identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DD87997445FFD15DC8EA0AFBE7FB95.text	03DD87997445FFD15DC8EA0AFBE7FB95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leeuwenia Karny 1912	<div><p>Leeuwenia Karny</p><p>Leeuwenia Karny, 1912: 161 . Type species Leeuwenia gladiatrix Karny.</p><p>The only thorough diagnosis of this genus was provided by Okajima (2006). However, that diagnosis must now be interpreted in the light of the structural diversity among the included species that is indicated below. In particular, the presence of prosternal basantra is remarkable, not just for any members of this genus but for most members of the Liothrips -lineage (Mound &amp; Marullo 1996). Members of the genus Leeuwenia are found widely in the tropical forests of Asia and Australia, with the most Western species being from Mauritius (Mound 2004). The galls known to be induced by these species range from irregularly distorted leaves (Fig. 4) to neatly rolled and even brightly coloured structures (Fig. 6). Host records are not available for many of the 30 species now listed in the genus, but 11 Leeuwenia species are recorded from trees in the family Myrtaceae, including two of the new species described below. In contrast, seven described Leeuwenia species are associated each with a tree in a different plant family (Mound 2004), including four of the species known only from Australia ( L. diospyrae — Ebenaceae; L. polyosmae — Grossulariaceae; L. scolopiae — Flacourtiaceae; L. tetrastigmae — Vitaceae).</p></div>	https://treatment.plazi.org/id/03DD87997445FFD15DC8EA0AFBE7FB95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mound, Laurence A.;Tree, Desley J.	Mound, Laurence A., Tree, Desley J. (2021): Structural variation among species of Leeuwenia (Thysanoptera, Phlaeothripinae) with three new species from Australia. Zootaxa 4903 (3): 439-447, DOI: 10.11646/zootaxa.4903.3.9
03DD87997446FFD25DC8E962FA4BFBC1.text	03DD87997446FFD25DC8E962FA4BFBC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leeuwenia Karny 1912	<div><p>Key to Leeuwenia species from Australia</p><p>1. Tube no more than 1.5 times as long as head................................................................ 2</p><p>-. Tube at least 2.0 times as long as head..................................................................... 4</p><p>2. Tergite IX with setae S1 and S2 unusually elongate, between 2.0 and 3.0 times as long as basal width of tube (Fig. 12)................................................................................................ aliceae sp.n.</p><p>-. Tergite IX setae S1 and S2 shorter, less than 1.5 times as long as basal width of tube................................ 3</p><p>3. Fore tarsus with prominent lateral tooth in both sexes.................................................. polyosmae</p><p>-. Fore tarsus without a prominent lateral tooth........................................................... diospyri</p><p>4. Antennal segment III at least 3.5 times as long as maximum width; prosternal basantra absent........................ 5</p><p>-. Antennal segment III less than 3.0 times as long as maximum width (Fig. 16); prosternal basantra present (Fig. 20) but sometimes weakly sclerotised (Fig. 19)........................................................................ 6</p><p>5. All femora and tibiae clear yellow; head about 1.6 times as long as width across eyes; antennal segment III less than 4.0 times as long as maximum width; tube less than 9 times as long as basal width................................ tetrastigmae</p><p>-. Mid and hind femora as brown as body, mid and hind tibiae yellow but brown at base; head 2.0 times as long as width across eyes; antennal segment III at least 5.0 times as long as maximum width; tube at least 13 times as long as basal width.................................................................................................. convergens</p><p>6. Pelta wider than long (Fig. 21); prosternal basantra well developed with several setae (Fig. 20); metanotum with several pairs of setae anterior to median pair; tergal wing-retaining setae slender and pointed............................. scolopiae</p><p>-. Pelta longer than basal width (Figs 17, 18); prosternal basantra weak and without setae (Fig. 19); metanotum without scattered setae anterior to major median setae; tergal wing-retaining setae broadly flattened.................................. 7</p><p>7. Bicoloured (Fig. 2), head yellow medially, pronotum and abdominal segments II‒VII largely yellow; mid and hind legs with femora brown but tibiae yellow; pronotum posterior fifth weakly reticulate (Fig. 7); male with large circular pore plate on sternite VIII............................................................................... cameroni sp.n.</p><p>-. Head, thorax and abdomen uniformly light brown, head with small pale area antero-medially (Fig. 3); mid and hind legs with femora and tibiae brown; pronotum posterior fifth finely tuberculate (Fig. 8); male with pore plate extending almost fully across sternite VIII............................................................................... irukandji sp.n.</p></div>	https://treatment.plazi.org/id/03DD87997446FFD25DC8E962FA4BFBC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mound, Laurence A.;Tree, Desley J.	Mound, Laurence A., Tree, Desley J. (2021): Structural variation among species of Leeuwenia (Thysanoptera, Phlaeothripinae) with three new species from Australia. Zootaxa 4903 (3): 439-447, DOI: 10.11646/zootaxa.4903.3.9
03DD87997446FFD35DC8EC91FC61FF5D.text	03DD87997446FFD35DC8EC91FC61FF5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leeuwenia aliceae Mound & Tree 2021	<div><p>Leeuwenia aliceae sp.n.</p><p>[Figs 1, 9‒12, 15]</p><p>Female macroptera. Body dark brown with yellow longitudinal stripe medially between compound eyes (Figs 1, 9); coxae and femora brown, tibiae yellow but washed with brown basally, tarsi yellow; fore wings pale with median longitudinal brown stripe; antennal segments I‒II brown, III‒VIII yellow. Head with genae parallel-sided; vertex with narrow transverse reticulation; postocular setae weakly capitate, small but distinct from minor setae, post ocellar setae slender and acute; maxillary stylets one third of head width apart, not deeply retracted into head (Fig. 9). Antennal segment III slender, almost 4.0 times as long as maximum width; III with one sense cone, IV with two, these sense cones slightly longer than apical width of their segment. Pronotum reticulate (Fig. 10), reticles with internal markings; am setae pointed and longer than discal setae, the other major setae capitate; notopleural sutures not fully complete. Mesonotal lateral setae small. Metanotum reticulate (Fig. 11), reticles with internal markings, with one pair of pointed setae medially and one pair of smaller setae near anterior margin. Prosternal basantra not developed, ferna transverse, mesopresternum complete; sternopleural sutures not present. Fore tarsi without tooth. Pelta wider than long (Fig. 11); tergites II‒VII each with two pairs of slender wing-retaining setae; tergite IX setae S1 slender with bluntly pointed apices, S2 longer and finely acute (Fig. 12); tube about 5.3 times as long as basal width.</p><p>Measurements (holotype female in microns). Body length 2900. Head, length 325; width across genae 220; postocular setae 25. Pronotum, length 175; width 325; major setae am 25, aa 50, ml 40, epim 65, pa 35. Fore wing length 1100; sub-basal setae S1‒ S3 35, 50, 50. Tergite IX setae S1 210, S2 275. Tube 480. Antennal segments III‒VIII length, 95, 80, 80, 75, 55, 45.</p><p>Material studied. Holotype female, Australia, Queensland, Fishery Falls [17°11’S; 145°53’E], 30km south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.88333&amp;materialsCitation.latitude=-17.183332" title="Search Plazi for locations around (long 145.88333/lat -17.183332)">Cairns</a>, from? Myrsine leaves, 10.xi.2007 (Alice Wells).</p><p>Paratypes: 8 females collected with holotype .</p><p>Comments. This species is remarkable for the elongate, pointed major setae on the ninth abdominal tergite, but it shares a relatively short tube with both L. dyospyri and L. polyosmae .</p></div>	https://treatment.plazi.org/id/03DD87997446FFD35DC8EC91FC61FF5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mound, Laurence A.;Tree, Desley J.	Mound, Laurence A., Tree, Desley J. (2021): Structural variation among species of Leeuwenia (Thysanoptera, Phlaeothripinae) with three new species from Australia. Zootaxa 4903 (3): 439-447, DOI: 10.11646/zootaxa.4903.3.9
03DD87997447FFD35DC8E962FEE8FA65.text	03DD87997447FFD35DC8E962FEE8FA65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leeuwenia cameroni Mound & Tree 2021	<div><p>Leeuwenia cameroni sp.n.</p><p>[Figs 2, 7, 13, 16, 17]</p><p>Female macroptera. Body bicoloured (Fig. 2), pterothorax and all coxae and femora brown, tibiae and tarsi yellow; head and pronotum yellow with brown lateral margins; abdominal segments II-VII largely yellow with variable brown markings medially, VIII light brown, IX and tube uniformly darker; fore wing pale without basal shading but with faint median longitudinal shaded line; antennal segments I‒II brown, III‒VIII yellow. Head with genae convex, sharply constricted behind compound eyes (Fig. 7); vertex with numerous small tubercles on lines of reticulation; postocular setae not distinguished from minor setae, post ocellar setae small; maxillary stylets about one fifth of head width apart, retracted to postocular region (Fig. 7). Antennal segment III less than 3.0 times as long as maximum width (Fig. 16); III with one sense cone equal in length to segment width, IV with two sense cones about 1.5 times as long as segment apical width. Pronotum reticulate, with median longitudinal band of reticles; setae am, ml and pa not distinguished from discal setae, aa and epim broadly capitate; notopleural sutures not complete. Mesonotal lateral setae minute. Metanotum (Fig. 17) reticulate, reticles with internal markings, sometimes with one or more pairs of small setae lateral to the median pair. Prosternal basantra weakly developed, ferna transverse and complete medially, mesopresternum complete; sternopleural sutures not present, meta-epimera swollen. Fore tarsi without tooth. Pelta rectangular, longer than wide with lateral margins constricted (Fig. 17); tergites II‒VII each with two pairs of broadly flattened wing-retaining setae (Fig. 13); tergite IX setae S1 and S2 short and blunt to weakly capitate; tube at least 12 times as long as basal width.</p><p>Measurements (holotype female in microns). Body length 2500. Head, length 250; width across genae 185; postocular setae 10. Pronotum, length 150; width 250; major setae am 10, aa 25, ml 10, epim 55, pa 10. Fore wing length 700; sub-basal setae S1‒ S3 10, 10, 10. Tergite IX setae S1 15, S2 40. Tube 600. Antennal segments III‒VIII length, 60, 50, 55, 50, 45, 40.</p><p>Male macroptera. Closely similar to female in colour and structure but smaller; tergite IX setae S2 shorter than S1; sternite VIII with large circular pore plate medially.</p><p>Material studied. Holotype female, Australia, Queensland, Mt Lewis [16°35’S; 145°18’E], 60km northwest of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.3&amp;materialsCitation.latitude=-16.583334" title="Search Plazi for locations around (long 145.3/lat -16.583334)">Cairns</a>, from leaf galls on Syzygium sp., 1.x.2015 (LAM 6171) [on slide with paratype male].</p><p>Paratypes: 16 females, 10 males collected with holotype; same locality, 5 females, 4 males from leaf galls on Syzygium alatoramulum, 18.ix.2013 (DJT 1652) .</p><p>Comments. As indicated, this species is very similar in many structural details to L. irukandji but is noticeably different in colour. It is named in recognition of the extensive support provided to thrips studies in Queensland by Cameron Tree.</p></div>	https://treatment.plazi.org/id/03DD87997447FFD35DC8E962FEE8FA65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mound, Laurence A.;Tree, Desley J.	Mound, Laurence A., Tree, Desley J. (2021): Structural variation among species of Leeuwenia (Thysanoptera, Phlaeothripinae) with three new species from Australia. Zootaxa 4903 (3): 439-447, DOI: 10.11646/zootaxa.4903.3.9
03DD87997447FFD35DC8EE3AFE69F919.text	03DD87997447FFD35DC8EE3AFE69F919.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leeuwenia convergens Hood 1918	<div><p>Leeuwenia convergens Hood, 1918</p><p>This species remains known only from two females collected in Queensland near Cairns from an unknown plant (Mound 2004). It has a distinctively elongate head with a transverse row of small setae across the vertex, and the tube is exceptionally long.</p></div>	https://treatment.plazi.org/id/03DD87997447FFD35DC8EE3AFE69F919	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mound, Laurence A.;Tree, Desley J.	Mound, Laurence A., Tree, Desley J. (2021): Structural variation among species of Leeuwenia (Thysanoptera, Phlaeothripinae) with three new species from Australia. Zootaxa 4903 (3): 439-447, DOI: 10.11646/zootaxa.4903.3.9
03DD87997447FFD35DC8EF26FC18F879.text	03DD87997447FFD35DC8EF26FC18F879.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leeuwenia diospyri Mound 2004	<div><p>Leeuwenia diospyri Mound, 2004</p><p>This species was described from a series of both sexes taken in curled leaf galls on Diospyros pentamera [ Ebenaceae] in coastal rain forest at Taree, New South Wales. However, it was found subsequently in similar leaf-roll galls on the same plant species at Lamington in southeast Queensland. It is unusual amongst members of the genus in having the tube rather short, scarcely six times as long as the basal width.</p></div>	https://treatment.plazi.org/id/03DD87997447FFD35DC8EF26FC18F879	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mound, Laurence A.;Tree, Desley J.	Mound, Laurence A., Tree, Desley J. (2021): Structural variation among species of Leeuwenia (Thysanoptera, Phlaeothripinae) with three new species from Australia. Zootaxa 4903 (3): 439-447, DOI: 10.11646/zootaxa.4903.3.9
03DD87997442FFD65DC8E8BEFEF5FB4D.text	03DD87997442FFD65DC8E8BEFEF5FB4D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leeuwenia irukandji Mound & Tree 2021	<div><p>Leeuwenia irukandji sp.n.</p><p>[Figs 3, 8, 14, 18, 19]</p><p>Female macroptera. Body light brown with yellow longitudinal stripe medially between compound eyes (Figs 3, 8); tarsi yellow, but all coxae also mid and hind femora and tibiae brown, fore tibiae and most of fore femora yellow; fore wings pale, shaded at base and with weak median longitudinal darker stripe; antennal segments I‒II brown, III‒VIII yellow. Head with genae convex, constricted behind large compound eyes (Fig. 8); vertex with numerous small tubercles; postocular setae not distinguished from minor setae, post ocellar setae small; maxillary stylets less than one third of head width apart, retracted to postocular region (Fig. 8). Antennal segment III less than 3.0 times as long as maximum width; III with one sense cone, IV with two, these sense cones distinctly longer than apical width of their segment. Pronotum reticulate, posterior fifth with transverse band of minute tubercles (Fig. 8); setae am, ml and pa not distinguished from discal setae, aa and epim broadly capitate; notopleural sutures not complete. Mesonotal lateral setae minute. Metanotum reticulate, reticles with internal markings, with one pair of pointed setae medially. Prosternal basantra weakly developed, ferna transverse and complete medially (Fig. 19), mesopresternum complete; sternopleural sutures not present, meta-epimera swollen. Fore tarsi without tooth. Pelta rectangular, longer than wide with lateral margins slightly constricted (Fig. 18); tergites II‒VII each with two pairs of broadly flattened wing-retaining setae (Fig. 14); tergite IX setae S1 and S2 short and blunt to weakly capitate; tube at least 12 times as long as basal width.</p><p>Measurements (holotype female in microns). Body length 2500. Head, length 250; width across genae 200; postocular setae 10. Pronotum, length 135; width 250; major setae am 10, aa 25, ml 10, epim 45, pa 10. Fore wing length 800; sub-basal setae S1‒ S3 10, 10, 20. Tergite IX setae S1 25, S2 70. Tube 760. Antennal segments III‒VIII length, 60, 50, 55, 50, 45, 35.</p><p>Male macroptera. Closely similar to female but smaller; tergite IX setae S2 shorter than S1; sternite VIII fully occupied by pore plate.</p><p>Material studied. Holotype female, Australia, Queensland, Cairns, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.66667&amp;materialsCitation.latitude=-16.866667" title="Search Plazi for locations around (long 145.66667/lat -16.866667)">Stoney Creek</a> Road [16°52’S; 145°40’E], from Syzygium tierneyanum leaves, 6.xi.2008 (LAM 5202).</p><p>Paratypes: 2 females, 6 males collected with holotype; 2 females, 1 male, same locality and tree, 4.x.2012 .</p><p>Comments. This species is structurally very similar to L. cameroni, and it lives on the leaves of a tree in the same genus, Syzygium . The two species are strikingly different in colour (Figs 2, 3), but also differ in head shape and the character states indicated in the key above. The name refers to the language of the original people of northern Queensland.</p></div>	https://treatment.plazi.org/id/03DD87997442FFD65DC8E8BEFEF5FB4D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mound, Laurence A.;Tree, Desley J.	Mound, Laurence A., Tree, Desley J. (2021): Structural variation among species of Leeuwenia (Thysanoptera, Phlaeothripinae) with three new species from Australia. Zootaxa 4903 (3): 439-447, DOI: 10.11646/zootaxa.4903.3.9
03DD87997442FFD65DC8ED19FCE2FA5E.text	03DD87997442FFD65DC8ED19FCE2FA5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leeuwenia polyosmae Mound 2004	<div><p>Leeuwenia polyosmae Mound, 2004</p><p>Described originally from two females taken in a leaf gall on Polyosma cunninghamii [ Grossulariaceae] near Brisbane, Queensland, this thrips was found subsequently on the same plant species in rainforests at Lamington near the border with New South Wales. Some of the galls were a remarkable blue colour (Fig. 6), and the species is unusual within the genus in having a fore tarsal tooth, in both sexes.</p></div>	https://treatment.plazi.org/id/03DD87997442FFD65DC8ED19FCE2FA5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mound, Laurence A.;Tree, Desley J.	Mound, Laurence A., Tree, Desley J. (2021): Structural variation among species of Leeuwenia (Thysanoptera, Phlaeothripinae) with three new species from Australia. Zootaxa 4903 (3): 439-447, DOI: 10.11646/zootaxa.4903.3.9
03DD87997442FFD65DC8EE69FA88F942.text	03DD87997442FFD65DC8EE69FA88F942.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leeuwenia scolopiae Mound 2004	<div><p>Leeuwenia scolopiae Mound, 2004</p><p>This species is known from Lamington and Mt Tambourine, two rainforest localities south of Brisbane, Queensland, breeding in curled leaf-galls on Scolopia braunii [ Flacourtiaceae]. It is unusual within the genus in having the proternal basantra well-developed, and in having a number of minor setae on the anterior half of the metanotum.</p></div>	https://treatment.plazi.org/id/03DD87997442FFD65DC8EE69FA88F942	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mound, Laurence A.;Tree, Desley J.	Mound, Laurence A., Tree, Desley J. (2021): Structural variation among species of Leeuwenia (Thysanoptera, Phlaeothripinae) with three new species from Australia. Zootaxa 4903 (3): 439-447, DOI: 10.11646/zootaxa.4903.3.9
03DD87997442FFD65DC8EF1BFEB6F85E.text	03DD87997442FFD65DC8EF1BFEB6F85E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leeuwenia tetrastigmae Mound 2004	<div><p>Leeuwenia tetrastigmae Mound, 2004</p><p>The host plant of this thrips, Tetrastigma nitens [ Vitaceae], is widespread in rainforest areas of the Australian East coast from Sydney northwards. However, populations of the thrips and the distorted leaf-galls it induces have a patchy distribution. Adults are smaller than those of convergens, with the head and the tube shorter, and the legs uniformly yellow.</p></div>	https://treatment.plazi.org/id/03DD87997442FFD65DC8EF1BFEB6F85E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mound, Laurence A.;Tree, Desley J.	Mound, Laurence A., Tree, Desley J. (2021): Structural variation among species of Leeuwenia (Thysanoptera, Phlaeothripinae) with three new species from Australia. Zootaxa 4903 (3): 439-447, DOI: 10.11646/zootaxa.4903.3.9
