identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DD8F18613BFFB4A7BCFCE1FE618D62.text	03DD8F18613BFFB4A7BCFCE1FE618D62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Noctuidae	<div><p>Noctuidae: Noctuinae</p><p>Checklist in alphabetical order. Species found in Thailand are shown in bold face</p><p>E. alleni Holloway, 2009</p><p>E. cryptodisca Hampson, 1926</p><p>E. discophora Hampson, 1926</p><p>E. dulgistriga Walker, 1858</p><p>E. elliptica Holloway, 2009</p><p>E. grisescens sp. n.</p><p>E. lingulata Holloway, 2009</p><p>E. longinquua Swinhoe, 1890</p><p>E. melas Bethune-Baker, 1906</p><p>E. obscura Holloway, 2009</p><p>E. ovifera Holloway, 2009</p><p>E. pauli Holloway, 1976</p><p>E. philippinensis Wileman &amp; West, 1929</p><p>E. sciachroa Hampson, 1926</p><p>E. spiculivalva sp. n.</p><p>E. susanae Holloway, 1976</p><p>E. triangulata Holloway, 2009</p></div>	https://treatment.plazi.org/id/03DD8F18613BFFB4A7BCFCE1FE618D62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pellinen, Markku J.;Mutanen, Marko	Pellinen, Markku J., Mutanen, Marko (2019): Two new species of Ecpatia Turner, 1902, and the first records of Ecpatia sciachroa Hampson, 1926 and Ecpatia obscura Holloway, 2009 from Thailand (Lepidoptera: Noctuidae, Noctuinae). Zootaxa 4609 (3): 574-582, DOI: 10.11646/zootaxa.4609.3.11
03DD8F18613BFFB7A7BCF92DFCAD8C1A.text	03DD8F18613BFFB7A7BCF92DFCAD8C1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ecpatia grisescens Pellinen & Mutanen 2019	<div><p>Ecpatia grisescens sp. n. Pellinen</p><p>Barcode Index Number: BOLD:ADL2288</p><p>(Figs 1 a–b, 6 a–b and 7)</p><p>Type material. Holotype male: Thailand, Lampang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.545555&amp;materialsCitation.latitude=18.723612" title="Search Plazi for locations around (long 99.545555/lat 18.723612)">Chae Hom</a>, 09.xi. 2016, 350 m, 18°43’25’’N, 99°32’44’’E, M. J. Pellinen leg., genitalia slide MJP1005. In coll. Pellinen (Finland). The holotype will be deposited in a public museum . Paratypes: 30 males and 8 females, all specimens collected by first author. Males: Chiangmai, Thoeng Dang, 02.iv. 2006, 500 m, 19°39’52’’N, 99°12’14’’E, Lampang, Phraya Chae, 20.x. 2009, 350 m, 18°17’75’’N, 99°56’02’’E, genitalia slide MJP174, Lampang, Muban Phichai, 21.ix. 2010, 240 m, 18°18’15’’N, 99°31’06’’E, idem. 20.v.2011, idem., 29.v.2013, idem. genitalia slide MJP812, idem., 09.vi.2014, Lampang, Chae Hom, 27.ix. 2014, 350 m, 18°43’25’’N, 99°32’44’’E, genitalia slide MJP838, idem., 07.x.2014, idem., 17.x.2014, idem., 01.xi.2014, idem., 14. iii.2015, genitalia slide MJP829, idem., 14.iii.2015, genitalia slide MJP994, idem., 09.v.2015, idem., 26.vi.2015, idem., 26.vi.2015, idem., 26.vi.2015, genitalia slide MJP730, idem., 26.viii.2015, Lampang, Phraya Chae, 23.xi. 2016, 350 m, 18°17’75’’N, 99°56’02’’E, Lampang, Chae Hom, 26.xi. 2016, 350 m, 18°43’25’’N, 99°32’44’’E, idem., 26.xi.2016, genitalia slide MJP1004, idem., 17.xii.2016, genitalia slide 1012, idem., 29.vii.2017, idem., 09.ix.2017, genitalia slide 1252, idem., 18.xi.2017, idem., 24.ii.2018, genitalia slide MJP1256, idem., 14.iv.2018, idem., 25.v.2018, idem., 25.v.2018, idem., 14.x.2018, idem., 27.x.2018,. Females: Lampang, Muban Phichai, 21.iii. 2011, 240 m, 18°18’15’’N, 99°31’06’’E, Lampang, Chae Hom, 04.v. 2015, 350 m, 18°43’25’’N, 99°32’44’’E, M. J. Pellinen leg., genitalia slide MJP825,, idem., 26.vi.2015, idem., 26.vi.2015, genitalia slide MJP731, idem., 18.vii.2015, Lampang, Chae Hom, 12.ix. 2015, 350 m, 18°43’25’’N, 99°32’44’’E, M. J. Pellinen leg., idem., 17.x.2015 ., idem., 27.x.2018 . Paratypes in coll. Pellinen (Finland). All type specimens have red or green rectangle label, indicating the type status (holotype / paratype) .</p><p>Morphological diagnosis. Ecpatia grisescens is slightly smaller than other Ecpatia species. Externally it differs in having dark hindwings; all other known Ecpatia species have at least some part of the hindwing white. It has forewing markings, with a small faint red blotch between orbicular and reniform stigma, unlike other Ecpatia species. As in other congenera, the penicular process bears strong setae, separated from the sacculus by a distinct constriction. Ecpatia grisescens is bilaterally asymmetrical with respect to the harpe, as in E. elliptica . Female genitalia have no signum in the corpus bursae (otherwise a common feature to other Ecpatica species), which is pyriform and partly corrugated.</p><p>Genetic characterization. Sequences of the two barcoded specimens of E. grisescens are identical. Among five species occurring in Thailand, the barcode sequence of E. grisescens is most similar to that of E. spiculivalva (6.73%). Among 10 species of Ecpatia (described or putative) available in to us BOLD, the most similar are E. dulcistriga and E. melas (both differing by 6.06% from E. grisescens).</p><p>Description. External characters. Wingspan of both sexes 22–24 mm. Frons smooth, lower half whitish with narrow scales and upper half grey with wide scales. Labial palps upturned, medium size, covered with grey-brown scales mixed with some white, third segment less than half the length of second segment. Antennae of males fasciculate and filiform in females, cilia in males about 1.5x the length of flagellomere. Otherwise sexes are externally similar. Thorax grey-brown with broad scales, patagia and tegulae of same color. Legs grey-brown, with whitish tips on segments. Forewing grey with dark brown patches between black lines. Orbicular stigma small, reniform obscure with posterior reddish spot. Postmedial line wavy and double, cilia grey. Hindwing uniformly grey-brown, except for small faint dot dorsally, terminal line blackish, cilia grey. Underside pale greyish-brown. Vein M2 on hindwing present, almost equidistant between M1 and M3, weaker compared to other veins, arising from close dstans of mcell. Abdomen similar in color to the wings, with tuft of scales on tergites 4–7, underside color grey, mixed with white scales. Male genitalia. (Figs. 5 a–b) Uncus narrow, bent, apically sharp, penicular process in tegumen obtuse triangular, juxta roundish, valvae with distally notched plate at the basal area of the costal margin, sacculus terminating in strong harpe, bifid on the right side only, cucullus axe-shaped with setae, separated distally from the sacculus by a narrow constriction. Saccus v-shape, narrow. Aedeagus: vesica with two cornuti and small round patch. Female genitalia. (Fig. 6) Papillae anales triangular, sharp, setose, apophyses posteriores slightly longer than apophyses anteriores, ostium bursae broad, ventrally cleft, ductus bursae short, corpus bursae pyriform, corrugated, without basal appendix.</p><p>Etymology. The species name refers to grey color of the adults.</p><p>Distribution. North Thailand. Specimens were collected in various biotopes between 240–350 m, all year round. It is most commonly encountered Ecpatia species in Northern Thailand.</p><p>Biology. Pre-imaginal stages of the new species are unknown.</p><p>Remark. Figure of this species is illustrated in the Moths of Thailand Vol. 3, part two (Kononenko &amp; Pinratana 2013) erroneously as Beana nitida Tams, plate 21, fig. 51.</p></div>	https://treatment.plazi.org/id/03DD8F18613BFFB7A7BCF92DFCAD8C1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pellinen, Markku J.;Mutanen, Marko	Pellinen, Markku J., Mutanen, Marko (2019): Two new species of Ecpatia Turner, 1902, and the first records of Ecpatia sciachroa Hampson, 1926 and Ecpatia obscura Holloway, 2009 from Thailand (Lepidoptera: Noctuidae, Noctuinae). Zootaxa 4609 (3): 574-582, DOI: 10.11646/zootaxa.4609.3.11
03DD8F186138FFB6A7BCF8F5FD578D8E.text	03DD8F186138FFB6A7BCF8F5FD578D8E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ecpatia spiculivalva Pellinen & Mutanen 2019	<div><p>Ecpatia spiculivalva sp. n. Pellinen</p><p>Barcode Index Number: BOLD:ADL2493</p><p>(Figs 2 a–b, 8a–b and 9)</p><p>Type material. Holotype male: Thailand, Lampang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.51833&amp;materialsCitation.latitude=18.304167" title="Search Plazi for locations around (long 99.51833/lat 18.304167)">Muban Phichai</a>, 09.vii. 2009, 240 m, 18°18’15’’N, 99°31’06’’E, M. J. Pellinen leg., genitalia slide MJP109. In coll. Pellinen (Finland). The holotype will be deposited in a public museum . Paratypes: 8 males and 4 females, all specimens collected by first author. Males: Chiangmai, Ban Thun Sala, 07.iv. 2005 , 350 m, 18°40’32’’N, 98°51’20’’E, Lampang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.85555&amp;materialsCitation.latitude=18.675554" title="Search Plazi for locations around (long 98.85555/lat 18.675554)">Muban Phichai</a>, 09.vi. 2007 , 240 m, 18°18’15’’N, 99°31’06’’E, idem., 09.vii.2008, idem., 27.iv.2009, idem., 12.vi.2013, Lampang, Chae Hom, 14.x. 2015 , 350 m, 18°43’25’’N, 99°32’44’’E, idem., 26.xi.2016, Lampang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.545555&amp;materialsCitation.latitude=18.723612" title="Search Plazi for locations around (long 99.545555/lat 18.723612)">Muban Phichai</a>, 19.v. 2017 , 240 m, 18°18’15’’N, 99°31’06’’E. Females: Lampang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.51833&amp;materialsCitation.latitude=18.304167" title="Search Plazi for locations around (long 99.51833/lat 18.304167)">Muban Phichai</a>, 29.vi. 2011 , 240 m, 18°18’15’’N, 99°31’06’’E, M. J. Pellinen leg., genitalia slide MJP110, Lampang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.51833&amp;materialsCitation.latitude=18.304167" title="Search Plazi for locations around (long 99.51833/lat 18.304167)">Chae Hom</a>, 23.v. 2015 , 350 m, 18°43’25’’N, 99°32’44’’E, genitalia slide MJP1020, idem., 14.iv.2018, genitalia slide MJP1242, idem., 09.xii.2018 . All type specimens have red or green rectangle label, indicating the type status (holotype / paratype).</p><p>Morphological diagnosis. The new species is equal in size to other Ecpatia species except to the smaller E. grisescens . Forewing color brown-grey, slightly paler than most other species and also differing by two red blotches on the postmedial line, one close to costa and the other close to the tornus. There is also a red blotch next to subbasal line. White spot in hindwing is small and roundish and, as in E. obscura, two white sections on the fringes are clearly visible. Male genitalia differ from those of other congeners in having spikes at the base of the sacculus.</p><p>Genetic characterization. The DNA barcode sequence of E. spiculivalva differs from that of E. dulcistriga by 4.9%. Of the species occurring in Thailand, that with the most similar barcode sequence is E. grisescens (at 6.73%).</p><p>Description. External characters (Fig. 2 a–b). Wingspan of both sexes 24–25 mm. Frons smooth, lower half whitish with narrow scales and upper half with reddish wide scales. Labial palpi upturned, medium size, covered with grey-brown scales mixed with some white, third segment less than half of the second segment. Antennae of males fasciculate, cilia about twice the length of flagellomere and filiform in females. Otherwise sexes externally alike. Thorax grey-brown, patagia and tegulae of the same color with wide scales. Legs grey-brown, with whitish tips on segments. Forewing brownish grey with darker brown patches between black lines. Orbicular stigma small and reniform reasonably clear, paler than general color of the forewing. Medial line unclear, wavy and partly double postmedial, with two reddish blotches, one close to costa and the other near tornus, third reddish blotch anterior to subbasal line. Cilia dark grey. Hindwing with clear small white centrum and two white sections on the fringe. Underside dark with white bands in the median area of both wings. Vein M2 on hindwing present, weaker compared to other veins. Abdomen same color as wings, with tufts of scales on tergites 4–7, underside color grey mixed with white scales. Male genitalia. (Figs. 8 a–b) Uncus narrow bended, apically sharp, penicular process in tegumen blunt drop-shaped, juxta unclear rectangle, valvae without clear plate basally on costa, sacculus terminating in strong sharp harpe, cucullus almost the same width as valva with setae and strong spines, two to three strong spikes rise from middle part of sacculus with setae and strong spines, two to three strong spikes arose from middle part of sacculus. Saccus dull v-shape. Aedeagus: vesica with one strong cornutus, a spatulate sclerite and two patches of deciduous spicules. Female genitalia. (Fig. 9) Papillae anales setose, concave, apophyses narrow, posteriores longer than anteriores, ostium bursae ventrally cleft, slightly broader than medium long ductus bursae, corpus bursae pyriform with bended appendix posteriorly, moderately corrugated, without basal appendix.</p><p>Etymology. The species name refers of the strong spikes in valvae of the male genitalia.</p><p>Distribution. North Thailand. The species was collected in various biotopes between 240–350 m, in April–July and November.</p><p>Biology. Preimaginal stages of the new species are unknown.</p><p>Remark. Figures of this species are illustrated in the Moths of Thailand Vol. 3, part two (Kononenko &amp; Pinratana 2013) as unidentified sp. Plate 42, no 51–52.</p></div>	https://treatment.plazi.org/id/03DD8F186138FFB6A7BCF8F5FD578D8E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pellinen, Markku J.;Mutanen, Marko	Pellinen, Markku J., Mutanen, Marko (2019): Two new species of Ecpatia Turner, 1902, and the first records of Ecpatia sciachroa Hampson, 1926 and Ecpatia obscura Holloway, 2009 from Thailand (Lepidoptera: Noctuidae, Noctuinae). Zootaxa 4609 (3): 574-582, DOI: 10.11646/zootaxa.4609.3.11
03DD8F186139FFB1A7BCF899FD578D1E.text	03DD8F186139FFB1A7BCF899FD578D1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ecpatia sciachroa Hampson 1926	<div><p>Ecpatia sciachroa Hampson, 1926</p><p>Barcode Index Number: BOLD:ACB3991</p><p>(Figs. 3 a–b, 10a–b and 11)</p><p>This species was described from Sri Lanka and has been recorded from the Philippines Hampson, 1926. Thai specimens fit with E. sciachroa, though the concept of this one will however have to be revised because of the fairly wide geographic distribution currently shown by the taxon (from Sri Lanka to the Philippines) (A. Zilli NHM, personal comment).</p><p>Specimens found in Thailand: 1 male, Lampang, Muban Phichai, 30.v. 2009, 240 m, 18°18’15’’N, 99°31’06’’E, M. J. Pellinen leg., genitalia slide MJP111. 8 females, Lampang, Muban Phichai, 21.viii. 2007, 240 m, 18°18’15’’N, 99°31’06’’E, idem., 29.viii.2007, Lampang, Mon Kao Kaew, 04.ii. 2008, 300 m, 20°24’00’’N, 99°32’44’’E, Lampang, Muban Phichai, 03.v. 2009, 240m, 18°18’15’’N, 99°31’06’’E, idem., 12.v.2009, idem., 09.vi.2009, Lampang, Phraya Chae, 27.ix. 2009, 350 m, 18°17’75’’N, 99°56’02’’E, genitalia slide MJP112, Lampang, Phraya Chae, 23.xi. 2016, 350 m, 18°17’75’’N, 99°56’02’’E, genitalia slide MJP1156.All specimens collected by first author.</p><p>Morphological diagnosis. This species is slightly larger than most other Ecpatia species, with somewhat narrower forewings. Externally it differs from other Ecpatia species in its more hazy brown color and reniform stigma which is positioned diagonally from middle part of the costa to tornus. White area in hindwing is triangular, reaching almost the base. Male genitalia of E. sciachroa (Fig. 8.) differ from E. longinquua by the broader saccus and juxta (Fig. 13.). (Gen. slide from Teruko Fukuda)</p><p>Genetic characterization. The species with the barcode sequence most similar to that of E. sciachroa is E. spiculivalva (6.9%). The sequence of E. longinquua shows no difference from that of E. sciachroa in the barcode region, but we cannot verify the identity of the available specimen of E. longinquua in BOLD (mined from GenBank) as it is not associated with a specimen image.</p><p>Description. External characters. Wingspan of both sexes 25–26 mm. Frons smooth, lower half whitish with narrow scales and upper half dull red with wide scales. Labial palps upturned, medium size, covered with light grey and dark scales, third segment about bone third of the second segment. Antennae of males fasciculate, cilia quite short, only same length than flagellomere and filiform in females. Thorax, patagia and tegulae dark grey with wide scales. Legs dark grey, with reddish and grey tips on segments. Forewing variegated brown-grey with yellowish brown patches between postmedial and submarginal lines. Orbicular stigma small, reniform, diagonally stretched toward dorsum, yellowish brown. Postmedial line wavy, almost zigzag-like and partly double, cilia grey brown. Submarginal line unclear white, marginal line black and fragmentary. Hindwings dark-grey, with clear round white area. Underside dark grey, with narrow white sections in the middle parts. Hindwing venation same as in preceding species. Abdomen same color as wings, with tuft of scales on tergites 4–7, underside color grey mixed with white scales. Male genitalia. (Figs. 9 a–b). Uncus not especially narrow, bent, apically sharp, penicular process in tegumen large quadrangular, juxta unclear triangular, valvae with arrow-shape plate basally on costal margin, sacculus terminating in strong bifid harpe, cucullus small axe-shape, with setae and stronger spines, separated distally with narrow constriction from sacculus. Saccus obtuse-angled triangle. Vesica of aedeagus corrugated with small broad based cornutus and spatulate sclerite. Female genitalia. (Figs. 10) Papillae anales round, apophyses posteriores slightly longer than apophyses anteriores, narrow. Ostium bursae round, posteriorly v-shaped, broader than medium-long ductus bursae. Corpus bursae only slightly corrugated, pyriform with unclear posterior appendix.</p><p>The species was collected in Northern Thailand from various biotopes between 240– 350 m. in May–June and August–November.</p><p>Biology. Preimaginal stages of the new species are unknown.</p><p>Remark. Figures of this species are illustrated in the Moths of Thailand Vol. 3, part two (Kononenko &amp; Pinratana 2013) as unidentified sp. Plate 42, no 53–54.</p></div>	https://treatment.plazi.org/id/03DD8F186139FFB1A7BCF899FD578D1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pellinen, Markku J.;Mutanen, Marko	Pellinen, Markku J., Mutanen, Marko (2019): Two new species of Ecpatia Turner, 1902, and the first records of Ecpatia sciachroa Hampson, 1926 and Ecpatia obscura Holloway, 2009 from Thailand (Lepidoptera: Noctuidae, Noctuinae). Zootaxa 4609 (3): 574-582, DOI: 10.11646/zootaxa.4609.3.11
03DD8F18613EFFB1A7BCF909FC8A8C8A.text	03DD8F18613EFFB1A7BCF909FC8A8C8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ecpatia obscura Holloway 2009	<div><p>Ecpatia obscura Holloway 2009</p><p>Barcode Index Number: not available at present</p><p>The presence of E. obscura (Figs. 4 and 12) Holloway in Thailand is confirmed, from northern Thailand: province Lampang). It was identified by comparing its genitalia (slide no MJP1021) to fig.82. in Moths of Borneo, Part 13. One specimen was detected from Thailand: province Lampang, Muban Phichai, 02.iii.2011. Externally, E. obscura is closest to E. longinquua (Fig 5).</p><p>Genetic characterization. E. obscura shows a highly characteristic DNA barcode, the closest sequences available to us in BOLD are those of E. dulcistriga (7.52%).</p></div>	https://treatment.plazi.org/id/03DD8F18613EFFB1A7BCF909FC8A8C8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pellinen, Markku J.;Mutanen, Marko	Pellinen, Markku J., Mutanen, Marko (2019): Two new species of Ecpatia Turner, 1902, and the first records of Ecpatia sciachroa Hampson, 1926 and Ecpatia obscura Holloway, 2009 from Thailand (Lepidoptera: Noctuidae, Noctuinae). Zootaxa 4609 (3): 574-582, DOI: 10.11646/zootaxa.4609.3.11
