identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4F1FA810C39BC570692482188FFC2F59.text	4F1FA810C39BC570692482188FFC2F59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liolaemus chavin	<div><p>Liolaemus chavin sp. n. Figure 8</p><p>Liolaemus alticolor Lehr 2002</p><p>Liolaemus incaicus Lobo, Quinteros and Díaz Gómez 2007</p><p>Liolaemus aff. walkeri Langstroth 2011</p><p>Holotype.</p><p>MUSM 25417, adult male collected at Conococha, Recuay Province, Ancash Department, Peru, 10.123S, 77.293W, elevation 4100 m, on 31 March 2006 by Mikael Lundberg.</p><p>Paratypes.</p><p>Three males (MUSM 20141, 20143, 20146) and twelve females (MUSM 25324, 25327, 25328, 25331, 25333, 25334, 25340, 25423, 25412, 30812, 30813, BYU 50192) from the same locality as the holotype. One male (MUSM 20147) from Carpa, Recuay Province, Ancash Department, on 28 February 2001 by Edgar Lehr and César Aguilar (see Data resources for elevation and coordinates). One female (MUSM 20201) from La Unión, Huánuco Department, on 3 March 1997 by Edgar Lehr (see Data resources for elevation and coordinates). Seven males (CORBIDI 10439, 10450, 10452, 10442, 10441, 10443, 10437) and six females (CORBIDI 10444, 10451, 10440, 10438, 10445, 10449) from Pampas de Huamani, San Marcos District, Huari Province, Ancash Department, on 12 February 2012 by Pablo J. Venegas (see Data resources for elevation and coordinates).</p><p>Diagnosis.</p><p>Small (61.7 mm maximum SVL), slender Liolaemus closely related to Liolaemus walkeri, Liolaemus tacnae, Liolaemus pachacutec sp. n. and Liolaemus wari sp. n. (described below) (Fig. 1). It differs from Liolaemus walkeri, Liolaemus pachacutec sp. n. and Liolaemus wari sp. n. in the absence of precloacal pores in males. It differs from Liolaemus tacnae in having a melanistic belly in adult males (not melanistic in adult Liolaemus tacnae males).In comparison with other species assigned to the Liolaemus alticolor group, Liolaemus chavin sp. n. differs from Liolaemus bitaeniatus and Liolaemus pagaburoi in having a smooth dorsal surface of the head (rough to slightly rough dorsal surface). It differs from Liolaemus alticolor, Liolaemus aparicioi, Liolaemus incaicus, Liolaemus paulinae, Liolaemus pyriphlogos, Liolaemus puna, and Liolaemus variegatus in the absence of precloacal pores in males. Liolaemus chaltin also lacks precloacal pores in males, but Liolaemus chavin sp. n. differs in having also a melanistic belly in adult males.</p><p>Description of holotype.</p><p>Adult male; SVL 56.8 mm; head length 13.7 mm; head width 11.3 mm; head height 7.7 mm; axilla-groin 21.0 mm (37% of SVL); foot length 10.3 mm (18.3% of SVL); tail length (regenerated) 35.2 mm (0.6 times SVL).</p><p>Fifteen dorsal head scales (from a line drawn horizontally between anterior edges of external auditory meatus to anterior border of rostral). Dorsal head scales smooth except for the interparietal and surrounding scales, scale organs more abundant in prefrontal, internasal, and supralabial regions. Five scale organs on postrostral. Nasal scale in contact with rostral, separated from first supralabial by one scale, nasal bordered by eight scales; canthus separated from nasal by one scale. Six supralabials. Six lorilabial scales, three in contact with the subocular. Six infralabials. Auditory meatus oval (height 2.3 mm, width 1.2 mm), with three small, projecting scales on anterior margin. Seven convex, smooth temporals. Orbit–auditory meatus distance 4.9 mm. Orbit–anterior margin of rostral distance 6.3 mm. Rostral almost three times wider than high (width 2.9 mm; height 1.2 mm). Mental subpentagonal, about two times as wide as high (width 3.2 mm; height 1.7 mm). Interparietal pentagonal with an elongated posterior apex, bordered by eight scales, the parietal slightly smaller. Frontal quadrangular. Supraorbital semicircles complete on both sides. Semicircles formed by 6 scales. Four enlarged supraoculars. Six distinctly imbricate superciliaries on both sides. Eleven upper and ten lower ciliaries. Subocular elongate, 3.8 mm, longer than eye diameter (2.9 mm), separated from supralabials by a single, but interrupted row of lorilabials. Second supralabial elongate, 1.9 mm. Six lorilabials with single and double rows of scale organs. Sixth, fifth and fourth lorilabials contacting subocular. Preocular small, separated from lorilabial row by one scale. Postocular as large as preocular. Mental in contact with four scales: first infralabials (on each side) and two enlarged chin shields. Chin shields forming a longitudinal row of three enlarged scales separated one from the other by seven smaller scales. Scales of throat round, flat, and imbricate. Twenty-four gulars between auditory meatus. Longitudinal neck fold without keeled scales, that are similar to dorsal in size scales. Antehumeral pocket and antehumeral neck fold well developed. Forty-two scales between auditory meatus and shoulder (counting along postauricular and longitudinal neck fold), thirty-two scales between auditory meatus and antehumeral neck fold. Gular folds absent.</p><p>Dorsal scales rhomboidal, keeled, and imbricate. Sixty-six dorsal scales between occiput and level of groin. Sixty-two scales around midbody. Thirty rows of keeled scales on dorsum at midtrunk. Scales become smooth along flank and toward belly. Ventral scales slightly wider than dorsals. Eighty-two ventral scales between mental and cloaca; no precloacal pores. Supracarpals laminar, round, and smooth. Subdigital lamellae of fingers with three keels, in number I: 6; II: 11; III: 14; IV: 15; V: 10 (right hand). Claws moderately long. Supradigital lamellae convex, smooth, and imbricate. Infracarpals and infratarsals keeled, distinctly imbricate. Supratarsals smooth. Subdigital lamellae of toes I: 13; II: 13; III: 13; IV: 12; V: 6 (right foot).</p><p>Color pattern in preservation.</p><p>Dorsal background color from occiput to base of tail greenish brown. Black continuous vertebral stripe present. Dark paravertebral marks. Paravertebral and vertebral fields of same background color. Dorsolateral stripes distinctly cream-color. Small dark cream-colored markings scattered in lateral field. Cream ventrolateral stripe, beginning on the upper auricular meatus, continuing across the longitudinal neck fold, through the shoulders, ending in the groin. Dark and small cream-colored marks in the ventral field. Black ventral color from about second third of head to femur, tibia and first third of tail. Dark and cream-colored small markings in first third of ventral head and two posterior thirds of tail.</p><p>Color pattern in life.</p><p>Head dorsally brown with black and light brown dots. Subocular cream colored, dorsum bisected by a dark vertebral line. Vertebral field not conspicuous, bordering the vertebral line with a tenuous yellowish line. Paravertebral field with dark marks, bordered dorsally by a yellowish cream dorsolateral stripe. Lateral field with black and yellow reticulated pattern and white dots. Inconspicuous ventrolateral stripe, beginning on upper margin of auricular meatus, continuing from the longitudinal neck fold, through the shoulders, ending in the groin. Ventrolateral similar to lateral field but with more white dots. Fore and hind limbs same color as the paravertebral field, with diffuse dorsal markings. Dark, melanistic ventral color from about second third of head to femur, tibia and first third of tail. Dark and white dots in first third of ventral head and two posterior thirds of tail.</p><p>Variation.</p><p>Variation in characters is summarized in Tables 1-4. There is sexual dichromatism. Adult males exhibit melanistic belly, cloacal region and throat, or mela nistic belly only; adult females exhibit black and white spots on belly, cloacal region and throat, or yellowish belly and tail.</p><p>Etymology.</p><p>The specific epithet chavin refers to the pre-Inca culture Chavin, which had its center close to the type locality and frequently depicted reptile figures on some of its most remarkable sculptures. The species name is in the nominative singular.</p><p>Distribution and natural history.</p><p>Liolaemus chavin sp. n. is known from four localities in the central Andes, at elevations of 3535-4450 m in Ancash and Huánuco Departments in western central Peru (Fig. 11). It is the northernmost species of the subgenus Liolaemus .</p><p>Liolaemus chavin sp. n. was found active and under rocks in grassland and shrubland habitats at higher and lower elevations respectively (Fig. 8). In Pampas de Huamani the new species was usually found basking on grass up to 60 cm above the ground, and when they were disturbed they escaped into the base of grass clumps. Individuals basking on rocks were very rare in all localities. On cloudy days we found this species inactive hidden in the base of grass clumps, although some individuals were also found inactive under rocks. This species is viviparous; one female showed two uterine chambers per side with developed embryos, yolk and no visible shell in either chamber, and three females showed two uterine chambers per side with yolk, without developed embryos and no visible shell in each chamber. At the type locality no sympatric species of reptiles were found, but four amphibians are known: Pleurodema marmoratum ( Duméril &amp; Bibron, 1840), Telmatobius mayoloi Salas &amp; Sinsch, 1996, Gastrotheca peruana and Rhinella (Bufo) spinulosa (Wiegmann, 1834) (Lehr 2002; personal observations). Sympatric species at Catac include the anurans Gastrotheca peruana, Rhinella (Bufo) spinulosa, Telmatobius rimac Schmidt, 1954, Telmatobius mayoloi, and the lizard Stenocercus chrysopygus Boulenger, 1900; at Carpa, Gastrotheca peruana (Boulenger, 1900), Rhinella (Bufo) spinulosa and Pleurodema marmoratum; at Pampas de Huamani, Gastrotheca peruana, Pleurodema marmoratum and Rhinella (Bufo) spinulosa; and at La Unión, Gastrotheca griswoldi Shreve, 1941, Gastrotheca peruana, Rhinella (Bufo) spinulosa and Stenocercus chrysopygus (Lehr, 2002).</p></div>	https://treatment.plazi.org/id/4F1FA810C39BC570692482188FFC2F59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aguilar, Cesar;Wood Jr, Perry L.;Cusi, Juan C.;Guzman, Alfredo;Huari, Frank;Lundberg, Mikael;Mortensen, Emma;Ramirez, Cesar;Robles, Daniel;Suarez, Juana;Ticona, Andres;Vargas, Victor J.;Venegas, Pablo J.;Sites Jr, Jack W.	Aguilar, Cesar, Wood Jr, Perry L., Cusi, Juan C., Guzman, Alfredo, Huari, Frank, Lundberg, Mikael, Mortensen, Emma, Ramirez, Cesar, Robles, Daniel, Suarez, Juana, Ticona, Andres, Vargas, Victor J., Venegas, Pablo J., Sites Jr, Jack W. (2013): Integrative taxonomy and preliminary assessment of species limits in the Liolaemus walkeri complex (Squamata, Liolaemidae) with descriptions of three new species from Peru. ZooKeys 364: 47-91, DOI: http://dx.doi.org/10.3897/zookeys.364.6109, URL: http://dx.doi.org/10.3897/zookeys.364.6109
732E9F3E22FC322D109D0DA372864F20.text	732E9F3E22FC322D109D0DA372864F20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liolaemus pachacutec	<div><p>Liolaemus pachacutec sp. n. Figure 9</p><p>Holotype.</p><p>MUSM 29683, adult male collected at Challabamba, Paucartambo Province, Cusco Department, Peru, 13.254S, 71.838W, elevation 4364 m, on 1 April 2009 by César Ramírez .</p><p>Paratypes.</p><p>Three males (MUSM 29681, 29687, 29678) and four females (MUSM 29679, 29689, 29680, 29682) from the same locality as the holotype. Two males MUSM (29665, 29668) and one female (MUSM 29669) from Lamay, Calca Province, Cusco Department, on 12 October 2009 by César Ramírez (see Data resources for elevations and coordinates). One male (MUSM 29664), two females (MUSM 29688, BYU 50237) and one juvenile (MUSM 31412) from Pisac, Calca Province, Cusco Department, on 4 July and 11 October 2009 by César Ramírez, and on 28 June 2012 by César Aguilar, Perry Wood and Juan Carlos Cusi (see Data resources for elevations and coordinates). One male (MUSM 31540), two females (MUSM 31538-39) and one juvenile (MUSM 31537) from Tiaparo, Pocohuanca District, Aymaraes Province, Apurímac Department, on 11 June 2013 by Alfredo Guzmán (see Data resources for elevations and coordinates).</p><p>Diagnosis.</p><p>Small (51.9 mm maximum SVL) Liolaemus closely related to Liolaemus chavin sp. n., Liolaemus tacnae, Liolaemus walkeri, and Liolaemus wari sp. n. (described below) (Fig. 1). It differs from Liolaemus chavin sp. n. and Liolaemus tacnae in having precloacal pores (males). Liolaemus pachacutec differs from Liolaemus wari sp. n. in having a partial or complete melanistic belly in adult males and in lacking a ringed pattern in ventral tail. Liolaemus pachacutec differs from most individuals (90%) of Liolaemus walkeri in lacking spots in the lateral field. In comparison with other species assigned to the Liolaemus alticolor group, Liolaemus pachacutec differs from Liolaemus chaltin in having precloacal pores in males. It differs from Liolaemus paulinae in the presence of a vertebral line and smooth neck scales. It differs from Liolaemus puna, Liolaemus alticolor and Liolaemus incaicus in having a partial or complete melanistic belly in adult males. It differs from Liolaemus aparicioi in lacking keeled temporal scales. It differs from Liolaemus bitaeniatus and Liolaemus pagaburoi in having a smooth dorsal surface of the head. It differs from Liolaemus pyriphlogos in the absence of red marks in lateral fields. It differs from Liolaemus variegatus in lacking keeled temporal scales, rugose dorsal head scales, and precloacal pores in females.</p><p>Description of holotype.</p><p>Adult male; SVL 44.8 mm; head length 11.0 mm; head width 8.2 mm; head height 6.2 mm; axilla-groin distance 18.4 mm (41.1% of SVL); foot length 13.6 mm (30.4% of SVL); tail length 74.9 mm. (1.7 times SVL).</p><p>Dorsal head scales 16, dorsal head scales smooth, scale organs more abundant in loreal and supralabial regions. Two scale organs on postrostral. Nasal scale in contact with rostral, separated from first supralabial by one scale, nasal bordered by six scales; canthus separated from nasal by one scale. Four supralabials. Four lorilabials scales and one in contact with the subocular. Five infralabials. Auditory meatus oval (height 2.0 mm, width 1.0 mm), with two small, projecting scales on anterior margin. Six convex, smooth temporals (counting vertically from buccal commissure to posterior corner of orbit). Orbit–auditory meatus distance 3.9 mm. Orbit–anterior margin of rostral distance 4.3 mm. Rostral about two times wider than high (width 2.3 mm; height 1.0 mm). Mental subpentagonal, about two times as wide as high (width 2.5 mm; height 1.0 mm). Interparietal pentagonal with an elongated posterior apex, bordered by five scales, the parietal of similar size. Frontal trapezoidal.</p><p>Supraorbital semicircles complete on both sides. Semicircles formed by six scales. Five enlarged supraoculars. Six distinctly imbricate superciliaries on both sides. Eleven upper and lower ciliaries. Subocular elongate, 2.8 mm, longer than eye diameter (2.1 mm; measured between anterior and posterior commissure of ciliaries), separated from supralabials by a single, but interrupted row of lorilabials. Fourth supralabial elongate, 2.0 mm. Four lorilabials with single row of scale organs. Fourth lorilabial contacting subocular. Preocular small, separated from lorilabial row by one scale. Postocular as large as preocular. Mental in contact with four scales: first infralabials (on each side) and two enlarged chin shields. Chin shields forming a longitudinal row of four enlarged scales separated one from the other by six smaller scales. Scales of throat round, flat, and imbricate. Twenty-two gulars between auditory meatus. Longitudinal neck fold without keeled scales and smaller in size than dorsal scales. Antehumeral pocket and antehumeral neck fold well developed. Thirty-six scales between auditory meatus and shoulder (counting along postauricular and longitudinal neck fold), twenty-six scales between auditory meatus and antehumeral neck fold. Gular folds absent.</p><p>Dorsal scales rhomboidal, keeled, and imbricate. Forty-two dorsal scales between occiput and level of groin. Forty-five scales around midbody. Nineteen rows of keeled scales on dorsum at midtrunk. Scales becoming smooth along flank and toward belly. Ventral scales slightly wider than dorsals. Seventy-seven ventral scales between mental and precloacal pores. Five precloacal pores. Supracarpals laminar, round, and smooth. Subdigital lamellae of fingers with three keels, in number I: 8; II: 12; III: 16; IV: 18; V: 12 (right fingers). Claws moderately long. Supradigital lamellae convex, smooth, and imbricate. Infracarpals and infratarsals keeled, distinctly imbricate. Supratarsals smooth. Subdigital lamellae of toes I: 10; II: 14; III: 18; IV: 22; V: 15 (right toes).</p><p>Color in preservation.</p><p>Dorsal background color from occiput to base of tail brownish-green. Black thin continuous vertebral line present. No dark paravertebral marks. Paravertebral and vertebral fields with same background color. Distinct cream dorsalateral stripes. No marks in lateral field. Cream ventrolateral stripes, beginning on the posterior corner of the eye, continuing across the upper auricular meatus, the longitudinal neck fold, through the shoulders, ending in the groin. No marks in the ventral field. Melanistic venter on throat, femur, tibia, and belly. Small and scattered dark marks in chin area and ventrolateraly. Ventral tail melanistic near the cloaca, with a thin longitudinal stripe, first half with small marks lateral to the stripe.</p><p>Color pattern in life.</p><p>Head dorsally brown with scattered black dots. Subocular white. Thin and faint black vertebral line. Paravertebral field without dark marks. Creamy dorsolateral stripes. Lateral field without marks. Faint cream-white ventrolateral stripe, beginning on upper margin of eye, continuing from auricular meatus, the longitudinal neck fold, through the shoulders, ending in the groin. Ventral field yellow. Forelimbs and chin scales white with scattered black dots. Melanistic belly, hind limbs, posterior two thirds of throat. Belly with scattered yellow dots laterally. Tail with a black region close to the cloaca, black longitudinal stripe and dots at each side of the stripe.</p><p>Variation.</p><p>Variation in characters is summarized in Table 1-4. There is sexual dichromatism. Males have a complete or partial melanistic belly and throat, while females have a white or yellow belly and black spots on throat. Some males have orange and yellow dots on lateral belly and yellow dots on chin scales, and ventral field with orange and black dots.</p><p>Etymology.</p><p>The specific epithet pachacutec refers to one of most important Inca rulers, Pachacutec, who built the best known Inca ruins, including Machu Picchu and Pisac, this last site at a higher elevation just above the type locality. The species name is in the nominative singular.</p><p>Distribution and natural history.</p><p>Liolaemus pachacutec sp. n. is known from four localities in the central Andes, at elevations of 4023-4972 m in the departments of Cusco and Apurímac in southeastern Peru (Fig. 11). The species was found under rocks in grassland habitats (Fig. 9). It was found in sympatry at similar elevations with Liolaemus ortizi Laurent, 1982 and Tachymenis peruviana Wiegmann, 1835. This species is probably viviparous; two females showed one or two uterine chambers per side, with an embryo and abundant yolk in each chamber, but without a visible shell.</p></div>	https://treatment.plazi.org/id/732E9F3E22FC322D109D0DA372864F20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aguilar, Cesar;Wood Jr, Perry L.;Cusi, Juan C.;Guzman, Alfredo;Huari, Frank;Lundberg, Mikael;Mortensen, Emma;Ramirez, Cesar;Robles, Daniel;Suarez, Juana;Ticona, Andres;Vargas, Victor J.;Venegas, Pablo J.;Sites Jr, Jack W.	Aguilar, Cesar, Wood Jr, Perry L., Cusi, Juan C., Guzman, Alfredo, Huari, Frank, Lundberg, Mikael, Mortensen, Emma, Ramirez, Cesar, Robles, Daniel, Suarez, Juana, Ticona, Andres, Vargas, Victor J., Venegas, Pablo J., Sites Jr, Jack W. (2013): Integrative taxonomy and preliminary assessment of species limits in the Liolaemus walkeri complex (Squamata, Liolaemidae) with descriptions of three new species from Peru. ZooKeys 364: 47-91, DOI: http://dx.doi.org/10.3897/zookeys.364.6109, URL: http://dx.doi.org/10.3897/zookeys.364.6109
C9BE151058350344E145DB61A66C8827.text	C9BE151058350344E145DB61A66C8827.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liolaemus wari	<div><p>Liolaemus wari sp. n. Figure 10</p><p>Liolaemus walkeri Lobo and Espinoza 1999</p><p>Liolaemus walkeri Martínez Oliver and Lobo 2002</p><p>Liolaemus walkeri Lobo, Quinteros and Díaz Gómez 2007</p><p>Liolaemus walkeri Quinteros 2012</p><p>Liolaemus walkeri Ocampo, Aguilar-Kirigin and Quinteros 2012</p><p>Holotype.</p><p>MUSM 30837, adult male collected at Abra Toccto, Huamanga Province, Ayacucho Department, Peru, 13.345S, 74.167W, elevation 4231 m, on 4 June 2012 by César Aguilar and Víctor Vargas.</p><p>Paratypes.</p><p>Three males (MUSM 30823, BYU 50184, 50185) and ten females (MUSM 30824, 30825, 30826, 30827, 30828, 30831, BYU 50186, 50187, 50191, 50243) from the same locality as the holotype. Two males (MUSM 30830, 30834) and three females (MUSM 30829, BYU 50188, 50190) from high area above the Historic Sanctuary Pampas, Huamanga Province, Ayacucho Department, on 3 June 2012 by César Aguilar and Víctor Vargas (see Data resources for elevations and coordinates). Two males (MUSM 25703, 25704) and one female (MUSM 25702) from Yanacocha Lake, La Mar Province, Ayacucho Department, on 24 November 2010 by Margarita Medina (see Data resources for elevations and coordinates). Two females (MUSM 25719, BYU 50189) from Huaychao, Huamanga Province, Ayacucho Department, on 1 December 2010 by Margarita Medina (see Data resources for elevations and coordinates). Two females (MUSM 30243, 30244) from Tambo, San Miguel Province, Ayacucho Department, by Michael Harvey. One male (MUSM 31411) and two juveniles (BYU 50235-36) from about 45 Km west Puquio-Cusco roadway, Lucanas Province, Ayacucho Department, on 11 June 2012 by César Aguilar and Víctor Vargas (see Data resources for elevations and coordinates).</p><p>Diagnosis.</p><p>Small (61.4 mm maximum SVL), slender Liolaemus, closely related to Liolaemus chavin sp. n., Liolaemus pachacutec sp. n., Liolaemus tacnae and Liolaemus walkeri (Fig. 1). It differs from Liolaemus chavin sp. n., Liolaemus pachacutec sp. n. and Liolaemus walkeri in having a ringed pattern on the ventral tail of adult males. It differs from Liolaemus pachacutec sp. n. in having spots in the lateral fields. Liolaemus wari differs from Liolaemus tacnae and Liolaemus chavin in having precloacal pores in males. In comparison with other species assigned to the Liolaemus alticolor group, Liolaemus wari sp. n. differs from Liolaemus chaltin in having precloacal pores in males. It differs from Liolaemus paulinae in lacking keeled neck scales. It differs from Liolaemus puna, Liolaemus alticolor and Liolaemus incaicus in having black spots on belly of adult males. It differs from Liolaemus aparicioi in lacking keeled temporal scales. It differs from Liolaemus bitaeniatus and Liolaemus pagaburoi in having a smooth dorsal surface of the head (rough to slightly dorsal surface of the head). It differs from Liolaemus pyriphlogos in the absence of red marks in the lateral field (red marks in the lateral fields present). It differs from Liolaemus variegatus in the absence of keeled temporal scales, rugose dorsal head scales and precloacal pores in females.</p><p>Description of holotype.</p><p>Adult male; SVL 55.4 mm; head length 11.4 mm; head width 9.8 mm; head height 6.2 mm; axilla–groin distance 23.3 mm (42% of SVL); foot length 15.0 mm. (27.1% of SVL); tail length 83.7 mm. (1.5 times SVL).</p><p>Dorsal head scales 14, dorsal head scales smooth, scale organs more abundant in loreal and supralabial regions. Five scale organs on postrostral. Nasal scale in contact with rostral, separated from first supralabial by one scale, nasal bordered by seven scales; canthus separated from nasal by one scale. Four supralabials. Five lorilabials scales and two in contact with the subocular. Four infralabials. Auditory meatus oval (height 2.0 mm, width 1.9 mm), with two small, projecting scales on anterior margin. Seven convex, smooth temporals (counting vertically from buccal commissure to posterior corner of orbit). Orbit–auditory meatus distance 4.6 mm. Orbit–anterior margin of rostral distance 7.9 mm. Rostral almost three times wider than high (width 2.7 mm; height 1.0 mm). Mental subpentagonal, about two times as wide as high (width 2.6 mm; height 1.2 mm). Interparietal pentagonal with an elongated posterior apex, bordered by seven scales, the parietal slightly smaller. Frontal trapezoidal. Supraorbital semicircles complete on both sides. Semicircles formed by 6 scales. Four enlarged supraoculars. Five distinctly imbricate superciliaries on both sides. Eleven upper and lower ciliaries. Subocular elongate, 3.2 mm, longer than eye diameter (2.3 mm; measured between anterior and posterior commissure of ciliaries), separated from supralabials by a single, but interrupted row of lorilabials. Second supralabial elongate, 1.6 mm. Five lorilabials with single and double rows of scale organs. Fifth and fourth lorilabials contacting subocular. Preocular small, separated from lorilabial row by one scale. Postocular as large as preocular. Mental in contact with four scales: first infralabials (on each side) and two enlarged chin shields. Chin shields forming a longitudinal row of three enlarged scales separated one from the other by six smaller scales. Scales of throat round, flat, and imbricate. Twenty-one gulars between auditory meatus. Longitudinal neck fold without keeled scales and smaller in size than dorsal scales. Antehumeral pocket and antehumeral neck fold well developed. Twenty-nine scales between auditory meatus and shoulder (counting along postauricular and longitudinal neck fold), 21 scales between auditory meatus and antehumeral neck fold. Gular folds absent.</p><p>Dorsal scales rhomboidal, keeled, and imbricate. Forty-four dorsal scales between occiput and level of groin. Fifty-three scales around midbody. Twenty-two rows of keeled scales on dorsum at midtrunk. Scales becoming smooth along flank and toward belly. Ventral scales slightly wider than dorsals. Seventy-three ventral scales between mental and precloacal pores. Five precloacal pores. Supracarpals laminar, round, and smooth. Subdigital lamellae of fingers with three keels, in number I: 8; II: 12; III: 16; IV: 16; V: 10 (right fingers). Claws moderately long. Supradigital lamellae convex, smooth, and imbricate. Infracarpals and infratarsals keeled, distinctly imbricate. Supratarsals smooth. Subdigital lamellae of toes I: 8; II: 12; III: 16; IV: 20; V: 13 (left toes).</p><p>Color pattern in preservation.</p><p>Dorsal background color from occiput to base of tail brownish-green. Black continuous vertebral line present. Dark paravertebral marks. Paravertebral and vertebral fields with same background color. Highly distinct creamy-yellow dorsalateral stripes. Large dark and small cream marks in lateral field. Cream ventrolateral stripe, beginning on the posterior corner of the eye, continuing across the upper auricular meatus, the longitudinal neck fold, through the shoulders, ending in the groin. Dark and cream small marks in the ventral field. Black spots on throat, femur, tibia, posterior third of belly and laterally in anterior two thirds of belly. Small and scattered dark marks in chest and anterior two thirds of belly. Tail with dark horizontal rows.</p><p>Color pattern in life.</p><p>Head dorsally brown with black dots. Subocular cream. A black vertebral band with a thin yellow stripe on the middle. The vertebral band has a thin white stripe on each side. Paravertebral field with dark marks with posterior white dots. Creamy-yellow dorsolateral stripes. Lateral field with black marks separated by cream diagonal stripes. Yellowhish-white ventrolateral stripe, beginning on upper margin of eye, continuing from auricular meatus, the longitudinal neck fold, through the shoulders, ending in the groin. Ventrolateral similar to lateral field and same color as the paravertebral field, with diffuse dorsal markings. Forelimbs, chest and belly yellowish-white with scattered and diffuse black dots. Black marks on hind limbs, throat, and posterior third of belly. Tail with black horizontal bands separated by white bands.</p><p>Variation.</p><p>The variation in morphological characters is shown in Tables 1-4. There is sexual dichromatism. Males have white or yellow belly and throat covered completely with black spots, yellowish belly and throat with black spots on posterior third of belly, or a melanistic belly on posterior third and cloacal region, with black dots on a white throat; females have white belly and yellowish throat with faint black dots, yellowish belly and throat with faint black spots, or yellowish belly and throat without spots. Adult males have white, yellowish and yellow tails with a conspicuous ringed pattern; adult females have white, yellowish or reddish ventral tails with or without a faint ringed pattern.</p><p>Etymology.</p><p>The specific epithet wari refers to the pre-Inca culture Wari (600-850 AD), which had its center close to the type locality. The species name is in the nominative singular.</p><p>Distribution and natural history.</p><p>Liolaemus wari sp. n. is known from seven localities in the central Andes, at elevations of 3768-4246 m in Ayacucho Department in eastern southern Peru (Fig. 11).</p><p>Liolaemus wari sp. n. was active on the ground or found under rocks in grassland (Fig. 10) and shrubland habitats. It was found in sympatry with another Liolaemus species belonging to the Liolaemus montanus series and the snake Tachymenis peruviana . This species is probably viviparous; three females each showed three uterine chambers per side; each chamber showed yolk, but with no developed embryos or visible shell.</p></div>	https://treatment.plazi.org/id/C9BE151058350344E145DB61A66C8827	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aguilar, Cesar;Wood Jr, Perry L.;Cusi, Juan C.;Guzman, Alfredo;Huari, Frank;Lundberg, Mikael;Mortensen, Emma;Ramirez, Cesar;Robles, Daniel;Suarez, Juana;Ticona, Andres;Vargas, Victor J.;Venegas, Pablo J.;Sites Jr, Jack W.	Aguilar, Cesar, Wood Jr, Perry L., Cusi, Juan C., Guzman, Alfredo, Huari, Frank, Lundberg, Mikael, Mortensen, Emma, Ramirez, Cesar, Robles, Daniel, Suarez, Juana, Ticona, Andres, Vargas, Victor J., Venegas, Pablo J., Sites Jr, Jack W. (2013): Integrative taxonomy and preliminary assessment of species limits in the Liolaemus walkeri complex (Squamata, Liolaemidae) with descriptions of three new species from Peru. ZooKeys 364: 47-91, DOI: http://dx.doi.org/10.3897/zookeys.364.6109, URL: http://dx.doi.org/10.3897/zookeys.364.6109
