taxonID	type	description	language	source
03DB87C58D768C03ECAAFBA3FD9CF8D5.taxon	type_taxon	Type species: Aedes hoffeinsorum Szadziewski, 1998 (by monotypy).	en	Zakrzewska, Marta, Perkovsky, Evgeny E., Harbach, Ralph E., Giłka, Wojciech (2025): Aetheapnomyia Harbach & Greenwalt (Diptera: Culicidae: Culicinae: Aedini) revisited based on a new record from Rovno amber. Zootaxa 5706 (2): 175-188, DOI: 10.11646/zootaxa.5706.2.2, URL: https://doi.org/10.11646/zootaxa.5706.2.2
03DB87C58D768C03ECAAFBA3FD9CF8D5.taxon	diagnosis	Diagnosis (based on Szadziewski 1998; Harbach 2024, 2025, and current observations, amended). Adult male: Antenna longer than proboscis, pedicel enlarged, terminal flagellomere with apical stylet-like or conical prolongation; maxillary palpus shorter than proboscis, comprised of 4 (?) or 5 palpomeres (see species description and Discussion), terminal palpomere very short, about 0.3 length of preceding palpomere; proboscis noticeably swollen distally; paratergal, postspiracular and lower mesepimeral setae present; wing with cell R 2 much shorter than vein R 2 + 3; fore- and midungues long, unequal, simple, borne well before apex of tarsomere 5.	en	Zakrzewska, Marta, Perkovsky, Evgeny E., Harbach, Ralph E., Giłka, Wojciech (2025): Aetheapnomyia Harbach & Greenwalt (Diptera: Culicidae: Culicinae: Aedini) revisited based on a new record from Rovno amber. Zootaxa 5706 (2): 175-188, DOI: 10.11646/zootaxa.5706.2.2, URL: https://doi.org/10.11646/zootaxa.5706.2.2
03DB87C58D768C03ECAAFBA3FD9CF8D5.taxon	discussion	Notes. Aetheapnomyia, like most generic-level taxa of Aedini (Reinert et al. 2009), is mainly a polythetic taxon that is diagnosed by a unique combination of characters. At present, the only possible unique (wholly diagnostic) character may be the preapical attachment of the fore- and midungues. Aetheapnomyia hoffeinsorum was originally described as a species of Aedes (Finlaya), but, as explained by Harbach & Greenwalt (2012), the absence of spotted wings precludes it from being placed in the subgenus Finlaya, and the unique combination of short maxillary palpi, greatly reduced palpomere 5, proboscis swollen distally, cell R 2 of the wing much shorter than vein R 2 + 3 and the preapical attachment of the fore- and midungues preclude it from being placed in the extant genus Aedes. Szadziewski (1998) described the antenna as shorter than the proboscis, which is inconsistent with detailed measurements in his paper. The genitalia bear some resemblance to certain extant species of the Ochlerotatus Group of subgenera of the genus Aedes (see the Discussion).	en	Zakrzewska, Marta, Perkovsky, Evgeny E., Harbach, Ralph E., Giłka, Wojciech (2025): Aetheapnomyia Harbach & Greenwalt (Diptera: Culicidae: Culicinae: Aedini) revisited based on a new record from Rovno amber. Zootaxa 5706 (2): 175-188, DOI: 10.11646/zootaxa.5706.2.2, URL: https://doi.org/10.11646/zootaxa.5706.2.2
03DB87C58D768C06ECAAF8B5FC95FB60.taxon	description	(Figs 1 – 4)	en	Zakrzewska, Marta, Perkovsky, Evgeny E., Harbach, Ralph E., Giłka, Wojciech (2025): Aetheapnomyia Harbach & Greenwalt (Diptera: Culicidae: Culicinae: Aedini) revisited based on a new record from Rovno amber. Zootaxa 5706 (2): 175-188, DOI: 10.11646/zootaxa.5706.2.2, URL: https://doi.org/10.11646/zootaxa.5706.2.2
03DB87C58D768C06ECAAF8B5FC95FB60.taxon	materials_examined	Material examined. Adult male, complete specimen (visible only from lateral side, visibility of head and genitalia limited by resin layers between which the specimen is located; right midleg attached, remaining legs present but detached, lying next to specimen), preserved in a piece of amber 46 × 20 × 17 mm (Eocene, Priabonian, 33.9 – 37.8 Mya), Zarechnoye District, Rovno Region, Ukraine (SIZK Z- 9; Fig. 1 A – C). Syninclusions: Coleoptera incertae sedis, 1 specimen; Hymenoptera incertae sedis, 1 specimen.	en	Zakrzewska, Marta, Perkovsky, Evgeny E., Harbach, Ralph E., Giłka, Wojciech (2025): Aetheapnomyia Harbach & Greenwalt (Diptera: Culicidae: Culicinae: Aedini) revisited based on a new record from Rovno amber. Zootaxa 5706 (2): 175-188, DOI: 10.11646/zootaxa.5706.2.2, URL: https://doi.org/10.11646/zootaxa.5706.2.2
03DB87C58D768C06ECAAF8B5FC95FB60.taxon	description	Description. Adult male. Colour: Body, mouthparts and legs brown to dark brown or black; wing membrane uniformly coloured, without spots; scales of thorax and wing pale, scales darker on mouthparts, legs and abdomen. Size: Body length, excluding proboscis, 4.4 mm, body + proboscis 5.9 mm, thorax + abdomen 3.9 mm. Antenna (Figs 1 C, 2 A – C): Length including pedicel 2.14 mm; flagellum 1.99 mm, comprised of 13 flagellomeres, flagellomeres 12 and 13 longest (0.39 and 0.58 mm, respectively), their combined length slightly less than total length of preceding flagellomeres (ratio 0.95); flagellar whorls well-developed, flagellomeres 1 – 12 each with whorl of many long setae, flagellomere 13 with whorl of several shorter setae and conical apical prolongation; antenna about 1.43 × length of proboscis. Proboscis (Figs 1 C, 2 A, D): Length 1.50 mm, longer than maxillary palpus (ratio 1.30), flattened dorsoventrally and gradually broadened toward apex; labella of typical aedine form, with well-defined proximal and distal sclerites, proximal sclerite crescentic in lateral view, dorsal sclerite slightly longer than proximal sclerite, shaped like convex surface of bowl of a spoon, both sclerites largely covered with setae. Maxillary palpus (Fig. 2 A, E): Length 1.15 mm, shorter than proboscis (ratio 0.77), with 5 (?) palpomeres (possibly only 4, see Discussion) combined length of palpomeres 1 – 3 882 μm, about 0.59 length of proboscis; palpomere 4 (209 μm) more than 3 times longer than palpomere 5 (67 μm), ratio 3.12; palpomere 1 (= palpifer?, see Discussion) short, somewhat bulbous; palpomeres 2 – 4 slender, each gently broadening toward apex; palpomere 5 greatly reduced, ovoid with rounded apex, bearing up to 5 long setae distally. Thorax (Figs 1 C, 2 A): Integument dark brown (some areas obscured by milky amber); paratergite with pale scales; postspiracular, prealar, upper and lower mesokatepisternal and upper and lower mesepimeral setae present; some pale scales on upper mesokatepisternal, lower mesokatepisternal, and upper and lower mesepimeral areas. Wing (Fig. 3 A, B): Length 2.60 mm (base to tip), 2.29 mm (arculus to tip), maximum width 0.63 mm; scales spatulate, slender, apically rounded or slightly apically triangulate, apices minutely pectinate; cells R 2 and R 3 distinctly shorter than vein R 2 + 3 (R 2 / R 2 + 3 = 0.80, R 3 / R 2 + 3 = 0.90); arrangement of veins in area of radiomedial and mediocubital crossveins as shown in Fig. 3; mediocubital crossvein [base of M 3 + 4 of Harbach & Knight 1980] unusually short; cubitus posterior (plical vein of Belkin 1962) developed as strongly as cubitus anterior, with scales at least on proximal part; anal vein (1 A) ending 1 / 3 (0.35) of distance between intersection of mediocubital crossvein and cubitus and base of M 3 + 4 toward apex of wing. Legs (Figs 1 C, 4 A – D): Entirely dark-scaled; fore- and midungues borne well before apex of tarsomere 5, long, curved, unequal on foreleg (lengths 90 μm and 120 μm) and midleg (longer unguis 110 μm, shorter unguis unmeasurable), short and equal on hindleg (~ 60 μm); all ungues simple, without teeth. Lengths of segments of legs in Table 2. Abdomen (Fig. 1 C): Laterotergite of segment I apparently without scales, with setae only; tergum II with a few basolateral pale scales; terga III ‒ VIII apparently without pale bands, entirely dark-scaled; scaling of sterna uncertain, but some dark scales evident on lateral areas of sterna III ‒ VI. Genitalia (Fig. 4 E, F): Observable in dorsolateral view (in part obscured by layer of milky amber); gonocoxite elongate, length ~ 300 µm; gonostylus slender, slightly arched, about half length of gonocoxite, with long, slender terminal gonostylar claw; claspette moderately long with narrow columnar stem and terminal flattened filament (see Szadziewski 1998: fig. 3). Scales (Fig. 4 G – Q): Variable in shape, depending on location on wing veins (Fig. 4 G – J), proboscis (Fig. 4 K), maxillary palpus (Fig. 4 L), thorax (Fig. 4 M), legs (Fig. 4 N, O) or abdomen (Fig. 4 P, Q); ranging from broadly spatulate (twice as long as wide at most) with apex slightly triangulate or rounded, to slender (several times as long as wide) with apex more or less convex; apices pectinate.	en	Zakrzewska, Marta, Perkovsky, Evgeny E., Harbach, Ralph E., Giłka, Wojciech (2025): Aetheapnomyia Harbach & Greenwalt (Diptera: Culicidae: Culicinae: Aedini) revisited based on a new record from Rovno amber. Zootaxa 5706 (2): 175-188, DOI: 10.11646/zootaxa.5706.2.2, URL: https://doi.org/10.11646/zootaxa.5706.2.2
