identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DA1A1EFF9FFFA1FF053ECCE2B63DC9.text	03DA1A1EFF9FFFA1FF053ECCE2B63DC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius Beier 1951	<div><p>Genus Lagynochthonius Beier, 1951</p><p>Type species: Chthonius johni Redikorzev, 1922, by original designation.</p></div>	https://treatment.plazi.org/id/03DA1A1EFF9FFFA1FF053ECCE2B63DC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFF9FFFAEFF053F79E3C23D46.text	03DA1A1EFF9FFFAEFF053F79E3C23D46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius biyunensis Hou & Feng & Zhang 2023	<div><p>Lagynochthonius biyunensis sp. nov. (ỠỄṄṁae)</p><p>Figs 2–5</p><p>Type material. Holotype ♂ (Ps.- MHBU-GZC190805-01): China, Guizhou Province, Panzhou City, Chengguan Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.64&amp;materialsCitation.latitude=25.77192" title="Search Plazi for locations around (long 104.64/lat 25.77192)">Nanmen Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.64&amp;materialsCitation.latitude=25.77192" title="Search Plazi for locations around (long 104.64/lat 25.77192)">Biyun Park</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.64&amp;materialsCitation.latitude=25.77192" title="Search Plazi for locations around (long 104.64/lat 25.77192)">Biyun Cave</a>, under the detritus and on the walls in the deep zone [25°46′18.91″N, 104°38′24.0″E], 1496 m a.s.l., 5 August 2019, Zegang Feng, Zhaoyi Li &amp; Chen Zhang leg. Paratypes: 4 ♂ (Ps.- MHBU-HBUARA#2022-498 - 01 –HBUARA#2022-498-04), 2 ♀ (Ps.- MSWU-HBUARA#2022-498 - 05 &amp; HBUARA#2022-498-06), with the same location as the holotype, 2 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg.</p><p>Etymology. Named after the type locality, Biyun Cave.</p><p>Diagnosis (♂ ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with two setae; tergites Ⅰ– Ⅴ each with four setae. Pedipalps slender, femur 8.06 (♂), 7.82–7.89 (♀) times longer than broad; chela 6.85–7.08 (♂), 7.27–7.43 (♀) times longer than broad; chelal fingers without intercalary teeth; chemosensory setae (sc) present on dorsum of chelal hand .</p><p>Description. Males (holotype and paratypes) (Figs 2F, 3A, 4A–F, 5).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 4C, 5A): carapace 1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with ten terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 5C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 4D, 5B): large, about as long as carapace, 2.40–2.47 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 15–17 teeth, distal one largest; movable finger with 21–22 retrorse contiguous small teeth; galea absent (Fig. 5B). Serrula exterior with 22–23 blades and serrula interior with 14–15 blades. Rallum with nine blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 5D).</p><p>Pedipalp (Figs 4A–B, E, 5E–G): long and slender, trochanter 1.20–1.26, femur 8.06, patella 2.35, chela 6.85– 7.08, hand 2.70–2.81 times longer than broad; femur 2.74 times longer than patella; movable chelal finger 1.49–1.52 times longer than hand and 0.59–0.60 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs 4E, 5E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb situated midway between b and st; b and t situated subdistally, t situated distal to it; est situated distal to b (Fig. 5F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 29–30 macrodenticles, slightly retrorse and pointed; movable chelal finger with 27–29 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed (Fig. 5F). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 4B, 5G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 4: 4: 4: 4: 4: 4–5: 4: 4–5: 4–5: 4: T2T: 0, tergites VIII–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 13–14: 12–14: 9: 9: 8–9: 9: 9: 9: 0: 2. Anterior genital operculum with ten setae, genital opening slit-like, with 11–13 marginal setae on each side, 34 in total (Fig. 4F).</p><p>Legs (Fig. 5H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of trochanter IV, femur Ⅰ, IV and patella IV. Femur of leg Ⅰ 1.79–1.86 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.38–2.47 times longer than tibia. Femoropatella of leg IV 4.33–4.38 times longer than deep; tibia 6.55–6.64 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.78–4.00 times longer than deep (TS = 0.32–0.34), tarsus 14.50–14.83 times longer than deep and 2.62–2.72 times longer than metatarsus (TS = 0.27–0.33). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 3B, 4G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, Ⅰ 3, II 3–4, III 5, IV 5; tergal chaetotaxy Ⅰ–XII: 4: 4: 4: 4: 4: 4: 4: 4: 4: 4: T2T: 0; sternal chaetotaxy IV–XII: 12–13: 10: 9: 9: 9: 9: 9: 0: 2; anterior genital operculum with ten setae, posterior margin with 12–14 marginal setae, 22–24 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 4.00–4.63 times longer than deep (TS = 0.32– 0.34), tarsus 14.17–16.17 times longer than deep and 2.62–2.66 times longer than metatarsus (TS = 0.22–0.28).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.02–2.10. Pedipalps: trochanter 0.24/0.19–0.20 (1.20–1.26), femur 1.29/0.16 (8.06), patella 0.47/0.20 (2.35), chela 1.84–1.85/0.26–0.27 (6.85–7.08), hand 0.73/0.26–0.27 (2.70–2.81), movable chelal finger length 1.09–1.11. Chelicera 0.72–0.74/0.30 (2.40–2.47), movable finger length 0.39–0.40. Carapace 0.60/0.60 (1.00). Leg Ⅰ: trochanter 0.19/0.16 (1.19), femur 0.69–0.70/0.09 (7.67–7.78), patella 0.37–0.39/0.08 (4.63–4.88), tibia 0.32/0.06 (5.33), tarsus 0.76–0.79/0.05–0.06 (13.17–15.20). Leg IV: trochanter 0.27–0.29/0.16 (1.69–1.81), femoropatella 1.04–1.05/0.24 (4.33–4.38), tibia 0.72–0.73/0.11 (6.55–6.64), metatarsus 0.32–0.34/0.08–0.09 (3.78–4.00), tarsus 0.87–0.89/0.06 (14.50–14.83).</p><p>Females: body length 2.19–2.27. Pedipalps: trochanter 0.24–0.27/0.20–0.21 (1.20–1.29), femur 1.33– 1.42/0.17–0.18 (7.82–7.89), patella 0.46–0.51/0.20–0.22 (2.30–2.32), chela 1.89–2.08/0.26–0.28 (7.27–7.43), hand 0.75–0.82/0.26–0.28 (2.88–2.93), movable chelal finger length 1.13–1.24. Chelicera 0.76–0.83/0.33–0.34 (2.30– 2.44), movable finger length 0.41–0.45. Carapace 0.64/0.64–0.72 (0.89–1.00). Leg Ⅰ: trochanter 0.18–0.19/0.16 (1.13–1.19), femur 0.69–0.80/0.09 (7.67–8.89), patella 0.39–0.43/0.08 (4.88–5.38), tibia 0.32–0.36/0.06–0.07 (5.14–5.33), tarsus 0.79–0.86/0.06 (13.17–14.33). Leg IV: trochanter 0.28–0.30/0.16 (1.75–1.88), femoropatella 1.03–1.11/0.23–0.24 (4.48–4.63), tibia 0.69–0.75/0.11–0.12 (6.25–6.27), metatarsus 0.32–0.37/0.08 (4.00–4.63), tarsus 0.85–0.97/0.06 (14.17–16.17).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/03DA1A1EFF9FFFAEFF053F79E3C23D46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFF90FFABFF053E97E3C23BF6.text	03DA1A1EFF90FFABFF053E97E3C23BF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius guanniuensis Hou & Feng & Zhang 2023	<div><p>Lagynochthonius guanniuensis sp. nov. (ΪϮṄṁae)</p><p>Figs 6–9</p><p>Type material. Holotype ♀ (Ps.-MHBU-GZXF-20-02-01): China, Guizhou Province, Xifeng County, Wenquan Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.871666&amp;materialsCitation.latitude=27.223333" title="Search Plazi for locations around (long 106.871666/lat 27.223333)">Wenquan Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.871666&amp;materialsCitation.latitude=27.223333" title="Search Plazi for locations around (long 106.871666/lat 27.223333)">Guanniu Cave</a>, under the detritus in the deep zone [27°13'24.00″N, 106°52'18.00″E], 853 m a.s.l., 15 August 2020, Zegang Feng, Hongru Xu &amp; Yanmeng Hou leg. Paratype: 1 ♀ (Ps.-MSWU-GZXF-20-02- 02), with the same data as the holotype .</p><p>Etymology. Named after the type locality, Guanniu Cave.</p><p>Diagnosis (♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed and small, triangular; posterior margin of carapace with two setae; tergites Ⅰ–IV each with two setae. Pedipalps slender, femur 8.19–8.47 times longer than broad; chela 6.86–7.03 times longer than broad; chelal fingers with intercalary teeth; chelal fingers markedly curved in dorsal view; chemosensory setae (sc) present on dorsum of chelal hand.</p><p>Description. Females (holotype and paratype), male unknown (Figs 6D, 7–9).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 8C, 9A): carapace 0.95–0.97 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 9C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 8D–E, 9B): large, about as long as carapace, 2.48–2.50 times longer than broad; five setae (right chelicera with six setae) and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 11–12 teeth, distal one largest; movable finger with 15–17 retrorse contiguous small teeth; galea absent (Fig. 9B). Serrula exterior with 21–23 blades and serrula interior with 13–14 blades. Rallum with eight blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 9D).</p><p>Pedipalp (Figs8A–B, G, 9E–G):long and slender, trochanter1.23–1.33, femur 8.19–8.47,patella 2.39–2.52, chela 6.86–7.03, hand 2.57–2.62 times longer than broad; femur 2.47–2.62 times longer than patella; movable chelal finger 1.65–1.67 times longer than hand and 0.62 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs 8G, 9E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming an oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb situated midway between b and st; b and t situated subdistally, t situated at the same level as it and distal to b; est situated distal to b (Fig. 9F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 33 macrodenticles, slightly retrorse and pointed, plus 25– 27 intercalary microdenticles, 58–60 in total; movable chelal finger with 29 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 25 intercalary microdenticles and 6–7 vestigial, rounded and contiguous basal teeth, 60–61 in total (Fig. 9F). Chelal fingers markedly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 8B, 9G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 2: 2: 2: 2: 4: 4: 4: 4: 5: 2: T2T: 0, tergite IX with an unpaired median seta. Sternal chaetotaxy IV–XII: 11–14: 8–9: 7: 7: 7–8: 7: 7: 0: 2. Anterior genital operculum with 10–11 setae, posterior margin with 12–13 marginal setae, 23 in total (Fig. 8F).</p><p>Legs (Fig. 9H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur Ⅰ, IV and patella IV. Femur of leg Ⅰ 1.91–1.95 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.50–2.54 times longer than tibia. Femoropatella of leg IV 4.32–4.76 times longer than deep; tibia 6.18–7.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.22–4.63 times longer than deep (TS = 0.26–0.27), tarsus 14.57–15.67 times longer than deep and 2.54–2.68 times longer than metatarsus (TS = 0.30–0.31). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Females: body length 2.20–2.55. Pedipalps: trochanter 0.27–0.28/0.21–0.22 (1.23–1.33), femur 1.31–1.44/0.16–0.17 (8.19– 8.47), patella 0.53–0.55/0.21–0.23 (2.39–2.52), chela 1.92–2.04/0.28–0.29 (6.86–7.03), hand 0.72–0.76/0.28–0.29 (2.57–2.62), movable chelal finger length 1.19–1.27. Chelicera 0.75–0.77/0.30–0.31 (2.48–2.50), movable finger length 0.40. Carapace 0.63–0.67/0.66–0.69 (0.95–0.97). Leg Ⅰ: trochanter 0.19–0.22/0.17 (1.12–1.29), femur 0.78–0.86/0.09–0.10 (8.60–8.67), patella 0.40–0.45/0.08–0.09 (5.00), tibia 0.32–0.35/0.06–0.07 (5.00–5.33), tarsus 0.80–0.89/0.06–0.07 (12.17–13.33). Leg IV: trochanter 0.30–0.31/0.17 (1.72–1.82), femoropatella 1.08–1.19/0.25 (4.32–4.76), tibia 0.68–0.77/0.11 (6.18–7.00), metatarsus 0.37–0.38/0.08–0.09 (4.22–4.63), tarsus 0.94–1.02/0.06– 0.07 (14.57–15.67).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/03DA1A1EFF90FFABFF053E97E3C23BF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFF95FFB0FF05394AE2953C0A.text	03DA1A1EFF95FFB0FF05394AE2953C0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius latipectus Hou & Feng & Zhang 2023	<div><p>Lagynochthonius latipectus sp. nov. (ȐṞṄṁae)</p><p>Figs 10–14</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-525 - 01): China, Guizhou Province, Jiangkou County, Taiping Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.81547&amp;materialsCitation.latitude=27.748589" title="Search Plazi for locations around (long 108.81547/lat 27.748589)">Yunshe Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.81547&amp;materialsCitation.latitude=27.748589" title="Search Plazi for locations around (long 108.81547/lat 27.748589)">Xianren Cave</a>, under stones and detritus in deep zone [27°44′54.92″N, 108°48’55.66″E], 517 m a.s.l., 15August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratypes: 5 ♂ (Ps.- MHBU-HBUARA#2022-525 - 02 –HBUARA#2022-525-06), 10 ♀ (Ps.- MSWU-HBUARA#2022-525 - 07 –HBUARA#2022-525-16), all with the same data as the holotype; 3 ♀ (Ps.- MSWU-HBUARA#2022-306 - 01 –HBUARA#2022-306-03), Jiangkou County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.85925&amp;materialsCitation.latitude=27.827532" title="Search Plazi for locations around (long 108.85925/lat 27.827532)">Nuxi Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.85925&amp;materialsCitation.latitude=27.827532" title="Search Plazi for locations around (long 108.85925/lat 27.827532)">Mengjiatun Village</a>, Shenxian Cave, [27°49′39.11″N, 108°51′33.31″E], 452 m a.s.l., 4 July 2022, Yanmeng Hou, Nana Zhan, Long Lin &amp; Jianzhou Sun leg.</p><p>Etymology. The specific name is derived from a combination of the Latin words “ latus ” and “ pectus ”, and refers to its broad cephalothorax.</p><p>Diagnosis (♂ ♀). Moderately sized troglomorphic species with elongated appendages; carapace with a pair of anterior eyespots only; anterior margin of carapace thin, finely denticulated, epistome obtuse and small, round; posterior margin of carapace with two setae; tergites Ⅰ–II each with four setae. Pedipalps slender, femur 6.92–7.23 (♂), 6.80–7.31 (♀) times longer than broad; chela 7.33–7.67 (♂), 6.71–7.50 (♀) times longer than broad; chelal fingers without intercalary teeth but fixed chelal finger with a modified accessory tooth (td) on retrolateral face; chemosensory setae (sc) present on dorsum of chelal hand.</p><p>Description. Males (holotype and paratypes) (Figs 12A, 13A–F, 14).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 13C, E, 14A): carapace 0.96–1.02 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; anterior eyes reduced into eyespots, posterior eyes reduced completely; anterior margin slightly serrate; epistome obtuse and small, round, with two setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 4–5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with 9–11 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 14C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 13D, 14B): large, about as long as carapace, 2.26–2.30 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 14–16 teeth, distal one largest; movable finger with 13 retrorse contiguous small teeth; galea absent (Fig. 14B). Serrula exterior with 20–22 blades and serrula interior with 12–13 blades. Rallum with eight blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 14D).</p><p>Pedipalp (Figs 13A–B, H, 14E–G): long and slender, trochanter 1.36, femur 6.92–7.23, patella 2.27–2.43, chela 7.33–7.67, hand 2.78–2.83 times longer than broad; femur 2.65–2.76 times longer than patella; movable chelal finger 1.68–1.71 times longer than hand and 0.63–0.64 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs 13H, 14E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to it and b; est slightly distal to b (Fig. 14F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 27–29 macrodenticles, slightly retrorse and pointed, plus a modified accessory tooth on retrolateral face (td, close to dx), 28–30 in total; movable chelal finger with 13 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 16–17 vestigial, rounded and contiguous basal teeth, 29–30 in total (Fig. 14F). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 13B, 14G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 4: 4: 4: 4: 4: 4: 4: 4–5: 5: 4: T2T: 0, tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 13–14: 12–15: 9: 9: 9: 9: 10–11: 10: 0: 2. Anterior genital operculum with ten setae, genital opening slit-like, with 12–14 marginal setae on each side, 36 in total (Fig. 13F).</p><p>Legs (Fig. 14H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg Ⅰ 1.82–1.85 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.33 times longer than tibia. Femoropatella of leg IV 3.04–3.13 times longer than deep; tibia 5.67–5.89 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.43–3.57 times longer than deep (TS = 0.32–0.33), tarsus 12.00–12.20 times longer than deep and 2.44–2.50 times longer than metatarsus (TS = 0.25–0.34). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 11D, 12B, 13G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5; tergal chaetotaxy Ⅰ–XII: 4: 4: 3–4: 4: 4: 4–5: 4–5: 5: 5–6: 4: T2T: 0; sternal chaetotaxy IV– XII: 11–13: 9–10: 9: 9: 9–10: 10–12: 9–10: 0: 2; anterior genital operculum with 9–10 setae, posterior margin with 12–13 marginal setae, 21–23 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 2.88–3.57 times longer than deep (TS = 0.30–0.38), tarsus 10.33–11.80 times longer than deep and 2.48–2.81 times longer than metatarsus (TS = 0.27–0.32).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.71–1.88. Pedipalps: trochanter 0.19/0.14 (1.36), femur 0.90–0.94/0.13 (6.92–7.63), patella 0.34/0.14–0.15 (2.27–2.43), chela 1.32–1.38/0.18 (7.33–7.67), hand 0.50–0.51/0.18 (2.78–2.83), movable chelal finger length 0.84– 0.87. Chelicera 0.52–0.53/0.23 (2.26–2.30), movable finger length 0.28–0.29. Carapace 0.50/0.49–0.52 (0.96–1.02). Leg Ⅰ: trochanter 0.15–0.16/0.13 (1.15–1.23), femur 0.50–0.51/0.08 (6.25–6.38), patella 0.27–0.28/0.07 (3.86–4.00), tibia 0.24/0.05–0.06 (4.00–4.80), tarsus 0.56/0.05 (11.20). Leg IV: trochanter 0.23–0.24/0.14–0.15 (1.60–1.64), femoropatella 0.73–0.75/0.24 (3.04–3.13), tibia 0.51–0.53/0.09 (5.67–5.89), metatarsus 0.24–0.25/0.07 (3.43–3.57), tarsus 0.60–0.61/0.05 (12.00–12.20).</p><p>Females: body length 1.84–2.32. Pedipalps: trochanter 0.18–0.22/0.14–0.16 (1.25–1.43), femur 0.95– 1.02/0.13–0.15 (6.80–7.31), patella 0.35–0.40/0.14–0.17 (2.25–2.57), chela 1.35–1.50/0.18–0.22 (6.71–7.50), hand 0.50–0.56/0.18–0.22 (2.55–2.78), movable chelal finger length 0.86–0.94. Chelicera 0.53–0.62/0.24–0.29 (2.14–2.32), movable finger length 0.28–0.33. Carapace 0.50–0.57/0.52–0.61 (0.93–0.98). Leg Ⅰ: trochanter 0.14– 0.16/0.13–0.14 (1.00–1.23), femur 0.51–0.57/0.08–0.09 (6.11–7.13), patella 0.26–0.30/0.07–0.08 (3.63–3.75), tibia 0.23–0.28/0.06 (3.83–4.67), tarsus 0.55–0.65/0.05–0.06 (10.00–11.40). Leg IV: trochanter 0.22–0.25/0.13–0.16 (1.50–1.69), femoropatella 0.73–0.79/0.21–0.26 (3.04–3.48), tibia 0.51–0.56/0.09–0.11 (5.09–5.89), metatarsus 0.21–0.26/0.07–0.08 (2.88–3.57), tarsus 0.59–0.66/0.05–0.06 (10.33–11.80).</p><p>Distribution. Known only from the Xianren and Shenxian caves.</p></div>	https://treatment.plazi.org/id/03DA1A1EFF95FFB0FF05394AE2953C0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFF8EFFBDFF053FDBE3C23BCA.text	03DA1A1EFF8EFFBDFF053FDBE3C23BCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius longedentatus Hou & Feng & Zhang 2023	<div><p>Lagynochthonius longedentatus sp. nov. (KfflṄṁae)</p><p>Figs 15–18</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2021-433 - 01): China, Guizhou Province, Ceheng County, Rongdu Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.717766&amp;materialsCitation.latitude=25.07842" title="Search Plazi for locations around (long 105.717766/lat 25.07842)">Hongbaituo Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.717766&amp;materialsCitation.latitude=25.07842" title="Search Plazi for locations around (long 105.717766/lat 25.07842)">Hongbaituo Cave</a>, under stones in the deep zone [25°4′42.31″N, 105°43′3.96″E], 1033 m a.s.l., 10 October 2021, Zegang Feng, Yanmeng Hou, Lu Zhang &amp; Liu Fu leg. Paratype: 1 ♀ (Ps.- MSWU-HBUARA#2021-433 - 02), with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin words “ longus ” and “ dentatus ”, meaning elongate and toothed, respectively, and referring to the elongated intercalary teeth on the fixed chelal finger.</p><p>Diagnosis (♂ ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with two setae; tergites III–VII each with four setae. Pedipalps slender, femur 7.15 (♂), 7.42 (♀) times longer than broad; chela robust, 6.65 (♂), 6.63 (♀) times longer than broad; chelal fingers with intercalary teeth; fixed chelal finger with a row of macrodenticles that are distinctly larger than movable chelal finger teeth, pointed and slightly retrorse, fixed chelal finger intercalary teeth almost as large as movable finger teeth; chemosensory setae (sc) present on dorsum of chelal hand.</p><p>Description. Male (holotype) (Figs 20A, 17A–F, 18).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 17C, 18A): carapace 1.00 times longer than broad, squarish, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small, pointed, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with seven terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 18C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 17D, 18B): large, about as long as carapace, 2.48 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 17 teeth, distal one largest; movable finger with 17 retrorse contiguous small teeth; galea absent (Fig. 18B). Serrula exterior with 20 blades and serrula interior with 12 blades. Rallum with eight blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 18D).</p><p>Pedipalp (Figs 17A–B, E, 18E–G): long and slender, trochanter 1.07, femur 7.15, patella 2.20, chela 6.65, hand 2.50 times longer than broad; femur 2.82 times longer than patella; movable chelal finger 1.66 times longer than hand and 0.62 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs 17E, 18E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming an oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and it; est situated at same level as b (Fig. 18F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 28 macrodenticles, slightly retrorse and pointed, plus 19 intercalary microdenticles, 47 in total; movable chelal finger with 24 macrodenticles (markedly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 12 intercalary microdenticles (extending roughly to the middle of sb and b) and nine vestigial, rounded and contiguous basal teeth, 45 in total (Fig. 18F). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 17B, 18G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 2: 4: 4: 4: 4: 4: 4: 5: 5: 4: T2T: 0, tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 12: 9: 9: 9: 9: 10: 9: 7: 0: 2. Anterior genital operculum with 11 setae, genital opening slit-like, with 13–15 marginal setae on each side, 39 in total (Fig. 17F).</p><p>Legs (Fig. 18H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg Ⅰ 1.77 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.46 times longer than tibia. Femoropatella of leg IV 3.48 times longer than deep; tibia 5.67 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.71 times longer than deep (TS = 0.35), tarsus 14.00 times longer than deep and 2.69 times longer than metatarsus (TS = 0.23). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult female (paratype) (Figs 15D, 16B, 17G). Mostly same as male; with same chaetotaxy of coxae as male; tergal chaetotaxy Ⅰ–XII: 4: 3: 4: 4: 4: 4: 4: 4: 5: 4: T2T: 0; sternal chaetotaxy IV–XII: 11: 9: 8: 9: 9: 9: 8: 0: 2; anterior genital operculum with ten setae, posterior margin with 12 marginal setae, 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.43 times longer than deep (TS = 0.38), tarsus 12.60 times longer than deep and 2.63 times longer than metatarsus (TS = 0.29).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Male: body length 1.42. Pedipalps: trochanter 0.16/0.15 (1.07), femur 0.93/0.13 (7.15), patella 0.33/0.15 (2.20), chela 1.33/0.20 (6.65), hand 0.50/0.20 (2.50), movable chelal finger length 0.83. Chelicera 0.57/0.23 (2.48), movable finger length 0.30. Carapace 0.49/0.49 (1.00). Leg Ⅰ: trochanter 0.15/0.14 (1.07), femur 0.53/0.07 (7.57), patella 0.30/0.06 (5.00), tibia 0.24/0.05 (4.80), tarsus 0.59/0.05 (11.80). Leg IV: trochanter 0.22/0.13 (1.69), femoropatella 0.80/0.23 (3.48), tibia 0.51/0.09 (5.67), metatarsus 0.26/0.07 (3.71), tarsus 0.70/0.05 (14.00).</p><p>Female: body length 1.59. Pedipalps: trochanter 0.18/0.14 (1.29), femur 0.89/0.12 (7.42), patella 0.30/0.15 (2.00), chela 1.26/0.19 (6.63), hand 0.49/0.19 (2.58), movable chelal finger length 0.76. Chelicera 0.55/0.22 (2.50), movable finger length 0.29. Carapace 0.47/0.48 (0.98). Leg Ⅰ: trochanter 0.14/0.12 (1.17), femur 0.52/0.07 (7.43), patella 0.28/0.06 (4.67), tibia 0.23/0.05 (4.60), tarsus 0.55/0.05 (11.00). Leg IV: trochanter 0.22/0.14 (1.57), femoropatella 0.71/0.20 (3.55), tibia 0.46/0.08 (5.75), metatarsus 0.24/0.07 (3.43), tarsus 0.63/0.05 (12.60).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/03DA1A1EFF8EFFBDFF053FDBE3C23BCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFF83FFBAFF05391BE3C2385E.text	03DA1A1EFF83FFBAFF05391BE3C2385E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius longyanensis Hou & Feng & Zhang 2023	<div><p>Lagynochthonius longyanensis sp. nov. (ỂẆṄṁae)</p><p>Figs 19–22</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2021-153 - 01): China, Guizhou Province, Bozhou District, Fengxiang Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.501915&amp;materialsCitation.latitude=27.630383" title="Search Plazi for locations around (long 106.501915/lat 27.630383)">Qingkeng Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.501915&amp;materialsCitation.latitude=27.630383" title="Search Plazi for locations around (long 106.501915/lat 27.630383)">Longyan Cave</a>, under stones in the deep zone [27°37′49.38″N, 106°30′6.90″E], 1082 m a.s.l., 6 July 2021, Zegang Feng, Hongru Xu, Liu Fu &amp; Nana Zhan leg. Paratypes: 2 ♂ (Ps.- MSWU-HBUARA#2021-153 - 02 &amp; HBUARA#2021-153-03), 1 ♀ (Ps.- MSWU-HBUARA#2021-153 - 04), all with the same data as the holotype .</p><p>Etymology. Named after the type locality, Longyan Cave.</p><p>Diagnosis (♂ ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome obtuse and small, round; posterior margin of carapace with two setae; tergites Ⅰ–Ⅴ each with four setae. Pedipalps slender, femur 6.79–7.50 (♂), 7.07 (♀) times longer than broad; chela 6.63–6.95 (♂), 5.75 (♀) times longer than broad; chelal fingers without intercalary teeth but fixed chelal finger with a modified accessory tooth (td) on retrolateral face; chemosensory setae (sc) present on dorsum of chelal hand.</p><p>Description. Males (holotype and paratypes) (Figs 19D, 20A, 21A–F, 22).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 21C, 22A): carapace 0.94–1.04 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome obtuse and small, round; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with 7–9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 22C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 21D, 22B): large, about as long as carapace, 2.19–2.35 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 8–10 teeth, distal one largest; movable finger with 13–14 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 22B). Serrula exterior with 19–20 blades and serrula interior with 12–13 blades. Rallum with eight blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 22D).</p><p>Pedipalp (Figs 21A–B, E, 22E–G): long and slender, trochanter 1.29–1.44, femur 6.79–7.50, patella 2.21–2.50, chela 6.63–6.95, hand 2.79–2.84 times longer than broad; femur 2.71–2.90 times longer than patella; movable chelal finger 1.43–1.48 times longer than hand and 0.60–0.61 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs 21E, 22E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb situated midway between b and st; b and t situated subdistally, t situated distal to it and b; est situated at same level as b (Fig. 22F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 24–25 macrodenticles, slightly retrorse and pointed, plus a modified accessory tooth on retrolateral face (td, close to dx), 25–26 in total; movable chelal finger with ten macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 16–17 vestigial, rounded and contiguous basal teeth, 26–27 in total (Fig. 22F). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 21B, 22G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 4: 4: 4: 4: 4: 4: 4–5: 5: 5: 4: T2T: 0, tergites VII–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 14–16: 12–13: 10: 8–9: 9: 9: 9: 9: 0: 2. Anterior genital operculum with 10–11 setae, genital opening slit-like, with 12–15 marginal setae on each side, 35–39 in total (Fig. 21F).</p><p>Legs (Fig. 22H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg Ⅰ 1.96 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.24–2.25 times longer than tibia. Femoropatella of leg IV 3.13–3.33 times longer than deep; tibia 5.22–5.67 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.29–3.57 times longer than deep (TS = 0.39–0.40), tarsus 11.20–12.60 times longer than deep and 2.43–2.52 times longer than metatarsus (TS = 0.29). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult female (paratype) (Figs 20B, 21G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, Ⅰ 3, II 3–4, III 5, IV 5; tergal chaetotaxy Ⅰ–XII: 4: 4: 4: 4: 4: 5: 5: 5: 5: 4: T2T: 0; sternal chaetotaxy IV–XII: 12: 10: 9: 9: 9: 10: 9: 0: 2; anterior genital operculum with ten setae, posterior margin with 12 marginal setae, 22 in total; tarsal and tibial segments of leg Ⅰ missing; leg IV with a long tactile seta on both tarsal segments: metatarsus 4.50 times longer than deep (TS = 0.41), tarsus 12.80 times longer than deep and 2.37 times longer than metatarsus (TS = 0.31).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.76–1.95. Pedipalps: trochanter 0.18–0.23/0.14–0.16 (1.29–1.44), femur 0.90–0.95/0.12–0.14 (6.79–7.50), patella 0.31–0.35/0.14 (2.21–2.50), chela 1.26–1.32/0.19 (6.63–6.95), hand 0.53–0.54/0.19 (2.79–2.84), movable chelal finger length 0.76–0.80. Chelicera 0.46–0.47/0.20–0.21 (2.19–2.35), movable finger length 0.25. Carapace 0.47–0.49/0.47–0.50 (0.94–1.04). Leg Ⅰ: trochanter 0.16–0.17/0.12–0.13 (1.31–1.33), femur 0.51–0.53/0.07 (7.29– 7.57), patella 0.26–0.27/0.07 (3.71–3.86), tibia 0.24–0.25/0.05–0.06 (4.17–4.80), tarsus 0.54–0.56/0.05 (10.80– 11.20). Leg IV: trochanter 0.22–0.23/0.13–0.14 (1.57–1.77), femoropatella 0.70–0.75/0.21–0.24 (3.13–3.33), tibia 0.47–0.51/0.09 (5.22–5.67), metatarsus 0.23–0.25/0.07 (3.29–3.57), tarsus 0.56–0.63/0.05 (11.20–12.60).</p><p>Female: body length 1.93. Pedipalps: trochanter 0.20/0.15 (1.33), femur 0.99/0.14 (7.07), patella 0.36/0.15 (2.40), chela 1.38/0.24 (5.75), hand 0.59/0.24 (2.46), movable chelal finger length 0.80. Chelicera 0.53/0.23 (2.30), movable finger length 0.29. Carapace 0.51/0.51 (1.00). Leg Ⅰ: trochanter 0.17/0.14 (1.21), femur 0.59/0.08 (7.38), patella 0.28/0.07 (4.00). Leg IV: trochanter 0.20/0.12 (1.67), femoropatella 0.79/0.20 (3.95), tibia 0.54/0.09 (6.00), metatarsus 0.27/0.06 (4.50), tarsus 0.64/0.05 (12.80).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/03DA1A1EFF83FFBAFF05391BE3C2385E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFF84FF86FF053BEFE3C23CA6.text	03DA1A1EFF84FF86FF053BEFE3C23CA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius maanensis Hou & Feng & Zhang 2023	<div><p>Lagynochthonius maanensis sp. nov. (ḢḊṄṁae)</p><p>Figs 23–26</p><p>Type material. Holotype ♀ (Ps.-MHBU-GZZA-20-18-01): China, Guizhou Province, Zheng’an County, Hexi Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.44218&amp;materialsCitation.latitude=28.410517" title="Search Plazi for locations around (long 107.44218/lat 28.410517)">Maan Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.44218&amp;materialsCitation.latitude=28.410517" title="Search Plazi for locations around (long 107.44218/lat 28.410517)">Pianxiao Cave</a>, under stones in the deep zone [28°24′37.86″N, 107°26′31.84″E], 932 m a.s.l., 25 August 2020, Zegang Feng, Hongru Xu &amp; Yanmeng Hou leg. Paratype: 1 ♀ (Ps.-MSWU-GZZA-20-18-02), with the same data as the holotype .</p><p>Etymology. Named after the village of Maan, near the type locality.</p><p>Diagnosis (♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome obtuse and small, round; posterior margin of carapace with two setae; tergites Ⅰ–II each with two setae, tergites III–Ⅴ each with four setae. Pedipalps slender, femur 5.43–5.75 times longer than broad; chela 5.84–5.88 times longer than broad; chelal fingers without intercalary teeth but fixed chelal finger with a modified accessory tooth (td) on retrolateral face; chemosensory setae (sc) present on dorsum of chelal hand.</p><p>Description. Females (holotype and paratype), male unknown (Figs 23E, 24–26).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 25C, 26A): carapace 0.93–0.96 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome obtuse and small, round, with two setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 26C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 25D, 26B): large, about as long as carapace, 2.14–2.17 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 11–13 teeth, distal one largest; movable finger with 11–12 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 26B). Serrula exterior with 17–18 blades and serrula interior with 13–14 blades. Rallum with eight blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 26D).</p><p>Pedipalp (Figs 25A–B, F, 26E–G): long and slender, trochanter 1.18–1.38, femur 5.43–5.75, patella 2.00–2.08, chela 5.84–5.88, hand 2.29–2.32 times longer than broad; femur 2.71–2.76 times longer than patella; movable chelal finger 1.55–1.56 times longer than hand and 0.61 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs 25F, 26E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming an oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b; it and est situated between t and b (Fig. 26F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 25 macrodenticles, slightly retrorse and pointed, plus a modified accessory tooth on retrolateral face (td, close to dx), 26 in total; movable chelal finger with 11 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 14–15 vestigial, rounded and contiguous basal teeth, 25–26 in total (Fig. 26F). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 25B, 26G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 2: 2: 4: 4: 4: 4–5: 5: 5–6: 5–6: 4: T2T: 0. Sternal chaetotaxy IV–XII: 13–14: 9–10: 8: 9–10: 9: 10–12: 10–11: 0: 2. Anterior genital operculum with ten setae, posterior margin with 14–15 marginal setae, 24–25 in total (Fig. 25E).</p><p>Legs (Fig. 26H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg Ⅰ 2.00–2.06 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.35–2.37 times longer than tibia. Femoropatella of leg IV 2.80–2.94 times longer than deep; tibia 4.63–5.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.00 times longer than deep (TS = 0.33–0.39), tarsus 10.00–11.50 times longer than deep and 2.22–2.56 times longer than metatarsus (TS = 0.30–0.35). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Females: body length 1.64–1.83. Pedipalps: trochanter 0.13–0.18/0.11–0.13 (1.18–1.38), femur 0.69–0.76/0.12–0.14 (5.43– 5.75), patella 0.25–0.28/0.12–0.14 (2.00–2.08), chela 1.00–1.11/0.17–0.19 (5.84–5.88), hand 0.39–0.44/0.17–0.19 (2.29–2.32), movable chelal finger length 0.61–0.68. Chelicera 0.45–0.50/0.21–0.23 (2.14–2.17), movable finger length 0.24–0.26. Carapace 0.40–0.47/0.43–0.49 (0.93–0.96). Leg Ⅰ: trochanter 0.11/0.10–0.11 (1.00–1.10), femur 0.37–0.40/0.06–0.07 (5.71–6.17), patella 0.18–0.20/0.06 (3.00–3.33), tibia 0.17–0.19/0.04 (4.25–4.75), tarsus 0.40–0.45/0.04 (10.00–11.25). Leg IV: trochanter 0.17–0.19/0.11–0.12 (1.56), femoropatella 0.50–0.56/0.17–0.20 (2.80–2.94), tibia 0.37–0.40/0.08 (4.63–5.00), metatarsus 0.18/0.06 (3.00), tarsus 0.40–0.46/0.04 (10.00–11.50).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/03DA1A1EFF84FF86FF053BEFE3C23CA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFFB8FF83FF053FB7E3C23D9A.text	03DA1A1EFFB8FF83FF053FB7E3C23D9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius mawangensis Hou & Feng & Zhang 2023	<div><p>Lagynochthonius mawangensis sp. nov. (¤王Ṅṁae)</p><p>Figs 27–30</p><p>Type material. Holotype ♂ (Ps.-MHBU-GZWC-20-16-01): China, Guizhou Province, Wuchuan County, Duru sub-district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.88268&amp;materialsCitation.latitude=28.529003" title="Search Plazi for locations around (long 107.88268/lat 28.529003)">Mawang Cave</a>, under stones in the deep zone [28°31′44.41″N, 107°52′57.64″E], 706 m a.s.l., 24 August 2020, Zegang Feng, Hongru Xu &amp; Yanmeng Hou leg. Paratypes: 8 ♂ (Ps.-MHBU-GZWC-20-16-02–GZWC-20- 16-09), 2 ♀ (Ps.-MSWU-GZWC-20-16-10 &amp; GZWC-20-16-11), all with the same data as the holotype .</p><p>Etymology. Named after the type locality, Mawang Cave.</p><p>Diagnosis (♂ ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome obtuse and small, round; posterior margin of carapace with two setae; tergites Ⅰ–Ⅴ each with four setae. Pedipalps slender, femur 7.18–7.40 (♂), 6.67–7.27 (♀) times longer than broad; chela 6.87–6.94 (♂), 6.22–6.33 (♀) times longer than broad; chelal fingers without intercalary teeth and fixed chelal finger with a modified accessory tooth (td) on retrolateral face; chemosensory setae (sc) present on dorsum of chelal hand.</p><p>Description. Males (holotype and paratypes) (Figs 27D, 28A, 29A–F, 30).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 29C, 30A): carapace 1.07–1.08 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome obtuse and small, round, with two setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 4–5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with nine terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 30C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 29D, 30B): large, about as long as carapace, 2.17–2.24 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 10–11 teeth, distal one largest; movable finger with 9–11 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 30B). Serrula exterior with 19–20 blades and serrula interior with 11–12 blades. Rallum with eight blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 30D).</p><p>Pedipalp (Figs 29A–B, E, 30E–G): long and slender, trochanter 1.18–1.33, femur 7.18–7.40, patella 2.25–2.36, chela 6.87–6.94, hand 2.80–2.81 times longer than broad; femur 2.85–2.93 times longer than patella; movable chelal finger 1.49–1.50 times longer than hand and 0.60–0.61 times longer than chela. Setae generally long and acuminate; one distal lyrifissures present on patella (Figs 29E, 30E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b; it and est situated between t and b (Fig. 30F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 25–27 macrodenticles, slightly retrorse and pointed, plus a modified accessory tooth on retrolateral face (td, close to dx), 26–28 in total; movable chelal finger with ten macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 14–15 vestigial, rounded and contiguous basal teeth, 24–25 in total (Fig. 30F). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 29B, 30G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 4: 4: 4: 4: 4: 4: 5: 5: 5: 4–5: T2T: 0, tergites VII–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 13–15: 14: 9–10: 10: 9–10: 11: 10–12: 8–10: 0: 2. Anterior genital operculum with 12 setae, genital opening slit-like, with 11–14 marginal setae on each side, 35–39 in total (Fig. 29F).</p><p>Legs (Fig. 30H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg Ⅰ 2.00–2.16 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.45–2.61 times longer than tibia. Femoropatella of leg IV 2.90–3.05 times longer than deep; tibia 5.38–6.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.00–3.33 times longer than deep (TS = 0.40–0.44), tarsus 10.40–12.00 times longer than deep and 2.60–2.67 times longer than metatarsus (TS = 0.29–0.31). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 28B, 29G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5–6; tergal chaetotaxy Ⅰ–XII: 4: 4: 4: 4: 4: 4–5: 5: 5: 5: 4–5: T2 T: 0; sternal chaetotaxy IV–XII: 12–13: 9–11: 10: 9: 9–10: 11: 8–10: 0: 2; anterior genital operculum with ten setae, posterior margin with 11–13 marginal setae, 21–23 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.50 times longer than deep (TS = 0.33–0.38), tarsus 12.25–12.75 times longer than deep and 2.33–2.43 times longer than metatarsus (TS = 0.29–0.31) .</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.59–1.66. Pedipalps: trochanter 0.13–0.16/0.11–0.12 (1.18–1.33), femur 0.74–0.79/0.10–0.11 (7.18–7.40), patella 0.26–0.27/0.11–0.12 (2.25–2.36), chela 1.03–1.11/0.15–0.16 (6.87–6.94), hand 0.42–0.45/0.15–0.16 (2.80– 2.81), movable chelal finger length 0.63–0.67. Chelicera 0.38–0.39/0.17–0.18 (2.17–2.24), movable finger length 0.20. Carapace 0.42–0.44/0.39–0.41 (1.07–1.08). Leg Ⅰ: trochanter 0.10–0.12/0.09–0.11 (1.09–1.11), femur 0.41– 0.44/0.06–0.07 (6.29–6.83), patella 0.19–0.22/0.06 (3.17–3.67), tibia 0.18–0.20/0.04–0.05 (4.00–4.50), tarsus 0.47– 0.49/0.04–0.05 (9.80–11.75). Leg IV: trochanter 0.18–0.19/0.13 (1.38–1.46), femoropatella 0.58–0.61/0.19–0.21 (2.90–3.05), tibia 0.42–0.43/0.07–0.08 (5.38–6.00), metatarsus 0.18–0.20/0.06 (3.00–3.33), tarsus 0.48–0.52/0.04– 0.05 (10.40–12.00).</p><p>Females: body length 1.64–1.73. Pedipalps: trochanter 0.15–0.16/0.12 (1.25–1.33), femur 0.80/0.11–0.12 (6.67– 7.27), patella 0.28–0.29/0.12–0.13 (2.23–2.33), chela 1.12–1.14/0.18 (6.22–6.33), hand 0.45/0.18 (2.50), movable chelal finger length 0.69. Chelicera 0.41–0.42/0.19–0.20 (2.05–2.21), movable finger length 0.21–0.22. Carapace 0.42–0.44/0.43–0.47 (0.94–0.98). Leg Ⅰ: trochanter 0.11–0.13/0.11–0.12 (0.92–1.18), femur 0.42–0.43/0.07 (6.00– 6.14), patella 0.21–0.22/0.06 (3.50–3.67), tibia 0.20/0.05 (4.00), tarsus 0.48–0.49/0.05 (9.60–10.80). Leg IV: trochanter 0.19–0.20/0.11–0.13 (1.54–1.73), femoropatella 0.59–0.61/0.18–0.19 (3.21–3.28), tibia 0.40–0.43/0.08 (5.00–5.38), metatarsus 0.21/0.06 (3.50), tarsus 0.49–0.51/0.04 (12.25–12.75).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/03DA1A1EFFB8FF83FF053FB7E3C23D9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFFBDFF89FF053EABE29538F6.text	03DA1A1EFFBDFF89FF053EABE29538F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius nigriculus Hou & Feng & Zhang 2023	<div><p>Lagynochthonius nigriculus sp. nov. (ḆṘṄṁae)</p><p>Figs 31–35</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2021-423 - 01): China, Guizhou Province, Weining County, Yangjie Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.33991&amp;materialsCitation.latitude=26.989408" title="Search Plazi for locations around (long 104.33991/lat 26.989408)">Songshan Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.33991&amp;materialsCitation.latitude=26.989408" title="Search Plazi for locations around (long 104.33991/lat 26.989408)">Yuqiyan Cave</a>, under stones in the deep zone [26°59′21.87″N, 104°20′23.68″E], 2143 m a.s.l., 4 October 2021, Zegang Feng, Yanmeng Hou, Lu Zhang &amp; Liu Fu leg. Paratypes: 4 ♀ (Ps.- MHBU-HBUARA#2021-423 - 02 –HBUARA#2021-423-05), all with the same data as the holotype; 6 ♀ (Ps.- MSWU-HBUARA#2021-425 - 01 –HBUARA#2021-425-06), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.3426&amp;materialsCitation.latitude=26.987408" title="Search Plazi for locations around (long 104.3426/lat 26.987408)">Songshan Village</a>, unnamed cave, [26°59′14.67″N, 104°20′33.35″E], 2165 m a.s.l., with the same collection date and collectors as the holotype; 1 ♂ (Ps.- MSWU-HBUARA#2022-495 - 01) , 4 ♀ (Ps.- MSWU-HBUARA#2022-495 - 02 –HBUARA#2022-495-05), Songshan Village, unnamed cave, 31 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg.</p><p>Etymology. The specific name is derived from the Latin adjective “ nigriculus ”, meaning blackish, which refers to the black tergites.</p><p>Diagnosis (♂ ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with two setae; tergites Ⅰ–II each with two setae. Pedipalps slender, femur 8.93–9.00 (♂), 8.24– 8.67 (♀) times longer than broad; chela 8.00–8.14 (♂), 7.25–7.68 (♀) times longer than broad; chelal fingers without intercalary teeth but fixed chelal finger with a modified accessory tooth (td) on retrolateral face; chemosensory setae (sc) present on dorsum of chelal hand.</p><p>Description. Males (holotype and paratype) (Figs 31D, 33A, 34A–F, 35).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 34C, 35A): carapace 0.97–1.05 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with two setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 3–4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with 13–15 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 35C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 34D, 35B): large, about as long as carapace, 2.43–2.48 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 19–20 teeth, distal one largest; movable finger with 21–23 retrorse contiguous small teeth; galea absent (Fig. 35B). Serrula exterior with 23 blades and serrula interior with 12 blades. Rallum with eight blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 35D).</p><p>Pedipalp (Figs 34A–B, E, 35E–G): long and slender, trochanter 1.32–1.44, femur 8.93–9.00, patella 2.56–2.67, chela 8.00–8.14, hand 3.33–3.41 times longer than broad; femur 2.81–3.05 times longer than patella; movable chelal finger 1.37–1.39 times longer than hand and 0.58 times longer than chela. Setae generally long and acuminate; one distal lyrifissures present on patella (Figs 34E, 35E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b; b distal to est; it situated between t and b (Fig. 35F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 29–30 macrodenticles, slightly retrorse and pointed, plus a modified accessory tooth on retrolateral face (td, close to dx), 30–31 in total; movable chelal finger with 20–21 macrodenticles (smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 7–9 vestigial, rounded and contiguous basal teeth, 28–29 in total (Fig. 35F). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 34B, 35G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 2: 2: 2–4: 4: 4: 4–6: 4–5: 5: 5: 2: T2T: 0, tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 13–14: 11–12: 8–9: 7–8: 7–9: 8–9: 7–9: 7–8: 0: 2. Anterior genital operculum with 11 setae, genital opening slit-like, with 12–15 marginal setae on each side, 36–39 in total (Fig. 35F).</p><p>Legs (Fig. 35H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg Ⅰ 1.83–1.92 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.59 times longer than tibia. Femoropatella of leg IV 4.68–4.95 times longer than deep; tibia 6.90–7.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.88–4.14 times longer than deep (TS = 0.26–0.28), tarsus 14.83–15.40 times longer than deep and 2.66–2.87 times longer than metatarsus (TS = 0.34–0.36). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 31E, 33B, 34G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, Ⅰ 3–4, II 4, III 5, IV 5; tergal chaetotaxy Ⅰ–XII: 2: 2: 2–4: 3–4: 4: 4: 4: 5: 5: 2: T2T: 0; sternal chaetotaxy IV–XII: 12–14: 8–10: 8–10: 8–9: 8–9: 7–9: 7–8: 0: 2; anterior genital operculum with 10–12 setae, posterior margin with 13–15 marginal setae, 24–26 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.63–3.88 times longer than deep (TS = 0.23–0.31), tarsus 14.50–17.20 times longer than deep and 2.77–2.97 times longer than metatarsus (TS = 0.28–0.34).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.00. Pedipalps: trochanter 0.23–0.25/0.16–0.19 (1.32–1.44), femur 1.25–1.35/0.14–0.15 (8.93–9.00), patella 0.41–0.48/0.16–0.18 (2.56–2.67), chela 1.79–1.92/0.22–0.24 (8.00–8.14), hand 0.75–0.80/0.22–0.24 (3.33– 3.41), movable chelal finger length 1.03–1.11. Chelicera 0.67–0.73/0.27–0.30 (2.43–2.48), movable finger length 0.37–0.40. Carapace 0.56–0.64/0.58–0.61 (0.97–1.05). Leg Ⅰ: trochanter 0.16–0.20/0.13–0.16 (1.23–1.25), femur 0.69–0.73/0.08–0.09 (8.11–8.63), patella 0.36–0.40/0.07–0.08 (5.00–5.14), tibia 0.29–0.32/0.06–0.07 (4.57–4.83), tarsus 0.75–0.83/0.06 (12.50–13.83). Leg IV: trochanter 0.26–0.29/0.15–0.16 (1.73–1.81), femoropatella 0.94– 1.03/0.19–0.22 (4.68–4.95), tibia 0.63–0.69/0.09–0.10 (6.90–7.00), metatarsus 0.29–0.31/0.07–0.08 (3.88–4.14), tarsus 0.77–0.89/0.05–0.06 (14.83–15.40).</p><p>Females: body length 2.17–2.74. Pedipalps: trochanter 0.23–0.26/0.18–0.21 (1.19–1.37), femur 1.30–1.40/0.15– 0.17 (8.24–8.67), patella 0.44–0.48/0.18–0.20 (2.40–2.67), chela 1.84–2.03/0.24–0.28 (7.25–7.68), hand 0.78– 0.84/0.24–0.28 (3.00–3.25), movable chelal finger length 1.06–1.20. Chelicera 0.73–0.81/0.29–0.33 (2.38–2.52), movable finger length 0.40–0.44. Carapace 0.61–0.72/0.64–0.68 (0.93–1.06). Leg Ⅰ: trochanter 0.18–0.22/0.15– 0.18 (1.19–1.25), femur 0.71–0.78/0.08–0.10 (7.80–9.13), patella 0.37–0.42/0.08–0.10 (4.67–5.29), tibia 0.30– 0.34/0.06–0.07 (4.86–5.33), tarsus 0.77–0.89/0.06–0.07 (12.71–13.33). Leg IV: trochanter 0.27–0.31/0.15–0.17 (1.71–1.94), femoropatella 1.00–1.10/0.20–0.22 (4.59–5.00), tibia 0.65–0.71/0.10–0.11 (6.18–6.50), metatarsus 0.29–0.34/0.08–0.09 (3.63–3.88), tarsus 0.83–0.95/0.05–0.06 (14.50–17.20).</p><p>Distribution. Known only from the Yuqiyan and unnamed caves.</p></div>	https://treatment.plazi.org/id/03DA1A1EFFBDFF89FF053EABE29538F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFFB7FF95FF053847E3C23EAE.text	03DA1A1EFFB7FF95FF053847E3C23EAE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius tuoluoensis Hou & Feng & Zhang 2023	<div><p>Lagynochthonius tuoluoensis sp. nov. (ŔOEṄṁae)</p><p>Figs 36–39</p><p>Type material. Holotype ♂ (Ps.- MHBU-GZC190806-01): China, Guizhou Province, Shuicheng District, Duge Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.69559&amp;materialsCitation.latitude=26.308266" title="Search Plazi for locations around (long 104.69559/lat 26.308266)">Hongguang Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.69559&amp;materialsCitation.latitude=26.308266" title="Search Plazi for locations around (long 104.69559/lat 26.308266)">Tuoluodadong Cave</a>, under stones in the deep zone [26°18′29.76″N, 104°41′44.11″E], 1435 m a.s.l., 6 August 2019, Zegang Feng, Zhaoyi Li &amp; Chen Zhang leg. Paratypes: 1 ♂ (Ps.- MHBU-GZC190806- 02), 6 ♀ (Ps.- MSWU-GZC190806-03 – GZC190806-08), all with the same data as the holotype .</p><p>Etymology. Named after the type locality, Tuoluodadong Cave.</p><p>Diagnosis (♂ ♀). Small-sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin and finely denticulated, epistome obtuse and small, round; posterior margin of the carapace with two setae; tergites Ⅰ–II each with two setae, tergites III–IV each with four setae. Pedipalps slender, femur 6.67–6.89 (♂), 6.45–6.80 (♀) times longer than broad; chela 5.87–6.21 (♂), 5.76–5.88 (♀) times longer than broad; chelal fingers without intercalary teeth but fixed chelal finger with a modified accessory tooth (td) on retrolateral face; chemosensory setae (sc) present on dorsum of chelal hand.</p><p>Description. Males (holotype and paratype) (Figs 37A, 38A–F, 39).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 38C, 39A): carapace 1.06–1.09 times longer than broad, markedly narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome obtuse and small, round, with two setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with 8–10 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 39C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 38D, 39B): large, about as long as carapace, 2.13–2.33 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with ten teeth, distal one largest; movable finger with 8–9 retrorse contiguous small teeth; galea absent (Fig. 39B). Serrula exterior with 16–17 blades and serrula interior with 10–11 blades. Rallum with eight blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 39D).</p><p>Pedipalp (Figs 38A–B, E, 39E–G): long and slender, trochanter 1.33–1.50, femur 6.67–6.89, patella 2.20–2.33, chela 5.87–6.21, hand 2.27–2.43 times longer than broad; femur 2.82–2.86 times longer than patella; movable chelal finger 1.56–1.59 times longer than hand and 0.60–0.62 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs 38E, 39E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b; it and est situated between b and t (Fig. 39F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 26–27 macrodenticles, slightly retrorse and pointed, plus a modified accessory tooth on retrolateral face (td, near tip), 27–28 in total; movable chelal finger with 11 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 16–17 vestigial, rounded and contiguous basal teeth, 27–28 in total (Fig. 39F). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 38B, 39G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 2: 2: 4: 4: 4: 4: 4–6: 5–6: 5–6: 4–5: T2T: 0. Sternal chaetotaxy III–XII: 12–14: 14: 8–10: 8–9: 8–10: 8–10: 9–11: 9: 0: 2. Anterior genital operculum with 11–12 setae, genital opening slit-like, with 13–16 marginal setae on each side, 38–43 in total (Fig. 38F).</p><p>Legs (Fig. 39H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg Ⅰ 1.94–2.06 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.38–2.47 times longer than tibia. Femoropatella of leg IV 2.33–2.59 times longer than deep; tibia 4.71 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.20 times longer than deep (TS = 0.38), tarsus 9.25–9.50 times longer than deep and 2.31–2.38 times longer than metatarsus (TS = 0.27–0.29). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 36C, 37B, 38G). Mostly same as males, but a little larger on average; with same coxal chaetotaxy as males; tergal chaetotaxy Ⅰ–XII: 2: 2: 4: 4: 5–6: 5–6: 5–6: 6–7: 6–7: 3–4: T2T: 0; sternal chaetotaxy IV–XII: 11–12: 9–11: 8–10: 9–11: 10–12: 11: 11: 0: 2; anterior genital operculum with 12 setae, posterior margin with 12–13 marginal setae, 24–25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.00–3.40 times longer than deep (TS = 0.33–0.35), tarsus 10.00–10.25 times longer than deep and 2.28–2.35 times longer than metatarsus (TS = 0.24–0.28).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.14–1.28. Pedipalps: trochanter 0.12–0.15/0.09–0.10 (1.33–1.50), femur 0.60–0.62/0.09 (6.67–6.89), patella 0.21–0.22/0.09–0.10 (2.20–2.33), chela 0.87–0.88/0.14–0.15 (5.87–6.21), hand 0.34/0.14–0.15 (2.27–2.43), movable chelal finger length 0.53–0.54. Chelicera 0.34–0.35/0.15–0.16 (2.13–2.33), movable finger length 0.18– 0.19. Carapace 0.37–0.38/0.35 (1.06–1.09). Leg Ⅰ: trochanter 0.08–0.10/0.09 (0.89–1.11), femur 0.31–0.33/0.05– 0.06 (5.50–6.20), patella 0.16/0.05 (3.20), tibia 0.15–0.16/0.04–0.05 (3.00–4.00), tarsus 0.37–0.38/0.04 (9.25– 9.50). Leg IV: trochanter 0.12–0.15/0.09–0.10 (1.20–1.67), femoropatella 0.42–0.44/0.17–0.18 (2.33–2.59), tibia 0.33/0.07 (4.71), metatarsus 0.16/0.05 (3.20), tarsus 0.37–0.38/0.04 (9.25–9.50).</p><p>Females: body length 1.44–1.64. Pedipalps: trochanter 0.13/0.10–0.11 (1.18–1.30), femur 0.68–0.71/0.10–0.11 (6.45–6.80), patella 0.24–0.25/0.12 (2.00–2.08), chela 0.94–0.98/0.16–0.17 (5.76–5.88), hand 0.38–0.40/0.16–0.17 (2.35–2.38), movable chelal finger length 0.57–0.59. Chelicera 0.39–0.41/0.17–0.19 (2.16–2.29), movable finger length 0.20–0.22. Carapace 0.40–0.43/0.40–0.43 (1.00). Leg Ⅰ: trochanter 0.11/0.08–0.10 (1.10–1.38), femur 0.35– 0.39/0.06 (5.83–6.50), patella 0.19/0.05–0.06 (3.17–3.80), tibia 0.17–0.18/0.04–0.05 (3.60–4.25), tarsus 0.40– 0.43/0.04 (10.00–10.75). Leg IV: trochanter 0.13–0.15/0.11–0.12 (1.18–1.25), femoropatella 0.47–0.50/0.18–0.19 (2.47–2.78), tibia 0.35/0.07–0.08 (4.38–5.00), metatarsus 0.17–0.18/0.05–0.06 (3.00–3.40), tarsus 0.40–0.41/0.04 (10.00–10.25).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/03DA1A1EFFB7FF95FF053847E3C23EAE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFFA8FF92FF053AC1E3C23A2E.text	03DA1A1EFFA8FF92FF053AC1E3C23A2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius xiaoensis Hou & Feng & Zhang 2023	<div><p>Lagynochthonius xiaoensis sp. nov. (OiffiṄṁae)</p><p>Figs 40–43</p><p>Type material. Holotype ♀ (Ps.- MHBU-GZC190727-01 - 01): China, Guizhou Province, Suiyang County, Yangchuan Sub-district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.175514&amp;materialsCitation.latitude=27.999336" title="Search Plazi for locations around (long 107.175514/lat 27.999336)">Xinglong Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.175514&amp;materialsCitation.latitude=27.999336" title="Search Plazi for locations around (long 107.175514/lat 27.999336)">Xiao Cave</a>, under stones in the deep zone [27°59′57.61″N, 107°10′31.85″E], 976 m a.s.l., 27 July 2019, Zegang Feng, Zhaoyi Li &amp; Chen Zhang leg. Paratype: 1 ♀ (Ps.- MSWU-GZC190727-01 - 02), with the same data as the holotype .</p><p>Etymology. Named after the type locality, Xiao Cave.</p><p>Diagnosis (♀). Small-sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome obtuse and small, round; posterior margin of carapace with two setae; tergites Ⅰ–VI each with four setae. Pedipalps slender, femur 5.91–6.09 times longer than broad; chela 6.33–6.47 times longer than broad; chelal fingers without intercalary teeth but fixed chelal finger with a modified accessory tooth (td) on retrolateral face; chemosensory setae (sc) present on dorsum of chelal hand.</p><p>Description. Females (holotype and paratype), male unknown (Figs 40D, 41–43).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 42C, 43A): carapace 0.93 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome obtuse and small, round, with two setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with 7–8 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 43C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 42D, 43B): large, about as long as carapace, 2.15 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12–14 teeth, distal one largest; movable finger with 11 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 43B). Serrula exterior with 15–16 blades and serrula interior with 11–12 blades. Rallum with nine blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 43D).</p><p>Pedipalp (Figs 42A–B, F, 43E–G): long and slender, trochanter 1.27, femur 5.91–6.09, patella 1.92–2.08, chela 6.33–6.47, hand 2.53 times longer than broad; femur 2.68–2.83 times longer than patella; movable chelal finger 1.50–1.58 times longer than hand and 0.60–0.62 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs 42F, 43E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming an oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb situated midway between st and b; b and t situated subdistally, t situated distal to b and between it and est; est distal to b (Fig. 43F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 24–25 macrodenticles, slightly retrorse and pointed, plus a modified accessory tooth on retrolateral face (td, close to dx), 25–26 in total; movable chelal finger with ten macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 14–16 vestigial, rounded and contiguous basal teeth, 24–26 in total (Fig. 43F). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 42B, 43G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 4: 4: 4: 4: 4: 4: 5: 5–6: 5–6: 4: T2T: 0. Sternal chaetotaxy IV–XII: 12–13: 10: 8–9: 8: 10–11: 10: 9–10: 0: 2. Anterior genital operculum with 10–11 setae, posterior margin with 13–16 marginal setae, 24–26 in total (Fig. 42E).</p><p>Legs (Fig. 43H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg Ⅰ 1.83–1.89 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.24–2.31 times longer than tibia. Femoropatella of leg IV 2.82 times longer than deep; tibia 4.25–4.86 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 2.83–3.00 times longer than deep (TS = 0.35–0.39), tarsus 9.50 times longer than deep and 2.11–2.24 times longer than metatarsus (TS = 0.32). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Females: body length 1.25–1.47. Pedipalps: trochanter 0.14/0.11 (1.27), femur 0.65–0.67/0.11 (5.91–6.09), patella 0.23–0.25/0.12 (1.92–2.08), chela 0.95–0.97/0.15 (6.33–6.47), hand 0.38/0.15 (2.53), movable chelal finger length 0.57–0.60. Chelicera 0.43/0.20 (2.15), movable finger length 0.23. Carapace 0.40/0.43 (0.93). Leg Ⅰ: trochanter 0.11–0.12/0.10 (1.10–1.20), femur 0.33–0.34/0.06 (5.50–5.67), patella 0.18/0.05–0.06 (3.00–3.60), tibia 0.16–0.17/0.04 (4.00– 4.25), tarsus 0.37–0.38/0.04 (9.25–9.50). Leg IV: trochanter 0.16–0.17/0.11 (1.45–1.55), femoropatella 0.48/0.17 (2.82), tibia 0.34/0.07–0.08 (4.25–4.86), metatarsus 0.17–0.18/0.06 (2.83–3.00), tarsus 0.38/0.04 (9.50).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/03DA1A1EFFA8FF92FF053AC1E3C23A2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
03DA1A1EFFACFF98FF053900E4AF3C46.text	03DA1A1EFFACFF98FF053900E4AF3C46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagynochthonius zhijinensis Hou & Feng & Zhang 2023	<div><p>Lagynochthonius zhijinensis sp. nov. (ṂĤṄṁae)</p><p>Figs 44–47</p><p>Type material. Holotype ♀ (Ps.-MHBU-GZZJ-19-04-01): China, Guizhou Province, Zhijin County, Chengguan Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.78786&amp;materialsCitation.latitude=26.67934" title="Search Plazi for locations around (long 105.78786/lat 26.67934)">Yangliu Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.78786&amp;materialsCitation.latitude=26.67934" title="Search Plazi for locations around (long 105.78786/lat 26.67934)">Long Cave</a>, under stones in the deep zone [26°40′45.63″N, 105°47′16.30″E], 1345 m a.s.l., 2 October 2019, Zegang Feng &amp; Lingchen Zhao leg. Paratype: 1 ♀ (Ps.-MSWU-GZZJ-19-04-02), with the same data as the holotype .</p><p>Etymology. Named after the Zhijin County, near the type locality.</p><p>Diagnosis (♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed and small, triangular; posterior margin of carapace with two setae; tergites II–III each with two setae, tergites IV–VII each with four setae. Pedipalps slender, femur 8.47–8.53 times longer than broad; chela 7.46–7.71 times longer than broad; only fixed chelal finger with intercalary teeth and a modified accessory tooth (td) on retrolateral face; chelal fingers slightly curved in dorsal view; chemosensory setae (sc) present on dorsum of chelal hand.</p><p>Description. Females (holotype and paratype), male unknown (Figs 45–47).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 46C, 47A): carapace 0.93–0.98 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5–6; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with 11–12 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 47C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 46D, 47B): large, about as long as carapace, 2.29–2.31 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 20–24 teeth, distal one largest; movable finger with 18–19 retrorse contiguous small teeth; galea absent (Fig. 47B). Serrula exterior with 20–21 blades and serrula interior with 13–15 blades. Rallum with seven blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 47D).</p><p>Pedipalp (Figs 46A–B, F, 47E–G): long and slender, trochanter 1.25–1.32, femur 8.47–8.53, patella 2.21–2.37, chela 7.46–7.71, hand 3.04–3.17 times longer than broad; femur 2.84–3.02 times longer than patella; movable chelal finger 1.42–1.45 times longer than hand and 0.58–0.59 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs 46F, 47E). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb situated midway between b and st; b and t situated subdistally and situated at the same level as est and it, respectively (Fig. 47F). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 23 macrodenticles, slightly retrorse and pointed, plus 11– 12 intercalary microdenticles, 34–35 in total; movable chelal finger with 21–22 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 7–8 vestigial, rounded and contiguous basal teeth, 28–30 in total (Fig. 47F). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 46B, 47G).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 2–3: 2: 2: 4: 4: 4: 4: 5: 5: 2: T2T: 0, tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy IV–XII: 13: 9: 9–10: 8–9: 9: 9: 7–8: 0: 2. Anterior genital operculum with 9–10 setae, posterior margin with 12–14 marginal setae, 21–24 in total (Fig. 46E).</p><p>Legs (Fig. 47H–I): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg Ⅰ 1.76–1.79 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.48–2.56 times longer than tibia. Femoropatella of leg IV 4.12–4.13 times longer than deep; tibia 6.27–6.36 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.56 times longer than deep (TS = 0.28–0.34), tarsus 14.17–14.33 times longer than deep and 2.66–2.69 times longer than metatarsus (TS = 0.31–0.34). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Females: body length 2.17–2.36. Pedipalps: trochanter 0.25/0.19–0.20 (1.25–1.32), femur 1.27–1.28/0.15 (8.47–8.53), patella 0.42–0.45/0.19 (2.21–2.37), chela 1.79–1.85/0.24 (7.46–7.71), hand 0.73–0.76/0.24 (3.04–3.17), movable chelal finger length 1.06–1.08. Chelicera 0.71–0.74/0.31–0.32 (2.29–2.31), movable finger length 0.37–0.38. Carapace 0.62–0.65/0.66–0.67 (0.93–0.98). Leg Ⅰ: trochanter 0.19–0.21/0.16 (1.19–1.31), femur 0.70–0.72/0.09 (7.78–8.00), patella 0.39–0.41/0.08–0.09 (4.56–4.88), tibia 0.31–0.32/0.06 (5.17–5.33), tarsus 0.77–0.82/0.06–0.07 (11.71– 12.83). Leg IV: trochanter 0.28–0.29/0.16–0.18 (1.61–1.75), femoropatella 0.99–1.03/0.24–0.25 (4.12–4.13), tibia 0.69–0.70/0.11 (6.27–6.36), metatarsus 0.32/0.09 (3.56), tarsus 0.85–0.86/0.06 (14.17–14.33).</p><p>Distribution. Known only from the type locality.</p><p>Key to the species of Lagynochthonius from China</p><p>1 Carapace without antero-median setae; trichobothrium sb closer to b than to st ..................................... 2</p><p>- Carapace with antero-median setae; trichobothrium sb closer to st than to b or midway between st and b ................ 3</p><p>2 Chela 7.03 (♂) times as long as broad (length 2.04 mm); epistome pointed and small, triangular............................................................................................ L. xinjiaoensis Hou, Gao &amp; Zhang, 2022</p><p>- Chela 6.44–6.46 (♂), 5.87–6.04 (♀) times as long as broad (length 1.55–1.61 (♂), 1.69–1.76 (♀) mm); epistome small and obtuse, rounded..................................................... L. magnidentatus Hou, Gao &amp; Zhang, 2022</p><p>3 Carapace without eyes or eyespots........................................................................ 4</p><p>- Carapace with eyes or eyespots......................................................................... 13</p><p>4 At least one chela finger with intercalary teeth.............................................................. 5</p><p>- Both chelal fingers without intercalary teeth............................................................... 21</p><p>5 Only fixed chelal finger with intercalary teeth............................................................... 6</p><p>- Both chelal fingers with intercalary teeth................................................................... 8</p><p>6 Trichobothrium sb closer to st than to b; tergite Ⅹ with four setae............................................... 7</p><p>- Trichobothrium sb situated midway between st and b; tergite Ⅹ with two setae..................... L. zhijinensis sp. nov.</p><p>7 The intercalary teeth round and obtuse; the teeth on movable chelal fingers retrorse and contiguous; chela length 1.57–1.61 (♂), 1.64–1.69 (♀) mm; movable chelal finger length 0.93–0.96 (♂), 0.99–1.01 (♀) mm.... L. retrorsus Hou, Gao &amp; Zhang, 2022</p><p>- The intercalary teeth small and pointed; the teeth on movable chelal fingers pointed and well-spaced; chela length 2.07–2.16 (♂), 2.06–2.19 (♀) mm; movable chelal finger length 1.24–1.25 (♂), 1.20–1.26 (♀) mm.................................................................................................. L. spinulentus Hou, Gao &amp; Zhang, 2022</p><p>8 Tergites II–III and Ⅹ each with four setae.................................................................. 9</p><p>- Tergites II–III and Ⅹ each with two setae.................................................................. 10</p><p>9 Trichobothrium sb closer to st than to b; body length 1.42 (♂), 1.59 (♀) mm; chela 6.65 (♂), 6.63 (♀) times as long as broad (length 1.33 (♂), 1.26 (♀) mm); palpal femur 7.15 (♂), 7.42 (♀) times as long as broad (length 0.93 (♂), 0.89 (♀) mm); coxae II with seven terminally indented coxal spines on each side................................. L. longedentatus sp. nov.</p><p>- Trichobothrium sb situated midway between st and b; body length 1.63 (♂), 1.81–2.20 (♀) mm; chela 8.76 (♂), 8.30–8.32 (♀) times as long as broad (length 1.84 (♂), 1.83–1.91 (♀) mm); palpal femur 10.46 (♂), 10.07–10.38 (♀) times as long as broad (length 1.36 (♂), 1.35–1.41 (♀) mm); coxae II with 9–11 terminally indented coxal spines on each side.................................................................................. L. yaowangguensis Hou, Gao &amp; Zhang, 2022</p><p>10 Chelal fingers straight or slightly curved in dorsal view; fixed chelal finger with max. 47 teeth, movable chelal finger with max. 42 teeth............................................................................................ 11</p><p>- Chelal fingers markedly curved in dorsal view; fixed chelal finger with 58–60 teeth, movable chelal finger with 60–61 teeth.................................................................................. L. guanniuensis sp. nov.</p><p>11 Palpal femur 8.07–8.93 (♂) times as long as broad (length 1.21–1.25 mm), chela length 1.68–1.76 (♂) mm; trichobothrium sb closer to st than to b .................................................................................. 12</p><p>- Palpal femur 7.57–7.64 (♂) times as long as broad (length 1.06–1.07 mm), chela length 1.53–1.54 (♂) mm; trichobothrium sb situated midway between st and b .............................................. L. fengi Hou, Gao &amp; Zhang, 2022</p><p>12 Carapace markedly narrowed posteriorly; chela 8.38–8.84 (♂) times as long as broad, hand 3.38–3.58 (♂) times longer than broad; proximal half of movable chelal finger without vestigial teeth; trichobothrium sb of movable chelal finger 1.22–1.24 times as far from st as from b ............................................... L. serratus Hou, Gao &amp; Zhang, 2022</p><p>- Carapace gently narrowed posteriorly; chela 7.00 (♂) times as long as broad, hand 2.92 (♂) times longer than broad; proximal half of movable chelal finger with six vestigial teeth; trichobothrium sb of movable chelal finger 1.67 times as far from st as from b ............................................................... L. xiaolinensis Hou, Gao &amp; Zhang, 2022</p><p>13 Troglomorphic species with a larger body size and elongate appendages: body length not less than 1.71 (♂, ♀) mm, chela length not less than 1.32 (♂, ♀) mm; without corneate eyes, only anterior pair of eyespots present.......................... 14</p><p>- Epigean species with smaller body size and without elongate appendages: body length not more than 1.60 (♂, ♀) mm, chela length not more than 1.00 (♂, ♀) mm; with corneate eyes, anterior pair of eyes well developed....................... 15</p><p>14 Chela 7.00–7.14 (♂), 6.40–6.46 (♀) times as long as broad (length 1.54–1.57 (♂), 1.55–1.60 (♀) mm); palpal femur 8.21–8.50 (♂), 7.50–7.87 (♀) times as long as broad (length 1.15–1.19 (♂), 1.18–1.20 (♀) mm); coxae II with 8–9 terminally indented coxal spines on each side................................................... L. crassus Hou, Gao &amp; Zhang, 2022</p><p>- Chela 7.33–7.67 (♂), 6.71–6.82 (♀) times as long as broad (length 1.32–1.38 (♂), 1.41–1.50 (♀) mm); palpal femur 6.92–7.23 (♂), 6.80–7.29 (♀) times as long as broad (length 0.90–0.94 (♂), 1.02 (♀) mm); coxae II with 9–11 terminally indented coxal spines on each side.................................................................... L. latipectus sp. nov.</p><p>15 Two corneate eyes, posterior pair of eyes reduced to eyespots; tergites Ⅰ and II each with four setae................... 16</p><p>- Four corneate eyes, posterior pair of eyes well developed; tergites Ⅰ and II each with two setae................................................................................................... L. medog Zhang &amp; Zhang, 2014</p><p>16 Carapace without epistome or with extremely indistinct epistome.............................................. 17</p><p>- Carapace with a distinct triangular epistome........................................... L. niger Hu &amp; Zhang, 2012</p><p>17 At least one finger of chela with intercalary teeth........................................................... 18</p><p>- Both chelal fingers without intercalary teeth............................................................... 19</p><p>18 Only fixed chelal finger with intercalary teeth........................................ L. harveyi Hu &amp; Zhang, 2014</p><p>- Both chelal fingers with intercalary teeth.............................................. L. tonkinensis (Beier, 1951)</p><p>19 Movable chelal finger more than 1.20 (♂, ♀) times longer than hand; proximal half of movable finger with rounded vestigial teeth.............................................................................................. 20</p><p>- Movable chelal finger 1.03 (♂) times, 0.97–1.00 (♀) times longer than hand; proximal half of movable chelal finger without vestigial teeth........................................................ L. brachydigitatus Zhang &amp; Zhang, 2014</p><p>20 Chelal hand 2.50–2.87 (♂) times, 2.39–2.75 (♀) as long as broad; trichobothrium sb of movable chelal finger two times as far from st as from b; tergites Ⅰ–VII with four setae................................... L. leptopalpus Hu &amp; Zhang, 2012</p><p>- Chelal hand 2.00 (♂) times as long as broad; trichobothrium sb of movable chelal finger 1.5 times as far from st as from b; only the tergites Ⅰ–II with four setae......................................................... L. sinensis (Beier, 1967)</p><p>21 Tergites Ⅰ–II each with two setae........................................................................ 22</p><p>- Tergites Ⅰ–II each with three or four setae................................................................. 25</p><p>22 Carapace with a round and obtuse epistome; body length max. 1.83 mm, chelal length max. 1.11 mm .................. 23</p><p>- Carapace with a distinct triangular epistome; body length min. 1.85 mm, chelal length min. 1.51 mm .................. 24</p><p>23 Palpal femur 6.45–6.80 (♀) times as long as broad; chelal finger 1.48–1.50 (♀) times longer than hand; tergite Ⅴ with 5–6 setae; anterior genital operculum with 12 setae; trichobothrium ist situated distal to st ................... L. tuoluoensis sp. nov.</p><p>- Palpal femur 5.43–5.75 (♀) times as long as broad; chelal finger 1.55–1.56 (♀) times longer than hand; tergite Ⅴ with four setae; anterior genital operculum with ten setae; trichobothrium ist situated proximal st .............. L. maanensis sp. nov .</p><p>24 Tergite Ⅹ with two setae; body length min. 2.00 (♂), 2.17 (♀) mm, chelal length min. 1.79 (♂), 1.84 (♀) mm, palpal femur min. 1.25 (♂), 1.30 (♀) mm; coxae II with 13–15 terminally indented coxal spines on each side........... L. nigriculus sp. nov.</p><p>- Tergite Ⅹ with four setae; body length max. 1.78 (♂), 2.05 (♀) mm, chelal length max. 1.62 (♂), 1.66 (♀) mm, palpal femur max. 1.11 (♂), 1.17 (♀) mm; coxae II with nine terminally indented coxal spines on each side.......................................................................................... L. laoxueyanensis Hou, Gao &amp; Zhang, 2022</p><p>25 Carapace with a distinct triangular epistome............................................................... 26</p><p>- Carapace with a round and obtuse epistome............................................................... 27</p><p>26 Chelal finger 1.51 (♀) times longer than hand; trichobothrium sb situated midway between st and b; coxae II with ten terminally indented coxal spines on each side; rallum with nine blades; movable chelal finger with 27–29 teeth... L. biyunensis sp. nov.</p><p>- Chelal finger 1.24 (♀) times longer than hand; trichobothrium sb closer to st than to b; coxae II with 12 terminally indented coxal spines on each side; rallum with seven blades; movable chelal finger with 42 teeth.................................................................................................. L. xibaiensis Hou, Gao and Zhang, 2022</p><p>27 Smaller: body length max. 1.23 (♂), 1.47 (♀) mm; chela length max. 0.85 (♂), 0.97 (♀) mm; palpal femur length max. 0.60 (♂), 0.67 (♀) mm.................................................................................... 28</p><p>- Larger: body length min. 1.59 (♂), 1.64 (♀) mm; chela length min. 1.03 (♂), 1.12 (♀) mm; palpal femur length min. 0.74 (♂), 0.80 (♀) mm........................................................................................ 29</p><p>28 Fixed chelal finger with 24–25 teeth; trichobothrium sb situated midway between st and b; rallum with nine blades............................................................................................ L. xiaoensis sp. nov.</p><p>- Fixed chelal finger with 35–37 teeth; trichobothrium sb closer to st than to b; rallum with eight blades........................................................................................ L. minimus Hou, Gao and Zhang, 2022</p><p>29 Larger: body length min. 1.76 (♂), 1.93 (♀) mm; chela length min. 1.26 (♂), 1.38 (♀) mm; palpal femur length min. 0.90 (♂), 0.99 (♀) mm........................................................................................ 30</p><p>- Smaller: body length max. 1.66 (♂), 1.73 (♀) mm; chela length max. 1.11(♂), 1.14 (♀) mm; palpal femur length max. 0.79 (♂), 0.80 (♀) mm...................................................................... L. mawangensis sp. nov.</p><p>30 Trichobothrium sb situated midway between st and b; body length 1.76–1.95 (♂), 1.93 (♀) mm, chela 6.63–6.95 (♂), 5.75 (♀) times longer than broad (length 1.26–1.32 (♂), 1.38 (♀) mm)................................ L. longyanensis sp. nov.</p><p>- Trichobothrium sb closer to st than to b; body length 2.55–2.92 (♂), 2.72–2.95 (♀) mm, chela 8.50–10.20 (♂) times longer than broad (length 1.85–2.22 (♂), 1.94–2.03 (♀) mm).............................. L. bailongtanensis Li, Liu &amp; Shi, 2019</p></div>	https://treatment.plazi.org/id/03DA1A1EFFACFF98FF053900E4AF3C46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China. Zootaxa 5309 (1): 1-64, DOI: 10.11646/zootaxa.5309.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5309.1.1
