taxonID	type	description	language	source
03DF8784B134FFA9FF63924BC938298D.taxon	materials_examined	Type locality. Argentina, Tierra del Fuego, near Rio Grande (53 º 36 ’ S, 68 º 04 ’ W). Type material. Holotype female, Tierra de Fuego, near Rio Grande, Argentina, Chiliotrichum dense scrub, with Festuca gracillima and Chiliotrichum diffusum (after Collantes et al. 1999), S. M. Bonaventura and V. Mascitti leg., February 2001. Paratypes: All specimens from same locality as holotype, 44 females, 23 males and 46 juveniles. Holotype and 10 paratypes deposited in MACN, 10 paratypes in MNCN, and 93 paratypes in JIAP.	en	Arbea, Javier I. (2016): Three new species and one new record of Tullbergiidae (Collembola: Onychiuroidea) from Tierra del Fuego. Zootaxa 4093 (1), DOI: 10.11646/zootaxa.4093.1.4
03DF8784B134FFA9FF63924BC938298D.taxon	description	Description. Body shape elongate. Length 1.25 mm (female holotype), 1.1 – 1.4 mm (female paratypes), 0.9 – 1.0 mm (male paratypes). Colour white. Cuticular granulation of whole body relatively uniform but larger on posterior dorsum of Abd VI. Ungranulated cuticular ridges on dorsolateral anterior margins of Th III to Abd IV. Dorsal chaetae well differentiated into micro- and macrochaetae (Fig. 27). Antennae slightly shorter than head length (Fig. 27), ratio of lengths 1: 1.1. Ant I, II, and III with 7, 11, and 18 ordinary chaetae respectively. AIIIO consists of three large sensory clubs with papillate protecting integumentary fold covering two small sensory rods between two outermost of these clubs; single sensillum located ventrally on Ant III. Ant IV with five thickened and curved sensilla (S 1, S 4, S 7, S 8 and S 9 after D’Haese (2003) = e, c, b, a and d respectively after Rusek (1971 )), one dorsoexternal microsensillum and one small subapical organite; apical vesicle absent (Fig. 32). PAO elongate, consisting on about 20 simple vesicles lying in two parallel rows. Coarse secondary cuticular granules form anterior margin of PAO (Fig. 29). Labial palp with only papilla B (papillae A, C, D and E absent), 6 proximal chaetae and 3 hypostomal chaetae; basal parts of labium with 4 + 4 and 5 + 5 chaetae (Fig. 33). Maxillary palp simple, without sublobal hair (Fig. 33). Pso semielliptical to circular, with crescent shaped opening with five distinct ridges (type III after Weiner & Najt 1991) (Fig. 29). Pso formula 11 / 011 / 11111 (Fig. 27). Dorsal chaetotaxy as in Fig. 27. Th pleurites I, II, III with 2, 3, 3 chaetae respectively. Chaetae s on Th and Abd slightly thickened, similar in form to ordinary chaetae. Th II and III with basally thickened chaeta s in position m 7 and microsensillum ms adjacent to it. Abd I – IV with chaeta s in position p 3. One chaeta s also present on Abd II – III as a 7, on Abd IV – V as p 5 and a pair lateral to genital opening. Full description of dorsal chaetotaxy is given in Table 6. Abd VI with two large, curved anal spines placed on low papillae. Ratio anal spine: claw III as 1 – 1.25: 1. Seven spiniform processes on Abd VI, located as follows: 1 + 1 dorso-medial (dm), 2 + 2 latero-dorsal (ld) and 1 unpaired medio-ventral (mv) (Fig. 35). Ventral chaetotaxy as in Fig. 28. Head with 3 + 3 postlabial chaetae. Th I – III sterna with 0,1 + 1, 1 + 1 chaetae respectively. Ventral tube with 4 + 4 distal and 2 + 2 basal chaetae. Abd V sternum with 1 + 1 slightly thickened chaeta s lateral to genital plate. Chaetotaxy of anal lobes as in Fig. 34. Legs chaetotaxy from I to III, coxae, 3,4 + 7,4 + 7; trochanters 6,6 – 7,6 – 7; femora, 11,11,11; tibiotarsi with 15 (7 + 8), 15 (7 + 8), 14 (7 + 7) chaetae respectively (A 1 to 7, B 1 to 7 and M present, B 7 absent on metatibiotarsus). Pretarsus with two chaetae each. Claw untoothed. Empodial appendage 0.15 – 0.25 the length of claw (Figs 30 – 31).	en	Arbea, Javier I. (2016): Three new species and one new record of Tullbergiidae (Collembola: Onychiuroidea) from Tierra del Fuego. Zootaxa 4093 (1), DOI: 10.11646/zootaxa.4093.1.4
03DF8784B134FFA9FF63924BC938298D.taxon	etymology	Etymology. The new species is dedicated to the late Chilean poet Pablo Neruda for his 111 birthday (July 12, 1904).	en	Arbea, Javier I. (2016): Three new species and one new record of Tullbergiidae (Collembola: Onychiuroidea) from Tierra del Fuego. Zootaxa 4093 (1), DOI: 10.11646/zootaxa.4093.1.4
03DF8784B134FFA9FF63924BC938298D.taxon	discussion	Discussion. The new species belongs to the group of species with pso formula 11 / 011 / 11111, Abd VI with 7 spiniform processes and elongate PAO, D. chazeaui Weiner & Najt, 1991, D. harrisoni Bagnall, 1948, D. laterospina Salmon, 1941, D. matileorum Najt, 1988, D. pefauri Rubio & Najt, 1979 and D. spinosissima (Wahlgren, 1906) (Table 8). They differ in the number of vesicles on PAO (more than 26 vesicles in D. laterospina and D. pefauri), the apical vesicle on Ant IV (present and conspicuous in D. laterospina, D. pefauri, D. harrisoni and D. spinosissima) and the dorsal chaetotaxy (Table 8). Dinaphorura najtae sp. nov. Figs 36 – 43; Table 7	en	Arbea, Javier I. (2016): Three new species and one new record of Tullbergiidae (Collembola: Onychiuroidea) from Tierra del Fuego. Zootaxa 4093 (1), DOI: 10.11646/zootaxa.4093.1.4
03DF8784B134FFA9FF63924BC938298D.taxon	materials_examined	Type locality. Argentina, Tierra del Fuego, near Rio Grande (53 º 36 ’ S, 68 º 04 ’ W). Type material. Holotype female, Tierra de Fuego, near Rio Grande, Argentina, Chiliotrichum dense scrub, with Festuca gracillima and Chiliotrichum diffusum (after Collantes et al. 1999), S. M. Bonaventura and V. Mascitti leg., February 2001. Paratypes: All specimens from same locality as holotype, 41 females, 7 males and 6 juveniles. Holotype and 10 paratypes deposited in MACN, 10 paratypes in MNCN, and 34 paratypes in JIAP.	en	Arbea, Javier I. (2016): Three new species and one new record of Tullbergiidae (Collembola: Onychiuroidea) from Tierra del Fuego. Zootaxa 4093 (1), DOI: 10.11646/zootaxa.4093.1.4
03DF8784B134FFA9FF63924BC938298D.taxon	description	Description. Body shape elongate. Length 0.7 mm (female holotype), 0.6 – 0.9 mm (female paratypes), 0.6 – 0.7 mm (male paratypes). Colour white. Cuticular granulation of whole body relatively uniform but larger on posterior dorsum of Abd VI. Ungranulated cuticular ridges on dorsolateral anterior margins of Th III to Abd IV. Dorsal chaetae well differentiated into micro- and macrochaetae (Fig. 36). TABLE 8. Main diagnοstic characters οf Dinaphorura nerudai sp. nov., Dinaphorura najtae sp. nov. and related species in the genus Dinaphorura with 11 ⁄ 011 ⁄ 11111 dοrsal psο. "? " ‾‾ denοtes ∶ nο data available. (1) SP-Spinifοrm prοcesses οn Abd vI ∶ N ‾‾ Τοtal number (anal spines nοt included), dm ‾‾ dοrsοmedial, ld ‾‾ laterοdοrsal and mv ‾‾ mediοventral prοcesses, a ∶ p ‾‾ Ratiο anteriοr ∶ pοsteriοr laterοdοrsal prοcesses; (2) PAΟ-Fοrm, E ‾‾ elοngate, Τ ‾‾ triangular, and Number οf vesicles; (3) Av-Apical vesicle οn Ant Iv, B ‾‾ bilοbed, S ‾‾ simple, Τ ‾‾ trilοbed; (4) AS ∶ C-Ratiο anal spine ∶ claw length; (5) Dοrsal chaetοtaxy ∶ m, ca, p chaetae series. (*) Abd vI with 2 large curved anal hοrns οn papillae and nine shοrt fat spines after οriginal descriptiοn, but οnly 7 spinifοrm prοcesses are shοwn in the οriginal drawing (Fig. 134). Antennae slightly shorter than head length (Fig. 36), ratio of lengths 1: 1.15. Ant I, II, and III with 7, 11, and 18 ordinary chaetae respectively. AIIIO consists of three large sensory clubs with papillate protecting integumentary fold covering two small sensory rods between the two outermost of these clubs; single sensillum located ventrally on Ant III. Ant IV with five thickened and curved sensilla (S 1, S 4, S 7, s 8 and S 9 after D’Haese (2003) = e, c, b, a and d respectively after Rusek (1971 )), one dorsoexternal microsensillum and one small subapical organite; apical vesicle absent (Fig. 42). PAO triangular, consisting on 18 – 20 irregularly shaped vesicles in two rows. Coarse secondary cuticular granules form anterior margin of PAO (Fig. 43). Labial palp with only papilla B (papillae A, C, D and E absent), 6 proximal chaetae and 3 hypostomal chaetae; basal parts of labium with 4 + 4 and 5 + 5 chaetae (Fig. 41). Maxillary palp simple, without sublobal hair (Fig. 41). Pso semielliptical to circular, with crescent shaped opening with five distinct ridges (type III after Weiner & Najt 1991) (Fig. 43). Pso formula 11 / 011 / 11111 (Fig. 36). Dorsal chaetotaxy as in Fig. 36. Th pleurites I, II, III with 2, 3, 3 chaetae respectively. Chaetae s on Th and Abd slightly thickened, similar in form to ordinary chaetae. Th II and III with basally thickened chaeta s in position m 7 and microsensillum ms adjacent to it. Abd I – IV with chaeta s in position p 3. One chaeta s also present on Abd II – III as a 7, on Abd IV – V as p 5 and a pair lateral to genital opening. Full description of dorsal chaetotaxy is given in Table 7. Abd VI with two large, curved anal spines placed on low papillae. Ratio anal spine: claw III as 1 – 1.17: 1. Seven spiniform processes on Abd VI, located as follows: 1 + 1 dorso-medial (dm), 2 + 2 latero-dorsal (ld) and 1 unpaired medio-ventral (mv) (Fig. 39). Ventral chaetotaxy as in Fig. 37. Head with 3 + 3 postlabial chaetae. Th I – III sterna with 0,1 + 1, 1 + 1 chaetae respectively. Ventral tube with 4 + 4 distal and 2 + 2 basal chaetae. Abd V sternum with 1 + 1 slightly thickened chaeta s lateral to genital plate. Chaetotaxy of anal lobes as in Fig. 38. Legs chaetotaxy from I to III, coxae, 3,4 + 7,4 + 7; trochanters 6,6,6; femora, 11,11,11; tibiotarsi with 15 (7 + 8), 15 (7 + 8), 14 (7 + 7) chaetae respectively (A 1 to 7, B 1 to 7 and M present, B 7 absent on metatibiotarsus). Pretarsus with two chaetae each. Claw untoothed. Empodial appendage 0.23 – 0.28 the length of claw (Fig. 40).	en	Arbea, Javier I. (2016): Three new species and one new record of Tullbergiidae (Collembola: Onychiuroidea) from Tierra del Fuego. Zootaxa 4093 (1), DOI: 10.11646/zootaxa.4093.1.4
03DF8784B134FFA9FF63924BC938298D.taxon	etymology	Etymology. The new species is dedicated to the late Judith Najt, a prominent Argentinean researcher of collembolan fauna, who passed away in 2014.	en	Arbea, Javier I. (2016): Three new species and one new record of Tullbergiidae (Collembola: Onychiuroidea) from Tierra del Fuego. Zootaxa 4093 (1), DOI: 10.11646/zootaxa.4093.1.4
03DF8784B134FFA9FF63924BC938298D.taxon	discussion	Discussion. The new species is most similar to D. americana Rapoport, 1962 and D. jarai Najt & Rubio, 1978, they share the pso formula 11 / 011 / 11111 and triangular PAO (Table 8), but they differ in the number of vesicles on PAO (12 – 15 in D. americana, 10 - 11 in D. jarai vs 18 – 20 in the new species), the number of spiniform processes on Abd VI (5 in D. jarai vs 7 in D. americana and D. najtae sp. nov.), the apical vesicle on Ant IV (present in D. jarai and D. americana vs absent in the new species) and the dorsal chaetotaxy (Table 8).	en	Arbea, Javier I. (2016): Three new species and one new record of Tullbergiidae (Collembola: Onychiuroidea) from Tierra del Fuego. Zootaxa 4093 (1), DOI: 10.11646/zootaxa.4093.1.4
03DF8784B137FFB1FF6393A6C91C2DA6.taxon	materials_examined	Material examined. Argentina, Tierra de Fuego, near Rio Grande, Chiliotrichum dense scrub, with Festuca gracillima and Chiliotrichum diffusum (after Collantes et al. 1999), S. M. Bonaventura and V. Mascitti leg., February 2001, 9 specimens. Description of specimens from Tierra del Fuego. Habitus as in Figs 20 – 21. Body length (without antennae) 0.50 – 0.62 mm. Body white. Cuticular granulation of whole body relatively uniform. Antennal bases not marked. Dorsal chaetae well differentiated into meso- and macrochaetae (Fig. 20). Antennae slightly shorter than head length (Fig. 20), ratio of lengths 1: 1.2. Ant I, II, and III with 7, 11, and 17 ordinary chaetae respectively. AIIIO consists of two sensory clubs (inner sensillum larger than outer one) and two small sensory rods between them; single sensillum located ventrally on Ant III. Ant IV with five thickened and curved sensilla (S 1, S 4, S 7, S 8 and S 9 after D’Haese (2003) = e, c, b, a and d respectively after Rusek (1971 )), one dorsoexternal microsensillum and one small subapical organite; with a small, simple apical vesicle (Figs 22 – 23). PAO narrow elliptical, in shallow depression, 4 – 5 times longer than diameter of the nearest pso, and slightly shorter than width of the Ant I, consisting of 36 – 52 simple vesicles lying in two parallel rows (Fig. 25). Labial palp with 5 papillae A, B, C, D and E, 6 proximal chaetae and 3 hypostomal chaetae; basal parts of labium with 4 + 4 and 5 + 5 chaetae (Fig. 24). Maxillary palp simple, with 1 sublobal hair. Pso circular, stellate (type I after Weiner & Najt 1991). Pso formula: 11 / 111 / 11121 (Fig. 20). Dorsal chaetotaxy as in Fig. 20. Th pleurites I, II, III with 2, 3, 3 chaetae respectively. Chaetae s on Th and Abd simple and slightly thickened, similar in form to ordinary chaetae. Th II and III with long, basally thickened chaeta s in position m 7 and microsensillum ms adjacent to it. Abd I – IV with chaeta s in position p 3. One chaeta s also present on Abd II – III as a 7, on Abd IV – V as p 5 and a pair lateral to genital opening. Full description of dorsal chaetotaxy is given in Table 5 Abd VI with two short, curved anal spines placed on papillae. Ratio anal spine: claw III as 0.69 – 0.72: 1. Ventral chaetotaxy as in Fig. 21. Head with 3 + 3 postlabial chaetae. Th I – III sterna with 0,1 + 1, 1 + 1 chaetae respectively. Ventral tube with 4 + 4 distal and 2 + 2 basal chaetae. Abd V sternum with 1 + 1 slightly thickened chaeta s lateral to genital plate. Tibiotarsi with 15 (7 + 8), 15 (7 + 8), 14 (7 + 7) chaetae respectively (A 1 to 7, B 1 to 7 and M present, B 7 absent on metatibiotarsus). Pretarsus with two chaetae each. Claw untoothed. Empodial appendage 0.2 – 0.23 the length of claw on leg III (Fig. 26).	en	Arbea, Javier I. (2016): Three new species and one new record of Tullbergiidae (Collembola: Onychiuroidea) from Tierra del Fuego. Zootaxa 4093 (1), DOI: 10.11646/zootaxa.4093.1.4
03DF8784B137FFB1FF6393A6C91C2DA6.taxon	discussion	Remarks. Specimens from Tierra del Fuego conform to the original description by Cassagnau & Rapoport (1962) from Patagonia (Argentina) and differ to the description by Barra (1995) from Natal, South Africa: number of vesicles in PAO: 20 – 22 in the specimens from Natal, 36 – 52 in the specimens from Tierra del Fuego, 30 – 50 in the types from Patagonia; tibiotarsus III with 14 chaetae in the specimens from Tierra del Fuego, 12 chaetae in the specimens from Natal; shape of PAO identical in specimens from Tierra del Fuego and Patagonia, different in the specimens from Natal; position of pso on Abd IV egal in the specimens from Tierra del Fuego and Patagonia ,, different in the specimens from Natal; Ant. IV with sensilla S 7 and S 9 on the same level in the specimens from Tierra del Fuego and Patagonia, on the different level in the specimens from Natal; shape of guard papillae in AIIIO the same in the specimens from Tierra del Fuego and Patagonia, different in the specimens from Natal. Specimens from Natal are almost certainly a different species, closely related to T. meridionalis. The geographical distribution of these species corroborates the idea that southern South America shares a common history with the other austral areas, reflecting the existence of an ancient austral biota (Rapoport 1971).	en	Arbea, Javier I. (2016): Three new species and one new record of Tullbergiidae (Collembola: Onychiuroidea) from Tierra del Fuego. Zootaxa 4093 (1), DOI: 10.11646/zootaxa.4093.1.4
