taxonID	type	description	language	source
03DE87B4FFA9FF940F84683FFCFBF979.taxon	discussion	The subgenus Euandrena Hedicke, 1933 is taxonomically complex, as many cryptic taxa are present, and morphological recognition is highly challenging, even in comparatively well-studied regions such as Europe (e. g., Praz et al. 2019). The situation around the Himalayas (from the western to eastern extent) is currently unsatisfactory due to the large number of available names of uncertain status. Currently, seven red-marked species of Euandrena are known, specifically 1) Andrena familiaris Smith, 1878 (described from East Turkestan in what is now western China), 2) A. communis Smith, 1879 (described from Mussoorie in northern India), 3) A. mephistophelica Cameron, 1897 (described from Mussoorie in northern India), 4) A. murreensis Cockerell, 1923 (described from Murree in northern Pakistan), 5) A. euphorbiacea Scheuchl, 2005 (described from Yunnan in southern China), 6) A. humlaensis Scheuchl, 2005 (described from Nepal), and 7) A. kathmanduensis Tadauchi & Matsumura, 2007 (described from Nepal). Modern descriptions (Grünwaldt et al. 2005; Tadauchi & Matsumura 2007) unfortunately do not provide diagnoses against these older Himalayan taxa, and it is not clear if these authors were aware of these available names, their likely identities, or indeed how any of these names relate to each other. It is necessary to deal with the four oldest available names, and definitively locate the location of type material as the listings in Gusenleitner & Schwarz (2002) are unfortunately not always correct due to historical uncertainty, and the huge size of the genus which has not always allowed for each reported type locality to be specifically investigated. The first name, A. familiaris, was described from East Turkestan [neighbourhood of Yárkand = Yarkant County, Xinjiang, western China] (Smith 1878: 2). The species was described in the male sex only, and is illustrated (Smith 1878: fig. 3). Although Gusenleitner & Schwarz (2002) indicate that the type of A. familiaris is present at the NHMUK, no such specimen matching this collection information is present and Donald Baker (unpublished thesis: 272 – 273) concludes that type material is likely to be in the ZSI. Examination of the ZSI website (https: // zsicollections. in / search? genus = Andrena) shows that the male type of A. familiaris is present there (reference specimen code ZSI 0000008548). However, in the NHMUK type catalogue, there is an entry for A. familiaris (B. M. type 17 a 1306). Inspection of this specimen (Fig. 1) shows that it was collected from Masuri [= Mussoorie] in northern India, and it is a female. A handwritten label (not in the handwriting of Smith) indicates “ Andrena familiaris (Type) Sm. ”. An additional handwritten note by Donald Baker correctly indicates that this is a false type, since it is a female, and it comes from Masuri rather than Yárkand. However, the specimen matches perfectly the collection information for A. communis which was described from Masuri, collected at an elevation of 7000 feet (Smith 1879: 51). Andrena communis was described only in the female sex, and as the NHMUK specimen matches Smith’s description, the specimen is here considered to be the holotype of A. communis. No other specimens labelled as “ A. communis ” are known in the NHMUK catalogue (type collection or main collection). Since Smith (1879) is titled “ Descriptions of new Hymenoptera species in the collection of the British Museum ”, and all other species of Andrena described in this publication are present in the NHMUK, it is considered unlikely that the type might actually be in another institution. Gusenleitner & Schwarz (2002) list the NHMUK as the type depository for A. communis, but these authors did not inspect this material. They doubtful associate A. communis with the subgenus Oreomelissa Hirashima & Tadauchi, 1975, probably on the basis of Bingham (1897) who confusingly uses a character of a punctate propodeum in his identification key, later leading to A. communis. Based on this holotype specimen, A. communis is clearly a member of the subgenus Euandrena, lacks distinct punctures on the propodeum, and is relatively small at 9 mm in length. In Bingham’s (1897) identification key, A. communis is placed close to A. mephistophelica, and is separated by colouration, with A. mephistophelica having the tergal margins and bases lightened, rather than the red markings also covering the disc of T 2, as in A. communis (Fig. 1 D). Andrena mephistophelica was also described from Mussouri [= Mussoorie], and Cameron (1897) indicates its body length as 11 – 12 mm. Gusenleitner & Schwarz (2002) again indicate that type material is at the NHMUK, but it is actually in the Rothney Collection (OUMNH). However, the exact location of Andrena type material from the Rothney collection is unclear, and it is not currently available for study (J. Hogan, in litt.). Based on the description and the redescription by Bingham (1897: 443), A. mephistophelica is very close to A. murreensis which was described from Murree in northern Pakistan with a female body length of 9 – 11 mm (Cockerell 1923). Examination of the NHMUK collection produced a specimen of A. murreensis labelled by Cockerell with his distinctive handwriting and a yellow “ co-type ” label, from Murree at an elevation of 7500 feet. The specimen was collected by Dutt and has an additional label of “ 156 ”. This corresponds to Cockerell (1923: 265), who listed eight numbered specimens from the Fletcher collection. The specimen is hereby designated as a lectotype (Fig. 2; see below for full specimen details). An additional specimen is present in the USNM collection with the same collecting information (https: // collections. nmnh. si. edu / search / ento /) with the number “ 50 ”. It is part of the syntypic series, but is not designated as the lectotype. Genetic analysis of five samples of red-marked Euandrena from northern Pakistan provisionally identified as A. communis (one female) and A. murreensis (three females, one male) plus two sequences of A. murreensis available from BOLD (PCYU, two female specimens identified by TJW) produced three lineages with strong posterior support (Fig. 3). Three “ A. murreensis ” sequences form a clade with bootstrap support of 98 separated from A. communis + the remaining three “ A. murreensis ” sequences, these latter three sequences forming a clade with bootstrap support of 99. Morphologically, examination of the two “ A. murreensis ” clades plus the lectotype of A. murreensis shows subtle but consistent differences. Specifically, 1) true A. murreensis has the clypeus distinctly punctate, but punctures are sparse and scattered, separated by 1 – 3 puncture diameters, particularly medially where large impunctate areas are present (Fig. 4 C), whereas the comparison clade shows a more evenly punctate clypeus, with punctures separated by 1 – 2 puncture diameters over the entire disc (Fig. 4 D); 2) true A. murreensis has the scutum predominantly shiny, shagreened laterally and anteriorly but shining over 60 % of the scutum (Fig. 4 E), whereas the comparison clade has the scutum strongly and consistently shagreened, with around 20 % of the disc weakly shining medially (Fig. 4 F); and 3) true A. murreensis has the propodeal triangle more strongly depressed and the dorsolateral parts of the propodeum more strongly rugose, thus reciprocally more strongly contrasting, whereas the comparison clade has both the propodeal triangle less strongly depressed and the dorsolateral parts of the propodeum less strongly rugose, so they only weakly contrast each other. It remains unclear if the other lineage can be called A. mephistophelica, or whether A. mephistophelica is the senior synonym of A. murreensis, and the remaining lineage refers to one of the more recently described species of red-marked Euandrena (Grünwaldt et al. 2005; Tadauchi & Matsumura 2007), or perhaps is even undescribed. Based on the description, the next best candidate for the non- murreensis clade would be A. humlaensis given its comparatively densely punctate clypeus (Grünwaldt et al. 2005: 363), whereas A. euphorbiacea and A. kathmanduensis appear to be more similar to A. murreensis based on the sparse clypeal punctures. For the present time, this second lineage is referred to as “ aff. A. murreensis ”. The two lineages can be found in direct sympatry, for example at Shogran (both historical and modern specimens, see below). The true A. murreensis clade was separated from the aff. A. murreensis clade by 7.20 % (range 7.14 – 7.31 %), and A. communis was separated from the true A. murreensis clade by 8.23 % (range 8.10 – 8.37 %) and the aff. A. murreensis clade by 6.85 %. It is important to note that these distances in and of themselves (the so called “ barcoding gap ”) should not be simply accepted as proof of species-level differences, as high intraspecific divergence can occur within widely distributed species (e. g., Hickerson et al. 2006; Collins & Cruickshank 2013; Zhang et al. 2017). However, the species of Euandrena here have only moderate ranges (restricted to the Himalayas), show consistent morphological differences, and present a tree topology that indicates that genetic distance is not the sole difference, with the sequence of A. communis sitting between the two more morphologically similar larger A. murreensis and aff. A. murreensis clades. On the basis of these consistent morphological differences, combined with these divergent DNA barcodes and tree topology, the conclusion is therefore drawn that at least three red-marked species of Euandrena can be found in the Western Himalayas. Two of these species can be confidently referred to as A. communis (the smaller individuals with all of T 2 red-marked; Fig. 1 D, contrast Figs 4 G – H) and A. murreensis (larger individuals with the base and apexes of the terga red-marked, but not the tergal discs, Fig. 4 G, the clypeus with deep scattered punctures, Fig. 4 B, and the scutum predominantly shiny, Fig. 4 E) on the basis of currently available type material. Full details on examined specimens are given below under each species entry, but in conclusion, 1) A. familiaris is currently only confidently known from the type series in the ZSI (probably a single male specimen) from the locus typicus in western China, 2) A. communis can be placed in the Euandrena and is known from northern India and Pakistan, 3) A. mephistophelica remains an unclear taxon, but is likely referable to one of the two large-bodied Euandrena genetic clades, and 4) A. murreensis is known from across northern Pakistan and India into Nepal (see below for examined material). The more recently described taxa from the Eastern Himalayas require type investigation, combined with additional genetic analysis from Nepal and southern China.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA7FF900E936B51FC04F88F.taxon	description	urn: lsid: zoobank. org: act: 19 CF 4125 - 2926 - 4 B 48 - B 6 DA- 327 D 82095 F 3 C Figs 5, 6 A, C, E, G	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA7FF900E936B51FC04F88F.taxon	diagnosis	Diagnosis Andrena kedarnatha Wood & Gautam sp. nov. can be recognised as a Melandrena Pérez, 1890 due to its large body size and robust shape (15 – 16 mm), dark integument and predominantly dark pubescence with partially infuscate wings (Fig. 5 A), long ocelloccipital distance (3 × diameter of lateral ocellus, Fig. 6 A), large propodeal triangle with roughly raised internal carinae (Fig. 5 C), and lack of other diagnostic characters. Due to the dark integument without metallic reflections, the almost entirely dark pubescence (without white bands or spots of pubescence), the broad facial foveae that occupy almost the entire distance between the compound eye and lateral ocellus (Fig. 6 A), and the clearly and densely punctate terga (Figs 5 D, 6 E, 6 G), it can be placed close to A. cussariensis Morawitz, 1886. This species was described from what is now Azerbaijan (type material illustrated by Astafurova et al. 2021) and is found from Crimea and European Russia through Turkey, Iran, the Caucasus to Central Asia, Siberia, and Mongolia (Osytshnjuk et al. 2008; Astafurova et al. 2021; Wood & Monfared 2022). Andrena cussariensis is currently not confidently recorded from India specifically or the Himalayan region more generally, and is not considered to be present (see Remarks). Morphologically, A. kedarnatha Wood & Gautam sp. nov. can be separated due to its larger body length of 15 – 16 mm (slightly but noticeably larger than the body length of A. cussariensis which averages 13 – 14 mm in length), its longer ocelloccipital distance of 3 – 3.5 × diameter of lateral ocellus with the vertex most strongly and coarsely punctate (Fig. 6 A; in A. cussariensis with shorter ocelloccipital distance of 2 – 2.5 × diameter of lateral ocellus, with the vertex more weakly and finely punctate, Fig. 6 B), by a more consistently dense punctation of the scutum with punctures separated by <0.5 – 0.5 puncture diameters, not becoming slightly but noticeably sparser posteromedially (Fig. 6 C; in A. cussariensis with the scutal punctures typically separated by 0.5 puncture diameters but in some specimens with punctures separated by 1 – 2 puncture diameters posteromedially, forming a small but distinct shining space, Fig. 6 D), terga with slightly but comparatively larger and coarser punctures, most visible on the discs of T 1 – 2 (Fig. 6 E; in A. cussariensis with comparatively smaller and finer punctures, Fig. 6 F), and T 2 laterally with the shallow foveae broadened and filled with reddish-bristles, surface densely punctate with punctures separated by 0.5 puncture diameters (Fig. 6 G; in A. cussariensis without a noticeable foveae present laterally on T 2, surrounding surface comparatively sparsely and weakly punctate, Fig. 6 H). To best appreciate the final characters, comparative material is required. There is some variation in colouration between the older museum specimens which have the wing venation orange-brown, the tergal margins slightly lightened brownish, the tibial scopa with off-white to light brownish hairs whereas recently collected specimens are uniformly darker (dark venation, dark terga, and dark scopal hairs). This is considered to represent variation, as sculptural characters do not differ between these specimens.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA7FF900E936B51FC04F88F.taxon	etymology	Etymology Taken from the name of the Kedarnath temple (Uttarakhand) which is found close to the modern sampling localities of this new species (Fig. 7). It is a noun in apposition.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA7FF900E936B51FC04F88F.taxon	materials_examined	Type material Holotype INDIA • ♀; Uttarakhand, Chaumasi Village; 30.6146 ° N, 79.0695 ° E; 2250 m a. s. l.; 8 May 2022; R. K. Gautam leg.; WII. Paratypes INDIA • 1 ♀; Uttarakhand, Chaumasi Village; 30.6146 ° N, 79.0695 ° E; 2250 m a. s. l.; 8 May 2022; R. K. Gautam leg.; WII • 1 ♀; Gulmarg [Jammu and Kashmir]; summer 1913; Lt-Col. F. W. Thomson leg.; NHMUK. NEPAL • 1 ♀; W-Nepal, Jumla, Jumla; 2850 m a. s. l.; 4 Oct. 1993; E. Hüttinger leg.; OÖLM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA7FF900E936B51FC04F88F.taxon	description	Description Female BODY. Length 15 – 16 mm (Fig. 5 A). HEAD. Dark, 1.2 × as wide as long (Fig. 5 B). Clypeus domed, densely punctate, punctures separated by 0.5 puncture diameters, underlying surface dull. Process of labrum broadly trapezoidal, 2.5 × as wide as long, lateral margins weakly emarginate, apical margin truncate. Gena broad, 1.2 – 1.5 × diameter of compound eye; ocelloccipital distance 3 – 3.5 × diameter of lateral ocellus. Vertex strongly and coarsely punctate, punctures behind ocellar triangle separated by 0.5 puncture diameters (Fig. 6 A). Foveae dorsally broad, occupying almost entire space between compound eye and lateral ocellus, separated from lateral ocellus by distance subequal to its diameter; foveae ventrally strongly passing level of antennal insertions ventrally, filled with dark brown hairs. Head with black to dark brown hairs, none equalling length of scape. Antennae dark, A 3 equalling A 4 + 5, shorter than A 4 + 5 + 6. MESOSOMA. Scutum and scutellum densely and regularly punctate, punctures separated by <0.5 puncture diameters, almost confluent, interspaces weakly shining medially, dull anteriorly and laterally (Fig. 6 C). Pronotum rounded. Mesepisternum densely but shallowly punctate, punctures separated by 0.5 puncture diameters, interspaces microreticulate, dull. Dorsolateral parts of propodeum densely and shallowly punctate, punctures separated by 0.5 puncture diameters, interspaces raised to form network of finely raised rugae; propodeal triangle large, laterally delineated by finely raised carinae, internal surface with dense network of raised carinae (Fig. 5 C). Mesosoma with abundant but relatively short black pubescence, longest on mesepisternum, becoming shorter but not squamous on scutum and scutellum. Propodeal corbicula incomplete, dorsal fringe composed of abundant and strongly plumose black hairs, internal surface covered with latitudinal weakly raised rugae, with abundant simple black hairs. Legs dark, apical tarsal segments lightened reddish brown; pubescence black to dark brown. Flocculus incomplete, weakly produced, composed of plumose black hairs; femoral and tibial scopae composed of off-white to light brownish or predominantly black simple hairs. Hind tarsal claws with strong inner tooth. Wings infuscate over majority of surface, becoming marginally weaker away from venation; stigma and venation dark orange-brown to black, nervulus interstitial. METASOMA. Tergal discs dark, apical margins narrowly lightened dark brown or almost entirely dark (Fig. 5 D). Tergal discs densely but slightly irregularly punctate, punctures typically separated by 0.5 – 1 puncture diameters (Fig. 6 E), up to 2 puncture diameters medially; punctures becoming progressively finer and weaker on marginal areas from T 1 – 4, on T 4 almost disappearing into underlying sculpture. Tergal discs obscurely shagreened, shining to weakly shining. T 2 laterally with shallow but distinct broad foveae, foveae subtly impressed, covered with reddish-brown to black bristles (Fig. 6 G). Tergal discs with scattered short dark hairs, not forming apical hairbands. Apical fringe of T 5 and hairs flanking pygidial plate dark brown. Pygidial plate large, rounded triangular, surface finely and evenly shagreened, weakly shining. Male Unknown.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA7FF900E936B51FC04F88F.taxon	discussion	Remarks Andrena cussariensis was listed for the Indian fauna by Meena & Dey (2019) who mentioned it from Kohtak [= Kohat, Pakistan], Musooree [= Mussoorie], Shimla, and Punjab. The listing of this species from India is complicated, and requires dissection. The listing from Kohat refers to the record of Cockerell (1917) who reported A. cussariensis from “ Kohat, N. W. Provinces, India ”, and described the subspecies A. cussariensis kohatensis Cockerell, 1917. Kohat is today found in the state of Pakistan. Moreover, the material identified by Cockerell actually belongs to A. fuscosa, and A. cussariensis kohatensis has now been synonymised with A. fuscosa (Wood 2024 a). Records from low-elevation areas of India therefore would therefore seem most likely to be A. fuscosa. It is impossible to say whether the records from Mussoorie refer to A. kedarnatha Wood & Gautam sp. nov. or not without examination of specimens (which are not precisely cited or referenced), and so the presence of A. cussariensis in India is considered to be unproven given its known global distribution and the risk of confusion with A. kedarnatha.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA7FF900E936B51FC04F88F.taxon	distribution	Distribution Northern India (Jammu & Kashmir, Uttarakhand) and Nepal.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA3FF890E896B0AFCC2FAA1.taxon	description	urn: lsid: zoobank. org: act: 60275075 - 8392 - 48 A 1 - A 5 F 1 - C 4537 B 36 B 109 Figs 8 A – C, E, G – H, 9	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA3FF890E896B0AFCC2FAA1.taxon	diagnosis	Diagnosis Andrena tungnatha Wood & Gautam sp. nov. can quickly be recognised as part of the subgenus Simandrena Hedicke, 1933 due to the propodeal corbicula which is perfect (possessing strongly developed dorsal and anterior fringes), with the internal surface (the lateral faces of the propodeum) glabrous, the short male A 3 (clearly shorter than A 4), the long second tarsal segment of the hind leg in the male sex, and the simple genital capsule. Andrena tungnatha is similar to many Central Asian species of Simandrena in the female sex due to the fine and dense tergal punctation combined with clear apical tergal hairbands, e. g., A. quadrifasciata Morawitz, 1876 (Kazakhstan, Uzbekistan, Tajikistan), A. sarta Morawitz, 1876 (Kazakhstan, Uzbekistan), and A. iliana Shebl & Tadauchi, 2011 (Kazakhstan). However, in each comparison species, the tergal hairbands are thick, long, and dense, clearly extending from the base of the marginal area to beyond its apical rim, whereas in A. tungnatha the tergal hairbands are narrow, covering only the apical rim of the marginal area and barely extending beyond it (Fig. 8 H). Moreover, the scutum is completely dull (Fig. 8 D; partially shining in A. quadrifasciata and A. iliana, completely shining in A. sarta), and the clypeus is strongly and densely punctate with large punctures with relatively narrow shining interspaces (Fig. 8 B; clypeus shagreened to dull in A. quadrifasciata, shiny with fine and scattered punctures in A. sarta, and only weakly and finely punctate in A. iliana). The main comparison should be with A. gorkhana Tadauchi & Matsumura, 2007, which is the only other species of Simandrena known from the Western Himalayas (A. (Simandrena) metuoensis Xu & Tadauchi, 2001 is known from Xizang [Tibet] in China, but this species has T 1 – 3 lightened red, the scutellum is shiny and broadly impunctate, and the terga are impunctate), and both species display fine and narrow tergal hairbands, finely and densely punctate terga, and show a densely punctate clypeus with shining interspaces. Andrena tungnatha can be separated from A. gorkhana by its slightly larger body size of 9 mm (7 – 8 mm in A. gorkhana), by the more strongly sculptured scutum, at most weakly shining but predominantly dull (Fig. 8 C; in A. gorkhana with the scutum with extremely weak shagreen, almost smooth and shining over its entire area, Fig. 8 D), by the hairs of the scutum and scutellum that are moderately long and orange, of a uniform consistency and colour between the scutum and scutellum (Fig. 8 E, in A. gorkhana with scutal hairs short and pale whitish-brown, scutal hairs orange and distinctly longer, forming a clear tuft, Fig. 8 F), and by the hind tibiae lightened orange with orange-brown scopal hairs (Fig. 8 G; in A. gorkhana with hind tibiae dark, scopal hairs whitish-light brownish). Separation in the male sex can also be aided by size, as males of A. tungnatha Wood & Gautam sp. nov. are 8 – 8.5 mm long (Fig. 9 A) compared to 6 mm long in the newly discovered male of A. gorkhana (Fig. 10 A), having been described in the female sex only. This male of A. gorkhana is formally described below. However, since only one male specimen of A. gorkhana is known, and this may be an unusually small specimen, structural separation should and can be made. Specifically, A. tungnatha has the clypeus densely punctate, punctures separated by 0.5 – 1 puncture diameter, with a weak impunctate longitudinal mid-line (Fig. 9 B; in A. gorkhana with the clypeus more irregularly punctate, punctures separated by 0.5 – 2 puncture diameters, without an impunctate longitudinal mid-line, Fig. 10 B), scutum entirely dull, with strong and rough microsculpture, punctures obscure and hardly visible (Fig. 9 D, in A. gorkhana scutum anteriorly with microsculpture, but becoming smooth and shining medially, punctures clearly visible against underlying surface, Fig. 10 D), and genital capsule with the penis valves relatively wide (Fig. 9 F; in A. gorkhana with the penis valves relatively narrow, Fig. 10 F).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA3FF890E896B0AFCC2FAA1.taxon	etymology	Etymology Taken from the name of the Tungnath temple (Rudraprayag, Uttarakhand) which is found close to the modern sampling localities of this new species (Fig. 7). It is a noun in apposition.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA3FF890E896B0AFCC2FAA1.taxon	materials_examined	Type material Holotype INDIA • ♀; Uttarakhand, Hudu Village, 4.5 km W of Chopta; 30.4953 ° N, 79.1559 ° E; 2098 m a. s. l.; 6 Apr. 2022; R. K. Gautam leg.; WII. Paratypes INDIA • 1 ♂; Uttarakhand, Hudu Village, 4.5 km W of Chopta; 30.4953 ° N, 79.1559 ° E; 20 Mar. 2021; R. K. Gautam leg.; WII • 3 ♂♂; Uttarakhand, Ukhimath, Gari Village; 1586 m a. s. l.; 20 Mar. 2021; R. K. Gautam leg.; WII • 2 ♀♀; Simla [Shimla]; 1 – 31 May 1897; C. G. Nurse leg.; NHMUK.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA3FF890E896B0AFCC2FAA1.taxon	description	Description Female BODY. Length: 9 mm (Fig. 8 A). HEAD. Dark, 1.3 × as wide as long (Fig. 8 B). Clypeus moderately domed but medially somewhat flattened, surface uneven, irregularly punctate with deep punctures of variable size, punctures separated by <0.5 – 2 puncture diameters; underlying surface smooth and shiny medially, becoming shagreened laterally. Process of labrum rounded trapezoidal, slightly more than twice as wide as long, surface with obscure transverse striations. Gena slightly exceeding width of compound eye; ocelloccipital distance slightly exceeding diameter of lateral ocellus. Foveae dorsally broad, occupying ¾ of space between compound eye and lateral ocellus, separated from lateral ocellus by distance sub-equal to diameter of lateral ocellus; foveae slightly narrowing ventrally, ventrally extending slightly beyond lower margin of antennal insertions; foveae dorsally filled with dark brown hairs, hairs becoming pale brown ventrally. Face, gena, and scale with light brown hairs, frons dorsally and vertex with longer dark brown to black hairs, none equalling length of scape. Antennae dark basally, A 5 – 12 ventrally lightened by presence of orange scales; A 3 exceeding length of A 4, shorter than A 4 + 5. MESOSOMA. Scutum with strong microsculpture, microreticulate, weakly shining, densely punctate over majority of area, punctures separated by 0.5 – 1 puncture diameters except medially, here separated by 1 – 2 puncture diameters (Fig. 8 C). Scutellum strongly punctate, punctures confluent to separated by 1 puncture diameter, interspaces smooth and shining medially. Pronotum rounded. Mesepisternum finely shagreened, predominantly shining, surface with dense hair-bearing punctures, punctures separated by 1 puncture diameter. Dorsolateral parts of propodeum finely microreticulate, fine sculpture overlain by network of delicately raised rugosity; propodeal triangle laterally delineated by raised carinae, internal surface with network of finely raised rugae in basal ⅔, surface dull to weakly shining. Mesepisternum with abundant long and finely plumose hairs, whitish ventrally, becoming light brown dorsally, longest hairs not equalling length of scape. Scutum and scutellum with shorter densely plumose light brownlight orange hairs, hairs appearing sub-squamous (Fig. 8 E). Propodeal corbicula complete, perfect, dorsal and anterior fringes composed of long finely plumose light brown hairs, internal surface finely shagreened and shining, glabrous. Legs basally dark, apical tarsal segment of fore and mid legs reddish brown, hind tibiae and tarsi lightened orange; pubescence light brown to brownish (Fig. 8 G). Flocculus incomplete, composed of brown densely plumose hairs; femoral scopae composed of golden simple hairs, tibial scopae composed of simple hairs, golden ventrally, brownish dorsally, dorsal fringe with scattered weakly plumose hairs. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma and venation orange, nervulus weakly antefurcal. METASOMA. Tergal discs dark, apical rims narrowly lightened hyaline-yellow (Fig. 8 H). Tergal discs uniformly finely and densely punctate, punctures separated by 0.5 puncture diameters, punctures sparser only laterally on T 1, here separated by 2 – 3 puncture diameters; underlying surface finely shagreened, weakly shining to shining. T 1 latero-apically with weak hair fringes, not forming clear hairband, hairs extending onto basolateral parts of T 2; T 2 – 4 apically with dense apical hairbands of yellowish hairs, hairbands narrow, only slightly extending beyond the apex of apical rim, obscuring underlying surface. Apical fringe of T 5 and hairs flanking the pygidial plate dark brown to black. Pygidial plate narrow, rounded triangular, surface dull. Male BODY. Length: 8 – 8.5 mm (Fig. 9 A). HEAD. Dark, 1.25 × as wide as long (Fig. 9 B). Clypeus weakly domed, more or less flattened medially, regularly punctate with deep punctures, punctures separated by 0.5 – 1 puncture diameters with exception of impunctate longitudinal mid-line, mid-line extending ventrally to mid-point of clypeus; underlying surface smooth and shining. Process of labrum rounded rectangular, 2.5 × as wide as long. Gena equalling width of compound eye; ocelloccipital distance 1.5 × diameter of lateral ocellus. Head covered with mixture of long black and light brown hairs, black hairs covering majority of face with light brown hairs restricted to antennal insertions and apex of clypeus, gena and vertex with predominantly light brown hairs; longest hairs exceeding length of scape. Antennae dark, A 3 clearly shorter than A 4, A 4 – 13 rectangular, clearly longer than broad (Fig. 9 C). MESOSOMA. Scutum with rough microsculpture, dull, punctures abundant but shallow and obscure, with margins raised and blending into network of raised rugosity (Fig. 9 D). Scutellum less strongly sculptured, surface weakly shining medially, punctures without raised rims, separated by 0.5 – 1 puncture diameters. Pronotum rounded. Mesepisternum microreticulate, overlain by network of weakly raised rugosity, weakly shining. Propodeum with strong microreticulation overlain by dense network of raised rugosity, propodeal triangle broad, laterally defined by raised carinae, internal surface with finely raised rugosity. Mesosoma with long light brown finely plumose hairs, majority exceeding length of scape. Legs dark, tarsal segments dark reddish, pubescence light brown. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma and venation orange, nervulus weakly antefurcal. METASOMA. Tergal discs dark, apical rims narrowly lightened hyaline-yellow (Fig. 9 E). Tergal sculpture as in female. T 1 apico-laterally with weak fringe of long hairs, T 2 – 4 with complete hairbands of whitish hairs, rapidly degraded, not obscuring underlying surface. T 6 – 7 with long dark brown hairs overlying pseudopygidial plate of T 7. S 8 columnar, apex slightly broadened, rounded; ventral surface with lateral fan of brown hairs. Genital capsule slightly elongate, gonocoxae weakly produced into rounded teeth apically, gonostyli basally narrow, broadening apically, spatulate, with inner margin raised (Fig. 9 F). Penis valves weakly broadened basally, occupying ½ space between gonostyli, narrowing apically.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA3FF890E896B0AFCC2FAA1.taxon	discussion	Remarks Two undetermined specimens of this new species were found in undetermined material collected by C. G. Nurse in the NHMUK collection, where they seemingly have been overlooked by previous workers.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFA3FF890E896B0AFCC2FAA1.taxon	distribution	Distribution Northern India (Himachal Pradesh, Uttarakhand). Species considered to be part of the Indian fauna	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB9FF890ED06F3BFCE7F85B.taxon	materials_examined	Material examined INDIA • 1 ♀; Uttarakhand, 4 km N of Chaumasi; 30.6368 ° N, 79.0725 ° E; 2588 m a. s. l.; 9 May 2022; R. K. Gautam leg.; WII • 1 ♂; Uttarakhand, Kanda Village; 30.4987 ° N, 79.1457 ° E; 1920 m a. s. l.; 7 Apr. 2022; R. K. Gautam leg.; WII • 5 ♂♂, 4 ♀♀; Uttarakhand, Chhopta, Bhujgyali Bugyal; 30.4921 ° N, 79.2033 ° E; 2996 m a. s. l.; 17 Mar. 2021; R. K. Gautam leg.; WII • 6 ♂♂, 1 ♀; Uttarakhand, Chopta; 30.487 ° N, 79.2004 ° E; 2874 m a. s. l.; 6 Mar. 2021; R. K. Gautam leg.; WII • 1 ♂; Uttarakhand, Kanchula Kharg; 30.4596 ° N, 79.229 ° E; 2680 m a. s. l.; 24 Feb. 2021; R. K. Gautam leg.; WII • 6 ♀♀; Sikkim; C. T. Bingham leg.; ZMHB.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB9FF890ED06F3BFCE7F85B.taxon	discussion	Remarks This is the first Andrena (Andrena) species reported with confidence from India (see below). It was frequently encountered at high altitude sites in the spring.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB9FF890ED06F3BFCE7F85B.taxon	distribution	Distribution India * (Uttarakhand, Sikkim), China (Xizang) (Wu 1982).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB8FF880EE26B0AFEA5FD0A.taxon	materials_examined	Material examined INDIA • 1 ♀; Uttarakhand, Hudu Village; 30.5044 ° N, 79.1504 ° E; 1800 m a. s. l.; 19 Nov. 2019; R. K. Gautam; WII.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB8FF880EE26B0AFEA5FD0A.taxon	distribution	Distribution India * (Uttarakhand), China (Xizang, Yunnan) (Tadauchi & Xu 2002, as A. carinigera, incorrect spelling).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB8FF880E4868B3FAF9FAF7.taxon	materials_examined	Material examined INDIA • 1 ♂; Darjeeling; Juni [likely June 1902]; H. Fruhstorfer leg.; NMW • 1 ♀; Rungaroom [Darjeeling]; [undated]; C. T. Bingham leg.; ZMHB • 1 ♀; Sikkim; [undated]; C. T. Bingham leg.; ZMHB.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB8FF880E4868B3FAF9FAF7.taxon	discussion	Remarks Wood (2024 b) described the male of A. granulitergorum, and reported it from Nepal for the first time. Examination of material in the NMW collection revealed another male specimen, this time from the northern part of West Bengal. It was collected by Hans Fruhstorfer (1866 – 1922), a German explorer, insect trader, and entomologist. Between 1899 – 1902, he went on a three year round the world expedition through North America and Eastern Asia, returning via India. We therefore estimate that this specimen was collected in 1902. There were also two specimens from this region collected by Bingham, but the specimens are undated. It can be estimated that the specimens were collected in the 1890 s based on Bingham’s most active collecting period.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB8FF880E4868B3FAF9FAF7.taxon	distribution	Distribution India * (West Bengal, Sikkim), Nepal, and China (Sichuan, Yunnan) (Tadauchi & Xu 2002; Wood 2024).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB8FF8B0EA66F9CFAF9FED1.taxon	discussion	Remarks Tadauchi & Xu (2002) placed A. chagyabensis in combination with A. kishidai Yasumatsu, 1935. Wood (2024 b) reported A. kishidai chagyabensis as new for India, Bhutan, and Nepal, suggesting that A. k. chagyabensis may not only be distinct based on its disjunct distribution (A. kishidai s. str. is found in northern and north-eastern China), but may have been previously described from northern India by earlier workers (e. g., A. gracillima Cameron, 1897, see below). However, type material is currently unavailable for study to confirm or disprove this.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB8FF8B0EA66F9CFAF9FED1.taxon	distribution	Distribution (subspecies chagyabensis) India (Himachal Pradesh), Bhutan, Nepal, China (Xizang, Yunnan) (Xu & Tadauchi 2002; Wood 2024 b).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBBFF8B0EE36B7BFC12FC0E.taxon	materials_examined	Material examined (see also Wood 2024 b) INDIA • 7 ♀♀; Ladakh, Leh, garden; 3535 m a. s. l.; 12 Aug. 1983; I. Tangelder leg.; RMNH; ZMA. INS. 5103695 to ZMA. INS. 5103701 • 4 ♂♂, same data as for preceding; ZMA; ZMA. INS. 5103702 to ZMA. INS. 5103705. Listed by Wood (2024 b).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBBFF8B0EE36B7BFC12FC0E.taxon	distribution	Distribution Kazakhstan, Uzbekistan, Tajikistan, Kyrgyzstan, Afghanistan, India (Jammu & Kashmir, Ladakh) (Osytshnjuk et al. 2005; Astafurova et al. 2022; Wood 2024 b).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBBFF8A0ECF69B7FD7BFE36.taxon	materials_examined	Material examined INDIA • 1 ♀; Simla [Shimla]; 1 – 31 Aug. 1898; C. G. Nurse leg.; NHMUK (holotype of A. arima) • 3 ♀♀; NW India, U. P. [Uttar Pradesh, now Uttarakhand], Kumaon, Naini Tal [Nainital]; 2000 – 2480 m a. s. l.; 3 – 11 Oct. 1978; M. A. Lieftinck leg.; RMNH; RMNH. INS. 1266112 to RMNH. INS. 1266114. NEPAL • 1 ♀; Langtang-Ghora Tabela; 18 Jul. 1975; M. Kraus leg.; OÖLM • 1 ♀; Langtangtal; 14 Jul. 1975; M. Kraus leg.; OÖLM • 1 ♀; Nawakot, Langtang Khola, Ghora Tabela; 3200 m a. s. l.; 3 Oct. 1982; C. Holzschuh leg.; OÖLM • 3 ♀♀; Kathmandu, Pulchoki, SE Godavari; 2760 m a. s. l.; 14 Sep. 1995; E. and J. Hüttinger leg.; OÖLM • 1 ♂, 1 ♀; Kathmandu, SE Kathmandu, Pulchoki; 1860 – 2500 m a. s. l.; 23 Sep. 1993; E. Hüttinger leg.; OÖLM • 2 ♀♀; Kathmandu, SE Kathmandu, Pulchoki; 2610 m a. s. l.; 20 Sep. 1993; E. Hüttinger leg.; OÖLM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBBFF8A0ECF69B7FD7BFE36.taxon	discussion	Remarks Gusenleitner & Schwarz (2002) did not confidently place A. arima into a subgenus, whereas Tadauchi & Matsumura (2007) placed it in the subgenus Ptilandrena Robertson, 1902. Inspection of the type along with additional material from Nepal suggests that A. arima is associated with the subgenus Euandrena due to the simple scopal hairs, pronotum rounded and without a humeral angle, facial foveae weakly dropshaped, weakly sculptured terga, and lack of any other distinctive characters. The use of the subgenus Ptilandrena by Tadauchi & Matsumura indicates that they drew essentially the same conclusions, as Ptilandrena and Euandrena are sister subgenera, but following the revision of Pisanty et al. (2022 b), the use of Ptilandrena is restricted to North America. A specimen of the undescribed male was found in the OÖLM collection, but unfortunately it is stylopised and does not represent a typical individual, so it is not described here. A genetic analysis is required to confirm the phylogenetic position of A. arima, though it appears likely to fall in the derived group of subgenera around or close to Euandrena.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBBFF8A0ECF69B7FD7BFE36.taxon	distribution	Distribution India (Himachal Pradesh, Uttarakhand), Nepal *.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBAFF8A0ECD6BDCFDD6F984.taxon	materials_examined	Material examined INDIA • 1 ♀; Masuri [= Mussoorie]; 7000 ft a. s. l.; NHMUK (holotype of A. communis) • 1 ♀; Uttarakhand, Kanda Village; 30.4987 ° N, 79.1457 ° E; 1920 m a. s. l.; 7 Apr. 2022; R. K. Gautam leg.; WII • 1 ♀; Uttarakhand, Trijuginarayan; 30.6365 ° N, 78.9688 ° E; 2418 m a. s. l.; 2 Jun. 2022; R. K. Gautam leg.; WII • 1 ♂; Uttarakhand, Hudu; 30.5044 ° N, 79.1504 ° E; 1800 m a. s. l.; 19 Nov. 2019; R. K. Gautam leg.; WII. PAKISTAN • 1 ♀; Azad Kashmir, Paras, Shogran; 25 May 2019; M. Kafka leg.; BOLD: WPATW 979 - 23; TJWC • 1 ♀; Hazara, Kawai, Shogran; 14 May 1978; C. Holzschuh leg.; OÖLM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBAFF8A0ECD6BDCFDD6F984.taxon	discussion	Remarks Smith (1879) described A. communis from Masuri [= Mussoorie] in Uttarakhand, and Cameron (1897) described A. maligna from Mussouri [= Mussoorie] as well. Cameron (1897: 114) wrote that his species could “ hardly be the ♂ of A. communis ” but Bingham (1897: 443) nevertheless synonymised the two species, without any justification. The type material of A. maligna is in the Rothney Collection (OUMNH), but is not currently available for study. This synonymy is maintained pending further investigations, though it is plausible given that the two taxa share the same locus typicus. As discussed above, the specimen labelled as “ Andrena familiaris (Type) Sm. ” in the NHMUK collection is considered to be the holotype of A. communis.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBAFF8A0ECD6BDCFDD6F984.taxon	distribution	Distribution India (Uttarakhand) and Pakistan *.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBAFF850EDB6C29FE5BFA84.taxon	materials_examined	Material examined INDIA • 9 ♀♀; Uttarakhand, Chhopta, Bhujgyali Bugyal; 30.4921 ° N, 79.2033 ° E; 2996 m a. s. l.; 8 Jun. 2021; R. K. Gautam leg.; WII • 2 ♀♀; Uttarakhand, Chhopta, Bhujgyali Bugyal; 30.4921 ° N, 79.2033 ° E; 2996 m a. s. l.; 17 Mar. 2021; R. K. Gautam leg.; WII • 9 ♀♀; Uttarakhand, Chopta; 30.487 ° N, 79.2004 ° E; 2874 m a. s. l.; 6 Mar. 2021; R. K. Gautam leg.; WII • 1 ♀; Uttarakhand, Ukhimath, Gari Village 30.5288 ° N, 79.1001 ° E; 1586 m a. s. l.; 2 Mar. 2021; R. K. Gautam leg.; WII. KAZAKHSTAN • 1 ♀; Ketman Mts; ~ 2100 m a. s. l.; 14 Jun. 1988; V. Gurko leg.; OÖLM • 1 ♀; Arslambob, Kyzyl-Suu; 1590 m a. s. l.; 14 Jun. 2017; M. Kafka leg.; OÖLM • 1 ♂, 1 ♀; Frunze env. [Bishkek] (30 km) Orto-Sai [Orto-Say]; 28 May 1980; Z. Pádr leg.; OÖLM • 2 ♀♀; Jalal Abad, 5 km SE of Arslanbob; 1600 m a. s. l.; 2 – 3 Jun. 2019; J. and L. Halada leg.; OÖLM / TJWC • 1 ♀; Jalal-abad Oblast, 5 km W of KashkaSuu, Jylamysh R.; 1774 m a. s. l.; 27 May 2009; L. R. Best leg.; PCYU • 1 ♀; Moldo-Too NR, S slope, Teke-Uyuk rav.; 2300 m a. s. l.; 30 Jun. 1999; D. Milko leg.; OÖLM • 2 ♀♀; Tschuy, Dist. Moskau, Kirgis Geb. S. Sosnovka; 1600 – 1750 m a. s. l.; 23 May 2008; H. and R. Rausch leg.; OÖLM / TJWC. UZBEKISTAN • 2 ♀♀; Daganadzham env., I, near Surxondaryo; 973 m a. s. l.; 9 – 11 Apr. 2022; D. Benda leg.; NMPC / TJWC • 2 ♀♀; Daganadzham env., II, near Surxondaryo; 973 m a. s. l.; 9 Apr. 2022; D. Benda leg.; NMPC / TJWC • 1 ♀; Kishlyk env., III, Kashkadarya reg.; 1065 m a. s. l.; 7 – 10 Apr. 2022; D. Benda and T. Fraňková leg.; NMPC • 1 ♀; Myk env., II, NP Zaamin, Jizzakh reg.; 1828 m a. s. l.; 19 Apr. 2022; D. Benda and T. Fraňková leg.; NMPC.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBAFF850EDB6C29FE5BFA84.taxon	discussion	Remarks Andrena flavitarsis was described from what is now Uzbekistan, and most authors reported it only from this country (Astafurova et al. 2022). However, Osytshnjuk et al. (2008: 48) report the species from Uzbekistan and “ Pendzhab (India) ”. ‘ Pandzhab’ is Polish for ‘ Punjab’, and may be an overlooked translation error, as the work of Osytshnjuk et al. was translated from Russian to English by Polish coauthors following the death of Anna Osytshnjuk in 1998. This listing has been overlooked by subsequent authors (e. g., Meena & Dey 2019; Astafurova et al. 2022), and indeed is difficult to believe without additional supporting evidence. Examination of new material shows that A. flavitarsis is widespread in Central Asia and extends south into the Western Himalayas in Uttarakhand. In this context, the mention of Punjab is more plausible, though presumably it would have to be from northern Punjab (e. g., close to Kashmir) since A. flavitarsis is found in mountainous habitats. Therefore, A. flavitarsis is not reported as new for India, though this is the first mention supported by detailed specimen records.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFBAFF850EDB6C29FE5BFA84.taxon	distribution	Distribution Kazakhstan *, Uzbekistan, Kyrgyzstan *, India (? Punjab, Uttarakhand) (Osytshnjuk et al. 2008; Astafurova et al. 2022).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB5FF850E8F6F2EFDFBF8E4.taxon	description	Bingham (1897); Cameron (1897); Bingham (1908); Meena & Dey (2019); Chandra et al. (2021); Ascher & Pickering (2023).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB5FF850E8F6F2EFDFBF8E4.taxon	discussion	Remarks As discussed above, A. mephistophelica is likely to be one of the larger-bodied red-marked Euandrena species present in the Himalayas. The exact location of the type material must be ascertained before further taxonomic action can be conducted.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB5FF850E8F6F2EFDFBF8E4.taxon	distribution	Distribution India (Uttarakhand) (Cameron 1897).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB5FF840EF36D89FB7CF8B5.taxon	materials_examined	Type material Lectotype (by present designation, Fig. 2) PAKISTAN • 1 ♀; Murree, 7500 ft, 18 Jun. 1920; “ 156 ”; Dutt leg. Syntype (examined by photograph) PAKISTAN • 1 ♀; Murree Hills, Punjab; 7500 ft; May. 1920; “ 50 ”; Dutt leg.; type no. 18260; USNM. Further material examined true A. murreensis INDIA • 1 ♀; Kumaon Himalaya, U. P. [Uttar Pradesh, now Uttarakhand], distr. Almora, Ranikhet – Chaubattia; 1800 m a. s. l.; 28 May 1981; C. Holzschuh leg.; OÖLM • 1 ♀; Simla [Shimla]; 1 – 31 May 1897; C. G. Nurse leg.; NHMUK • 2 ♀♀; Shillong, Assam; 1 – 30 Apr. 1903; R. Turner leg.; NHMUK • 1 ♀; NE India, Arunchal pr., Dirang vicinity; 1800 m a. s. l.; 8 – 22 May 2006; P. Pacholátko leg.; OÖLM • 4 ♀♀; NW India, U. P. [Uttar Pradesh, now Uttarakhand], Kumaon, Naini Tal [Nainital]; 2000 – 2480 m a. s. l.; 3 – 8 Oct. 1978; M. A. Lieftinck leg.; RMNH; RMNH. INS. 1266115 to RMNH. INS. 1266118 • 1 ♀; U. P. [Uttar Pradesh, now Uttarakhand], Garhwal Himalaya, Uttarkashi; 1300 m a. s. l.; 28 – 29 Apr. 1995; S. W. T. Batra leg.; SEMC; SEMC 0988949. NEPAL • 1 ♀; E Nepal, Dhankuta, Arun-Valley, Lamobagar Gola; 1000 – 1400 m a. s. l.; 27 May – 3 Jun. 1980; C. Holzschuh leg.; OÖLM • 1 ♀; Jiri, Bhodung [?]; 2400 m a. s. l.; 19 May 1962; G. Ebert and H. Falkner leg.; OÖLM • 1 ♀; Tatopani; 1200 m a. s. l.; 14 May 1984; C. Holzschuh leg.; OÖLM • 1 ♀; Kathmandu, Shivaburi [Shivapuri] whatersheet; 1800 – 1950 m a. s. l.; 17 Apr. 1994; E. and J. Hüttinger leg.; OÖLM. PAKISTAN • 1 ♀; Azad Kashmir, Paras, Shogran; 25 May 2019; M. Kafka leg.; BOLD: WPATW 982 - 23; TJWC • 1 ♀; Hazara, Kawai, Shogran; 14 May 1978; C. Holzschuh leg.; OÖLM. aff. A. murreensis, cryptic species NEPAL • 1 ♀; Kathmandu, Shivaburi [Shivapuri] whatersheet; 1800 – 1950 m a. s. l.; 17 Apr. 1994; E. and J. Hüttinger leg.; OÖLM • 1 ♀; Langtangtal; 14 Jul. 1975; M. Kraus leg.; OÖLM • 1 ♀; Trisuli Khola-Langtang Kh., Syabru Bensi [Syapru Besi]; 1600 m a. s. l.; 29 Sep. 1982; C. Holzschuh leg.; OÖLM. PAKISTAN • 5 ♂♂; Azad Kashmir, Paras, Shogran; 25 May 2019; M. Kafka leg.; BOLD: WPATW 981 - 23; OÖLM / TJWC • 34 ♀♀; same data as for preceding; BOLD: WPATW 983 - 23; OÖLM / TJWC • 1 ♂, 5 ♀♀; Azad Kashmir, Murree, Bhurban; 19 May 2019; M. Kafka leg.; BOLD: WPATW 980 - 23; OÖLM / TJWC • 1 ♀; Hazara, Kawai, Shogran; 14 May 1978; C. Holzschuh leg.; OÖLM • 1 ♀; Murree Hills, Bhurban; 1900 m a. s. l.; 3 May 1978; C. Holzschuh leg.; OÖLM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB5FF840EF36D89FB7CF8B5.taxon	discussion	Remarks As discussed above, A. murreensis is morphologically very close to a cryptic taxon for which the appropriate name is unclear. However, based on the designation of a lectotype for A. murreensis, this species can be recognised as having a range that extends across northern India to central Nepal.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB5FF840EF36D89FB7CF8B5.taxon	distribution	Distribution (true A. murreensis) Pakistan, India * (Himachal Pradesh, Uttarakhand, Meghalaya, Arunachal Pradesh), Nepal *.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB4FF870E8C6D5EFCA9FA42.taxon	materials_examined	Type material Syntype (examined by photograph) PAKISTAN • 1 ♀; Menserah [Mansehra]; 1 – 31 Mar. 1906; F. Benton leg.; type no. 23136; USNM. Other material examined INDIA • 1 ♀; Uttarakhand, Chopta; 2874 m a. s. l.; 6 Mar. 2021; R. K. Gautam leg.; WII. PAKISTAN • 4 ♀♀; Azad Kashmir, Paras, Shogran; 25 May 2019; M. Kafka leg.; OÖLM / TJWC.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB4FF870E8C6D5EFCA9FA42.taxon	discussion	Remarks Andrena subspinigera was described from Mansehra in what is now north-western Pakistan in the foothills of the Himalayas. Cockerell drew comparison with A. spinigera (Kirby, 1802) [= A. trimmerana (Kirby, 1802), see Wood et al. 2022], but considered the two taxa distinct. It was possible to generate a barcode sequence from a specimen collected 40 km north-east of the locus typicus (Shogran). Genetically, A. subspinigera is well differentiated from A. trimmerana, and instead is closest to A. nuptialis Pérez, 1902 (Fig. 11), though is clearly distinct with a genetic distance of 8.50 % (range 8.37 – 8.63 %). It is important to note that the type material of A. subspinigera has the discs of T 1 – 2 almost entirely red-marked, whereas in our examined material, only the marginal areas of T 1 – 2 and a narrow strip immediately anterior to the margins are lightened dull reddish. This is considered to represent variation, as members of the subgenus Hoplandrena Pérez, 1890 are well known for extensive variation in their tergal colouration, leading to the creation of a large number of synonyms (e. g., Gusenleitner & Schwarz 2002; Wood et al. 2022; Wood 2023 a). Meena & Dey (2019) listed A. subspinigera from central India, but provided no supporting evidence. We consider the presence of A. subspinigera in central India unlikely, since all confirmed records come from the Himalayas or close to them, and therefore our records represent the first confirmed records of the species in India.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB4FF870E8C6D5EFCA9FA42.taxon	distribution	Distribution India * (Uttarakhand) and Pakistan (Cockerell 1917).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB7FF860ECB6FEBFD69FDC9.taxon	discussion	Remarks Several authors, beginning with Makkar et al. (2016), reported A. agilissima for India when in fact they were referring to A. fuscosa (see below).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB7FF860ECB6FEBFD69FDC9.taxon	materials_examined	Material examined INDIA • 1 ♀; Ferozepore [Firozpur]; 1 – 31 Mar. 1898; C. G. Nurse leg.; NHMUK (syntype of Melanapis violaceipennis). PAKISTAN • 1 ♀; Quetta; 1 – 31 Mar. 1903; C. G. Nurse leg.; NHMUK (syntype of Melanapis rufifrons).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB7FF860ECB6FEBFD69FDC9.taxon	distribution	Distribution West and Central Palaearctic to Pakistan and India (Punjab, Delhi, Bihar) (Gusenleitner & Schwarz 2002; Ascher & Pickering 2023; Wood 2023 b).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB6FF810EAD6872FACCF981.taxon	materials_examined	Type material Syntypes CHINA • 1 ♀; Yangihissár [= Yengisar]; ZSI; ZSI 0000008643 • 1 ♀; Yarkand [= Yarkant]; NHMUK. Other material examined INDIA • 5 ♀♀; West Himalaya, Khalatse [Khalsi], Kashmir, 4000 – 5000 m a. s. l.; 12 – 23 May 1933; ZMHB.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB6FF810EAD6872FACCF981.taxon	discussion	Remarks It is difficult to find a primary listing for the presence of A. cineraria in India. Andrena cineraria has been treated as a broad taxon, but it represents a species complex including A. barbareae Panzer, 1805 and A. danuvia Stöckhert, 1950. In Asia, “ A. cineraria ” has been reported to be bivoltine (e. g., Osytshnjuk et al. 2008; Xu & Tadauchi 2009 a), and hence is suspected to be more closely related to A. barbareae rather than to A. cineraria (Wood 2023 a). Further study is required, along with genetic analysis (Gueuning et al. 2020). Additionally, the name A. peregrina (Smith, 1878) is available, and is currently listed as a synonym of A. cineraria (Gusenleitner & Schwarz 2002). Originally described as a Lamprocolletes Smith, 1853 (Colletidae), examination of a specimen in the NHMUK collection labelled as “ Type ” in the handwriting of Smith (Fig. 12) shows that this taxon is an Andrena (although the head is missing, the remaining body is clearly a species of Andrena), and is closely affiliated with A. cineraria. Indeed, Warncke (1967: 298) considered A. peregrina to be conspecific with A. cineraria, leading to the current listing of Gusenleitner & Schwarz. However, the published type locality is Yangihissár [Yengisar] in East Turkestan [= Xinjiang in western China], whereas the specimen is labelled as “ Yarkand ” [= Yarkant] which is approximately 100 km to the south-east of Yangihissár. Donald Baker (unpublished thesis: 273) suggested that additional type material of A. peregrina may be in the ZSI, and that this NMHUK specimen is presumably a syntype retained by Smith. Searching through the ZSI website produces a specimen of A. peregrina from Yangihissár (ZSI 0000008643) which is clearly syntypic, but is not automatically considered to be the holotype since the NHMUK specimen is also considered to be a syntype. At the present time, neither specimen is designated as a lectotype. Given the complexity of species delineation within this group of species (Gueuning et al. 2020), no taxonomic action is taken here. The status of A. basifusca Cockerell, 1930 (described from Uzbekistan and currently listed as a valid species by Gusenleitner & Schwarz 2002) must also be resolved in future molecular revisions focusing on Asia. Since Nurse (1904) mentioned that A. cineraria was “ fairly common in Kashmir ”, the species is retained on the Indian list of Andrena in a broad sensu lato, pending further investigations.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB6FF810EAD6872FACCF981.taxon	distribution	Distribution Unclear, possibly restricted to the West Palaearctic or perhaps more broadly across the Palaearctic.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB1FF830ECD6C2AFE85FC2E.taxon	materials_examined	Material examined INDIA • 1 ♀; Ferozepore [Firozpur]; 1 – 31 May 1898; C. G. Nurse leg.; NHMUK (syntype of A. levilabris) • 1 ♀; Ferozepore [Firozpur]; 1 – 31 Mar. 1898; C. G. Nurse leg.; NHMUK (syntype of A. punjaubensis) • 1 ♂; Darjeel. Kalimpong; 20 Apr. 1984; Bahadur leg.; OÖLM • 19 ♀♀; Kalimpong [West Bengal]; 1350 m a. s. l.; 7 May 1981; M. Kraus leg.; OÖLM • 1 ♀; Uttarakhand, 4 km N of Chaumasi; Ukhimath, Gari Village; 30.5288 ° N, 79.1001 ° E; 2588 m a. s. l.; 9 May 2022; R. K. Gautam leg.; WII • 7 ♂♂; Uttarakhand, Kanda Village; 30.4987 ° N, 79.1457 ° E; 1908 m a. s. l.; 16 Nov. 2019; R. K. Gautam leg.; WII • 1 ♂, 1 ♀; Uttarakhand, Chaumasi; 30.6146 ° N, 79.0695 ° E; 2260 m a. s. l.; 8 May 2022; R. K. Gautam leg.; WII • 1 ♀; Uttarakhand, Gwar Village; 30.4931 ° N, 79.1375 ° E; 1404 m a. s. l.; 8 Apr. 2022; R. K. Gautam leg.; WII • 1 ♂; Uttarakhand, Kanchula Kharg; 2680 m a. s. l.; 24 Feb. 2021; R. K. Gautam leg.; WII • 1 ♂; Uttarakhand, Trijuginarayan; 30.6365 ° N, 78.9688 ° E; 2418 m a. s. l.; 2 Jun. 2022; R. K. Gautam leg.; WII • 1 ♀; U. P. [Uttar Pradesh, now Uttarakhand], Himalayas, Harsil; 2700 m a. s. l.; 20 May 1990; S. W. T. Batra leg.; SEMC; SEMC 0988947 • 3 ♀♀; Sikkim; [undated]; C. T. Bingham leg.; ZMHB.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB1FF830ECD6C2AFE85FC2E.taxon	distribution	Distribution West and Central Palaearctic to Pakistan, India (Punjab, Kashmir, Himachal Pradesh, Uttarakhand, West Bengal, Sikkim, Arunachal Pradesh), and China (Gusenleitner & Schwarz 2002; Ascher & Pickering 2023).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB3FF820ED069D7FE0AFE18.taxon	materials_examined	Material examined INDIA • 1 ♂; Uttarakhand, 4 km N of Chaumasi; 30.6368 ° N, 79.0725 ° E; 2588 m a. s. l.; 9 May 2022; R. K. Gautam leg.; WII • 1 ♀; Uttarakhand, Hudu Village; 30.4953 ° N, 79.1559 ° E; 2070 m a. s. l.; 4 Jun. 2022; R. K. Gautam leg.; WII • 1 ♀; Uttarakhand, Duggalbitta; 30.4883 ° N, 79.1738 ° E; 2607 m a. s. l.; 10 Apr. 2022; R. K. Gautam leg.; WII • 1 ♀; Gulmarg; summer 1913; NHMUK • 1 ♀; Kashmir; 6000 – 7000 ft a. s. l.; 1 – 30 Aug. 1901; C. G. Nurse leg.; NHMUK (syntype of A. patella) • 1 ♀; Ladakh, Ronding; 14 Aug. 1977; M. Kraus leg.; OÖLM • 1 ♀; Matiana, Simla Hills; 8000 ft a. s. l.; 28 – 30 Apr. 1907; ZSI; ZSI 0000008660; (syntype of A. brunneipennis) • 1 ♀; Matiana, Simla Hills; 8000 ft a. s. l.; 28 – 30 Apr. 1907; ZMHB (syntype of A. brunneipennis) • 1 ♀; West Himalaya, Khalatse [Khalsi], Kashmir; 4000 – 5000 m a. s. l.; 6 May 1939; OÖLM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB3FF820ED069D7FE0AFE18.taxon	discussion	Remarks Wood & Monfared (2022) synonymised A. patella with A. induta. To date, confirmed Indian records come from high altitude sites in the Western Himalayas. The holotype depository for A. brunneipennis was unclear (see Wood 2024 b), but a syntype (marked as a paratype by Warncke) was examined in the ZMHB collection (Fig. 13), and examination of photographs from the ZSI website indicate that a syntype is present there as well. These specimens confirm that the currently accepted synonymy (see Warncke 1967) with A. induta is correct. A lectotype is currently undesignated, but the type depository for A. brunneipennis can be considered to be the ZSI based on the original intent of Bingham (1908).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB3FF820ED069D7FE0AFE18.taxon	distribution	Distribution Turkey, Armenia, Iran, Turkmenistan, Uzbekistan, Tajikistan, Afghanistan, India (Jammu & Kashmir, Ladakh, Himachal Pradesh, Uttarakhand) (Osytshnjuk et al. 2008; Astafurova et al. 2021; Wood & Monfared 2022).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB2FF820EAA6BBDFC8AFDC9.taxon	distribution	Distribution Northern India (Ladakh, Uttarakhand) and Nepal.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB2FF820E846872FDCCFBDC.taxon	materials_examined	Material examined INDIA • 2 ♂♂, 1 ♀; Uttarakhand, Dever Village; 1934 m a. s. l.; 9 Mar. 2021; R. K. Gautam leg.; WII • 3 ♀♀; Uttarakhand, Hudu Village; 30.4953 ° N, 79.1559 ° E; 2098 m a. s. l.; 20 Mar. 2021; R. K. Gautam leg.; WII • 6 ♂♂, 2 ♀♀; Uttarakhand, Kanda Village; 30.4987 ° N, 79.1457 ° E; 1908 m a. s. l.; 26 Feb. 2021; R. K. Gautam leg.; WII • 3 ♂♂, 9 ♀♀; Uttarakhand, Hudu Village; 30.4953 ° N, 79.1559 ° E; 2098 m a. s. l.; 13 Jul. 2021; R. K. Gautam leg.; WII.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB2FF820E846872FDCCFBDC.taxon	distribution	Distribution India * (Uttarakhand), Nepal, China, South Korea, Russia (Far East), Japan (Tadauchi & Matsumura 2007; Ascher & Pickering 2023).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB2FFBD0EE96E81FDD0FED1.taxon	materials_examined	Type material Holotype INDIA • ♀; W. India, Lonavala; 1 – 30 Nov. 1964; NHMUK. Paratypes INDIA • 1 ♂; W. India, Lonavala; 1 – 30 Nov. 1964; SEMC; SEMC 0977708 • 1 ♀; same data as for preceding; SEMC; SEMC 0977709. Other material examined INDIA • 2 ♀♀; W Ghats, Lonavala; 1 Jan. 1967; OÖLM • 4 ♂♂; W Maharashtra st. 70 km SSW of Pune; 1400 m a. s. l.; 30 Sep. 2005; J. Bezděk leg.; OÖLM • 1 ♂; W Ghats, Lonavala; 620 m a. s. l.; 9 Nov. 1961; SEMC; SEMC 0981603 • 1 ♀; same data as for preceding; SEMC; SEMC 0981604.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB2FFBD0EE96E81FDD0FED1.taxon	discussion	Remarks Placement in the subgenus Notandrena Pérez, 1890 requires further support (see Wood 2023 a).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FFB2FFBD0EE96E81FDD0FED1.taxon	distribution	Distribution India (Maharashtra) (Wood 2023 a).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8DFFBC0EF56B7AFDB8FED1.taxon	materials_examined	Type material Syntype INDIA • 1 ♂; Ferozepore [Firozpur]; 1 – 28 Feb. 1898; C. G. Nurse leg.; NHMUK.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8DFFBC0EF56B7AFDB8FED1.taxon	discussion	Remarks Examination of type material supports continued placement in Notandrena due to the yellow male clypeus, broadened gena, and pronotum with humeral angle (previously placed in the now synonymous subgenus Carandrena Warncke, 1968).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8DFFBC0EF56B7AFDB8FED1.taxon	distribution	Distribution India (Punjab) (Cameron 1908).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8CFFBC0EF76B7AFC0BFBDC.taxon	materials_examined	Material examined CHINA • 1 ♂, 1 ♀; Gyangtse [Gyantse]; 13 000 ft a. s. l.; 1 – 30 Jun. 1904; H. J. Walton; NHMUK • 2 ♂♂; Khamba Jong., Sikkim [Kampa Dzong, actually in Tibet]; 15 000 – 16 000 ft a. s. l.; 15 – 30 Jul. 1903; Tibet Exped. leg.; NHMUK. INDIA • 5 ♀♀; Thangu; 3800 m a. s. l.; 1 – 12 Jul. 1938; Schäfer. Exped. leg.; ZMHB • 2 ♂♂, 4 ♀♀; Thangu R.; 2 Jul. 1938; Schäfer. Exped. leg.; ZMHB • 8 ♂♂, 3 ♀♀; Sikkim, Teesta Vy., Tungu, [Thangu]; 13 000 – 14 000 ft a. s. l. [= 4000 – 4200 m a. s. l.]; 1 – 15 Jul. 1903; Tibet Expedition leg.; NHMUK.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8CFFBC0EF76B7AFC0BFBDC.taxon	discussion	Remarks Material in the ZMHB was separated as “ Andrena dictyonota n. sp. ”, an unpublished name. The location of “ Tungu ” as indicated on the NHMUK specimens from Sikkim is unclear, but based on the recorded altitude (ca 4000 – 4200 m a. s. l.), it likely refers to Thangu Valley which is found 3900 m a. s. l. Some specimens marked as “ Sikkim ” occur slightly further north in what is now Chinese Tibet.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8CFFBC0EF76B7AFC0BFBDC.taxon	distribution	Distribution India * (Sikkim) & China (Xizang, Yunnan) (Xu et al. 2000).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8CFFBC0E4C6E81FC20F97D.taxon	materials_examined	Material examined INDIA • 1 ♀; NW India, U. P. [Uttar Pradesh, now Uttarakhand], Kumaon, Naini Tal [Nainital]; 2000 – 2480 m a. s. l.; 3 – 11 Oct. 1978; M. A. Lieftinck leg.; RMNH; RMNH. INS. 1266150. NEPAL • 1 ♀; Trisuli Khola-Langtang Kh., Syabru Bensi [Syapru Besi]; 1600 m a. s. l.; 29 Sep. 1982; C. Holzschuh leg.; OÖLM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8CFFBC0E4C6E81FC20F97D.taxon	discussion	Remarks One specimen was found in the RMNH collection that matches material from central Nepal (Syapru Besi) that is relatively close (ca 100 km) to the locus typicus of Namche Bazar (Tadauchi & Masumura 2007).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8CFFBC0E4C6E81FC20F97D.taxon	distribution	Distribution India * (Uttarakhand) and Nepal (Tadauchi & Matsumura 2007).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8CFFBF0EE66D26FC79FC59.taxon	materials_examined	Type material Holotype INDIA • ♀; Masuri [Mussoorie]; NHMUK. Other material examined INDIA • 1 ♂; Simla [Shimla]; 1 – 30 Aug. 1898; NHMUK (holotype of A. simlaensis) • 1 ♂, 1 ♀; Simla [Shimla]; 1 – 30 Aug. 1898; ZMHB. NEPAL • 1 ♀; Anapurnareg., Ghorepani [Ghode Pani]; ca 2850 m a. s. l.; 12 Oct. 1999; W-H. Liebig leg.; WHLC • 1 ♀; Anapurnareg., Koto; ca 2600 m a. s. l.; 29 Sep. 1999; W-H. Liebig leg.; WHLC • 1 ♀; Kathmandu, Pulchoki, SE of Godavari; 2760 m a. s. l.; 14 Sep. 1995; E. and J. Hüttinger leg.; E. Scheuchl det.; OÖLM • 3 ♀♀; Kathmandu, SE of Kathmandul, Pulchoki; 1860 – 2500 m a. s. l.; 23 Sep. 1993; E. and J. Hüttinger leg.; E. Scheuchl det.; OÖLM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8CFFBF0EE66D26FC79FC59.taxon	distribution	Distribution India (Himachal Pradesh, Uttarakhand, Arunachal Pradesh) & Nepal *.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8FFFBF0ED86E02FB1CF88C.taxon	materials_examined	Material examined INDIA • 3 ♀♀; Uttarakhand, Nanu Chatti, Gaundhar; 30.61126 ° N, 79.2013 ° E; 2440 m a. s. l.; 5 Oct. 2021; R. K. Gautam leg.; WII • 1 ♂, 3 ♀♀; Thangu; 3800 m a. s. l.; 1 – 12 Jul. 1938; Schäfer. Exped. leg.; ZMHB • 1 ♀; Thangu R.; 2 Jul. 1938; Schäfer. Exped. leg.; ZMHB.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8FFFBF0ED86E02FB1CF88C.taxon	discussion	Remarks Specimens from Uttarakhand were collected from Geranium wallichianum Oliv. (Geraniaceae Juss.) (Fig. 14). Females of A. submontana present partially plumose tibial scopal hairs and long spines on the posterior face of the hind femorae, along with two other Himalayan species of Oreomelissa (Xu et al. 2000). This is reminiscent of the characters present in some members of the unrelated subgenus Avandrena Warncke, 1968 that are specialised on Erodium L'Hér. ex Aiton (Geraniaceae) (Wood 2023 b). Further study is required to assess whether these structures are functionally involved in pollen collection, and thus represent convergent evolution, since spined Avandrena and Oreomelissa are strongly separated phylogenetically (Pisanty et al. 2022 b). Specimens from Sikkim collected by the Schäfer Expedition were separated as “ Andrena adicyta n. sp. ”, an unpublished name.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8FFFBF0ED86E02FB1CF88C.taxon	distribution	Distribution India * (Uttarakhand, Sikkim) & China (Xizang, Qinghai, Yunnan) (Xu et al. 2000).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8FFFB90EDC6D31FC82FDE6.taxon	materials_examined	Type material Holotype INDIA • ♀; Masuri [Mussoorie]; NHMUK. Other material examined INDIA • 1 ♀; Simla, Matiana; ZSI; ZSI 0000008659 (syntype of A. burkelii). NEPAL • 1 ♀; Nawakot, Langtang Khola, Ghora Tabela; 3200 m a. s. l.; 3 Oct. 1982; C. Holzschuh leg.; OÖLM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8FFFB90EDC6D31FC82FDE6.taxon	discussion	Remarks Gusenleitner & Schwarz (2002) did not place A. morosa within a subgenus, and Tadauchi & Matsumura (2007) placed it within Andrena s. str. Fabricius, 1775. Examination of the holotype (Fig. 15) shows that it presents a strongly emarginate process of the labrum, and the ventral pollen-collecting hairs of the tibial scopae are long and strongly plumose. This strongly suggests placement in the subgenus Pallandrena Warncke, 1968, members of which are associated with the family Geraniaceae which has large pollen grains, necessitating sparse plumose scopal hairs. Indeed, the hind legs of the type specimen are covered in large grains that are visible to the naked eye (Fig. 15 D). We therefore place A. morosa into the subgenus Pallandrena, and hypothesise that it will be caught on Geranium or related genera in the future. Andrena burkelii was described from Matiana from an altitude of 8000 feet. Bingham mentioned that it had a body length of 12 mm, and that it was unlike any other species of Andrena he had examined. Possible type depositories were discussed by Wood (2024 b), but examination of the ZSI website revealed type material in that collection. Examination of the photographed female specimen (possibly automatically the holotype, but it is unclear how many specimens are included in the type series) shows that it displays the same characters (particularly the same body size, pubescence, and decoloured tergal margins) as A. morosa, and indeed the hind tibial scopae are completely covered in the same large pollen grains. Within the Western Himalayas, the locus typicus of Matiana is only 100 km to the north-west of Mussoorie, the locus typicus of A. morosa. We therefore synonymise A. burkelii with A. morosa syn. nov.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8FFFB90EDC6D31FC82FDE6.taxon	distribution	Distribution India (Himachal Pradesh, Uttarakhand) & Nepal *.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF89FFB80EFB688FFEC2FDF1.taxon	materials_examined	Material examined INDIA • 1 ♂; West Himalaya, Khalatse [Khalsi], Kashmir [Ladakh]; 4000 – 5000 m a. s. l.; 6 May 1939; K. Warncke det.; OÖLM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF89FFB80EFB688FFEC2FDF1.taxon	discussion	Remarks Members of the subgenus Plastandrena Hedicke, 1933 are fiendishly difficult to identify due to variable morphology without clear and consistent ‘ break points’ between species and lack of resolution using genetic barcoding (e. g., Wood 2023 b). Further study using more powerful genetic techniques (e. g., including nuclear DNA) is required, but A. mongolica can be reported from India based on a male specimen presenting the characteristic genital capsule of this species.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF89FFB80EFB688FFEC2FDF1.taxon	distribution	Distribution Turkey, Azerbaijan, Iran, Afghanistan, Kyrgyzstan, Kazakhstan, China, India * (Ladakh) (Astafurova et al. 2021).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF88FFB80EDD689AFAB9FAF7.taxon	materials_examined	Material examined PAKISTAN • 1 ♀; Peshin [Pishin]; 1 – 30 Apr. 1903; C. G. Nurse leg.; NHMUK (syntype of A. dolorosa) • 1 ♀; 1 – 31 Mar. 1902; C. G. Nurse leg.; NHMUK (syntype of A. dolorosa).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF88FFB80EDD689AFAB9FAF7.taxon	discussion	Remarks Nurse (1904) noted that this species was “ very common at all elevations in Kashmir ”. We have not been able to examine any material, but retain this listing as the distribution is plausible, since the species is widespread across Central Asia south to Pakistan, and Kashmir contains more Palaearctic faunal elements than the remaining parts of the Indian Himalayas.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF88FFB80EDD689AFAB9FAF7.taxon	distribution	Distribution West and Central Palaearctic to China (Gusenleitner & Schwarz 2002; Ascher & Pickering 2023).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF88FFBB0E456F9CFB80F94F.taxon	materials_examined	Material examined INDIA • 1 ♂, 9 ♀♀; Uttarakhand, Chandrabani; 30.2830 ° N, 77.9753 ° E; 662 m a. s. l.; 10 Apr. 2023; R. K. Gautam leg.; WII • 1 ♀; Punjab, Chandigarh; 17 Apr. 1965; S. W. T. Batra leg.; SEMC; SEMC 0981602 • 1 ♀; Uttarakhand, Dehradun, Forest Research Institute; 18 Apr. 2023; @ ashirwad; https: // www. inaturalist. org / observations / 155452620. NEPAL • 1 ♀; Kathmandu Valley, Nagarjong [Nagarjun]; 1500 – 1700 m a. s. l.; 1 May 1967; Dieri-Forster-Schacht leg.; OÖLM • 1 ♀; Kathmandu; 1800 m a. s. l.; 29 Apr. 1996; OÖLM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF88FFBB0E456F9CFB80F94F.taxon	description	Description Male BODY. Length: 6 mm (Fig. 10 A). HEAD. Dark, 1.2 × as wide as long (Fig. 10 B). Clypeus domed, irregularly punctate, punctures separated by 0.5 puncture diameters laterally, by 0.5 – 2 puncture diameters medially, underlying surface smooth and shining. Process of labrum rounded rectangular, 2 × as wide as long, fore margin emarginate, surface smooth and shining. Gena equalling width of compound eye; ocelloccipital distance 1.5 × diameter of lateral ocellus. Head medially with short pale pubescence, with scatter black hairs laterally; gena, vertex, and scape with pale hairs. Antennae dark, A 4 – 13 ventrally lightened by presence of dark orange-brown scales. A 3 much shorter than A 4, approximately ½ length, A 4 – 13 elongate, rectangular, 2 × as long as broad (Fig. 10 C). MESOSOMA. Scutum laterally and anteriorly microreticulate, dull, sculpture becoming weaker medially, here smooth and shining; surface punctate, punctures separated by 0.5 – 1 puncture diameters, obscure laterally, becoming more visible medially (Fig. 10 D). Scutellum with weaker sculpture, predominantly smooth and shining, punctation equal. Pronotum rounded. Mesepisternum microreticulate, overlain by network of moderately raised irregular rugosity, dull. Propodeum strong microreticulation overlain by dense network of raised rugosity, propodeal triangle broad, laterally defined by raised carinae, internal surface with raised rugosity, not strongly differentiated from sculpture of dorsolateral parts of propodeum. Mesosoma covered with pale, weakly plumose hairs. Legs dark, apical tarsal segments lightened brownish, pubescence whitish. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma orange-brown, venation dark orange, nervulus strongly antefurcal. METASOMA. Tergal discs dark, apical rims moderately lightened hyaline-brown yellow (Fig. 10 E). Tergal discs and margins finely and densely punctate, punctures separated on average by 1 puncture diameter, underlying surface finely shagreened, shining. Declivity of T 1 with erect white hairs, T 2 – 4 with weak apical hairbands of moderately long white hairs, condition degraded, widely separated on T 2 – 3, nominally complete on T 4. T 6 – 7 with light brown hairs overlying pseudopygidial plate of T 7. S 8 columnar, apex slightly broadened, rounded; ventral surface with lateral fan of brown hairs. Genital capsule somewhat elongate, gonocoxae weakly produced into rounded teeth apically, gonostyli basally narrow, broadening apically, spatulate, with inner margin raised (Fig. 10 F). Penis valves parallel sided, occupying ½ space between gonostyli.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF88FFBB0E456F9CFB80F94F.taxon	discussion	Remarks Newly recorded from India, including the first record of the male, which is described here. Tadauchi & Matsumura (2007) recorded the species from January and February only, and our late capture dates indicate that the species may be bivoltine, or at least sporadic in emergence. Diagnosis against A. tungnatha is provided above, and the male sex is described here.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF88FFBB0E456F9CFB80F94F.taxon	distribution	Distribution India * (Punjab, Uttarakhand) and Nepal (Tadauchi & Matsumura 2007).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8BFFBB0EA36CF4FC8DF800.taxon	distribution	Distribution Northern India (Himachal Pradesh, Uttarakhand).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8AFFBA0ECB6B0BFB2BFB94.taxon	materials_examined	Material examined INDIA • 1 ♀; Ferozepore [Firozpur]; 1 – 31 Mar. 1898; C. G. Nurse leg.; NHMUK (syntype of A. ilerda) • 1 ♂; Ferozepore [Firozpur]; 1 – 28 Feb. 1898; C. G. Nurse leg.; NHMUK (syntype of A. ferozeporensis) • 13 ♂♂; Rajasthan, Sawai Madhopur Rathambhore; 308 m a. s. l.; 15 Jan. 2005; D. Brzoska leg.; SEMC; SEMC 0648294 to SEMC 0648306 • 14 ♀♀; same data as for preceding; SEMC 0648307 to SEMC 0648320.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8AFFBA0ECB6B0BFB2BFB94.taxon	distribution	Distribution From the Canary Islands (Spain), across North Africa, France (Corsica), Italy (Sardinia and mainland), the Levant, the Arabian Peninsula, Iran, Iraq, Afghanistan, Pakistan, to India (Gujarat, Rajasthan, Punjab, Bihar, Arunachal Pradesh) (Gusenleitner & Schwarz 2002; Wood & Monfared 2022).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8AFFB50ED96E3EFD3EFC48.taxon	materials_examined	Type material Syntypes EGYPT • 3 ♀♀; Cairo; 13 Mar. – 4 Apr. 1899; F. Morice leg.; OÖLM / ZMHB. Other material examined INDIA • 1 ♂; Banhar, Bihar; 8 Apr. 1919; H. Inglis leg.; NHMUK (lectotype by present designation of A. comberima beharica, Fig. 16) • 1 ♀; Banhar, Bihar; 10 Apr. 1919; H. Inglis leg.; NHMUK (syntype of A. comberima beharica) • 1 ♀; Deesa [Gujarat]; 1 – 31 Mar. 1901; C. G. Nurse leg.; NHMUK. JORDAN • 1 ♀; Wadi Ramm [Wadi Rum]; 12 May 1995; K. Deneš leg.; OÖLM. OMAN • 1 ♀; Muscat, Bosher sand; 7 Feb. 2020; A. Al Jahdhami leg.; NHMUK. PAKISTAN • 1 ♀; Krchi [Karachi]; NHMUK (syntype of A. comberima). TUNISIA • 6 ♂♂, 1 ♀; Nefta [Naftah]; 20 – 22 May 1993; J. Batelka leg.; OÖLM • 2 ♂♂; Tozeur; 24 Mar. 1978; K. M. Guichard leg.; NHMUK • 2 ♀♀; SE Ben Guardane; 13 Apr. 2001; M. Halada leg.; E. Scheuchl det.; OÖLM • 1 ♂, 2 ♀♀; Hazoua, 30 km W of Nefta [Naftah]; 19 Apr. 1996; J. Batelka and H. Podrouzkova leg.; E. Scheuchl det.; OÖLM. UNITED ARAB EMIRATES • 2 ♂♂, 2 ♀♀; Al Ain Road; 1 Feb. 1985; J. N. B. Brown leg.; NHMUK • 1 ♂, 1 ♀; Sarjan Province, W of Dhaid; 120 m a. s. l.; 2 Mar. 2017; Ma. Halada leg.; OÖLM • 1 ♂, 1 ♀; Umm al Qaywayn, Biatah; 50 m a. s. l.; 3 Mar. 2017; Ma. Halada leg.; OÖLM • 1 ♂; Suweihan [Sweihan]; 21 Mar. 1986; I. L. Hamer leg.; NHMUK.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8AFFB50ED96E3EFD3EFC48.taxon	discussion	Remarks Andrena aegyptiaca is a complex of species that is currently undergoing revision (e. g., Pisanty et al. 2023). If the eastern populations are separated, A. comberima will be the priority name. For now, a broad A. aegyptiaca concept is maintained. As the type material of A. comberima beharica was located in the NHMUK collection, a male is here designated as the lectotype (Fig. 16).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF8AFFB50ED96E3EFD3EFC48.taxon	distribution	Distribution (A. aegyptiaca sensu lato) Spain (Canary Islands), Morocco, Algeria, Tunisia *, Egypt, Israel, Jordan *, Saudi Arabia, United Arab Emirates *, Oman *, Pakistan, India (Gujarat, Punjab, Himachal Pradesh, Bihar) (Gusenleitner & Schwarz 2002; Ascher & Pickering 2023).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF84FFB40EEF6B0BFDFBFD7B.taxon	description	Bingham (1897); Cameron (1897); Cockerell (1923); Meena & Dey (2019); Chandra et al. (2021); Ascher & Pickering (2023).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF84FFB40EEF6B0BFDFBFD7B.taxon	discussion	Remarks The identity of A. anonyma is completely unclear, as no-one has examined the type material. It must remain a mystery taxon until the type material becomes available for study.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF84FFB40EEF6B0BFDFBFD7B.taxon	distribution	Distribution India (Uttarakhand) (Cameron 1897).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF84FFB40ED46920FD99F941.taxon	materials_examined	Material examined Holotype INDIA • ♀; Leh; ZSI; ZSI 0000008658.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF84FFB40ED46920FD99F941.taxon	discussion	Remarks Gusenleitner & Schwarz (2002) indicate that the NHMUK is the type depository, but no entry exists in the type register and no specimens could be found here. Baker (unpublished thesis) indicates that type material could be in the ZSI, but he never saw any such material. Examination of the ZSI website indicates that a specimen is present in that collection. The specimen is in very poor condition with the metasoma having been partially glued back onto the mesosoma and with the hind legs missing, but a dorsal view of the head shows the presence of facial foveae, even though Smith described the species in the male sex. Examination of the illustrated plate provided by Smith (1878: fig. 4, as A. floricula sic) indicates that the type specimen may actually be female, since the hind legs (not preserved in the ZSI specimen) seem to present scopae. We therefore consider this specimen to be the holotype, and actually be a female despite Smith’s original description as a male specimen.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF84FFB40ED46920FD99F941.taxon	distribution	Distribution India (Ladakh) (Smith 1878).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF84FFB70EE86CEAFA83FDAB.taxon	discussion	Remarks This is a mystery taxon, as no-one appears to have examined the type material. Wood (2024 b) speculated, based on the description, its reported large body length of 15 mm, and the illustration provided by Cameron (1897: pl. 4, fig. 19), that it could potentially be the senior name for A. kishidai chagyabensis. However, type location and examination are required before any further action can be taken.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF84FFB70EE86CEAFA83FDAB.taxon	distribution	Distribution India (Uttarakhand) (Cameron 1897), doubtfully listed from West Bengal by Bingham (1908).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF87FFB70EDE6850FD12FA54.taxon	materials_examined	Type material Syntype INDIA • 1 ♀; Ferozepore [Firozpur]; 1 – 31 Mar. 1898; C. G. Nurse leg.; NHMUK. Other material examined INDIA • 1 ♀; Ferozepore [Firozpur]; 1 – 31 Mar. 1898; C. G. Nurse leg.; NHMUK (previously undetermined).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF87FFB70EDE6850FD12FA54.taxon	discussion	Remarks Although the type of A. leaena is conserved, its subgeneric affiliation remains unclear, and the male has never been described. Additional study is required to clarify the nature of this species. Though the specimen in the NHMUK type collection is labelled “ type ”, it is not automatically the holotype and is only a syntype as Cameron (1907) indicates that specimens come from both Ferozepore and Deesa, indicating that multiple individuals were used to describe the species.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF87FFB70EDE6850FD12FA54.taxon	distribution	Distribution India (Punjab, Gujarat) (Cameron 1907).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF87FFB70EEA6FF9FDD0F8B7.taxon	distribution	Distribution India (Maharashtra) (Wood 2023 a).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF87FFB60EFB6D5CFDC0FBC7.taxon	materials_examined	Type material Syntype (possibly automatically the holotype, type no. 23136, photograph examined) INDIA • 1 ♀; Kashmir, Ladakh, Rupshu; 16 000 ft a. s. l. [= ca 5000 m a. s. l.]; 22 Jul. 1897; W. L. Abbott leg.; USNM.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF87FFB60EFB6D5CFDC0FBC7.taxon	discussion	Remarks and diagnosis Andrena rupshuensis is a mysterious species that was previously known only from the type series (an unclear number of specimens, not specified in the original publication). It was collected at the extremely high altitude of nearly 5000 meters above sea level which would be one of if not the highest ever reported altitude for a species of Andrena. The morphology (photographs available on the Smithsonian website https: // collections. nmnh. si. edu / search / ento /) does not conform to any current species concept, as the malar space is clearly elongate and shining. Some of the morphology is reminiscent of the subgenus Taeniandrena Hedicke, 1933, specifically the broad facial foveae, the terga with clear apical hairbands, the general rounded shape of the body and ovoid metasoma, the shortish light brown hairs on the scutum contrasting white hairs on the mesepisternum, and the tarsi which are lightened orange. However, the clypeus appears to be shining with a broad impunctate mid-line which is not known from the subgenus. Further study is required, as well as new collections at high altitude around the locus typicus.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF87FFB60EFB6D5CFDC0FBC7.taxon	distribution	Distribution India (Ladakh) (Cockerell 1911 a).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF86FFB60EC36EA5FDBBF9F5.taxon	discussion	Remarks Andrena helvola is a predominantly West Palaearctic species, with some populations extending into the East Palaearctic in the Altai mountains of Kazakhstan and also into northern China (Heilongjiang; Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005; Xu & Tadauchi 2012; Ascher & Pickering 2023; Wood, unpublished data). It is therefore considered implausible that A. helvola is present in northern India, since even in Ladakh the nearest verified populations would be over 1500 km away in the Altai mountains. Meena & Dey (2019) present no evidence to support this listing, and so this species is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF86FFB10EC66C9EFAF8FEC6.taxon	discussion	Remarks Andrena praecox is a predominantly West Palaearctic species, with some populations extending into northern Kazakhstan. The south-eastern limit is the Caucasus region (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005; Ascher & Pickering 2023). It is considered implausible that this species of temperate woodland is present in Bihar, a state with a predominantly subtropical climate. Meena & Dey (2019) present no evidence to support this listing, and so this species is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF81FFB10D376B6FFAF8FD51.taxon	discussion	Remarks Andrena varians is a predominantly West Palaearctic species, with some populations extending into Kazakhstan (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005; Shebl & Tadauchi 2009; Ascher & Pickering 2023). It is considered implausible that this species is present in northern India. Meena & Dey (2019) present no evidence to support this listing, and so this species is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF81FFB10EC168FAFB95FBDC.taxon	discussion	Remarks Andrena florea is a West Palaearctic species extending east into Iran and southern Turkmenistan (Gusenleitner & Schwarz 2002; Wood & Monfared 2022; Ascher & Pickering 2023). It is considered implausible that this species is present in northern India. Meena & Dey (2019) present no evidence to support this listing, and so this species is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF81FFB10ED36E86FA9BFA67.taxon	description	Listed by Meena & Dey (2019): North India, Bihar.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF81FFB10ED36E86FA9BFA67.taxon	discussion	Remarks Andrena taraxaci is a West Palaearctic species, with the East Asian subspecies ssp. orienticola Strand, 1915 currently in the process of being elevated (Schwenninger 2015; Xu & Tadauchi 2002; Wood, in review). There is no evidence to suggest that this species as currently defined is present in India (or indeed any Chlorandrena species at all), and so this species is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF81FFB10ECC6C0CFCCDF838.taxon	description	Listed by Meena & Dey (2019): Punjab, North India.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF81FFB10ECC6C0CFCCDF838.taxon	discussion	Remarks Andrena bicolor is a widespread West and Central Palaearctic species that extends to parts of Central Asia (though due to its highly variable nature, several distinct, cryptic, and / or unrecognised species have been confused with it, see Praz et al. (2019) and Wood (2023 c) for examples). The exact distribution is unclear, but given the enormous confusion surrounding this taxon and its scarcity in Central Asia based on examined material, and the lack of specific supporting records by Meena & Dey (2019), true A. bicolor is considered unlikely to be present in India.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF80FFB00D3B6B0BFCFEFD10.taxon	discussion	Remarks Andrena rosae is a widespread Palaearctic species, found from Europe to Japan, including Central Asia (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008). However, in Central Asia it is found in mountainous regions only. Ecologically, it is very difficult to consider that it could be present in the lowlying state of Bihar (average elevation 53 m a. s. l.). Due to the lack of supporting records presented by Meena & Dey (2019), it is removed from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF80FFB00EE568B5FDC2FBBE.taxon	discussion	Remarks Andrena barbilabris is a widespread Holarctic species, found from Europe to North America (Ascher & Pickering 2023). However, in Asia it is found in the north, and in the Central Palaearctic is found only in Kazakhstan, Russia, and Mongolia (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008; Xu & Tadauchi 2009 b). Its reported presence in Bihar is therefore considered to be impossible, and the species is removed from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF80FFB00ED46E67FB92F9B2.taxon	discussion	Remarks Andrena ventralis is a trans-Palaearctic species of temperate woodland that ranges from Europe to China and Japan, including parts of Central Asia (Gusenleitner & Schwarz 2002; Xu & Tadauchi 2005; Ascher & Pickering 2023). In Central Asia, the species is sporadic in occurrence and more or less restricted to mountainous areas due to its ecological specialisation on Salix L. (Salicaceae Mirb.) that is principally found in areas with a temperate climate. Meena & Dey (2019) present no evidence to support the presence of this species in India. It is not impossible that it could be present in Kashmir, but without verified specimen records, A. ventralis is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF80FFB30EFF6C58FC8BFE18.taxon	discussion	Remarks As discussed above, the listing of Meena & Dey (2019) probably refers to multiple misidentified specimens including A. fuscosa and possibly A. kedarnatha Wood & Gautam sp. nov. There is currently no evidence that true A. cussariensis has a range that extends south of Central Asia.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF80FFB30EFF6C58FC8BFE18.taxon	distribution	Distribution Crimea, Russia (European part to the Urals), Turkey, the Caucasus (Georgia, Armenia, Azerbaijan), Iran, Central Asia (Kazakhstan, Uzbekistan, Tajikistan, Kyrgyzstan), Mongolia (Osytshnjuk et al. 2008; Astafurova et al. 2021; Wood & Monfared 2022).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF83FFB30EFB6BBDFD7AFD5C.taxon	discussion	Remarks Meena & Dey (2019: 444) introduced the name A. delrragarensis with no description or further details. It is clearly a nomen nudum and is ignored here.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF83FFB30ECA6901FD4CFBC4.taxon	discussion	Remarks There is no evidence supporting the listing of A. gravida in India. Andrena gravida is a West Palaearctic species of temperate woodland, with no records outside of this area (Gusenleitner & Schwarz 2002). It is therefore excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF83FFB30EC66E69FE56FA4F.taxon	discussion	Remarks Andrena limata is a widespread Palaearctic species, extending from the West Palaearctic into most countries in Central Asia (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008). It is possible that A. limata could be present in Kashmir, without available specimens it cannot be confidently listed as part of the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF83FFB30EE06FF4FC4DF820.taxon	discussion	Remarks The listing of A. nigroaenea in India may come from a misidentified A. induta; for example, Bingham (1908: 363) in the description of the junior synonym A. brunneipennis mentioned that it was: “ Nearest to probably a race of the European A. nigroaenea, Kirby, but larger; the colour of the pubescence and of the wings different ”. Andrena nigroaenea is a West Palaearctic species, with no records from Central Asia (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008). The listing from Bihar is not considered to be plausible, and the species is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF82FFB20ECC6B0BFBC7FDC9.taxon	discussion	Remarks Andrena nitida is a predominantly West Palaearctic species, extending into Central Asia only in Kazakhstan (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008). The listing from Bihar is not considered to be plausible, and the species is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF82FFB20E876873FD7AFC0E.taxon	discussion	Remarks Meena & Dey (2019: 444) introduced the name A. pantragarensis with no description or further details. It is clearly a nomen nudum and is ignored here.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF82FFB20ECC69B7FE0AFABC.taxon	discussion	Remarks Meena & Dey (2019) present no evidence to support a listing of this species from India. Andrena oralis is a species of open grasslands and steppe from Central Europe to Turkey and Central Asia (Gusenleitner & Schwarz 2002; Astafurova et al. 2022). It is not impossible that A. oralis could be present in Kashmir, but it is ecologically inconceivable that it will be present in Bihar, and so this species is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF82FFB20EF76F61FD0CF8B1.taxon	discussion	Remarks Makkar et al. (2016) reported A. agilissima as new for India based on specimens from Muktsar in western Punjab. They also provided a barcode for this species (Genbank: KT 960836). However, this specimen is A. fuscosa, both based on the morphology presented in photographs, and also based on the barcode which clusters with specimens from Spain and Israel (Wood 2023 b). Andrena agilissima (eastern limit in Hungary and Poland) is therefore excluded from the Indian fauna, with these Indian records referring instead to A. fuscosa.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF82FFAD0EEA6D5AFAEAFD0E.taxon	discussion	Remarks Andrena bimaculata is a widespread but taxonomically challenging species. There is a great deal of uncertainty concerning species boundaries in the subgenus Plastandrena, with enormous colour variation in females that is not associated with commensurate variation in male genital capsules. Some members of the Plastandrena, including A. bimaculata and A. tibialis, are also seemingly not always identifiable using barcodes (Schmidt et al. 2015; Wood 2023 b). In this context, A. bimaculata appears to have a predominantly Palaearctic distribution extending from the West Palaearctic to Mongolia, including Central Asia (Gusenleitner & Schwarz 2002; Ascher & Pickering 2023). However, this requires investigation using more powerful genetic techniques (e. g., using nuclear DNA). However, given the nominal distribution, the biogeography, the taxonomic uncertainty, and the lack of supporting specimen records, A. bimaculata cannot be accepted as present in Bihar, and is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9DFFAD0EF468B7FBFBFB6C.taxon	discussion	Remarks Cockerell (1920) described A. peridonea from Hangu which is now in the state of Pakistan (Khyber Pakhtunkhwa Province). Although Gusenleitner & Schwarz (2002) indicated that the type depository was unclear, the type material is actually in the USNM collection (USNMENT 00533689). Photographs on the Smithsonian website (https: // collections. nmnh. si. edu / search / ento /) indicate that the species is part of the subgenus Plastandrena, though since it was described in the female sex only it is very difficult to comment on its specific status due to the numerous taxonomic problems in this subgenus, particularly in the female sex (2023 b). Meena & Dey (2019) list the species from northern India, but without any specimen records. This listing likely refers to the concept of pre-partition British India (the title of Cockerell’s paper is “ Some Indian bees of the genus Andrena ”), and hence the species is excluded from the contemporary Indian fauna pending precise specimen records.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9DFFAD0ED76F11FED7F9B1.taxon	discussion	Remarks The situation for A. tibialis is the same as for A. bimaculata, and the species is also excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9DFFAD0ED56C5AFC32F818.taxon	discussion	Remarks Andrena scita is a predominantly West Palaearctic species typically of warm open grasslands, extending in the east only to Kazakhstan and Turkmenistan (Gusenleitner & Schwarz 2002). There is no evidence to support the listing of this highly distinctive species from India.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9CFFAC0EC86B0BFEDCFD33.taxon	discussion	Remarks Andrena dorsata is a widespread Palaearctic species extending from Europe into parts of Central Asia. Given that the Simandrena fauna of India is has been poorly known until now (with the addition of A. tungnatha Wood & Gautam sp. nov. and the relatively recently described A. gorkhana), the presence of A. dorsata in northern India is uncertain. It is best excluded from the Indian fauna pending precise occurrence records which can be examined and confirmed as either A. dorsata or a different species of Simandrena.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9CFFAC0EDE68D8FE88FB04.taxon	discussion	Remarks Meena & Dey (2019) present no evidence for the presence of A. ovatula in India. Moreover, it is more or less ecologically impossible for true A. ovatula to be present in India, as this taxon is confined to the western part of the West Palaearctic (Praz et al. 2022). It is not impossible that the taxon A. (Taeniandrena) afzeliella (Kirby, 1802) could be present in northern India, but this must be established robustly through verified specimens and ideally genetic work. Andrena ovatula is therefore excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9CFFAC0EDD6EAEFE0AF9F9.taxon	discussion	Remarks Nurse (1904) reported that A. convexiuscula (Kirby, 1802) was “ common ” in Kashmir, A. conveixuscula being a known synonym of A. wilkella (Gusenleitner & Schwarz 2002). The subgenus Taeniandrena is taxonomically complex and often defies morphological delineation (Praz et al. 2022), and historical species concepts cannot be used in a modern context. Whilst it is clear that Nurse was referring to a species of Taeniandrena, its identity must remain obscure until genetic sampling of the Indian Taeniandrena fauna is possible, as it is highly likely that true A. wilkella is not present. It is therefore excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9CFFAF0E886CA2FDB9FEE9.taxon	discussion	Remarks Meena & Dey (2019) present no evidence to support the presence of A. haemorrhoa in India. Andrena haemorrhoa is a trans-Palaearctic species of temperate woodland, with some marginal occurrences in North Africa (Gusenleitner & Schwarz 2002; Cherair et al. 2013; Ascher & Pickering 2023). In Central Asia, it is restricted to mountainous areas. It is not impossible that A. haemorrhoa could be present in Kashmir, but as no precise specimen records are available this is considered unproven. It is therefore excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9FFFAF0EE16B93FCE1FC97.taxon	discussion	Remarks Meena & Dey (2019) present no evidence to support the presence of this species in India. Andrena tscheki is a West Palaearctic species found in Europe to Turkey, the Caucasus, and the Levant (Gusenleitner & Schwarz 2002; Astafurova et al. 2021). Populations in the south-east of its range probably represent a distinct species (A. tscheki tritica Warncke, 1965), but it is ecologically inconceivable that populations reach to Bihar, and so it is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9FFFAF0D7E693DFD4FFB22.taxon	discussion	Remarks This species does not exist – Meena & Dey state that it was described by Cameron (1909: 130) from Ferozepur. However, the only species matching this information is A. punjaubensis which is a synonym of A. flavipes (see above). It is not clear where the name A. argada comes from, but since it is a nomen nudum it is excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9FFFAF0E776EC8FAB8F917.taxon	description	Listed by Gusenleitner & Schwarz (2002).	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9FFFAF0E776EC8FAB8F917.taxon	discussion	Remarks The name A. cameroni is a replacement name for A. caroli Cameron, 1909, a taxon described from Shimla (Cameron 1909: 130), but whose name is preoccupied by A. caroli Pérez, 1895 (junior primary homonym). Listed as an Andrena by Gusenleitner & Schwarz (2002) in line with its original description, the true identity of this taxon is actually that of a member of the genus Melitta Kirby 1802 (Melittidae Michener, 2000), to which it was moved by Michez & Eardley (2007). Interestingly, this was actually noticed by Nurse and pointed out to Cockerell (see Cockerell 1922: 247), but this did not achieve wider recognition within the Andrena literature. It is therefore excluded from the Indian Andrena fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
03DE87B4FF9FFFAE0D3E6CBCFABDFEE9.taxon	discussion	Remarks Andrena cara was described by Nurse from Peshin [= Pishin] in western Pakistan. It is a junior synonym of A. (incertae sedis) hieroglyphica Morawitz, 1876, and its complicated history of use was clarified by Wood & Monfared (2022). There is no evidence available to support a listing of A. hieroglyphica in Kashmir, as the species is found in dry desert habitats in Central Asia, typically not at high altitude (Astafurova et al. 2022; Wood & Monfared 2022). It is therefore excluded from the Indian fauna.	en	Gautam, R. K., Uniyal, V. P., Wood, Thomas J. (2024): A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand. European Journal of Taxonomy 948: 1-59, DOI: 10.5852/ejt.2024.948.2637, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143
