise moderately densely setose (Fig. 3). Aedeagus (Figs. 4 – 6) small, median lobe subparallel sided in middle portion, anteriorly narrowed into slightly asymmetrical apical portion with narrowly arcuate apex, on face adjacent to paramere, when paramere removed, with rather long median carina forming a fine hook in lateral view (Fig. 5). Apical portion of median lobe quite characteristic in lateral view (Fig. 5). Paramere (Figs. 4, 6) moderately long, slightly fusiform, with narrowly arcuate apex not reaching apex of median lobe; four fine setae at apical margin, median setae somewhat longer than lateral ones, two similar setae at each lateral margin below apex; sensory peg setae on underside of paramere numerous, forming two irregular, fairly long rows (Fig. 6). Female. First four segments of front tarsus simple, not dilated, without tenent setae ventrally. Tergite 10 of genital segment small, narrow, evenly narrowed toward arcuate apex, with several long setae at and near apex, otherwise only with a few fine setae (Fig. 7). Length 4.8 – 5.5 mm.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDD3F10FE67C1319CEF5DA2.taxon	etymology	Etymology. The specific epithet is partial name of the prefecture the species occurs in, a noun in apposition. Bionomics. The specimens of the original series were taken by sifting wet moss and litter at an elevation just below 3000 m. Geographical distribution. Quedius nujiang sp. nov. is at present known only from the type locality in Gaoligong Shan, west of Salween river, in westernmost Yunnan. Recognition and comments. Quedius nujiang sp. nov. is well characterized among the brachypterous species of the muscicola - group, in addition to the shape of the aedoeagus, by the relatively large size, the uniformly testaceous legs, the rather coarse punctation of elytra, and by the golden-yellow pubescence of the elytra and the abdomen. It only may be confused with Q. angustiarum sp. nov., but the latter differs by the entirely different shape of the aedoeagus, and by a few subtle external differences (see under Q. angustiarum sp. nov.).	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFD83F11FE72C6C29CCA5CC2.taxon	description	(Figs. 8 – 14)	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFD83F11FE72C6C29CCA5CC2.taxon	materials_examined	Type locality. People’s Republic of China, Yunnan, Nujiang Lisu Pref., Gaoligong Shan, ‘ Cloud Pass’ 21 km NW Liuku, 25 ° 58 ′ 21 ″ N 98 ° 41 ′ 01 ″ E, 3150 m a. s. l. Type material. HOLOTYPE: J (MSC, to be deposited in Naturhistorisches Museum in Berlin), ‘ CHINA (Yunnan) Nujiang Lisu Pref., Gaoligong Shan, “ Cloud Pass ” 3150 m, 21 km NW Liuku (shrubs, Vaccinium, bamboo, litter sifted) 25 ° 58 ′ 21 ″ N / 98 ° 41 ′ 01 ″ E 2. IX. 2009 D. W. Wrase [22 A] ’. ALLOTYPE: ♀ (ASC), same data as holotype. PARA- TYPES: 1 J, 2 ♀♀ (ASC, MSC), same data as holotype; 1 J 2 ♀♀ (ASC, MSC), same data as holotype but ‘ shrubs & bamboo, litter sifted, leg. M. Schülke [CH 09 - 22]. ’	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFD83F11FE72C6C29CCA5CC2.taxon	description	Description. In all characters quite similar to Q. nujiang sp. nov., but different by a few subtle external characters, and by the entirely differently shaped aedoeagus. Average size smaller, body shape slenderer. Antenna slenderer and slightly shorter. Metallic bronze lustre on head and pronotum markedly more pronounced. Microsculpture on head and pronotum somewhat less dense and coarser. Punctation of elytra similar, but finer and denser, transverse interspaces between punctures slightly smaller than diameters of punctures, surface between punctures without microscopic irregularities, elytra therefore appearing shinier. Punctation of abdominal tergites similar, but denser. Male. First four segments of front tarsus similar to those of Q. nujiang sp. nov., but slightly less dilated. Sternite 8 with two long setae on each side, medioapical emargination similar to that of Q. nujiang sp. nov., but deeper (Fig. 8). Genital segment with tergite 10 wider, with more numerous long setae on apical portion (Fig. 9); sternite 9 with basal portion narrower and longer, apical portion narrowly arcuate at apex, with a pair of apical and subapical long setae, otherwise only sparingly setose (Fig. 10). Aedoeagus (Figs. 11 – 13) narrow, elongate, median lobe subparallel-sided in middle portion, anteriorly slightly dilated and then narrowed into rather long apical portion with narrowly arcuate apex, on face adjacent to paramere, when paramere removed, with minute median carina situated far below apex of median lobe, forming a minute hook in lateral view (Fig. 12). Apical portion of median lobe in lateral view quite different from that of Q. nujiang sp. nov. (Fig. 12). Paramere very long, parallel-sided in middle portion, with narrowly arcuate apex not reaching apex of median lobe; four very fine setae at apical margin, median setae somewhat longer than lateral ones, two similar setae at each lateral margin below apex; sensory peg setae on underside of paramere numerous, forming two fairly regular, rather long rows (Fig. 13). Female. First four segments of front tarsus simple, not dilated. Tergite 10 of genital segment of different shape than that of Q. nujiang sp. nov., and with different setation (Fig. 14). Length 4.7 – 5.0 mm.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFD83F11FE72C6C29CCA5CC2.taxon	etymology	Etymology. The specific epithet is the genitive form of the Latin noun angustiae, - arum, (mountain pass), as noun in apposition. It refers to the occurrence of this species in ‘ Cloud Pass’ of the Gaoligong Shan. Bionomics. The specimens of the type series were taken by sifting litter under shrubs, Vaccinium and bamboo growths at an elevation 3150 m. Quedius angustiarum sp. nov. occurs in the same area of the Gaoligong Shan as Q. nujiang sp. nov., but in a different habitat at higher elevation. Geographical distribution. Quedius angustiarum sp. nov. is at present known only from the type locality in Gaoligong Shan, west of Salween river, in westernmost Yunnan. Recognition and comments. Quedius angustiarum sp. nov. may only be confused with Q. nujiang sp. nov., but it may be distinguished by the characters outlined above, particularly by the entirely different shape of the aedoeagus. The aedoeagus of Quedius angustiarum sp. nov. is similar to that of Q. microsauroides Smetana, 2008 from Tian Shan range in Xinjiang, but the latter differs by several external characters (e. g., impunctate scutellum, markedly smaller eyes, short antennae, etc.).	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFD93F13FE4BC7D29E2E5AC5.taxon	description	(Figs. 15 – 19)	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFD93F13FE4BC7D29E2E5AC5.taxon	materials_examined	Type locality. People’s Republic of China, W Sichuan, pass SE Barkam, between Zhuokeji-Lianghekou, 4100 m. Type material. HOLOTYPE: J (ASC, to be deposited in Muséum d’Histoire Naturelle, Genève), ‘ CHINA-W Sichuan Barkam, pass SE Barkam, pass between Zhuokeji-Lianghekou 4100 m, alpine zone 10 - 30. VI. 2004, leg. R. Fabbri’.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFD93F13FE4BC7D29E2E5AC5.taxon	description	Description. Head black, pronotum, elytra and abdomen piceous-black; head, pronotum and elytra with metallic bronze lustre, abdomen iridescent; both maxillary and labial palpi, antennae and legs testaceous, middle and hind tibiae markedly blackened. Head rounded, wider than long (ratio 1.15); eyes very large and convex, tempora very short, considerably shorter than length of eyes seen from above (ratio 0.12); no additional setiferous punctures between anterior frontal punctures; posterior frontal puncture touching posteriomedian margin of eye, one puncture between it and posterior margin of head; temporal puncture small, touching posterior margin of eye; surface of head with fine, moderately dense microsculpture of transverse and oblique waves. Antenna moderately long, segments 2 and 3 subequal in length, segments 4 - 7 longer than wide, gradually becoming shorter, segments 8 – 10 about as long as wide, segment 11 as long as two preceding segments combined. Pronotum about as long as wide, widely rounded basally, slightly narrowed anteriad, evenly transversely convex; dorsal rows left with four, right one with three punctures; sublateral rows each with three punctures, with posterior puncture situated slightly after level of large lateral puncture; surface of pronotum with microsculpture of transverse waves similar to that on head. Scutellum with punctures, exact number impossible to establish among surface microsculpture. Elytra quite short, at suture markedly shorter (ratio 0.66), at sides shorter (ratio 0.76) than pronotum at midline; punctation moderately fine, dense; transverse interspaces between punctures about as large as diameters of punctures; pubescence black; surface between punctures with microscopic irregularities. Wings apparently each reduced to nonfunctional, narrow stump. Abdomen with tergite 7 (fifth visible) without whitish apical seam of palisade setae; tergite 2 (in front of first fully visible tergite) entirely punctate and pubescent; punctation of tergites markedly finer than that on elytra, dense, becoming sparser toward posterior margin of each tergite and in general toward apex of abdomen; pubescence black; surface between punctures with excessively fine microsculpture of striae. Male. First four segments of front tarsus dilated, subbilobed, each with tenent setae ventrally, segment two somewhat narrower than apex of tibia (ratio 0.92); segment 4 narrower than preceding segments. Sternite 8 with three long setae on each side, apical margin with moderately wide and deep, obtusely triangular medioapical emargination, small area before emargination flattened and smooth (Fig. 15). Genital segment with tergite 10 evenly narrowed toward widely arcuate apex, sparingly setose, as in Fig. 16; sternite 9 with narrow basal portion, apical portion narrowly arcuate apically, without differentiated apical or subapical setae, setose as in Fig. 17. Aedoeagus (Figs. 18, 19) rather robust, median lobe subparallel-sided in middle portion, anteriorly narrowed into moderately long apical portion with subacute apex, on face adjacent to paramere, when paramere removed, with short carina far below apex of median lobe. Paramere rather large, widely subfusiform, in ventral view entirely covering median lobe except for very apex, with narrowly arcuate apex by far not reaching apex of median lobe; four minute setae at apical margin, median setae somewhat longer than lateral ones, two similar setae at each lateral margin below apex; sensory peg setae on underside of paramere situated as in Fig. 19. Female. Unknown. Length 4.0 mm.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFD93F13FE4BC7D29E2E5AC5.taxon	etymology	Etymology. The specific epithet is the Greek noun ορος, - εος (a mountain) in apposition, referring to the occurrence of this species high up in the mountains. Bionomics. No details are known about the collecting circumstances, except that it was taken in the alpine zone at 4100 m. Geographical distribution. Quedius oros sp. nov. is at present known only from the type locality in Qiongilai Shan in western Sichuan. Recognition and comments. Quedius oros sp. nov. is well characterized, in addition to the characteristic shape of the aedoeagus, by the markedly blackened middle and hind tibiae, combined with testaceous tarsi, and the uniformly black pubescence of the elytra and abdominal tergites.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDB3F1DFE73C1DA9C175D85.taxon	description	(Figs. 20 – 25)	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDB3F1DFE73C1DA9C175D85.taxon	materials_examined	Type locality. People’s Republic of China, Yunnan, Heishui, 35 km N Lijiang, 27 ° 13 ′ N 100 ° 19 ′ E. Type material. HOLOTYPE: J (ASC, to be deposited in Naturhistorisches Museum, Wien, Austria), ‘ CHINA, Yunnan prov. 1. - 19. 7. 1992; HEISHUI 35 km N Lijiang 27 ° 13 ′ N 100 ° 19 ′ E lgt. S. Becvar’. ALLOTYPE: ♀ (ASC), same data as holotype.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDB3F1DFE73C1DA9C175D85.taxon	description	Description. Head black, pronotum and elytra dark brown in male, piceous-black in female, abdomen piceous-black, slightly iridescent; both maxillary and labial palpi, antennae and legs testaceous (somewhat darker in female), hind tibiae slightly darkened. Head rounded, wider than long (ratio 1.20); eyes very large and convex, tempora very short, considerably shorter than length of eyes seen from above (ratio 0.11); no additional setiferous punctures between anterior frontal punctures; posterior frontal puncture touching posteriomedian margin of eye, one puncture between it and posterior margin of head; temporal puncture absent; surface of head with fine, moderately dense microsculpture of transverse and oblique waves becoming submeshed on clypeus. Antenna moderately long, segments 2 and 3 subequal in length, segments 4 – 8 longer than wide, gradually becoming shorter, segments 9 and 10 about as long as wide, segment 11 as long as two preceding segments combined. Pronotum about as long as wide, widely rounded basally, slightly narrowed anteriad, evenly transversely convex; dorsal rows with three punctures; sublateral rows each with two punctures, with posterior puncture situated slightly after level of large lateral puncture; surface of pronotum with microsculpture of transverse waves similar to those on head. Scutellum with six or seven punctures on apical portion, surface with very fine, dense microsculpture of transverse waves. Elytra short, at suture markedly shorter (ratio 0.71), at sides shorter (ratio 0.89) than pronotum at midline; punctation fine, dense, transverse interspaces between punctures mostly smaller than diameters of punctures; pubescence piceous; surface between punctures without microsculpture. Wings apparently each reduced to nonfunctional stump. Abdomen with tergite 7 (fifth visible) without whitish apical seam of palisade setae; tergite 2 (in front of first fully visible tergite) entirely, finely punctate and pubescent; punctation of tergites markedly finer than that on elytra, dense, becoming sparser toward posterior margin of each tergite and in general toward apex of abdomen; pubescence piceous; surface between punctures with excessively fine microsculpture of striae. Male. First four segments of front tarsus dilated, subbilobed, each with tenent setae ventrally, segment two about as wide as apex of tibia, segment 4 narrower than preceding segments. Sternite 8 with three long setae on each side, apical margin with moderately wide, not deep, obtusely triangular medioapical emargination, small area before emargination flattened and smooth (Fig. 20). Genital segment with tergite 10 narrow, evenly narrowed toward narrowly arcuate apex, setose as in Fig. 21; sternite 9 with rather long basal portion, apical portion arcuate apically, with two differentiated subapical setae, otherwise only sparingly setose (Fig. 22), but see Comments. Aedoeagus (Figs. 23, 24) narrow, elongate, medial lobe largely parallel-sided, except for apex entirely covered by paramere in ventral view, anteriorly narrowed into moderately long apical portion with subacute apex, on face adjacent to paramere with short medial carina below apex. Paramere narrow, elongate, largely parallel-sided, with narrowly arcuate apex not reaching apex of median lobe; four setae at apical margin, median setae markedly longer than lateral ones, two similar setae at each lateral margin below apex; sensory peg setae on underside of paramere numerous, forming two rather irregular longitudinal rows, as in Fig. 24. Female. First four segments of front tarsus simple, not dilated, without tenent setae ventrally. Tergite 10 of genital segment similar to that of Q. nujiang sp. nov., but with more numerous setae on apical portion (Fig. 25). Length 4.8 - 5.0 mm.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
03D39C15FFDB3F1DFE73C1DA9C175D85.taxon	etymology	Etymology. The specific epithet is the Greek noun χιων, - ονος (snow) in apposition, referring to the occurrence of this species in a very high mountain range. Bionomics. No details are known about the collecting circumstances of this species. Geographical distribution. Quedius chion sp. nov. is at present known only from the type locality in Yulong Xue Shan in Yunnan. It is likely endemic to that mountain range. Recognition and comments. Quedius chion sp. nov. is characterized, in addition to the shape of the aedoeagus, by the sexually dimorphic coloration of the pronotum and elytra (but this has to be confirmed on larger material), and by the entirely punctate and pubescent abdominal tergite 2. The setation of sternite 9 of male genital segment and tergite 10 of female genital segment is largely missing, except for apical portions. Only present setae are shown in Figs. 22 and 25. The holotype is missing segments 6 to 11 of left antenna and segments 5 to 11 of right antenna. The allotype is missing the last segment of right antenna.	en	Smetana, Aleš (2011): Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae 51 (1): 145-156, DOI: 10.5281/zenodo.5327302
