taxonID	type	description	language	source
03DB87CCFFBFFFC6B0C0FF1E7DE1604E.taxon	materials_examined	Type material. Male holotype, 5.0 mm (NHMUK 2015. 3058), Station A 2, Sea City, Kuwait, seaweeds and associates, 23 March 2010, M. Nithyanandan. Paratypes 32 males, 31 females (NHMUK 2015. 3059 – 3068), same data as holotype. Other material. 1 male, 2 females (NHMUK 2015. 3069 – 3071), Sea City, Kuwait, soft benthos, 24 October 2012, M. Nithyanandan.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFBFFFC6B0C0FF1E7DE1604E.taxon	etymology	Etymology. Named after the region of the type locality.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFBFFFC6B0C0FF1E7DE1604E.taxon	description	Description. Based on male holotype, 5.0 mm. Head. Head lobes truncate; eye round. Antenna 1 scarcely half-length of antenna 2; peduncular article 2 subequal with peduncular article 3. Antenna 2 less than half body length. Labium inner plates vestigial. Maxilla 1 inner plate with two stout distal setae; palp extending beyond bases of setae on outer plate. Maxilla 2 without oblique setal row. Mandible lacking palp; lacinia mobilis 5 - dentate. Maxilliped palp long; fourth article extending well beyond tip of outer plate; article 7 very long and whip-like. Pereon. Gnathopod 1 coxa large without posteroproximal shelf, posterior face with two very stout setae; basis expanded with moderately well developed anterodistal lobe; propodus elongate, longer than carpus, subrectangular; dactylus short, broad. Gnathopod 2 coxa large, subround with weak posteroproximal shelf; basis enlarged with strong anterodistal lobe; carpus very small, triangular; propodus greatly enlarged, subovoid, palm relatively short, only moderately oblique, defined by a strong, subacute spine; dactylus fitting palm. Pereopod 3 basis parallel-sided. Pereopod 5 basis subround with small mid posterior margin notch, crenulate on posterodistal part; dactylus well developed with small seta on posterodistal margin. Pereopod 6 basis subrectangular with small mid posterior margin notch, crenulate on posterodistal part; dactylus well developed with small seta on posterodistal margin. Pereopod 7 basis very expanded, subround, crenulate on posterior margin; dactylus well developed with small seta on posterodistal margin. Pleon. Epimeron 1 rounded. Epimeron 2 subrectangular. Epimeron 3 weakly extended on posterior distal margin into a triangular spine. Uropod 1 peduncle with strong distolateral robust seta; rami a little longer than peduncle, subequal in length. Uropod 2 peduncle short; inner ramus much longer than peduncle and longer than outer ramus. Uropod 3 peduncle more than twice length of single ramus. Telson unarmed. Female (sexually dimorphic characters). Gnathopod 2 basis expanded distally; carpus moderately large, subrectangular; propodus elongate, subrectangular, palm short, oblique; dactylus fitting palm.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFBFFFC6B0C0FF1E7DE1604E.taxon	discussion	Remarks. This species is similar to P. corallinicola (Hirayama, 1980), and to P. schmidti (Heller, 1866). It differs from both species, however, in the form of the male gnathopod 2. In P. a r ab i ca sp. nov., the palm is only moderately oblique reaching about one third the length of the posterior margin of the propodus, and is defined by a subacute spine. In P. corallinicola and in P. schmidti, the palm is very oblique, reaching at least half of the posterior margin of the propodus and is defined by an ill-defined rounded hump. It also differs from P. schmidti in the form of the maxilliped palp which is stout in P. schmidti but slender in P. arabica sp. nov.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFBFFFC6B0C0FF1E7DE1604E.taxon	distribution	Distribution. Kuwait.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFBAFFC5B0C0FBBE7DE1628B.taxon	materials_examined	Type material. Male holotype, 6.0 mm (NHMUK 2015. 3072), llBS A 1 25 May 2014, grab on bare sand benthos in phase A 1, M. Nithyanandan. Paratypes, (NHMUK 2015. 3073 – 3082), 4 males, 15 females, 2 juveniles, same date as holotype. Other material. 10 males, 12 females (NHMUK 2015. 3083 – 3092, A 3 S 1, sandy benthos in phase A 3 Sea City waterways, Ocklemann sledge, M. Nithyanandan, 20 October 2013; 1 male, 1 female (NHMUK 2015. 3093 - 3094), North Gate, from seaweed and hydroids growing on concrete sides of north tidal gate phase A 3 by diving, D. K. Raja, 19 April 2014; 1 male 3 females (NHMUK 2015. 3095 – - 3098), collected from tunicate samples taken by diving in Marina in phase A 2 1 April 2014, D. K. Raja.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFBAFFC5B0C0FBBE7DE1628B.taxon	etymology	Etymology. Named after the acute coxa 1 of the male gnathopod 1.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFBAFFC5B0C0FBBE7DE1628B.taxon	description	Description. Based on male holotype 6.0 mm. Head. Head lateral lobes moderately produced, truncate. Eye round. Mandible palp article 3 longer than 2, posterior margin almost straight with comb-setae. Maxilla 1 inner plate with a single, long, pectinate seta. Labium with acute mandibular processes. Antenna 1 longer than 2; peduncular article 2 longer than 1; primary flagellum longer than peduncle with 23 articles; accessory flagellum long, with 6 articles. Antenna 2 peduncular articles 4 and 5 subequal in length. Pereon. Segments 2 – 4 with strong ventral spines. Gnathopod 1 much larger than 2; coxa subrectangular, strongly produced anterodistally, subacute; basis about four times as long as broad; carpus reduced, cup-shaped; propodus elongate rectangular, posterodistal angle with strong acute tooth separated from short rounded palm by narrow sinus; dactylus greatly overlapping palm. Gnathopod 2 coxa subround; basis elongate, anterior margin weakly concave; carpus a little longer than propodus. Pereopods 3 – 7 of normal Bemlos form. Pleon. Epimera 2 – 3 with small posterodistal spine. Uropod 1 peduncle subequal with rami, with long, curved, distoventral spine; inner ramus scarcely longer than outer. Uropod 2 peduncle shorter than rami, with strong distoventral spine about two-thirds length of peduncle; inner ramus longer than outer. Uropod 3 rami much longer than peduncle, outer ramus longer than inner. Telson with each dorsolateral crest bearing a pair of long setae. Female (sexually dimorphic characters). Gnathopod 1 coxa rounded; carpus larger than that of male; propodus shorter than that of male, propodus posterodistal angle with blunt shelf with a strong robust seta at its base, palm, obsolescent.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFBAFFC5B0C0FBBE7DE1628B.taxon	discussion	Remarks. This species is very close to Bemlos aequimanus (Schellenberg, 1938), but it differs in several ways. In the male, gnathopod 1 coxa is markedly more produced antero-distally, and the spine on the posterodistal corner of the propodus is longer, reaching the level of the short palm (poorly produced and shorter than the palm margin in B. aequimanus). It also resembles B. teleporus (K. H. Barnard, 1955), but that species has a rounded male coxa 1 and a short posterodistal spine on the propodus of male gnathopod 1. It differs from both B. aequimanus and B. teleporus in the pereonal spines of the male. Bemlos acuticoxa sp. nov. has strong acute spines on segments 2 – 4 whereas B. aequimanus has short rounded spines on segments 1 – 3 and B. teleporus has an acute spine on segment 3 only (Ledoyer, 1983, Madagascar). In addition, the anterior margin of the basis of the male gnathopod 2 is weakly concave in B. acuticoxa sp. nov., but more or less straight in both B. aequimanus and B. teleporus.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFBAFFC5B0C0FBBE7DE1628B.taxon	distribution	Distribution. Kuwait.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB9FFC5B0C0FB41780E641C.taxon	materials_examined	Material. 1 male, 3 females (NHMUK 2015. 3116 – 3119), from rock and associated seaweeds and hydroids on sides of concrete north tidal gate, within Sea City waterways, 17.03.2014, D. K. Raja; 3 females (NHMUK 2015. 3120 – 3122), from sponges growing on concrete sides of north tidal gate, 8 April 2014, D. K. Raja; 2 males, 5 females (NHMUK 2015. 3123 – 3129), from tunicates attached to concrete sides of north tidal gate, Sea City, 20 April 2014, D. K. Raja; 3 females (NHMUK 2015. 3130 – 3132), from tunicate samples taken by diving in Marina in phase A 2, 1 April 2014, D. K. Raja.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB9FFC5B0C0FB41780E641C.taxon	discussion	Remarks. Krapp-Schickel (1982), Myers (1985) and Hughes & Lowry (2009) have pointed out that A. ramondi represents a species complex. It is not possible to elucidate this complex here, but present material closely agrees with that described from the Mediterranean Sea by Krapp-Schickel (1982).	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB9FFC5B0C0FDA27FDA60B1.taxon	materials_examined	Material. 5 males, 4 females (NHMUK 2015. 3099 – 3107), llBS, grab on bare sand benthos in phase A 1, M. Nithyanandan, 25 May 2014; 1 male, 7 females (NHMUK 2015. 3108 – 3115), tunicate samples taken by diving in Marina in phase A 2, D. K. Raja, 1 April 2014.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB9FFC5B0C0FDA27FDA60B1.taxon	discussion	Remarks. This apparently worldwide species may be a species complex.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB6FFCBB0C0FA7B7F2563A3.taxon	materials_examined	Material. 3 males, 7 females (NHMUK 2015. 3133 – 3142), grab sample on bare sand benthos in phase A 1, Sea City, 25 May 2014, M. Nithyanandan.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB6FFCBB0C0FA7B7F2563A3.taxon	discussion	Remarks. This species has long caused taxonomic confusion. Peart (2004), redescribed C. filosa and designated a neotype from the Mediterranean Sea (Sardinia). Present material agrees well with Mediterranean material and is therefore attributed to C. filosa Savigny.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB7FFCBB0C0FECA7D9A65CE.taxon	materials_examined	Type material. Male holotype, 2.2 mm (NHMUK 2015. 3143), samples from rock and associated seaweeds and hydroids on sides of concrete north tidal gate, within Sea City waterways, 17 March 2014, D. K. Raja. Paratypes, 4 males, 15 females (NHMUK 2015. 3144 – 3153), same data as holotype. Other material. 15 males, 61 females (NHMUK 2015. 3154 – 3163), from tunicates attached to concrete sides of north tidal gate in phase A 3 Sea City, 20 April 2014, D. K. Raja; 48 males, 102 females (NHMUK 2015. 3164 – 3173), from sponges growing on concrete sides of North Tidal Gate, phase A 3, Sea City, 8 April 2014, D. K. Raja.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB7FFCBB0C0FECA7D9A65CE.taxon	description	Description. Based on male holotype 2.2 mm. Head. Head with short triangular rostrum; Eyes subround. Antenna 1 peduncular articles 1 and 2 subequal, article 1 with two robust setae on the posterior proximal margin, the basal one recurved and two robust setae on the inner proximal margin; article 3 short; peduncular articles clothed with long fine setae; flagellum short, with three articles. Antenna 2 grossly enlarged; peduncular article 4 subrectangular with two strong spines on the posterodistal margin, a short inner one and a longer, recurved outer one; article 5 much more slender and about two thirds length of article 1, posterodistal margin with one spine, sub-distal margin with one spine; flagellum composed of two stout, setose articles. Maxilliped palp well developed, palp article 2 extending beyond end of outer plate. Pereon. Gnathopod 1, dactyl bifid, tip exceeding short, oblique palm. Gnathopod 2, carpus short, deep, posterodistal margin setose; propodus lacking palm; dactylus short, bidentate. Pereopods 3 – 4 short; basis broad; segment 4 broadened distally, slightly overhanging short article 5. Pereopods 5 – 6 short, similar in form but bases unequal in size; segment 5 short, with 2 posterolateral clusters of short hook spines. Pereopod 7 not elongate; basis medium broad; dactyl short. Pleon. Urosome segments 1 – 3 fused, plate-like. Uropod 1 peduncle broad elongate, rami much shorter than peduncle, inner ramus longer than outer, both rami with robust setae. Uropod 2 peduncle stout, short; outer ramus shorter than peduncle rounded, with robust and fine setae, inner ramus slender, subequal with peduncle with robust setae. Uropod 3 peduncle short with large outer lobe; ramus short subround. Telson small, rounded. Female up to 3.0 mm (sexually dimorphic characters). Antenna 2 peduncular article 4 much less robust than that of male, inner margin with one basal, one medio-distal and one distal robust seta; article 5 shorter and more slender than article 4, lacking spines or robust setae.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB7FFCBB0C0FECA7D9A65CE.taxon	discussion	Remarks. Laticorophium bifurcatum sp. nov. is very close to L. baconi Shoemaker, 1934. It differs in having the male antenna 2 with a strong spine on the posterior margin of article 5. In this respect it superficially resembles Apocorophium acutum (Chevreux, 1908) but unlike that species (and genus) it has a notched urosome, a bidentate dactylus on gnathopod 2 and a strongly lobate peduncle of uropod 3. It also differs from L. baconi in having two robust setae inserted on a swollen posteroproximal region of article 4 of the male antenna 2 as well as having the posterodistal spine on article 4 of the male antenna 2 divergent as opposed to recurved.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB7FFCBB0C0FECA7D9A65CE.taxon	distribution	Distribution. Kuwait	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB2FFCCB0C0F8827DE161F3.taxon	materials_examined	Type material. Male holotype, 3.5 mm (NHMUK 2015. 3174) from rock and associated seaweeds and hydroids on sides of concrete north tidal gate, within Sea City waterways, 17 March 2014, D. K. Raja. Paratypes, 11 males, 7 females (NHMUK 2015. 3175 – 3184), same data as holotype.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB2FFCCB0C0F8827DE161F3.taxon	etymology	Etymology. Named after one (Al-Mamlaha) of the three creeks upon which Sea City is based.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB2FFCCB0C0F8827DE161F3.taxon	description	Description. Based on male holotype 3.5 mm. Head. Eye large, subovoid. Antenna 1 about two thirds body length; peduncular article 1 short; article 2 long, more than twice length of article 1; article 3 shorter than article 2; flagellum subequal in length with peduncular article 3, with 5 articles. Antenna 2 longer than antenna 1 and almost as long as body; peduncular articles 4 and 5 greatly elongated and stout; 5 a little longer than 4; flagellum article 1 elongated; article 2 long but much shorter than article 1; articles 3 – 4 telescoped. Mandible palp with three articles; article 3 shorter than 2, parallel-sided with dense distal setae and a few scattered setae on posterior and anterior margins. Pereon. Segments 1 – 6 without dorsal spines or carinae. Segment 7 with small dorsal spine. Gnathopod 1 coxa weakly produced forward, subacute; basis slender; propodus a little longer than carpus, subovoid, palm convex, almost obsolete; dactylus greatly overlapping reduced palm, almost as long as propodus. Gnathopod 2 coxa subquadrangular; basis swollen, less than twice as long as broad; merus posterodistal corner weakly produced; carpus fused with propodus; propodus greatly enlarged, subovoid, palm convex, short, followed by two blunt spines on the posterodistal margin and one smaller blunt spine medially, posterior margin and inner lateral face clothed in long setae; dactylus curved, greatly overlapping palm, opposable to medial spine on posterior margin of propodus. Pereopods 3 – 7 basis short, subrectangular; dactylus stout. Pleon. Segments 1 – 2 with dorsal rounded spines that are folded back over the segment behind. Epimera 1 – 3 small, rounded. Urosome segment 1 elongate, longer than second and third segments combined. Uropod 1 peduncle and rami with many robust setae, inner ramus longer than outer and longer than peduncle. Uropod 2 peduncle short, inner ramus longer than outer. Uropod 3 very reduced with a small reduced ramus. Female (sexually dimorphic characters). Gnathopod 2 coxa deeper than broad; basis stout, anterior margin concave; merus expanded, protruding distally; carpus short, cup-shaped; propodus subovoid, palm and posterior margin weakly crenulate, posterior proximal margin with strong, triangular blunt spine bearing two robust setae; dactylus elongate, curved, opposable to proximal spine of propodus.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB2FFCCB0C0F8827DE161F3.taxon	discussion	Remarks. This species appears to be closest to P. zeylanicus Walker, 1904. It resembles that species in the general shape of the male gnathopod 2. It differs from that species, however, in the well developed flap-like rounded spines on the dorsum of pleon segments 1 – 2.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB2FFCCB0C0F8827DE161F3.taxon	distribution	Distribution. Kuwait.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB0FFCDB0C0FCFF7DE163A3.taxon	materials_examined	Type material. Male holotype 4.3 mm (NHMUK 2015. 3185), floating Sargassam, Sea City, D. A. Jones, 16 October 2011. Paratypes, 5 males, 1 female (NHMUK 2015. 3186 – 3191), same data as holotype. Other material. 18 males, 9 females (NHMUK 2015.3248 – 3257), mixed rubble, Ocklemann dredge on mixed rock / sand benthos at station NS 04 phase A 1 Sea City, M. Nithyanandan, 21.10.2014; 4 males, 2 females (NHMUK 2015. 3258 – 3263) tunicates collected by diving and removing tunicates attached to concrete sides of north tidal gate in phase A 3 Sea City, D. K. Raja, 20.4.2014.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB0FFCDB0C0FCFF7DE163A3.taxon	etymology	Etymology. Named for the shape of the male gnathopod 2 that superficially resembles that of some members of the genus Jassa (Ischyroceridae).	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB0FFCDB0C0FCFF7DE163A3.taxon	description	Description. Based on male holotype 4.3 mm. Head. Head elongate, lateral cephalic lobes strongly produced, rounded; eye lageniform. Mandible palp article 3 shorter than 2, spatulate with long distal setae. Maxilla 1 inner plate with setae along entire margin. Maxilla 2 with oblique setal row. Labium mandibular projections acute. Maxilliped palp elongate, second article extending beyond distal end of outer plate. Antenna 1 about half body length, peduncular article 2 a little longer than 1; primary flagellum subequal with peduncle, with 12 articles; accessory flagellum long with 5 articles. Antenna 2 unknown. Pereon. Gnathopod 1 coxa subsquare, weakly produced forward; basis enlarged, weakly excavated for reception of carpus when folded; carpus moderate, longer than broad; propodus longer than carpus, subrectangular, palm excavate, with blunt posterodistal spine. Gnathopod 2 coxa subrectangular; basis stout, excavated for reception of carpus when folded, strongly setose on anterior margin; carpus very short, cup-shaped; propodus grossly enlarged widening distally, with one strong posterodistal spine and one strong medial spine, anterior margin strongly setose; dactylus enlarged, overlapping palm. Pereopods 3 – 4 of normal Latigammaropsis form, dactylus short, much less than half length of propodus. Pereopod 5 basis moderately expanded. Pereopods 6 – 7 missing. Pleon. Uropod 1 peduncle with long, distal, interramal spine, half length of peduncle; inner ramus subequal with peduncle, outer ramus shorter than inner. Uropod 2 lacking interramal spine, peduncle shorter than rami, inner ramus a little longer than outer. Uropod 3 peduncle longer than broad; rami subequal, shorter than peduncle. Telson with each dorsolateral crest bearing a pair of robust setae Female (sexually dimorphic characters). Gnathopod 1 propodus longer than carpus, subovoid, palm oblique; dactylus fitting palm. Gnathopod 2 basis relatively slender; carpus short, cup-shaped; propodus subround with small posterodistal spine, palm evenly rounded; dactylus fitting palm.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB0FFCDB0C0FCFF7DE163A3.taxon	discussion	Remarks. Latigammaropsis pseudojassa sp. nov. shows no close relationship with any described species of the genus. The shape of the male gnathopod 2 is unique.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFB0FFCDB0C0FCFF7DE163A3.taxon	distribution	Distribution. Kuwait.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFAFFFD3B0C0FB7B7EAF6509.taxon	materials_examined	Material. 4 males, 2 females (NHMUK 2015. 3192 – 3197), A 2 S 4 Ocklemann sledge, benthos, Sea City, Kuwait, 22 October 2014, M. Nithyanandan.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFAFFFD3B0C0FB7B7EAF6509.taxon	discussion	Remarks. Despite the large distances that separate this material from the only other known locality (Hawaii), this material agrees well with the material described by J. L. Barnard (1955) and later (J. L. Barnard 1970). Anthropogenic dispersal cannot be ruled out.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFAAFFD5B0C0FF117DE1679E.taxon	materials_examined	Type material. Holotype male, 7.0 mm (NHMUK 2015. 3198) North Gate, collected from seaweed and hydroids growing on concrete sides of north tidal gate phase A 3 by diving. D. K. Raja, 19 April 2014. Paratypes, 6 males, 9 females (NHMUK 2015. 3199 – 3208), same data as holotype. Other material. 9 males, 7 females (NHMUK 2015. 3209 – 3218), llBS, grab on bare sand benthos in phase A 1, M. Nithyanandan, 25 May 2014; 7 males, 9 females (NHMUK 2015. 3219 – 3228), subtidal seagrass in phase A 1 Sea City, Ocklemann sledge and small grab, M. Nithyanandan, 14 October 2014; 2 males (NHMUK 2015. 3235 – 3236), from sandy benthos in phase A 3 Sea City waterways, Ocklemann sledge, M. Nithyanandan, 20 October 2013.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFAAFFD5B0C0FF117DE1679E.taxon	etymology	Etymology. Named after a creek (Al-Ama) in the type locality. Description. Based on male holotype 7.0 mm. Head. Eye lobe rounded with sub-ocular notch. Eye medium size, subround. Antenna 1 about two thirds body length; peduncular article 2 longer than 1, article 3 very short; accessory flagellum with 5 articles; primary flagellum longer than peduncle with 26 articles. Antenna 2 a little shorter than antenna 1; peduncular articles 4 and 5 subequal; flagellum with 24 articles. Maxilla 1 inner plate posterior margin fringed with setae. Mandible palp article 1 with acute spine at posterodistal corner; article 3 much shorter than 2 with long distal and marginal setae. Pereon. Gnathopod 1 coxa with very acute anterodistal corner; carpus and propodus subequal in length, strongly setose. Right gnathopod 2 greatly enlarged; coxa subquadrate; basis stout with spine at anterodistal corner; carpus short, cup-shaped; propodus massive, subovoid, palm with short overhanging shelf, followed by a shallow U-shaped excavation, followed by an oblique undefined margin; dactylus stout, curved, fitting palm. Left gnathopod 2 much smaller than right gnathopod 2, similar in form and size to that of female. Pereopod 6 basis posterodistal margin with multiple spines. Pereopod 7 basis posterodistal margin with a single acute spine. Pleon. Pleonal segments 1 – 3 strongly dorsally spinous with setae inserted between the spines. Epimera 1 – 3 posterior distal margins strongly spinous, with setae inserted between the spines, posterior margins with robust setae. Urosome segments 1 – 2 dorsally serrated with setae inserted between the spines. Uropod 1 peduncle a little longer than rami, with basofacial robust seta and with ventrodistal spine; inner ramus marginally longer than outer, slender with marginal and distal robust setae. Uropod 2 peduncle a little shorter than rami; inner ramus marginally longer than outer. Uropod 3 peduncle short; rami enlarged, broad, subequal in length. Telson deeply cleft, lobes strongly divergent.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFAAFFD5B0C0FF117DE1679E.taxon	description	Female (sexually dimorphic characters) Gnathopod 2 coxa elongate, rounded, unproduced; basis weakly expanded anteromedially; carpus well developed, about two thirds length of propodus; propodus elongate subovoid, palm oblique, defined by a robust seta; dactylus slender, fitting palm.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFAAFFD5B0C0FF117DE1679E.taxon	discussion	Remarks. Ceradocus (Denticeradocus) alama sp. nov. resembles C. (D.) serratus (Bate, 1862), as figured by J. L. Barnard 1972 (not Megamoera serrata Bate, 1862) from Australia in the male large gnathopod 2 having a short palmar shelf and in not having a posterodistal defining spine. That species, however, does not have an excavation in the palm of the large male gnathopod 2. It is similar to C. (D.) serratus as figured by Ledoyer (1983) (also not Megamoera serrata Bate, 1862), but that species has a palmar defining corner lacking in C. (D). alama sp. nov. Like C. (D). alama sp. nov., the C. (D). serratus of Ledoyer, 1983 has a palmar excavation, but that excavation is much narrower than in C. (D). alama sp. nov. Ceradocus (D.) serratus of Ledoyer (1983) also lacks the palmar shelf characteristic of C. (D). alama sp. nov.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFAAFFD5B0C0FF117DE1679E.taxon	distribution	Distribution. Kuwait.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFA9FFD5B0C0FA917F9065ED.taxon	materials_examined	Material. Male holotype and female paratype, mixed rubble, Ocklemann sledge on mixed rock / sand benthos at station NS 04 phase A 1 Sea City, M. Nithyanandan, 21 October 2014; 6 males, North Gate, collected from seaweed and hydroids growing on concrete sides of north tidal gate phase A 3 by diving, D. K. Raja, 19 April 2014; 11 males, 6 females, Nuwaiseeb sand beach 3 km south of Sea City at mid tide from sand core, M. Nithyanandan; 2 males, from subtidal seagrass in phase A 1 Sea City taken by Ocklemann sledge and small grab, M. Nithyanandan, 14 October 2014.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFA9FFD5B0C0FA917F9065ED.taxon	distribution	Distribution. Mediterranean Sea, Persian Gulf, Madagascar, India, Mauritius, Hawaii. Probably also but not confirmed: Suez Canal, Red Sea, East Africa, South Africa, Sri Lanka.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFA9FFD8B0C0F9787DE1621B.taxon	materials_examined	Type material. Male holotype, 5.0 mm (NHMUK 2015. 3237), Seagrass, Sea City, Kuwait, 20 October 2014, M. Nithyanandan; male paratype (NHMUK 2015. 3238), same data as holotype. Other material. 2 males (NHMUK 2015. 3239 – 3240), soft benthos, Sea City, Kuwait, 24 October 2012, M. Nithyanandan.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFA9FFD8B0C0F9787DE1621B.taxon	etymology	Etymology. Named after the irregular formation of the male gnathopod 2 palm.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFA9FFD8B0C0F9787DE1621B.taxon	description	Description. Based on male holotype, 5.0 mm. Head. Eye lobe rounded, eye large, ovate, with loose ommatidea. Antenna 1 peduncle elongate; article 2 longer than article 1; article 3 short, a little over one third length of article 2; accessory flagellum elongate with 5 articles; primary flagellum shorter than peduncle with 16 articles. Antenna 2 much shorter than antenna 1; peduncular article 5 shorter than article 4; flagellum subequal in length with peduncular article 5, with 6 articles. Pereon. Gnathopod 1 coxa weakly produced forward; basis moderately stout; carpus and propodus subequal in length; carpus with notch on anterior margin; propodus palm evenly convex; dactylus fitting palm. Gnathopod 2 enormous, basis stout, excavate on anterior margin for reception of carpus and propodus when folded; carpus short, cup-shaped; propodus four times length of carpus, subrectangular, palm irregular, with two proximal humps separated by a v-shaped excavation, followed by a sloping palp delineated by a small spine, followed by a stout spine; dactylus stout, curved, fitting palm, with long setae on anterior margin. Pereopods 3 – 7 missing. Pleon. Epimera 1 – 2 with small posterodistal spine; epimeron 3 with larger, triangular spine. Uropods 1 – 2, rami subequal, longer than peduncle. Uropod 3 large; peduncle short; rami twice length of peduncle, flattened, leaflike with numerous long setae marginally and distally. Telson almost completely divided, terminally notched, with three short robust setae on each side. Female (sexually dimorphic characters). Unknown.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFA9FFD8B0C0F9787DE1621B.taxon	discussion	Remarks. M aera irregularis sp. nov. has a male gnathopod 2 superficially similar in the shape to that of Maera lucinae Krapp-Schickel, 2004 from the Gulf of Mexico. In that species, however, the shape of the palm is subtly different. Also in M. lucinae, antenna 1 and 2 are subequal in length, the peduncle of uropod 3 is much longer and the telson has very long robust setae. It shows some similarity to M. edwardsi Chevreux, 1927 from Cape Verde and the Canary Islands and M. hirondellei Chevreux, 1900 from the Azores, but in both those species the male gnathopod palm is a different shape and in M. edwardsi the epimera 1 – 2 lack spines.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFA9FFD8B0C0F9787DE1621B.taxon	distribution	Distribution. Kuwait.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFA4FFD8B0C0FE5F7E86674B.taxon	materials_examined	Material. 3 males, 2 females (NHMUK 2015. 3241 – 3245), North Gate, rock and associated seaweeds and hydroids on sides of concrete north tidal gate, within Sea City waterways, collected by diving, D. K. Raja, 19 April 2014; 1 male (NHMUK 2015. 3246), mixed rock / sand, Ocklemann sledge on mixed rock / sand benthos in phase A 1 Sea City, M. Nithyanandan, 18 April 2014; 1 female (NHMUK 2015. 3247), mixed rubble, Ocklemann sledge on mixed rock / sand benthos at station NS 04 phase A 1 Sea City, M. Nithyanandan, 21 October 2014.	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
03DB87CCFFA4FFD8B0C0FE5F7E86674B.taxon	discussion	Remarks. Present material agrees well with the original description (Audouin 1826) as well as with a recent redescription in Lowry & Springthorpe (2007).	en	Myers, Alan A., Nithyanandan, Manickam (2016): The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea). Zootaxa 4072 (4): 401-429, DOI: 10.11646/zootaxa.4072.4.1
