taxonID	type	description	language	source
03DB878F1138196B638820C478CDF8E3.taxon	discussion	Remarks. The Metapseudidae, currently represented by 86 species classified in four subfamilies and 20 genera (Anderson 2013; Appeltans et al. 2013), is one of the most diverse families associated with coral reefs. From 79 species of tanaidaceans recorded globally from coral reefs, 28 are members of the Metapseudidae. The family was established by Lang (1970) to accommodate four genera: Apseudomorpha Miller, 1940, Cyclopoapseudes Menzies, 1953, Metapseudes Stephensen, 1927 and Synapseudes Miller 1940, which in those days included around 21 species. Two years later, Gutu (1972) defined two subfamilies, the Metapseudinae Lang 1970 and the Synapseudinae Gutu, 1972, based on the reduction (= fusion) of the pleonites with the pleotelson. Later on the family was supplemented by two other subfamilies: the Chondropodinae (Gutu 2008) and the Msangiinae (Gutu, 2006). Although the Metapseudidae currently includes four times as many species as when Lang (1970) established the family, its definition has never been updated. Most of the characters pinpointed by Lang are vague and allow only to distinguish the metapseudids from the members of the Kalliapseudidae (by having the dactylus not modified, without sensory setae) or some apseudids (by the character of the seta on the endite of the maxilliped). The metapseudids can be distinguished from the Sphyrapodidae by presence in the latter of six fully developed pereonites; from the Parapseudidae and most Apseudidae (except Frankapseudes and Atlantapseudes) by a reduced antennal squama; from the Whiteleggiidae by the lateral attachment or absence of pleopods, from the Sphaeromapseudidae and the Tanzanapseudidae by their elongated body; and from the Pagurapseudidae and the Numbakullidae by having the first pair of pereopods fossorial. Undoubtedly the Metapseudidae requires profound revision and new definition. The same is also true for some of metapseudid genera (e. g. Synapseudes), as well as for the subfamilies. It is very probable that the Msangiinae Gutu, 2006 or the Chondropodinae Gutu, 2008 might not be closely-related to the Synapseudinae Gutu, 1972 or the Metapseudinae Lang, 1970, and that they might need to be raised to family rank. A debate on the phyletic relationship within the Metapseudidae is beyond the scope of this paper, however a phylogenetical analysis based on morphological characters is demonstrating that all metapseudids, tanzanapseudids, pagurapseudids, whiteleggiids and sphaeromapseudids may constitute a monophyletic clade (studies in progress). Metapseudids are tropical and temperate crustaceans, with the most northerly records of the family coming from the English Channel in the Atlantic (Guţu 1989 a) and the East China Sea in the Pacific (Larsen & Shimomura 2006); in the southern hemisphere their most southern records are at Tierra del Fuego (Sieg 1986 a), waters off Southern Africa (Barnard 1914), New Zealand (Chilton 1882; Gardiner 1973) and SE Australia (Błażewicz- Paszkowycz & Bamber 2012). The Metapsudidae are generally shallow water crustaceans, which rarely occur below 100 m (Gardiner 1973; Gutu 1989 b). They are most often associated with coral rubble, but also with sponges (Shiino, 1951), algae (Menzies 1953), turf of Bryozoa (Menzies 1953, Bamber 2008) or occasionally with sand, mud or shell-sand (e. g. Gutu, 2006). Subfamily Chondropodinae Gu t u, 2008	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1138196B638820C478CDF8E3.taxon	description	Definition (amended after Gutu 2008): Body dorsoventrally flattened. Pleon with five short pleonites; pleotelson short with small acute lateral process. Antennule peduncle with at least one apophysis on inner margin of first article (except Bamberus); at least outer flagellum long, multiarticulated. Antenna second article long, with apophyses on inner margin (except Vestigiramus); squama small. Maxilliped palp article 3 longer than broad. Pereopod 1 fossorial; exopodite terminal article with usually more than four setae; basis thick with plumose setae on dorsal margin and usually with spiniform denticles (except Bamberus). Pereopods 2 to 6 slender, walking type; pereopods 3 to 6 with propodus longer than carpus, and usually subequal merus, slightly curved. Pleopods present, biramous, in five pairs. Uropod with multiarticulated rami. Dimorphism: usually well-developed, male with dimorphic chelipeds (except Calozodion). Genera included: Calozodion Gardiner, 1973; Chondropodus Gutu, 2006; Hoplomachus Gutu, 2002; Julmarichardia Gutu, 1989 b; Trichapseudes Barnard, 1920; Vestigiramus Guţu, 2009; Zaraza, Gutu, 2006; Bamberus n. gen.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1138196B638820C478CDF8E3.taxon	discussion	Remarks. According to Gutu (2008) one of the major features of the Chondropodinae is the presence of at least one apophysis on the antennule peduncle (as in Calozodion bacescui Gutu, 1996 or Calozodion singularis Gutu, 2002) and the presence of an apophysis on the upper margin of the first pereopod basis. Both characters are highly variable. The number of antennular apophyses can range from one to six in Calozodion and up to seven in some members of Julmarichardia while the number of apophyses on the pereopod can vary ontogenetically (Gutu 1984). The number of setae on the terminal article of the exopod of the first pereopod wasn’t considered to be valid diagnosic character. Nevertheless many of the chondropodinids (as well as many members of Parapseudidae Gutu, 1981) have four or more plumose setae in contrast to members of Apseudidae Leach, 1813.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F113B196A638825787DC3F831.taxon	diagnosis	Diagnosis. Body elongated, seven times as long as wide, integument smooth. Carapace little wider than long and as long as pereonites 1 and 2 together. Pleon about one-quarter of total body length, consisting of five free pleonites and pleotelson; pleotelson with one short apophysis on each side. Antennule biramous, peduncle first article without apophyses. Antennal squama small with three distal setae. Mandibular palp with three articles, article 1 with one seta. Maxillule inner endite grossly reduced with two distal setae; palp with three distal setae. Labium palp with two setae terminally. Cheliped with exopod, in female slender, in male enlarged. Pereopod 1 with exopod. Pereopods 1 – 3 basis without apophyses. Pleopods biramous; each ramus with six plumose setae. Uropod peduncle with strong apophysis dorsally; endopod as long as quarter of total body length, consisting of about 10 segments; exopod five-segmented.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F113B196A638825787DC3F831.taxon	materials_examined	Type species: Bamberus jinigudirus n. gen. n. sp., by monotypy.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F113B196A638825787DC3F831.taxon	etymology	Etymology. The name is given in honour of Dr Roger N. Bamber, our special friend, in recognition of his accomplished contribution to the knowledge of the Tanaidacea. Gender: masculine	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F113B196A638825787DC3F831.taxon	discussion	Remarks. The combination of the characters such as the elongate second article of the antenna armed with apophyses on its inner margin and with a small squama, a long third maxilliped-palp article, a first pereopod with an exopodite bearing five setae, and with a stout basis bearing plumose setae, pereopods 2 to 6 more slender (walking type) than the first pereopod and pereopods 3 to 6 with a propodus longer than the carpus, as well as much larger chelipeds in males than in females places Bamberus n. gen. in the family Chondropodinae. Bamberus in general view resembles three of the seven genera currently included in the Chondropodinae, i. e. Calozodion Gardiner, 1973, Chondropodus Gutu, 2006 and Hoplomachus Gutu, 2002. These four genera have a relatively elongated body, five well-developed pleonites with acute epimera, and a few setae often accompanied by a few robust spines on the basis of the first pereopod. Bamberus is most similar to Chondropodus — both of them lack plumose setae on pereopod 6 and have a slender cheliped in females (Table 2). In contrast to Chondropodus the new genus lacks a spine on the first article of the antennule and on the basis of the first pereopod. The new genus can be immediately distinguished from the other chondropodiins by having a smooth and poorly-developed rostrum, compared with a large, acute and serrated rostrum in Julmarichardia, a well-developed exopod (reduced or absent in Vestigiramus), by having only two segments in the accessory flagellum of the antennule (five segments in Zaraza) and by having only a few setae on the mandibular palp (numerous plumose setae in Trichapseudes). Bamberus n. gen. has a conspicuous apophysis on the uropod basis, apparently similar to that present in Vicinisyndes Gutu, 2007 and all members of the Whiteleggiidae Gutu, 1972. A distal apophysis on the basis of the uropod is characteristic for many metapseudids but only in Vicinisyndes, Bamberus and the whiteleggiids is situated dorsally. Vicinisyndes, with only one free pleonite, is a member of the subfamily Synapseudinae Gutu 1972, while the Whiteleggiidae is represented by three species in two genera which have a heavy and stronglysculptured integument.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F113D196363882281790DFE70.taxon	materials_examined	Material examined. Holotype: female (WAM C 54552), NR 09 - 70 B, 22 ° 38 ' 30.16 " S, 113 ° 41 ' 30.05 " E, Ningaloo, wreck of Cho Fu Kumar, sand and fine rubble in groove, outer reef edge, depth 8 – 12 m, 26 May 2009, coll. N. L. Bruce & L. Hughes. Allotype: male (WAM C 54553), NR 09 - 70 B, same locality as holotype. Paratypes: 5 females, 1 male (WAM C 54554), NR 09 - 70 B, same locality as holotype; 1 male (MTQ W 34257), NR 09 - 64 C, 22 ° 31 ' 2.21 " S, 113 ° 39 ' 47.63 " E, Ningaloo, wreck of Cho Fu Kumar, finger rubble on sand, outer reef edge, depth 9 m, 26 May 2009, coll. N. L. Bruce & L. Hughes; 1 female (MTQ W 34258), NIN 7 E, 21 ° 54 ' 43.44 " S, 113 ° 57 ' 6.30 " E, Ningaloo, lagoon patch reef, southern side of reef pass, sand, edge of large coral heads (small bommies), depth 4.5 – 5.5 m, 8 August 2008, coll. N. L. Bruce & M. Błażewicz-Paszkowycz; 1 female (W 34259), NIN 13 D, 21 ° 51 ' 8.88 " S, 114 ° 0 ' 12.24 " E, Ningaloo, off northern reef point, sand, adjacent to bommies, depth 8 – 10 m, 14 August 2008, coll. M. Błażewicz-Paszkowycz & N. L. Bruce; 3 males (WAM C 54555), 2 females (MTQ 34260), NIN 9 B, 21 ° 51 ' 23.94 " S, 114 ° 0 ' 9.12 " E, Ningaloo, edge of northern lagoon pass, dead Acropora, depth 4 – 5 m, 11 August 2008, coll. M. Błażewicz-Paszkowycz & C. Hass.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F113D196363882281790DFE70.taxon	etymology	Etymology. Jinigudirais is the local Aboriginal name for the Ningaloo area.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F113D196363882281790DFE70.taxon	description	Description of female. Body (Figs 1 A, B) 1.8 mm long, 6.9 times as long as wide. Carapace 15 % of total body length, with weakly-pointed rostrum. Pereonite 1 half as long as wide, anterolateral margin adhering to carapace; pereonite 2 trapezoidal; pereonites 3 and 4 0.8 times as long as wide; pereonite 5 0.6 times as long as pereonite 4; pereonite 6 0.6 times as long as pereonite 5; pereonites with two or three setae laterally. Pleon 15 % of total body length, each pleonite with ventral hyposphenium and setae laterally; pleonites 1 – 3 with one seta, pleonites 4 and 5 with two lateral setae; pleotelson with two setae laterally and two distally. Antennule (Fig. 3 B) peduncle article 1 2.7 times as long as wide, with four penicillate and three simple setae on outer margin, and with two penicillate setae and one simple seta on inner margin; article 2 0.2 times as long as article 1, bearing one simple seta on outer margin, and two simple and two penicillate setae on inner margin; article 3 0.8 times as long as article 2, with one simple seta on outer- and inner margin distally; article 4 naked; outer flagellum with five segments; segment 2 with two simple setae; segment 5 with one seta and aesthetasc; inner flagellum with two segments, both with simple setae. Antenna (Fig. 3 D) peduncle article 1 0.4 times as long as article 2, with rounded process bearing fine distal seta; article 2 1.4 times as long as wide, outer margin serrated, one simple seta on inner margin; squama small, little longer than wide, with three setae terminally; article 3 0.3 times longer than article 2, with one simple seta; articles 4 and 5 similar in length, each little shorter than article 2; article 4 with one penicillate seta; article 5 with two penicillate and two simple distal setae; flagellum with three segments; distal segment with three simple distal setae. Mouthparts. Mandible palp (Fig. 3 E) article 1 with long distal seta on inner margin; article 2 with three simple and three pinnate setae on inner margin; article 3 with four distally and two subdistally strong setae; molar process (Fig. 3 F) serrated distally; right incisor (Fig. 3 H) blunt, with straight edge; left incisor (Fig. 3 G) weakly serrated, lacina mobilis present on left mandible, with four teeth and with three trifurcated and one simple setae on setiferous lobe. Labium (Fig. 3 M) with two plumose setae terminally and with some fine setae along both margin. Maxillule outer endite (Fig. 3 J) with ten distal spines and with two simple setae on inner margin; inner endite (Fig. 3 K) with two fine setae distally; palp (Fig. 3 L) with two articles; article 2 with three distal setae. Maxilla (Fig. 3 I) outer lobe of movable endite margin serrated, with two simple subdistal (broken) and three distal setae; inner lobe of movable endite with some simple setae; outer lobe of fixed endite with numerous simple and three leaf-shaped setae distally, and one plumose seta subdistally; inner lobe of fixed endite with three teeth on margin, one plumose and three simple setae distally and row of nine setae subdistally. Maxilliped (Fig. 3 N) basis 0.7 times as long as wide, with distal seta on inner margin; palp article 1 0.6 times as long as basis, with simple seta on inner margin; article 2 1.1 times as long as basis, with spine on outer margin, and numerous simple setae along inner margin; article 3 0.8 times as long as article 2 with numerous simple setae on inner margin; article 4 with five simple distal and one pinnate subdistal setae; endite (Fig. 3 O) with numerous simple setae and spines along inner margin. Epignath (Fig. 3 P) slender. Cheliped (Fig. 4 A) basis almost twice as long as wide, with two midventral and two distoventrally setae; exopodite with three articles; distal article with four plumose setae; merus half as long as basis, with three fine simple setae along ventral margin; carpus elongate, 2.6 times as long as wide, 1.6 times as long as merus, with two simple ventral margin and with one fine dorsal-distal seta; propodus 0.8 times as long as carpus, with some simple setae near dactylus insertion; fixed finger with three simple ventral setae and three setae on cutting edge; dactylus 0.6 times as long as propodus, with two simple setae. Pereopod 1 (Fig. 4 E) basis 1.3 times as long as wide, with five plumose setae along dorsal margin, and with two simple setae and one distoventral spine; exopodite with three articles; distal article with five plumose setae; ischium 0.2 times as long as wide; merus 0.7 times as long as basis, with three simple ventral setae and with two spines – distodorsal and distoventral; carpus half length of merus, with two simple setae and one distoventral spines, and with three simple setae and one distodorsal spine; propodus 0.7 times as long as merus, with two simple setae and three spines on ventral margin and with two simple setae and two spines on dorsal margin; dactylus together with unguis 0.7 times as long as propodus. Pereopod 2 (Fig. 4 F) basis 3.5 times as long as wide, with three simple ventral setae and with two penicillate setae and one simple seta dorsally; ischium 0.9 times as long as wide, with two simple setae and one serrated spine distoventrally; merus 0.3 times as long as basis, with one distodorsal simple seta and one simple seta and one serrated spine ventrally; carpus similar in length to merus, with three serrated spines and one simple seta ventrally, and one simple dorsal seta; propodus 1.1 times as long as carpus, with three serrated spines and one pinnate seta on ventral margin and with one plumose, one penicillate seta and one spine on dorsal margin; dactylus together with unguis 0.7 times as long as propodus. Pereopod 3 (Fig. 4 G) similar to pereopod 2, but ischium without spines; carpus with two serrated spines and four simple setae; propodus with two simple setae and two spines near dactylus insertion. Pereopod 4 (Fig. 4 H) basis 3.7 times as long as wide, with two penicillate setae and two fine setae proximally; ischium with one long distoventral seta; merus 1.3 times as long as wide, with two simple setae and serrated spine ventrally and one strong dorsal seta; carpus 1.7 times as long as merus, with three pairs of serrated ventral spines; propodus 1.7 times as long as carpus, with one plumose dorsal seta and with six thick pinnate distal setae; dactylus little longer than unguis, with small distal seta; together with unguis 0.7 times as long as propodus. Pereopod 5 (Fig. 4 I) basis three times as long as wide, with two penicillate setae proximally and one simple distoventral seta; ischium with two short and one long distoventral seta; merus 0.3 times as long as basis, with two ventral and one dorsal simple seta; carpus subequal to merus, with three serrated spines and one simple seta ventrally and with one serrated spine and one simple seta dorsally; propodus 1.6 times as long as carpus, with serrated midventral spine, row of simple setae along ventral margin, and with two penicillate and one pinnate setae distodorsally; dactylus together with unguis 0.6 times as long as propodus, with fine seta; unguis 0.6 times as long as dactylus. Pereopod 6 (Fig. 4 J) similar to pereopod 5, but basis without penicillate setae, ischium with only one, long seta; propodus with simple midventral seta and with one dorsal plumose seta as well as row of simple setae near dactylus insertion. Pleopods (Fig. 4 C) in five pairs, basis 1.4 times as long as wide, with simple seta distally; endopod with four distal setae and one simple, one plumose seta at midlength; exopod with two articles; article 1 with plumose distal seta; article 2 with four distal and one subdistal setae. Uropod (Fig. 4 D) basis 2.5 times as long as wide, with forward-bent apophysis and with five simple distal setae; endopod with 14 segments, some armored with simple setae; distal segment with four terminal setae; exopod with five segments; distal segment with three simple setae. Description of male. Body (Figs. 2 A, B) 1.4 mm long, eight times as long as wide. Carapace 20 % of total body length. Pereonite 1 0.5 times as long as wide; pereonite 2 little shorter than pereonite 1, with one pair of setae anterolaterally; pereonites 3 to 6 similar in length, 0.6 times as long as wide, bearing two pair of lateral setae (anteriorly and posteriorly). Pleon and pleotelson similar to those of female. Antennule (Fig. 3 A) similar to that of female, but peduncle article 1 is 4.2 times as long as wide and article 2 is twice as long as wide; article 3 1.7 times as long as article 2. Antenna (Fig. 3 C) similar to that of female, but with longer peduncle articles: peduncle article 2 2.1 times as long as wide, article 3 1.2 times as long as wide, and article 4 and 5 are similar in length, and twice as long as wide. Cheliped (Fig. 4 B) left and right the same size, much stronger than that of female; propodus wide and flat, 0.8 times as long as wide, with two setae ventrally; fixed finger with two, large apophysis and row of simple setae on inner margin and four setae ventrally; dactylus inner margin serrated. Subfamily Synapseudinae Gu t u, 1972	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F113D196363882281790DFE70.taxon	discussion	Remarks. According to the diagnosis of Gutu (1972), the Synapseudidae includes metapseudids with fused pleonites. Two of the genera, Cryptapseudes Băcescu, 1976 and Curtipleon Băcescu, 1976, have all the pleonites fused with the pleotelson into one solid segment, while two others - Vicinisyndes Gutu 2007 and Creefs n. gen - have only one free pleonite. The number of pleonites fused with pleotelson varies in members of Synapseudes (Table 4). Usually members of this genus have two or three free pleonites, but the Subantarctic S. ideos Gardiner, 1973 has two fully-developed pleonites, the other three being apparently partially fused with the pleotelson. Another exception is S. shiinoi Riggio, 1973 that has only one free pleonite. Within the genus, these two species have also an unusual number of segments in both flagella of the antennule, a different number of articles in the antenna, unusual setation of the mandible palp, and a distinct number of segments in both rami of the uropods. A revision of the genus is beyond the scope of this paper but in all probability S. ideos and S. shiinoi are representatives of two distinct genera.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F11331963638823CA7E6FFB67.taxon	diagnosis	Diagnosis (amended after Larsen 2002). Body strongly calcified, tapering from anterior to posterior. Cephalothorax large, almost half as long as rest of body, usually with central carina that is bifurcated in front; branchial chambers swollen, wider than anterior part of carapace; six free pereonites with spines or tubercles dorsally, pereonite 4 little longer than others; pereonite 6 half as long as pereonite 5. Pleonites and pleotelson fused. Antennule well developed with a few strong apophyses on inner margin, three distal apophysis especially strong. Antenna small, first two articles with spines. Maxilliped endites very small, but retain setation typical for this part of appendages. Maxillula outer and inner endites with eight and five spines or setae respectively. Cheliped lacking exopodite, with strong apophysis on basis. Pereopods of walking type. Uropods uniramous, endopod with three to five segments. Male similar to female but chelipeds asymmetrical.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F11331963638823CA7E6FFB67.taxon	materials_examined	Type species: Synapseudes carinatoides Bâcescu, 1976 Species included: C. carinatoides (Bâcescu, 1976); C. carinatum (Makkavyeva, 1971); C. heterochelatum Larsen, 2002; C. loerzae Bamber, 2008 and C. chadi n. sp.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F11331963638823CA7E6FFB67.taxon	discussion	Remarks. The genus Curtipleon was erected by Bâcescu (1976 b) based mainly on the fully fused pleon, the absence of pleopods and the reduced uropod; later on Sieg (1983) designated C. carinatoides (Bâcescu, 1976 a) as the type species. All of the species come from the tropical Indian Ocean (Andaman Sea, Gulf of Aden, Tanzania) and East Australia. Three of them are associated with coral reefs (Makkavyeva 1971, Larsen 2002, this paper), C. carinatum with sand with rocks (Băcescu 1976 b), while C. loerzae was found among sponges, phoronids, bryozoans and algae (Bamber 2008).	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1133197B638826F17910FE00.taxon	materials_examined	Material examined: Holotype, female (MTQ W 34235), HI 10 - 55 B, 23 ° 26 ' 59.35 " S 151 ° 54 ' 45.18 " E, Heron I., Channel between Heron Island and Wistari Reef, reef channel, small coral rubble and sand, depth 30 m, 25 November 2010, coll. C. Buxton & M. Błażewicz-Paszkowycz. Allotype, male (MTQ W 34236), HI 10 - 55 A, 23 ° 26 ' 59.35 " S 151 ° 54 ' 45.18 " E, Channel between Heron Island and Wistari Reef, reef channel, large coral rubble, depth 30 m, 25 November 2010, coll. C. Buxton & M. Błażewicz-Paszkowycz. Paratypes, 4 females (MTQ W 34237), HI 10 - 55 B, the same locality; 5 females (MTQ W 34238), HI 10 - 55 A, the same locality; 1 female (MTQ W 34239), HI 10 - 009 E, 23 ° 25 ' 53.76 " S 152 ° 2 ' 57.48 " E, Sykes Reef W, coral rubble on sand, reef slope, depth 12 m, 14 November 2010, coll. C. Buxton; 1 female (Reg.), HI 10 - 31 B, 23 ° 15 ' 44.28 " S 151 ° 55 ' 32.16 " E, Broomfield, back reef, coral rubble in bommie crevasse, depth 6.5 m, 17 November 2010, coll. C. Buxton; 1 female (MTQ W 34241), HI 10 - 19 B, 23 ° 35 ' 35.52 " S 15 ° 23 ' 55.80 " E, Lamont Reef SE, reef wall, coarse sand and rubble at base of wall, depth 28 m, 15 November 2010, coll. M. Błażewicz-Paszkowycz & C. Buxton.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1133197B638826F17910FE00.taxon	diagnosis	Diagnosis. Frontal margin of carapace concave, smooth with one central spine and small spines on lateral edge. Antennule peduncle article 1 with numerous inner spines; distal process well-developed with strong spines. Antenna article 1 with large and heavily serrated process; article 2 with two or three spines on each of margins; cheliped basis with strong apophysis and disto-medially with prominent, distally serrated apophysis. Pereopods 1 to 6 basis dorsal margin with a few blunt apophysis; pereopods 1 to 3 three times as long as wide, with two rows of six ventral spines; maxillule palp with five setae; uropod endopod with five segments.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1133197B638826F17910FE00.taxon	etymology	Etymology. The species is named after Chad Buxton, who kindly helped to collect and sort the tanaidaceans in the field.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1133197B638826F17910FE00.taxon	description	Description of female. Body (Figs 5 A, B) 2.1 mm long, 3.3 times as long as wide, strongly calcified. Carapace 25 % of total body length, with median carina bifurcating to reach antennule insertion; anterior margin concave, smooth, with small, central spine and minute lateral spines; eye lobes present, eye with dark pigment. Pereon tapering from anterior to posterior; pereonite 1 0.3 times as long as wide, with median ridge, and with seven apophyses along anterior margin; pereonites 2 and 3 similar in length, with seven apophyses along anterior margin and two on posterior margin; pereonite 4 longest, 1.3 times as long as pereonite 3, with four apophyses along anterior margin, four apophyses in posterior half, and three median, red spots; pereonite 5 0.6 times as long as pereonite 4, disposition of apophyses as on pereonite 4; pereonite 6 shortest, 0.5 times as long as pereonite 5, with four apophyses. Antennule (Fig. 8 A) peduncle article 1 robust, 3.4 times as long as wide, with three strong, acute spine-like apophyses on outer margin and rows of spines along article, distal margin with robust spinose apophysis and three plumose setae; article 2 1.2 times as long as wide, 0.3 times as long as article 1, with one simple and one penicillate seta distally; article 3 much narrower, 0.3 times as narrow as article 2; article 4 with one simple and one plumose setae on outer margin; outer flagellum with 5 segments; segments 1, 2 and 3 each with three simple setae and one aesthetasc distally; segment 4 with one simple distal seta; segment 5 with six simple distal setae; inner flagellum with three segments; all segments with two or three simple distal setae. Antenna (Fig. 8 E) with five-articulated peduncle; article 1 square, with large rounded and heavily serrated distally process; article 2 almost twice as long as wide, with two spines on outer margin, and with three spines accompanied with two fine setae on inner margin; article 3 0.3 times as long as article 2, with one penicillate seta on outer margin; article 4 half as long as article 2, with one penicillate seta on inner distal corner; article 5 little shorter than article 4, with four penicillate setae distally; flagellum with one tiny segment, bearing two fine, simple setae Mouthparts. Labrum (Fig. 7 F) rounded, naked. Right mandible (Fig. 7 D) incisor with four blunt teeth, setiferous lobe with four trifurcate setae, molar (Fig. 7 C) serrated distally; palp (Fig. 7 A) three-articled; article 1 robust, with five dents and simple seta on inner margin and with two spines and one simple setae distally; article 2 with two rows of pinnate lateral setae and one simple subdistal seta; article 3 with row of pinnate setae along inner margin and with single distal seta. Left mandible (Fig. 7 B) with crenulated outer margin and with row of five trifurcated setae and three simple setae on setiferous lobe, lacinia mobilis armed with four teeth. Maxillule (Fig. 7 G) inner endite with five plumose setae distally; outer endite with nine spines, with both margins finely setullated; endites with minute setae on both margins; palp (Fig. 7 J) with two articles, distal article tipped with five terminally serrated setae. Maxilla (Fig. 7 H) outer lobe of moveable endite with five simple setae distally and two strong simple setae subdistally; inner lobe of moveable endite with several simple setae distally and two strong simple setae subdistally; outer lobe of fixed endite with six simple setae and three leaf-shaped setae distally and one subdistal plumose seta; inner lobe of fixed finger with six plumose distal setae and with row of about 25 simple subdistal setae. Epignath (Fig. 7 E) with long, plumose seta distally and four short, simple setae subdistally. Maxilliped (Fig. 7 I) palp article 1 with two fine setae in outer distal corner; article 2 with outer margin serrated and with one fine seta in outer distal corner, inner margin crenulated, with numerous simple setae and blunt, short spines; article 3 with two rows of strong, simple setae on inner margin; article 4 with eight plumose setae distally and five subdistally; endite (Fig. 7 K) with four couling-hooks and nine plumose setae on inner margin and simple setae along outer margin, and with five spines and four bifurcate setae distally, and two long plumose setae subdistally. Cheliped (Fig. 8 C) basis 1.6 times as long as wide ventral margin with four apophysis, dorsal margin with four apophysis and seven small spines; dorsodistal margin heavily serrated and supported with strong spinose process; exopodite absent; merus quadrangular, with four spines and six fine setae on ventral margin; carpus three times as long as wide, with three simple ventral setae, and one simple dorsodistal seta; propodus half as long as carpus, with one simple ventral seta and three simple setae on dorsal margin; fixed finger 0.75 times as long as propodus, with six fine ventral setae and with four fine median setae, cutting edge serrated, with six fine setae; dactylus with fine seta dorsally, and two simple setae near unguis; unguis 0.1 times as long as dactylus. Pereopod 1 (Fig. 9 A) basis 4.7 times as long as wide, with eight simple setae along ventral margin, and with four small and four large rounded linguiform apophyses, interspersed with simple setae on dorsal margin; ischium half as long as wide, naked; merus 2.3 times as long as wide, with three fine setae and one pinnate spine ventrally, and with fine seta on dorsal margin; carpus 1.2 times as long as merus, with two rows of six pinnate spines and two pinnate setae, and with three fine setae dorsally; propodus (Fig. 9 A’) similar in length to carpus, with two parallel rows of six and five respectively pinnate spines and with three plumose distal setae, two fine and one penicillate dorsal setae and tuft of pinnate setae distodorsally; dactylus half as long as propodus, with two simple dorsal setae and two simple ventral setae accompanied with apophysis. Pereopod 2 (Fig. 9 B) basis 5.2 times as long as wide, with four fine ventral setae and with six small and three large linguiform apophyses, interspersed with simple setae dorsally; ischium 0.7 times as long as wide, with fine seta ventrally; merus almost twice as long as wide, with four fine setae and one pinnate ventral spine and with two fine dorsal setae; carpus 1.3 times as long as merus, with five pinnate spines and two plumose setae on ventral margin, and with five simple dorsal setae; propodus similar in length to carpus, 2.6 times as long as wide, with six pinnate spines, three fine and one plumose ventral setae, and with two simple and one penicillate seta on dorsal margin as well as with two simple setae, near dactylus insertion; dactylus as on pereopod 1, but only one fine ventral seta. Pereopod 3 (Fig. 9 C, C’) similar to pereopod 2. Pereopod 4 (Fig. 9 D) basis 4.3 times as long as wide, with five simple and one penicillate seta on ventral margin, and with three large and one small blunt apophyses interspersed with simple setae, and three proximal penicillate setae on dorsal margin; ischium 0.6 times as long as wide, naked; merus almost twice as long as wide, with three simple setae and one pinnate spine ventrally, and with one simple seta dorsally; carpus 1.7 times as long as merus, with eight pinnate spines and four simple setae ventrally, and with two simple setae dorsally; propodus (Fig. 9 D’) similar in length to carpus, four times as long as wide, with six pinnate spines and six simple setae ventrally, and with one simple seta and one penicillate seta dorsally, and with tuft of pinnate setae distodorsally; dactylus as on pereopod 1. Pereopod 5 (Fig. 9 E) basis 5.7 times as long as wide, with five blunt apophyses and with seven fine ventral setae, one simple and three proximal penicillate setae on dorsal margin; ischium 1.3 times as long as wide, with two fine ventral setae; merus 2.2 times as long as wide, with two fine setae and two spines ventrally; carpus twice as long as merus, with six pinnate spines and four fine setae on ventral margin and with two simple dorsal setae; propodus 1.2 times as long as carpus, 4.2 times as long as wide, with five pinnate spines and four simple setae along ventral margin and with four simple and one penicillate seta along dorsal margin; dactylus as on pereopod 3. Pereopod 6 (Fig. 9 F) similar to pereopod 5, but two dorsal seta on dactylus. Uropods (Fig. 8 E) uniramous; basis 1.8 times as long as wide, endopod with five segments, segment 4 with two simple setae distally; segment 5 with four fine terminal setae. Description of male (Fig. 6 A, B) body similar to that of female, 2.5 mm long; antennule similar to that of female, but peduncle with fewer spines, and proximally-crenulated inner margin; cheliped (Fig. 8 D) larger, with only three spines on carpus.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1133197B638826F17910FE00.taxon	distribution	Distribution: Known only from the southern part of the Great Barrier Reef from the vicinity of Heron Island, in depths from 6 to 30 m, from coral rubble and coarse sand.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1133197B638826F17910FE00.taxon	discussion	Remarks. Curtipleon chadi n. sp. is the fifth member of the genus and the second species collected in Australian waters. The new species can be distinguished from the other members by its having five segments in the endopod of the uropods (Table 3). From C. loerzae, occurring in the East Australia, C. chadi differs by having numerous strong spines on the proximal peduncle article of the antennule and only a few strong spines on article 2 of the antennule. Additionally, C. chadi has an acute process in the distal part of the cheliped basis, which is not observed in any other species. Strong calcification of the cheliped may obstruct the observation of this feature; if only this process is absent in all other species it is a valuable species-specific character.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1133197B638826F17910FE00.taxon	diagnosis	Diagnosis. Body dorso-ventrally flattened, 6.5 times as long as wide. Eyes present. Pereonites wider than long. Pleon of one free pleonite and pleotelson. Antennule basis serrated, outer flagellum of three and inner of two segments. Antenna reduced, of two articles only; distal article rudimentary. Female cheliped slender, without exopodite. Pereopods 1 to 3 with hook-like apophyses on basis and serrated dorsal margin of carpus. Pereopods 4 to 6 propodus ventral margin with two simple setae. Pleopods absent. Uropod biramous. Male similar to female, but cheliped robust, and carpus of pereopod 1 dorsally expanded and hirsute.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1133197B638826F17910FE00.taxon	materials_examined	Type species: Creefs heronum n. gen n. sp. by monotypy Gender: masculine	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1133197B638826F17910FE00.taxon	etymology	Etymology. The name refers to the program Census of Coral Reef Ecosystem (CReefs) being a part of the Census of Marine Life (CoML) program, during which the type species was collected.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F1133197B638826F17910FE00.taxon	discussion	Remarks. With only one free pleonite Creefs n. gen. fits the subfamily Synapseudinae. At first glance Creefs n. gen. is most similar to Vicinisyndes Gutu, 2007, to which it is probably closely related (Table 4). The most striking character of the new genus is the biarticulated peduncle of the antenna, while that of Vicinisyndes consists of five articles. In addition, the new genus has serrated inner margins of the antennules, these being apparently smooth in Vicinisyndes, and it has a wide, dorsally serrated carpus on pereopods 1 to 3, while in Vicinisyndes these are apparently simple and slender. Additionally, Creefs has two simple setae on the ventral margin of the propodus of pereopods 4 to 6 in contrast to the spines present there in Vicinisyndes. character Cryptapseudes Curtipleon Synapseudes Vicinisyndes Creefs	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F112B1970638821FF7E19FC5E.taxon	materials_examined	Material examined: Holotype, female (MTQ W 34242), HI 10 - 009 E, 23 ° 25 ' 53.76 " S, 152 ° 2 ' 57.48 " E, Heron I., Sykes Reef W, coral rubble on sand, reef slope, depth 12 m, 13 November 2010, coll. C. Buxton. Allotype, male (MTQ W 34243), HI 10 - 009 E, the same locality as holotype. Paratypes, 1 female (MTQ W 34244), HI 10 - 020 B, 23 ° 36 ' 19.26 " S, 152 ° 1 ' 26.04 " E, Heron I., Lamont Reef, coarse sand, reef flat, depth 13 m, 16 November 2010, coll. C. Buxton; 1 female (MTQ W 34245), HI 10 - 055 B, 23 ° 26 ' 59.35 " S, 151 ° 54 ' 45.18 " E, Heron I., Channel between Heron Island and Wistari Reef, coral rubble on sand, reef channel, depth 30 m, 26 November 2010, coll. C. Buxton. 25 females (MTQ W 34246) HI 10 - 009 E, the same locatity as holotype; 23 females (MTQ W 34247) HI 10 - 009 D the same lacality as holotype, but reef flat, depth 14 m.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F112B1970638821FF7E19FC5E.taxon	diagnosis	Diagnosis. as for the genus, plus uropod endopod and exopod with four and three segments respectively	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F112B1970638821FF7E19FC5E.taxon	description	Description of female. Body (Fig. 10 A, B) 2 mm long, 6.5 times as long as wide. Carapace 20 % of total body length, without rostrum, eyes present, with pigment. Pereonite 1 0.4 times as long as wide, with short anterolateral seta; pereonites 2, 3 and 4 half as long as wide, each with short anterolateral seta, pereonite 5 half as long as wide, with two simple setae on each margin; pereonite 6 similar to pereonite 1. Pleon and pleotelson 16 % of total body length, pleonite 1 free, 0.2 times as long as wide, with two setae on each margin, other pleonites fused with pleotelson; pleotelson with a pair of lateral setae and with protruding rounded tip. Antennule (Fig. 12 A) peduncle article 1 4.5 times as long as wide, both margins serrated with three penicillate setae and two simple setae on outer margin, and with three simple setae and one spine on inner margin; article 2 1.3 times as long as wide, 0.2 times as long as article 1, with three teeth and three simple setae distally on outer margin, and with two simple setae on inner margin; article 3 elongated, 3.4 times as long as wide, with one simple seta on outer margin, and two simple setae on inner margin; article 4 with one simple distal seta on inner margin; outer flagellum with three segments, segment 1 with two distal setae; segment 2 with one simple seta and one aesthetasc distally; segment 3 with four simple distal setae; inner flagellum two-segmented, segment 1 with two, segment 2 with three simple distal setae. Antenna (Fig. 12 C) with two peduncle articles, article 1 robust, 1.6 times as long as wide, with two proximal spines and two penicillate distal setae on outer margin, and with three spines proximally, and three simple setae on inner margin; article 2 as long as wide, with two simple setae. Mouthparts. Epistome conspicuous (Fig. 12 A). Right mandible (Fig. 12 G) with finely serrated margin, setiferous lobe with one robust bifurcated seta and two pinnate setae; molar (Fig. 12 H) serrated distally, with two simple subdistal setae; palp (Fig. 12 F) with three articles, articles 1 and 2 4.4 times as long as wide, naked; article 3 0.4 times as long as article 2, with one short, one long pinnate distal setae. Left mandible (Fig. 12 E) with finely serrated inner margin, lacinia mobilis with five teeth, setiferous lobe with two simple setae. Maxillule (Fig. 12 J) outer endite with eight distal spines, inner endite with four simple setae; palp (Fig. 12 K) two-articled, article 2 with two simple distal seta. Maxilla outer margin serrated (Fig. 12 L) outer lobe of movable endite with four distal and two subdistal simple setae; inner lobe of movable endite with four simple setae; outer lobe of fixed endite with four simple and two trifurcated setae distally and with one plumose seta subdistally; inner lobe of fixed endite with six plumose and row of simple setae; inner margin with four spines. Epignath (Fig. 12 D) narrow, with setose margins and with one terminal seta. Maxilliped palp (Fig. 12 M) article 1 with one strong seta on outer margin; article 2 1.2 times as long as wide with serrated margin and with one distal spine, inner margin with row of nine simple setae; article 3 0.4 times as long as article 2, with row of six simple setae on inner margin; article 4 1.2 times as long as article 3, with 8 pinnate distal setae; endite (Fig. 12 I) with four couling-hooks and with three pinnate setae on inner margin, with five setulose spines and three pinnate distal setae and with row of simple setae on outer margin. Cheliped (Fig. 13 A) basis 1.4 times as long as wide, with one simple distal seta on ventral margin; merus 2.3 times as long as wide, with one fine ventral seta; carpus five times as long as wide, with two simple setae along ventral margin and one dorsal seta; propodus 2.3 times as long as wide, with one simple seta on ventral margin, and two plumose setae near dactylus insertion; fixed finger half as long as propodus, with three simple setae on ventral margin, cutting edge with four simple setae, serrated. Pereopod 1 (Fig. 14 A): basis 3.8 times as long as wide, with two simple setae and two hook-like apophyses on dorsal margin, and with one simple seta on ventral margin distally; ischium 0.2 times as long as wide, with one simple seta on ventral margin; merus 1.6 times as long as wide, with one fine seta dorsally, with three simple setae and one small spine on ventral margin; carpus 1.6 times as long as wide, both margin serrated with three simple setae and one spine on dorsal margin, with two simple setae and three spines ventrally; propodus similar in length to carpus, half as wide as carpus, with one penicillate dorsal seta, ventral margin serrated with four spines, one simple, one pinnate setae, near dactylus insertion one simple setae and two spines; dactylus together with unguis half as long as propodus, with one dorsal and one ventral fine seta; unguis half as long as dactylus. Pereopod 2 (Fig. 14 B) basis three times as long as wide, with two processes proximally, one hook-like apophysis in the middle of length and two simple and one penicillate seta dorsally and with one simple ventral seta; ischium 0.3 times as long as wide, with one simple ventral seta; merus 1.7 times as long as wide, with one dorsal and two ventral simple setae; carpus 1.6 times as long as wide, both margin serrated, with one simple dorsal seta and with four spines, with two ventral simple setae, and with two spines and one simple seta distally; propodus 3.3 times as long as wide, half as wide as carpus, with one penicillate seta on dorsal margin, with four spines and denticles on ventral margin and with two spines, one pinnate and one plumose seta distally; dactylus together with unguis half as long as propodus, with one dorsal seta and two ventral simple setae; unguis half as long as dactylus. Pereopod 3 (Fig. 14 C) similar to pereopod 2, but basis 4.5 times as long as wide, with only one proximal spine and with two penicillate dorsal setae. Pereopod 4 (Fig. 14 D) basis five times as long as wide, with two penicillate ventral setae; ischium 0.4 times as long as wide; merus 1.5 times as long as wide, with one simple dorsal seta and two simple ventral setae; carpus 2.5 times as long as wide, with one simple dorsal seta, and with two spines and four simple setae distally; propodus (Fig. 14 D’) 3.5 times as long as wide, similar in width to carpus, with eight pinnate setae and one simple seta distally, and four pinnate subdistal setae; dactylus together with unguis half as long as propodus, with one ventral and one dorsal simple seta; unguis 0.6 times as long as dactylus. Pereopod 5 (Fig. 14 E) basis four times as long as wide, with two penicillate setae and one simple setae dorsally, and with two simple ventral setae; ischium 0.3 times as long as wide, with one simple ventral seta; merus 2.6 times as long as wide, with one dorsal and ventral simple seta; carpus twice as long as wide, with one dorsal, two ventral simple setae and with one ventral spine; propodus 4.4 times as long as wide, similar in width to carpus, with one simple ventral and with three pinnate setae and three simple setae distally; dactylus together with unguis 0.7 times as long as propodus, with one ventral and one dorsal setae; unguis 0.3 times as long as dactylus. Pereopod 6 (Fig. 14 F) basis 3.3 times as long as wide, with one simple seta on ventral distal margin and penicillate dorsal seta; ischium 0.3 times as long as wide, with simple setae ventrally; merus twice as long as wide, with three simple distal seta; carpus 1.5 times as long as wide, with three long and one short distal setae; propodus 4 times as long as wide, 0.7 times as wide as carpus, with one penicillate seta on dorsal margin, with two simple setae along ventral margin and one fine setae distally; dactylus as in pereopod 5. Uropods (Fig. 13 D) basis 5.6 times as long as wide, with one simple distal seta and distodorsal hook-shape process; endopod with four segments, segment 2 with one simple and one penicillate setae distally; segment 3 with three, segment 4 with four simple setae distally; exopod with three segments, segment 3 with two simple distal setae. Description of male. Body (Fig. 11 A) 1.5 mm long. Generally similar to female, but differing in shape of antennule and cheliped. Antennule (Fig. 12 B) similar to that of female, but peduncle article 1 6.6 times as long as wide, without spines or denticles on inner margin; article 2 twice as long as wide; article 3 4.6 times as long as wide, with additional denticles on inner margin. Cheliped (Fig. 13 B) stronger than in female; basis 1.2 times as long as wide, with two simple setae on ventral margin, and with row of denticles on dorsal margin; merus twice as long as wide, with one simple ventral seta; carpus 1.8 times as long as wide, with five simple setae and denticles along ventral margin, and with one simple dorsodistal seta; propodus robust, as long as wide, with one simple seta on ventral margin and with two simple setae near dactylus insertion; fixed finger 0.6 times as long as propodus, with three setae ventrally and with four short and three long setae on cutting edge; dactylus 0.7 times as long as propodus, with two teeth and two short simple setae on inner margin. Pereopod 1 (Fig. 13 C) similar to female, but carpus more robust, 1.5 times as long as wide, with denticles and row of 18 setae along dorsal margin and with serrated ventral margin.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F112B1970638821FF7E19FC5E.taxon	distribution	Distribution. Known only from the type locality (Heron Island, GBR) from coral rubble on sand. Subfamily Msangiinae Gu t u, 2006	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F11201976638826007DAAFC14.taxon	materials_examined	Material examined: Holotype, female (MTQ W 34248), HI 10 - 31, 23 ° 15 ' 44.28 " S, 151 ° 55 ' 32.16 " E, Heron I., depth 6.4 m, 18 November 2010, coll. V. N. Ivanenko, M. Wakeford. Paratypes: 18 females (MTQ W 34249), HI 10 - 31; 2 female (MTQ W 34250), HI 10 - 009 E, 23 ° 25 ' 53.76 " S, 152 ° 2 ' 57.48 " E Heron I., Sykes Reef W, reef slope, coral rubble on sand, depth 12 m, 13 November 2010, coll. C. Buxton. 2 females (MTQ W 34251) HI 10 - 009 D the same lacality as holotype, but reef flat, depth 14 m.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F11201976638826007DAAFC14.taxon	diagnosis	Diagnosis. Body 10.6 times as long as wide, carapace 1.3 times as long as wide, eyes present with ommatidia. Pleonites shorter than pereonite 6. Antenna peduncle with four articles; inner margin of proximal antennular peduncle smooth. Chela twice as long as wide. Pereopod 1 - 3 robust, with merus wider than long; pereopod 1 - 2 with two and one spine on ventral margins. Uropod with four and three segments in endopod and exopod, respectively.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F11201976638826007DAAFC14.taxon	etymology	Etymology. The species name is given in relation to its habitat, which was a head of a live coral of the family Mussidae.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F11201976638826007DAAFC14.taxon	description	Description of female. Body (Figs. 15 A, B) 1 mm long, 10.6 times as long as wide. Carapace 14 % of total body length; rostrum rounded; eye with dark pigment. Pereonites rectangular; pereonite 1 half as long as wide, with three setae laterally; pereonites 2, 3 and 4 0.7 times as long as wide, with three simple lateral setae; pereonites 5 and 6 similar in length, 0.6 times as long as wide, with two simple lateral setae. Pleon 40 % of total body length; all pleonites similar in length, half as long as wide and shorter than pereonite 6, with one fine lateral seta; pleotelson 0.6 times as long as all pleonites together. Antennule (Fig. 17 A) peduncle article 1 margin smooth, three times as long as wide, with one outer and inner simple seta distally; article 2 0.3 times as long as article 1, with three simple setae on outer distal margin and with one simple seta on inner margin; article 3 with one simple setae on outer and inner margin distally; article 4 with one simple setae on inner margin. Outer flagellum with three segments; segment 1 with simple seta and one aesthetasc distally; segment 2 with two simple setae and one aesthetasc distally; segment 3 with three simple setae. Inner flagellum with two segments; segment 1 with two, segment 2 with four simple setae distally. Antenna (Fig. 17 B) peduncle with four articles; article 1 almost as long as wide, with rounded flat apophysis and with minute setae along outer margin; article 2 half as long as article 1, with minute setae along outer margin and in inner distal corner; article 3 2.2 times as long as wide, with one simple distal seta; article 4 partially fused to article 3 with one simple distal seta. Mouthparts. Right mandible (Fig. 16 A) bearing incisor with blunt tooth, setiferous lobe with two bifurcated and one robust trifurcated setae; palp (Fig. 16 C) with three articles; article 1 1.5 times as long as wide, naked; article 2 1.6 times as long as article 1, with two pinnate setae distally; article 3 0.3 times as long as article 2, with one long, pinnate seta, and three short, simple setae distally. Left mandible (Fig. 16 B) with blunt incisor; lacina mobilis with six denticulation; setiferous lobe with two pinnate setae. Molar not seen. Maxillule inner endite (Fig 16 E) with four simple setae; outer endite (Figs. 16 D, D’) with nine terminal spines. Maxilla (Fig. 16 F) outer lobe of movable endite with three simple setae distally and two simple setae subdistally; inner lobe of movable endite with five simple setae distally; outer lobe of fixed endite with four simple setae and two pinnate setae distally; inner lobe of fixed endite with some simple distal setae and row of five simple subdistal setae. Labium (Fig. 16 G) lobe with four combs of simple setae along outer margin; palp with fine lateral setules and one simple seta distally. Maxilliped palp (Fig. 16 I) article 1 0.8 times as long as wide, with one simple seta on inner margin and with one strong simple seta on outer margin distally; article 2 twice as long as article 1, with row of simple setae on inner margin and with one simple seta on outer distal margin; article 3 0.7 times as long as article 2, with row of five strong setae on inner margin; article 4 0.6 times as long as article 3, with row of six simple setae distally. Endite (Fig. 16 H) with three spines, one simple and one pinnate setae on inner margin, with two couling-hooks on outer margin, and with three simple setae, one spine, one forked and one pinnate setae distally. Cheliped (Fig. 17 D) basis 1.2 times as long as wide, with two simple ventral setae (one central, one distal); exopodite absent; merus 2.5 times as long as wide, with three simple setae on ventral margin; carpus 4.8 times as long as wide, with two simple setae ventrally and comb of five simple setae on distal ventral corner, and with two simple distodorsal setae; propodus 1.2 times as long as wide, with one ventral and two dorsal simple setae and one simple setae near dactylus insertion; fixed finger half as long as wide, with five simple setae on cutting edge and four simple setae on ventral margin; dactylus 0.6 times as long as propodus with crenulated inner margin; unguis 0.3 times as long as dactylus, with three processes on inner margin. Pereopod 1 (Fig. 18 A) basis 2.6 times as long as wide, with two penicillate setae on dorsal margin; ischium half as long as wide, with one simple ventral seta; merus 1.2 times as long as wide, with one simple distal seta on each of ventral and dorsal margins; carpus 0.6 times as long wide, with three spines and four simple setae on distal margin; propodus 1.4 times as long as carpus, with two spines and one pinnate setae on ventral margin, and with one penicillate seta, one simple seta and two spines on dorsal margin; dactylus together with unguis 1.2 times as long as propodus, with one simple seta on dorsal margin; unguis almost as long as dactylus. Pereopod 2 (Fig. 18 B) basis 2.6 times as long as wide, with two penicillate setae on dorsal margin and two simple setae distally; ischium twice as long as wide, with one simple ventral seta; merus 1.2 times as long as wide, with one simple seta on ventral margin; carpus 0.7 times as long as wide, with three spines and two simple setae distally; propodus 1.2 times as long as carpus, with two spines on ventral margin, and with two spines, one pinnate and one simple setae near dactylus insertion; dactylus together with unguis 1.2 times as long as propodus, with simple seta on dorsal margin, unguis 0.7 times as long as dactylus. Pereopod 3 (Fig. 18 C) similar to pereopod 2, but basis 2.3 times as long as wide; unguis 0.4 times as long as dactylus. Pereopod 4 (Fig. 18 D) basis 2.5 times as long as wide, with two penicillate setae on dorsal margin, and with one simple ventrodistal seta; ischium 0.6 times as long as wide, with one simple seta on each of ventral and dorsal distal margins; merus 1.2 times as long as wide, with one simple seta on dorsal margin, and with two simple setae on ventral margin; carpus as long as merus, with two spines and two simple setae distally; propodus as long as carpus, with one spine and one simple and four pinnate setae distally; dactylus together with unguis 1.1 times as long as propodus, with one ventral margin and one dorsal fine seta; unguis 0.8 times as long as dactylus. Pereopod 5 (Fig. 18 E) basis 2.6 times as long as wide, with two penicillate and two simple setae on dorsal margin and with two simple ventral setae; ischium as long as wide, with two dorsal and one ventral simple setae; merus 1.7 times as long as wide, with one dorsal and one ventral simple setae; carpus 1.3 times as long as wide, with one dorsal and one ventral simple seta; propodus 2.5 times as long as wide, 1.6 times as long as carpus, with one penicillate dorsal seta, one simple ventral seta and with four pinnate setae near dactylus insertion; dactylus together with unguis 0.6 times as long as propodus, with one dorsal and one ventral simple setae; unguis 0.3 times as long as dactylus. Pereopod 6 (Fig. 18 F) basis 2.5 times as long as wide, with one simple seta and two penicillate setae on dorsal margin, and with two simple setae on ventral margin; ischium 1.2 times as long as wide, with two dorsal and one ventral simple setae; merus twice as long as wide, with one dorsal and two ventral simple setae; carpus 1.8 times as long as wide, with one dorsal and three ventral simple setae; propodus 3.2 times as long as wide, 1.4 times as long as carpus, with one penicillate dorsal seta and with four distal pinnate setae; dactylus together with unguis 0.7 times as long as propodus, with one dorsal and one ventral seta; unguis 0.7 times as long as dactylus. Pleopods (Fig. 17 C) all similar; basis twice as long as wide, naked; endopod with two articles, first article with one distal simple seta, second article with two simple setae; exopod with one article, with three simple setae. Uropods (Fig. 17 E) basis 1.5 times as long as wide, with two simple distal setae on outer margin; endopod with four segments; segments 2 and 3 each with simple distal seta, segment 4 with three simple distal setae; exopod with three segments, segment 3 with two simple distal setae.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
03DB878F11201976638826007DAAFC14.taxon	discussion	Remarks. Msangia mussida n. sp. is the fourth member of the genus. M. larvoides Băcescu, 1976 occurring in the vicinity of Mbudya Island (Indian Ocean) has five segments in the endopod and two segments in the exopod of the uropod, while Msangia bacescui Gutu, 2006 inhabiting corral rubble in coasts of Tanzania has five and three segments in the uropod endopod and exopod respectively. The only species that has the same number of segments in the uropod, i. e. four in the endopod and three in the exopod, is M. tarangensis Gutu & Angsupanich, 2006, which was recorded from coral rubble on sand in the Andaman Sea (Gutu & Angsupanich 2006). From the last species M. mussida can be distinguished by the presence of three setae on the endopod of the pleopod (five in M. tarangensis). The new species also has quite robust pereopods, in which the carpus is wider than long in contrast to M. tarangensis. In addition, the new species has two and one spines respectively on the ventral margin of the carpus and the propodus of the first two pairs of pereopods, while there are three and two spines on those articles respectively in M. tarangensis.	en	Stępień, Anna, Błażewicz-Paszkowycz, Magdalena (2013): Four new species and two new genera of Metapseudidae (Crustacea: Tanaidacea: Apseudomorpha) from Australian coral reefs. Zootaxa 3717 (4): 559-592, DOI: 10.11646/zootaxa.3717.4.7
