taxonID	type	description	language	source
03DE1E7AFF956B7E9906AEFEFE02FE81.taxon	description	Description Male. Length 2.16. Maximum width 1.28. Head width across vertex 0.48; dorsal length 0.12; facial length 0.38; facial width 0.58. Anterior inter­ocular space 0.18; posterior space 0.40. Dorsal width of eye 0.15; maximum width 0.23; height 0.27. Height of gena 0.03. Ocellus width 0.03; interocellar space 0.10. Maximum width of pronotum 1.05; median length 0.32. Scutellum length 0.60; width 0.50. Cuneus length 0.50; width 0.32. Rostrum 1.28. Antennae I 0.08; II 0.46; III 0.36; IV missing. Elongate oval (figure 1). Head strongly vertical, from above about 4 x as broad as long; in front somewhat triangular, about 1.53 x as broad as high. Vertex, base of frons, lateral marginal band abutting on eyes, genae, lora, all dark­red to black; head disk otherwise pale whitish hyaline. Practically impunctate, pubescent with pale to yellow­white semi­erect and erect hairs arising from fine aciculate punctures; these dense and upturned on vertex and base of frons. Frons at base distinctly depressed; apex tumid, set with apical marginal blunt carina and another sub­apical somewhat weaker one. Eyes deep red, glabrous, emarginate behind ocelli. Ocelli whitish translucent, about 3.3 x width of each apart. Occiput, postgena, broadly exposed, pale hyaline. Antennae with segment I pale yellow; II and III pale yellow on ventral surface, darkened dorsally; IV missing. I about two­thirds as thick as long, glabrous; II somewhat narrowed medially, at base and apex about 1.2 x as thick as I, dense with semi­erect and erect yellow­brown hairs shorter than segment thickness; III subfusiform to linear, dense with semi­erect hairs longer than segment thickness. Rostrum yellow­brown, apical band on segment IV dark­red; reaching as far back as seventh abdominal sternite. Pronotum about 3.28 x as broad as long. Lateral margins distinctly explanate, hyaline; posterior margin bisinuate. Disk convexly raised, transversely rugose punctate except calli prominent, confluent, impunctate. Mesoscutum, scutellum strongly tumid, distinctly punctate with scutellum more coarsely so. Hemelytra glassy hyaline; corium, clavus finely punctate; embolium, cuneus impunctate; membrane smoky reddish­brown, glabrous, biareolate. Dorsum generally strongly shining, pale yellow hyaline; broad posterior band on pronotum dark reddish­brown; mesoscutum, basal two­thirds or so of scutellum, narrow basal and apical bands on clavus, basal extremity of corium, broad band covering apical half of corium and extending laterad to embolium, all contrasting piceous to black. Distinctly pubescent with semi­reclining to semi­erect golden yellow and reddish­brown hairs matching colour of background; these intermixed with dark reddish­brown bristles on corium, embolium and cuneus. Venter largely piceous to black; propleura pale yellow hyaline, impunctate. Prosternum, metasternum, abdominal sternites II to IV mid­ventrally, all yellow­brown with reddish tinge. Fore and mid legs generally pale yellow hyaline, tibiae tinged with reddish shading to distinct darkening on protibiae; metacoxae similarly pale, other leg units missing. Female. Unknown.	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF956B7E9906AEFEFE02FE81.taxon	discussion	Comments This new species belongs in the subgenus Jehania Distant and represents the third record of the subgenus in the Middle East; the two previously reported species being I. yemenensis Akingbohungbe and I. quadrivittatus Akingbohungbe, both from the Republic of Yemen. The new species can be readily separated from these two species by the distinctive colouration of the dorsum as described above. For example, the pronotal disk is not uniformly darkened and the cuneus lacks any dark band. Besides, the new species differs from all other known species of Jehania in having a weak blunt carina behind the usual apical marginal carina separating the frons from the base of the tylus.	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF956B7E9906AEFEFE02FE81.taxon	materials_examined	HOLOTYPE: male, Yemen, Sana’a: 12 km NW Manakhah, 27 March to 5 May, 2002, van Harten (LC).	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF976B789906AECBFE02FCE7.taxon	description	Description Male. Length 2.32; maximum width across hemelytra 1.36. Head width across vertex 0.46; dorsal length 0.10; facial length 0.44; facial width 0.66. Anterior interocular space 0.18; posterior space 0.46. Dorsal width of eye 0.14; maximum width 0.24; height 0.30. Height of gena 0.06. Ocellus width 0.06; interocellar space 0.06. Maximum width of pronotum 1.24; median length 0.40. Scutellum length 0.66; width 0.60. Cuneus length 0.48; width 0.34. Rostrum 1.08. Antennae I 0.10; II 0.52; III 0.34; IV 0.10. Ovate (figure 2). Head strongly vertical, from above 4.6 x as broad as long; in front somewhat ovoid (figure 3), about 1.5 x as broad as high. Vertex dark­red to black, sparsely finely punctate. Frons at base dark brown with reddish tinge, distinctly punctate; apex yellow­brown, coarsely rugose punctate, tumid convex with margin sharply carinate and sinuately convex. Entire head disk pubescent with upturned golden yellow short reclining hairs. Tylus dark reddish, depressed medially so base together with juga appearing as three pale tubercle­like prominences beneath frontal hood. Lora dark­red, strongly laterally compressed. Eyes deep red, weakly emarginate behind ocelli. Ocelli large, translucent, rather protuberant; each about equal in width to interocellar space. Occiput, postgena broadly exposed, translucent. Antennae mostly dark­red; tip and rather broad median band on ventral surface of segment II, basal and apical extremities of III, pale. I about half as thick as long; II slightly narrowed towards middle with base and apex about 1.6 x as thick as I; III and IV fusiform. II, III and IV pubescent with reddish brown hairs either shorter than or as long as segment thickness. Rostrum generally dark­red, ventral surface of segment I at base, entire base of IV pale; reaching as far back as third abdominal sternite. Pronotum about 3 x as broad as long, convexly raised at base, inclined forwards with sides rather strongly sloping laterad; lateral margins explanate, pale hyaline; posterior margin deeply bisinuate. Mesoscutum, scutellum concolourous dark­red to black with pronotal disk, tip of scutellum pale yellow. Hemelytra with clavus, base of corium together with narrow ental band adjoining claval suture dark­red to black; embolium, cuneus and remainder of corium reddish brown with a median somewhat triangular patch on corium much paler. Membrane smoky, glabrous, biareolate. Dorsum generally shiny; distinctly punctate, transversely rugose on pronotal disk, mesoscutum, scutellum and clavus; embolium and cuneus impunctate. Pronotal disk dense with reclining golden yellow somewhat sericeous, and semi­reclining reddish­brown hairs. Mesoscutum, scutellum and clavus similarly pubescent with semi­reclining and semi­erect reddish­brown hairs. Embolium, cuneus and corium dense with semi­erect reddish­brown hairs intermixed with erect and semi­erect dark­red bristles. Thoracic venter generally dark reddish­brown; prosternum, propleura pale yellowbrown with reddish tinge, mesosternum dark­red to black. Propleura coarsely rugose punctate, episternum more finely so. Abdomen with only segments II and III present, sternites dark reddish­brown with pale lateral extremities. Coxae dark reddish­brown to dark­red; trochanteral segment I pale hyaline on forecoxae, otherwise reddish­brown. Other leg units missing. Female. Unknown.	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF976B789906AECBFE02FCE7.taxon	discussion	Comments This new species shows affinity with Isometopus nigritulus Akingbohungbe known from Cyprus. However it differs from that species in several respects. The frons at the level of the ventral margin of the eyes in the new species lacks the ivory­white transverse stripes separated by distinct grooves which are distinctive features of I. nigritutlus. The interocellar space in the latter is 1.5 x as broad as each ocellus whereas in the new species, the ocelli are as broad as the interocellar space. Besides, the dorsum in the new species has extensive reddish­brown and pale areas in contrast to the rather uniformly piceous to black colouration in I. nigritulus. The holotype which was the only specimen available for study, has the abdomen and the legs excluding the coxae missing. It is nonetheless quite distinctive. The species is named in honour of its collector for his efforts in advancing our knowledge on the isometopine fauna of the Middle East.	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF976B789906AECBFE02FCE7.taxon	materials_examined	HOLOTYPE: male, Yemen, Sana’a 12 km NW Manakhah, 3 July to 21 August, 2001, van Harten (LC).	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF916B759906ACC8FE73FCE1.taxon	description	Description Female. Length 2.24. Maximum width across hemelytra 0.96. Head width across vertex 0.18; dorsal length 0.16; facial length 0.50; facial width 0.42. Minimum frontal inter­ocular space 0.04; anterior space 0.08; posterior space 0.32. Maximum width of eye 0.24; height 0.28. Height of gena 0.12; height of lorum 0.11. Ocellus width 0.02; interocellar space 0.04. Maximum width of pronotum 0.84; median length 0.28. Scutellum length 0.32; width 0.38. Cuneus length 0.34; width 0.28. Rostrum 0.96. Antennae I 0.08; II 0.50; III 0.18; IV missing. (0.14 in paratype). Elongate oval (figure 4). Head deflexed, vertex broadly horizontal, from above about 0.57 x as long as pronotum; in front obovate (figure 5), about 0.84 x as broad as high. Frons strongly tumid convex, weakly marginate apically. Gena sub­excavated, bluntly carinate laterally, about 0.43 x as high as eye with antennophore located at its apex distad of ventral eye margin. Lorum strongly depressed, about 0.92 x as high as gena. Postgena weakly tuberculate, broadly exposed laterally together with occiput. Entire head impunctate, darkred to black with occiput at base pale whitish; pubescent with dark reddish­brown hairs (largely rubbed in specimen). Eyes deep red, glabrous; distinctly separated mesally on frons (0.04 apart), more widely so behind vertex (0.06 apart); deeply emarginate behind ocelli. Ocelli dark translucent, twice width of each apart. Antennae with segment I darkred, about 2 x as long as its thickness. II with broad basal band dark­red to black, narrow sub­apical band brighter reddish, and remainder whitish; somewhat narrowed medially, about as thick at base and apex as I; clothed with semi­reclining hairs shorter than segment thickness. III subfusiform, pale suffused with reddish; clothed with semi­reclining hairs equal or subequal to segment thickness. Rostrum bright red on segment I, otherwise darkred to black; extending to eighth abdominal sternite. Pronotum trapeziform, about 3 x as long as broad; lateral margin sharply carinate, posterior bisinuate. Disk black, finely transversely rugose punctate; calli feebly delimited, confluent, impunctate; pubescent with short dark reddish­brown reclining hairs. Mesoscutum, basal two­fifths or so of scutellum black, scutellum otherwise whitish; both impunctate. Hemelytra pale to whitish ochraceous with base and apex of clavus, rather broad crescent­shaped band covering apical three­fifths of embolium and apical third of corium, apical half of cuneus, broad apical band on membrane, all smoky to dark reddish­brown. Generally impunctate, apparently dense (specimen somewhat rubbed) with short dark red bristle­like hairs intermixed with reddish­brown and golden yellow hairs; latter on whitish bands of scutellum, clavus, and cuneus. Membrane distinctly biareolate, glabrous. Venter generally dark­red to black, ovipositor base somewhat contrasting pale whitish. Propleura impunctate. Legs largely dark reddish­brown; basal trochanteral segments, tip of hind femora, apical band on tibiae, hind tarsi, all pale yellow to yellow­brown.	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF916B759906ACC8FE73FCE1.taxon	discussion	Comments This is the second species record of Myiomma Puton in the Middle East and it shares affinity with the first recorded species, M. maculata Akingbohungbe, also from the Republic of Yemen. The description of the new species as provided above is based primarily on the female holotype; the female paratype, apparently somewhat teneral at the time of collection, had undergone some shrinkage in storage; it is nonetheless quite distinctive. Very unfortunately, a single male specimen that was collected with the female holotype got damaged and lost during the process of microscopic examination. However, some pertinent measurements and notes had earlier been taken on the specimen. These are reproduced hereunder to allow for comparison, especially with M. maculata which was described based on a male specimen. Length 2.28. Maximum width across hemelytra 0.88. Head width across vertex 0.24; dorsal length 0.18; facial length 0.48; facial width 0.44. Minimum frontal inter­ocular space 0.02; anterior space 0.08; posterior space 0.32. Maximum width of eye 0.24; height 0.28. Height of gena 0.08; height of lorum 0.08. Ocellus width 0.03; inter­ocellar space 0.04. Maximum width of pronotum 0.86; median length 0.30. Scutellum length 0.36; width 0.42. Cuneus length 0.46; width 0.30. Rostrum 0.84. Antennae I 0.08; II 0.70; III 0.12; IV 0.12. In my 1996 key to Myiomma spp., the male specimen of the new species will run to couplet 14 just like M. maculata; here both key out as M. bredoi Akingbohungbe from Lumbumbashi, Republic of Congo. Both species are however relatively larger than M. bredoi; and while the coxae, trochanters, femora, and ostiolar peritreme are generally whitish in the latter species, they are generally reddish to dark reddish in the new species and M. maculata. The new species differs from M. maculata in having the pronotum about 2.87 x as broad as long compared to 2.56 x; antennal segment II of the male is generally dark red and clavate, whereas in male M. maculata it is narrowed towards the middle, and largely pale whitish with a subapical dark red band. Similarly, the hind femur in the new species is dark reddish­brown with contrasting whitish apical band whereas it is uniformly dark reddish­brown in M. maculata. The female of the new species will key out in couplet 30 in my 1996 key and there, it will fit more easily as M. albiscutellata Smith. It can however be readily separated from that species in the colouration of its dorsum, which differs in being largely dark brown to blackish. Besides, the head from above in M. albiscutellata is at least 0.70 x as long as the pronotum compared to 0.57 x in the new species; and similarly, antennal segment II is less than 2.50 x as long as III, compared to 2.78 x in the new species.	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF916B759906ACC8FE73FCE1.taxon	materials_examined	HOLOTYPE: female, Yemen, Hadhramaut, Ghail Ba Wazir, November / December, 2002, van Harten and Hubaishnan (LC). PARATYPE: female, Yemen, Ta’izz, Ta’izz­Al Turbah, 22 March, 1992, A. van Harten (LC).	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF9C6B779906ACDEFDC8FA99.taxon	description	Description Female. Length 2.02. Maximum width across hemelytra 1.68. Head width across vertex 0.50; dorsal length 0.16; facial length 0.46; facial width 0.70. Anterior inter­ocular space 0.22, posterior space 0.58. Dorsal width of eye 0.12; maximum width 0.22; height 0.30. Height of gena 0.14. Ocellus width 0.02; inter­ocellar space 0.16. Maximum width of pronotum 1.28; median length 0.44. Scutellum length 0.72; width 0.66. Cuneus length 0.48; width 0.42. Rostrum 0.96. Antennae I 0.12; II 0.42; III 0.28; IV 0.14. Strongly ovate (figure 6). Head strongly appressed, inclined forwards; from above 3.13 x as broad as long; in front more or less quadrate (figure 7), about 1.52 x as broad as high. Frons tumid towards apex, margin convex, finely carinate. Gena high, about 0.47 x as high as eye; broadly arcuate with margin finely carinate, forming prominent concavity on either side of frons. Entire head disk transversely rugose punctate, pubescent with very short reclining somewhat sericeous hairs; mostly golden to yellow­brown, vertex behind ocelli paler; genae dark red to black laterally. Eyes reddish, pubescent with obscure very short erect hairs. Ocelli deep red, surrounding annuli brighter red; about 8 x width of each apart. Antennal segment I dark red, about 0.33 x as thick as long, sparse with semi­erect hairs. II dark red, yellow­brown tinged with reddish along the middle; somewhat recurved, subequal in thickness at base to I, 1.5 x thicker towards apex; pubescent with reclining golden yellow hairs shorter than segment thickness. III yellow­brown, linear, 0.5 x as thick as I; pubescent with semi­erect golden yellow hairs generally longer than segment thickness. IV fusiform, similarly pubescent as III, black with apex whitish. Rostrum with segment I red, II pale reddish, III and IV dark­red; extending to sixth abdominal sternite. Pronotum about 2.90 x as broad as long; lateral margins strongly arcuate and explanate, posterior deeply bisinuate. Scutellum slightly longer (1.09 x) than broad. Dorsum generally strongly shiny; transversely rugose punctate, more coarsely so on pronotum; distinctly pubescent with short reclining golden yellow to dark golden hairs. General colouration ochraceous with extensive dark reddish areas as follows: broad somewhat crescent­shaped blotch covering most of pronotal disk, mesoscutum, scutellum excluding whitish tip, clavus and ental band on corium between radial vein and claval suture. Clavus at base ridged; embolium strongly explanate; costal margin broadly arcuate, forming open angle with cuneus. Venter largely reddish, mesosternum mostly dark red; propleura, lateral band on abdominal sternites II to VIII, ovipositor shaft and sternite IX, all ochraceous. Generally punctate except sternite IX impunctate, punctures more coarse and somewhat rugose on propleura. Abdominal sternites pubescent with short reclining golden yellow hairs; these on IX intermixed with medium sized semi­erect ones. Legs with coxae, fore­femora, mesofemora red to dark reddish; base and apex of fore­femora somewhat paler; metafemora, tibiae reddish­brown with tibiae paler towards apex; tarsi pale whitish. Male. Unknown.	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF9C6B779906ACDEFDC8FA99.taxon	discussion	Comments This new species is the first record of Paloniella Poppius in the Middle East. It is a relatively small species with a more extensive ochraceous dorsum than the generally observed dark chocolate­brown to dark­red in previously known species of the genus, with the only exception of P. flavicolor Akingbohungbe from South Africa. However, the latter is generally much lighter coloured than the new species, and it is much larger, being 2.60 in length. In terms of size, the new species shows some affinity with I. niger Linnavuori from the Sudan; but that species differs from it in several respects: head darkened with occipital margin strongly contrasting whitish; rostrum considerably more elongate, extending to ninth abdominal sternite; antennal segment II much less (1.25 x compared to 1.5 x) as long as III etc.	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF9C6B779906ACDEFDC8FA99.taxon	materials_examined	HOLOTYPE: female, Yemen: Sana’a, 12 km NW Manakhah, 27 March to 5 May, 2002, van Harten (LC).	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
03DE1E7AFF9E6B779906AAE6FCA3F896.taxon	description	Description Isometopus longirostris Akingbohungbe was described as new from the Sudan by me in 1996, but I was unaware of the paper by Josifov (1993) in which he described Isometopus longirostris as a new species from Bulgaria. Inadvertently therefore, a homonymy had been created and, a new name must be proposed for the species from the Sudan. Accordingly, Isometopus sudanicus is hereby proposed as a replacement name. Each species is quite distinctive and not synonymous, I. sudanicus belonging in the subgenus Jehania Distant while I. longirostris Josifov belongs in Isometopus sensu stricto as characterized in my 1996 paper referred to above.	en	Akingbohungbe, A. E. (2006): New species of Isometopinae (Hemiptera: Miridae) from Yemen with a new name for Isometopus longirostris Akingbohungbe from Sudan. Zootaxa 1210: 27-38, DOI: 10.5281/zenodo.273415
