taxonID	type	description	language	source
03DD045AFF8AFFE0FDFF4F6B2BB1FB7A.taxon	description	Stenotarsus is readily distinguished from other endomychids by the following set of characters — pseudotrimerous tarsi, with tarsomere II lobed and bearing a dense, spongy mat of setae underneath; antennal club 3 ­ segmented, segments not transverse; pronotum with broad, flat lateral bead; body distinctly pubescent, pubescence frequently dense, usually uniformly red­brown; body occasionally with a large black area on each elytron and the pronotum. Strohecker (1953) presents a key to the subfamilies and genera of the World’s Endomychidae. Since then, numerous taxonomic reorganizations have occurred. The first description of Stenotarsus larvae is in McHugh and Pakaluk (1997) based on two Australian taxa. Larvae of most Stenotarsus remain unknown. Published descriptions of adult beetles are brief and generally uninformative, unless one is familiar with the genus. Species identifications are based primarily upon comparison with properly identified material, and revisionary work is needed. Useful characters exist for identifying species of Stenotarsus, for example sexual dimorphism, the shape and color of antennomeres, body size and shape, vestiture color and density, and male genitalia. Species also are distinguishable by the presence or absence of black marks on the pronotum and elytra. Comparision of characters indicates that these marks are somewhat variable within a population, or are clinal (see following Remarks of Stenotarsus ovalis Arrow and S. panamanus Gorham). Not all species display sexual dimorphism, how­ ever. Males of some species can possess any of the following sexually dimorphic characters: spur on the metatrochanter (Tanaka 1986) or metafemur; metatibia modified, curved, with or without internal tubercles; metasternum with various structures on the anterior edge between the mesocoxae, usually a depression of some definable shape; abdomen sometimes with a dense patch of setae on one or all sternites. In addition, there can be different body and appendage proportions (Nedved and Windsor 1994 b). These dimorphisms are present in various combinations and degrees between species, and frequently offer the best characters for identification. Females of many species that lack such characters often are impossible to identify without an associated male. Many species of Stenotarsus occur in Panama. We were unable to identify some of the material available for study, and left these specimens for future work. The following key to eleven Panamanian species includes only those with published records, or specimens identified by Strohecker.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF89FFE2FE6A4DD62B15FD66.taxon	diagnosis	Diagnosis. Length 4.0 – 4.2 mm; width 2.5 – 2.8 mm. Body entirely red­orange, elongate (Fig. 8). Named for the color of the vestiture, yet no difference in color from that of many other species was noted. The male available for study is damaged; no sexual dimorphism was found.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF89FFE2FE6A4DD62B15FD66.taxon	discussion	Remarks. Some specimens in the type series were from Panama. A male and female from Panama (FSCA) were studied.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF89FFE2FE634EB32B81FC60.taxon	diagnosis	Diagnosis. Length 4.5 mm; width 3.0 mm. Body oval, red­orange; each elytron with a central mark of a metallic blue shine; pronotum with the lateral margin narrow (Fig. 4). Sexual dimorphism is unknown.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF89FFE2FE634EB32B81FC60.taxon	discussion	Remarks. This species was originally described from Panama. Only a female syntype (NHML) was studied. The narrow pronotal margin was commented upon by Arrow (1920), who transferred this species to Stenotarsus. Its placement here awaits further studies at the generic level.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF89FFE2FE024FB62C30FB56.taxon	diagnosis	Diagnosis. Length 5.5 – 6.1 mm; width 3.5 – 4.1 mm. Body oval, usually black, with only the elytral margins and suture red, but some specimens mostly red; pronotal setae purple­black; male metafemur with a basal spine, metatibia narrow, with internal row of tubercles (Fig. 11).	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF89FFE2FE024FB62C30FB56.taxon	discussion	Remarks. Based on various comments and discrepancies between localities cited in Gorham (1890), Arrow (1920), Blackwelder (1945), and Strohecker (1953), there is confusion as to the identity of specimens from southern Central America. Blackwelder (1945) states this taxon is in Panama, thus it is included here. Specimens studied were from Mexico and Central America (FSCA).	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF89FFE2FE6D48C32973FA6E.taxon	diagnosis	Diagnosis. Length 3.0 – 3.8 mm; width 2.0 – 2.2 mm. Body weakly elongate, red­orange; base of the pronotum without distinct margin (Fig. 5); pronotal base weakly concave over scutellum; no sexual dimorphism observed.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF89FFE2FE6D48C32973FA6E.taxon	discussion	Remarks. Specimens from Panama are present in Strohecker’s collection (FSCA).	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF89FFEDFE6F49BB2951FE83.taxon	diagnosis	Diagnosis. Length 5.0 – 6.0 mm; width 3.5 – 4.0 mm. Body red­orange, broadly oval; elytra with distinct coarse punctures near the humerus; male with spine on metatrochanter (Fig. 12), metasternum with a circular depression at anterior margin, hind tibia curved with internal row of tubercules, genitalia simple (Fig. 16), relatively short and broad.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF89FFEDFE6F49BB2951FE83.taxon	discussion	Remarks. Gorham (1890) identified specimens from Panama as this species, based on Gerstaecker’s description. We studied material from several South American countries (FSCA). Females were indistinguishable from those of S. ovalis, or the red specimens of S. panamanus, except by association with a male.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF86FFEDFE5C4D922B14F9A6.taxon	diagnosis	Diagnosis. Length 4.5 – 5.0 mm; width 3.0 – 3.5 mm. Body broadly oval, with color pattern or not; elytra with distinct coarse punctures near the humerus (Fig. 7); male with spine on metatrochanter, metasternum with transversely oval depression at anterior margin (Fig. 13), hind tibia curved, with row of internal tubercules, genitalia relatively short and broad, with accessory process (Fig. 17).	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF86FFEDFE5C4D922B14F9A6.taxon	discussion	Remarks. Arrow’s description of S. ovalis is based on Gorham’s apparent misidentification of S. orbicularis Gerstaecker from Guatemala (Gorham 1890). Later in the same paper, Arrow (1920) describes S. rotundus from a single Panamanian specimen identified by Gorham as S. pilatei Gorham. Strohecker’s collection contains his annotated copy of the Genera Insectorum for the Endomychidae (Strohecker 1953). In it, he notes that S. rotundus is a synonym of S. ovalis, which is a synonym of S. orbicularis. Upon studying a male syntype of S. ovalis from Guatemala (NHML) and the male holotype of S. rotundus from Panama (NHML), no morphological differences were found that would distinguish these nominate species. The only difference observed was that S. ovalis has a black mark on each elytron; while S. rotundus is entirely red. Such coloration is variable in other species of Stenotarsus. Additional specimens studied included Strohecker’s solid orange­red specimens of S. ovalis from Venezuela (FSCA) and a specimen of S. rotundus from Panama, identified by Strohecker (STRI). Strohecker’s notes also indicate he studied the type of S. orbicularis. Because we did not examine that specimen, no judgement can be made on possible synonymy of S. ovalis under S. orbicularis. Here we only synonymize S. rotundus under S. ovalis.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF86FFEDFE5C4D922B14F9A6.taxon	diagnosis	Diagnosis. Length 4.2 – 5.5 mm: width 3.0 – 4.0 mm. Body oval, usually with a color pattern (Fig. 9) (some solid orange), covered with golden pubescence; male genitalia long and narrow (Fig. 15). No sexual dimorphism noted.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF86FFEDFE5C4D922B14F9A6.taxon	discussion	Remarks. Stenotarsus panamanus and S. chiriquinus originally were described from Panama. The female holotype of S. chiriquinus (NHML) and specimens of S. panamanus from Panama (FSCA) were studied. No differences were discovered. Without a male to include in comparisons, we are hesitant to synonymize S. chiriquinus, which presumably lives at a higher elevation, and also is in the Central American biogeographic region, in contrast to eastern Panama (Kimsey 1992). These species usually have a color pattern. Mixed in with Strohecker’s normally colored S. panamanus (FSCA) are three solid red specimens from Barro Colorado Island, Panama, which he had identified as ‘‘ S. panamanus? ’’. These specimens, possibly teneral, differ from S. panamanus and S. chiriquinus only in their lack of the dark body marks. For this reason, S. panamanus appears in the key twice.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF87FFECFE1D4CF42B3CFE79.taxon	diagnosis	Diagnosis. Length 5.7 – 7.0 mm; width 4.0 – 4.8 mm. Body dull orange, elongate; antennae elongated, reaching elytral half; antennomere XI appearing curved (Fig. 2); no notable sexual dimorphism.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF87FFECFE1D4CF42B3CFE79.taxon	discussion	Remarks. Specimens studied are from Colombia and Costa Rica (FSCA), thus the species almost certainly occurs in Panama.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF87FFECFE544DA92A6BFBFD.taxon	description	Stenotarsus ‘‘ gigante ’’ (Nedved 1996: 3) Nomen Nudum	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF87FFECFE544DA92A6BFBFD.taxon	diagnosis	Diagnosis. Length 4.5 – 6.0 mm; width 3.5 – 4.2 mm. Body oval, elytra lacking distinct coarse punctures near humerus (Fig. 6), smoothly and evenly punctate; male with spine on metatrochanter, lacking depression on anterior margin of metasternum (Fig. 14), genitalia relatively short and broad (Fig. 18). Females also recognizable by elytral punctation.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF87FFECFE544DA92A6BFBFD.taxon	discussion	Remarks. Nedved’s (1996) use of the name S. ‘‘ gigante ’’ is not accompanied by a description nor any comments to set the identity of that population. Since we consider the Gigante population to be conspecific with S. subtilis, S. ‘‘ gigante ’’ is listed as a nomen nudum and placed as a synomym of S. subtilis. Specimens studied include three female syntypes of S. subtilis (NHML) from David (near sea level) and V. [Volcan] de Chiriquı´, Panama (approximately 1,200 m above sea level), as well as numerous specimens from both the Barro Colorado Island and Gigante Peninsula populations (approximately 100 m above sea level). Specimens from the aggregations collected during 1993 – 1999 will be deposited in various museums (STRI, NHML, FSCA, PESC, NMNH, OSUC, FEUP).	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF87FFECFE1148242B01FAD9.taxon	diagnosis	Diagnosis. Length 3.5 – 4.2 mm; width 2.8 – 3.5 mm. Body stout, red­orange; antennomeres III­VIII short, antennomeres IX­XI elongated and nearly parallel sided (Fig. 1); pronotal lateral edge with a small tooth at middle; no notable sexual dimorphism.	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
03DD045AFF87FFECFE1148242B01FAD9.taxon	discussion	Remarks. According to Arrow (1920: 49) the Panama specimens of Stenotarsus claviger Gerstaecker (1858) discussed by Gorham (1890) are actually S. validicornis. The specimens studied are from several South American countries (FSCA), with one from Panama (PESC).	en	Roubik, David W., Skelley, Paul E. (2001): Stenotarsus Subtilis Arrow, The Aggregating Fungus Beetle Of Barro Colorado Island Nature Monument, Panama (Coleoptera: Endomychidae). The Coleopterists Bulletin 55 (3): 249-263, DOI: 10.1649/0010-065X(2001)055[0249:SSATAF]2.0.CO;2, URL: http://dx.doi.org/10.1649/0010-065x(2001)055[0249:ssataf]2.0.co;2
