taxonID	type	description	language	source
03DD87C4FFD4FFFB1EFCDFF7FC37FC7C.taxon	description	Descriptions of the male and larva are available in Kohls (1950).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD4FFFB1EFCDE8FFE6CFA3C.taxon	description	The life cycle was described by Hoogstraal et al. (1975). It shows considerable variation in the timing of the different stages, depending on the climatic conditions in the area from which the ticks come. It can be found throughout the year, usually under the bark of trees where the hosts nest (McKilligan 1987). All life history stages of A. robertsi are described in Hoogstraal et al. (1968 a), with an adult female illustrated in Hoogstraal et al. (1975).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD5FFFA1EFCDDA7FC46FDE4.taxon	description	The description of an adult by Hirst (1929) is extremely brief. A key to the larvae of the then known Ornithodoros from the Eastern Hemisphere is provided by Sonenshine et al. (1966). Dumbleton (1958) provides illustrations of larval chaetotaxy, some larval morphology and the Haller’s organ of the adults. Klompen et al. (1995) provide a map of the distribution of O. batuensis and related species.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD5FFFA1EFCDFF7FF43FA8C.taxon	description	Continental Southeast Asian records of O. capensis are from Laridae, Sterna spp., in Thailand (coastal Chon Buri Province, Sattahip District) (Tanskul et al. 1983). These ticks tend to remain in the nesting places of their hosts outside the breeding season, allowing them to reinfest the hosts when they return to nest (Humphery-Smith and Moorhouse 1980). Heath (2006) provides some information on the life cycle and rearing of O. capensis on domestic chickens. The male, female, nymph and larva are illustrated in Yamaguti et al. (1971). Muñoz-Leal et al. (2017) describe the morphology of the larvae and provide micrographs. Larvae of this species are included in the key to species from the Eastern Hemisphere by Sonenshine et al. (1966). Some information on the adults is available in Kohls (1957).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD5FFF91EFCDB05FDEAFEBB.taxon	description	Durden and Keirans (1996) list this species as potentially endangered because one of its principal hosts, the wild pig Babyrousa celebensis (frequently listed as B. babyrussa, however this taxon has been divided into three separate species; Meijaard and Groves 2002 a, b), is listed as threatened by the International Union for the Conservation of Nature. All life history stages of this species are described in Keirans and Robbins (1987), and the male, female and nymph by Voltzit and Keirans (2002).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD6FFF91EFCDCCEFBEEFC7B.taxon	description	The male, female, nymph and larva are described in Voltzit and Keirans (2002).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD6FFF91EFCDE8EFC09FB1B.taxon	description	The male and female are described in Voltzit and Keirans (2002).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD6FFF91EFCD86EFBAAF9D7.taxon	description	The female, nymph and larva are described in Voltzit and Keirans (2002).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD6FFF91EFCDB22FCD4F8AF.taxon	description	The male and female are described and illustrated in Kaufman (1972, as Aponomma crassipes). Descriptions of the nymph and larva are in Roubaud and Colas-Belcour (1935).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD7FFF81EFCDDEEFC09FE57.taxon	description	The male and female are described in Voltzit and Keirans (2002).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD7FFF81EFCDCA2FC3BFC5F.taxon	description	The male, female and nymph are described in Roberts (1970, as Aponomma fimbriatum). Voltzit & Keirans (2002) treat A. fimbriatum (usually considered eyeless and with one spur on coxa I) as a synonym of A. helvolum (with eyes and two spurs on coxa I) without any further elaboration.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFD7FFF81EFCDEAAFEA6FAA7.taxon	description	In continental Southeast Asia, Am. geoemydae has been reported from Cambodia (Kolonin 2009), peninsular Malaysia, Myanmar, Thailand and Vietnam (Neumann 1908; Wilson 1966; Hoogstraal et al. 1968 b, 1972 b; Tanskul et al. 1983; Kolonin 1992; Petney and Keirans 1996 b; Robbins & Platt 2011). Frazier and Keirans (1990) consider it likely that this species is distributed on a variety of tortoise and lizard species from India and across Myanmar in addition to its known range. Nadchatram (1960) provides information on rearing this species using Malaysian specimens. The male, female, nymph and larva are described in Voltzit and Keirans (2002) and illustrated in Yamaguti et al. (1971).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC8FFE71EFCDC16FB40FD13.taxon	description	The male, female, nymph and larva are described in Voltzit and Keirans (2002). Voltzit & Keirans (2002) describe the nymph as having dentition 2 / 2 but the corresponding figure (14 c) shows dentition 3 / 3.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC8FFE71EFCD9B6FB2DF9F2.taxon	description	The male and female are described and illustrated in Kaufman (1972, as Aponomma pattoni). The nymph is briefly described in Schulze (1935), as Aponomma pseudolaeve. The larva is undescribed.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC8FFE61EFCDA08FBEEFD5B.taxon	description	Fujisaki et a 1. (1976) provide data on a laboratory colony of this species. Chao et al. (2017) provide information on the morphological and molecular identification of this species. The male, female, nymph and larva are described in Voltzit and Keirans (2002).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC9FFE61EFCDFAEFBC1FC33.taxon	description	The male and female are described in Roberts (1970, as Aponomma trimaculatum).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC9FFE51EFCDB4EFB01FF27.taxon	description	Dermacentor was the genus most commonly associated with intra-aural infestation in humans (otoacariasis) (99.7 %) in Malaysia (Mariana et al. 2008 b). Although four species of Dermacentor have been found in Malaysia, only three species were involved: D. atrosignatus, D. compactus and D. steini, with the first mentioned being the most frequently extracted. The fourth species, D. auratus, was not found in such infestations.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCAFFE51EFCDC35FBD4FD12.taxon	description	Descriptions of the male and female are available in Wassef and Hoogstraal (1984 b).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCAFFE51EFCDE59FBD6FAEF.taxon	description	Descriptions of the male and female are available in Wassef and Hoogstraal (1984 a).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCAFFE51EFCD8FDFB5FF892.taxon	description	Descriptions of all life history stages are available in Apanaskevich and Apanaskevich (2015 c).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCBFFE41EFCDFCDFAA9FC17.taxon	description	The male and female of D. filippovae are described in Apanaskevich and Apanaskevich (2015 a).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCBFFE41EFCD965FAA7FA8A.taxon	description	The male and female of D. limbooliati are described in Apanaskevich and Apanaskevich (2015 a).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCBFFE41EFCD8D1FC70F962.taxon	description	The male and female are described in Wassef and Hoogstraal (1986 b).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCBFFE31EFCDA69FB44FE72.taxon	description	Wassef and Hoogstraal (1986 a) described the male and female, and Kitaoka and Suzuki (1981) the immature stages. However, the descriptions of the larva and nymph of D. taiwanensis in Kitaoka & Suzuki (1981) are considered to be the first descriptions of the larva and nymph of D. bellulus according to Apanaskevich & Apanaskevich (2015 c). Apanaskevich and Apanaskevich (2015 b) redescribed the male and female.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCCFFE31EFCDF79FAA8FCE7.taxon	description	The male and female of D. tamokensis are described in Apanaskevich and Apanaskevich (2016).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCCFFE31EFCD9FFFC3DF9F4.taxon	description	All life history stages are described in Hoogstraal et al. (1971 b).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCDFFE21EFCDC16FB28FD12.taxon	description	The male, female, nymph and larva are described in Hoogstraal and Trapido (1966 a).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCDFFE21EFCDE66FC3FFB87.taxon	description	All life history stages are described in Hoogstraal et al. (1965 a).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCDFFE21EFCD9D2FC6EF9F0.taxon	description	All life history stages are described in Hoogstraal and Kohls (1965).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCEFFE11EFCDDEEFD2EFD13.taxon	description	The female neotype is described by Hoogstraal and Trapido (1966 b). Trapido et al. (1964 b) illustrate the larval capitulum and the nymphal capitulum, coxae and trochanters. Lim et al. (2017) describe the initiation of primary cell cultures from embryonic Ha. bispinosa.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCEFFE11EFCDE66FC10FBCF.taxon	description	Adults and nymphs are described in Hoogstraal and Wassef (1981).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCEFFE11EFCD91AFC0DFA43.taxon	description	The male, female and nymph are described by Hoogstraal (1971).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCEFFE01EFCDB92FE84FE58.taxon	description	This is a rare species known from Vietnam and southern and southwestern China (Hoogstraal and Wilson 1966; Phan Trong 1977; Kolonin 1992, 1995; Teng and Jiang 1971). Kolonin (2009) points out that its presence in northern Vietnam is uncertain and that a closely related species might occur there. In addition, he reports that females of this species from Bamun Island (Dao Ba Mun) are intermediate between Ha. colasbelcouri (as Ha. vietnamensis) and Ha. kitaokai, a primitive Haemaphysalis species from East Asia (Hoogstraal 1969; Chen et al. 2010). The female is described in Hoogstraal and Wilson (1966) and the male in Kolonin (1992), both as Ha. vietnamensis.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCFFFE01EFCDE66FCCDFC33.taxon	description	The male and female were described by Phan Trong (1977).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCFFFE01EFCD946FC72FAA7.taxon	description	The male and female are illustrated in Hoogstraal and Dhanda (1970).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCFFFE01EFCD8B2FC4AF93F.taxon	description	The male, female and nymph are described in Hoogstraal et al. (1970).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFCFFFEF1EFCDA4AFC7CFE57.taxon	description	All life history stages are illustrated in Hoogstraal and Wassef (1973).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC0FFEF1EFCDCA2FD57FD2F.taxon	description	The female is illustrated in Yamaguti et al. (1971).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC0FFEF1EFCD962FC15FA91.taxon	description	The male and female are described in Hoogstraal and Kohls (1968).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC0FFEF1EFCD8EAFC8DF9A1.taxon	description	All life history stages are described in Dhanda (1964).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC0FFEF1EFCDBBAFCD4F829.taxon	description	The male, female and nymph are described in Roberts (1960).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC1FFEE1EFCDE8EFC32FB37.taxon	description	The male and female are illustrated in Tanskul and Inlao (1989).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC2FFED1EFCDDEEFC93FD5B.taxon	description	Interestingly, Tsunoda and Tatsuzawa (2004) found that the questing height of this species (and Ha. longicornis) was related to the height of their sika deer (Cervus nippon) hosts, which varies between areas in Japan, as well as the height of the local vegetation. All stages are described in Saito and Hoogstraal (1973).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC2FFED1EFCDFAEFD09FBCF.taxon	description	Rajagopalan (1963) describes all life history stages; Trapido et al. (1964 b) illustrate the larval capitulum and the nymphal capitulum, coxae and trochanters.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC2FFED1EFCD8DEFBF2F8AF.taxon	description	The adults are described by Hoogstraal et al. (1966), and the immature stages by Hoogstraal et al. (1971 a). Guru et al. (1976) describe cell cultures derived from developing adults of Ha. obesa.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC3FFEC1EFCDEF2FC8DFBA3.taxon	description	The male and female are described in Kolonin (1992).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC3FFEC1EFCD9B6FCBBFA42.taxon	description	The male was described by Hoogstraal et al. (1966).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC4FFEB1EFCDCCEFE7AFB88.taxon	description	Bhat (1979, 1981) provides information on the life cycle under laboratory conditions as well as extremely useful information under field conditions. Trapido et al. (1964 b) illustrate the larval capitulum and the nymphal capitulum, coxae and trochanters. Banerjee et al. (1977) and Guru et al. (1976) demonstrated the growth of various arboviruses in cell cultures derived from Ha. spinigera.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC4FFEB1EFCD8DEFC46F9AA.taxon	description	The male, female, nymph and larva are illustrated in Hoogstraal (1964).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC4FFEB1EFCDBBEFBA8F867.taxon	description	The male and female are described in Hoogstraal and El Kammah (1971).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC5FFEA1EFCDB23FACDF84C.taxon	description	Descriptions of all active life history stages, together with illustrations of the male and female and line drawings of the nymphal and larval capitula, scuta and coxae, are available in Geevarghese & Dhanda (1987).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC6FFE91EFCD963FCD5FA60.taxon	description	All life history stages are illustrated in Yamaguti et al. (1971).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFC7FFE81EFCDCA2FB25FC17.taxon	description	The female is described in Kohls (1962), the female and male in Clifford et al. (1971).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFF8FFD71EFCDDEEFE6BFE73.taxon	description	The validity of this species is doubtful, with some authors treating it as a synonym of I. ovatus (Camicas et al. 1998; Kolonin 2009), but it is tentatively recognized by Guglielmone et al. (2009, 2014). There is thus a need to revise what is currently considered I. ovatus to determine whether I. siamensis is part of a species complex (Guglielmone et al. 2009).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFF8FFD71EFCD8DEFAB7F93F.taxon	description	The male and female of I. tanuki are illustrated in Yamaguti et al. (1971) and Clifford et al. (1975).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFF9FFD61EFCDBE5FC49F82C.taxon	description	The female, male and larva are described in Estrada-Peña et al. (2012).	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFFAFFD41EFCD896FAACFDC7.taxon	description	This group is being investigated by a number of institutions, and current information suggests that at least two main taxa are present: a temperate and a tropical one. Evidence from China indicates that the species found there is probably closely related to, but different from, that found in North America (Liu et al. 2013). Ticks classified as R. sanguineus from China were found to fall into either clade III, which includes R. guilhoni sequences, or clade IV, with R. turanicus sequences, in the R. sanguineus group as defined by Hekimoğlu et al. (2016). Currently, very little information is available on the genetic characterization of the Southeast Asia taxon (or taxa). Specimens from Thailand (labelled from both Bangkok and Phuket, and from a laboratory colony) fall into the tropical form, also found in Taiwan, Brazil, Mozambique and Cuba by molecular analysis using 12 S and 16 S rDNA sequences (Sanchez et al. 2016). Hekimoğlu et al. (2016) found that the tropical strain from South America showed a strong molecular similarity to R. camicasi from Africa. This species is clearly different from the temperate R. sanguineus, and the diseases transmitted by the two species may differ. Therefore, evidence for the transmission of tick-borne diseases should be accepted only for data from continental Southeast Asia.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
03DD87C4FFFBFFD41EFCDF12FB38FC17.taxon	description	All life history stages were described by Kitaoka and Suzuki (1983). The adults were described from moulted nymphs but were lost in transit. Petney and Keirans (1996 a) did not consider this species to be valid, but Guglielmone et al. (2014) consider it tentatively valid based on museum specimens of the immatures. The latter authors, however, are uncertain whether larvae found on larval hosts represent this species.	en	Petney, Trevor N., Boulanger, Nathalie, Saijuntha, Weerachai, Chitimia-Dobler, Lidia, Pfeffer, Martin, Eamudomkarn, Chatanun, Andrews, Ross H., Ahamad, Mariana, Putthasorn, Noppadon, Muders, Senta V., Petney, David A., Robbins, Richard G. (2019): Ticks (Argasidae, Ixodidae) and tick-borne diseases of continental Southeast Asia. Zootaxa 4558 (1): 1-89, DOI: 10.11646/zootaxa.4558.1.1
