taxonID	type	description	language	source
03DBD63AFFBCCD35FF17F89622A328F3.taxon	description	(Figs. 2 – 4)	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFBCCD35FF17F89622A328F3.taxon	description	In the morphological description of Eremias benguelensis, Bocage (1867 b) seemingly describes only one specimen and makes no reference to a type series, even though he provides a range of ventral scales. In an annotated type catalogue of the lacertids in the Museum für Naturkunde (ZMB), Berlin, Germany, Bauer & Günther (1995) note that the ZMB catalogue lists a syntype (ZMB 7762) of Eremias benguelensis collected in “ Maconjo ” and donated by Bocage, which could not be located by the authors and was considered lost. The locality “ Maconjo ” could not be located by Bauer & Günther (1995) and has been tentatively interpreted as “ Fazenda Mucungo, Namibe Province ” by Marques et al. (2018). However, Vaz Pinto et al. (2019) noted the existence of two historical collecting localities bearing the name “ Maconjo ” in Angola: one near Capangombe in Namibe Province, where Anchieta was known to collect; and another in Benguela Province, visited by William John Ansorge between 1903 and 1905, that these authors restricted to the “ vicinity of the streams Conjo, Conjo Pequeno, and Cocumba (12 º 52 ’ S, 13 º 21 ’ E, 355 m asl), 20 km south of Uche, Benguela Province ” (Vaz Pinto et al. 2019). Considering this, if a specimen bearing the locality “ Maconjo ” was indeed donated by Bocage to the ZMB, it most likely would have been collected by Anchieta in Namibe Province, near Capangombe. However, this specimen could not have been part of the original type series, as the description of Eremias benguelensis was published before Anchieta sent the first batch of specimens from Maconjo to Bocage in early October 1867 (Banha de Andrade 1985), and reference to that specific locality was only made in Bocage (1873). Nevertheless, this putative syntype has not since been located in the ZMB collections (Bauer & Günther 1995; LMPC pers. obs.) and its status could not be verified. Boulenger (1887) referred E. benguelensis to the synonymy of E. namaquensis without providing any justification or record of Angolan specimens, and this was followed by Bocage (1895), who referred his previous records to the synonymy of this species. However, Bocage’s (1895) interpretation of “ namaquensis ” could have included up to three distinct taxa, as his material at hand comprised specimens from “ Benguella ” and “ Catumbella ”, which likely represented true P. benguelensis, and specimens from “ l’intérieur de Mossamedes ” and “ Capangombe ”, which probably represented a different taxon. Except for a specimen collected by Anchieta in Capangombe (MHNCUP / REP 0231, Fig. 11, see below), this material was lost in the fire that engulfed and destroyed all the collections of Museu Bocage in 1978 (Marques et al. 2018), making it impossible to ascertain the taxonomic identity of these specimens. Based on material collected by Ansorge and deposited in the collections of the BMNH, Boulenger was the first author to recognize the existence of two distinct species in Angola, recovering E. benguelensis as a valid species, diagnosed by the presence of a “ large transparent disc formed of a single black-edged scale ” on the lower eyelid (Boulenger 1918), and assigning specimens characterized by a “ lower eyelid with a transparent disc formed of 2 (…) larger black-edged scales ”, from “ Maconjo ” and “ Huxe ”, in Benguela Province, to Eremias undata (Boulenger 1921). However, even though Bocage (1867 b) mentioned a “ transparent disc ” on the lower eyelid, he did not mention the number of scales it comprised. The interpretation of this character can be ambiguous, considering that Bocage might have ignored the taxonomic value of this character, as did Laurent (1964) when he included both specimens with one and two transparent scales on the lower eyelid in the material he assigned to Eremias undata. Additional specimens conforming to Boulenger’s (1921) description of E. undata were collected by Wulf Haacke in Benguela and Namibe provinces during the 1970 s and are deposited in the collections of the Ditsong National Museum of Natural History. Specimens conforming to Boulenger’s (1921) description of Eremias undata, with two transparent scales on the lower eyelid, were also collected by Conradie et al (2012 b). Boulenger’s (1918, 1921) diagnosis of E. benguelensis based on the condition of the lower eyelid was accepted by subsequent authors (e. g., Parker 1936; Monard 1937; Mertens 1971; Szczerbak 1975; Mayer 1989; Arnold 1991; Branch 1998), leading Conradie et al. (2012 b) to assign newly collected material from Namibe Province to this species. During recent surveys, additional material was collected in the coastal areas of Benguela Province, in the vicinity of the original type locality of “ Benguella ”, which allowed for a reassessment of the status and diagnosis of P. benguelensis. As the original type material was lost in the fire that destroyed the collections of Museu Bocage on 18 March 1978, and the putative syntype identified by Bauer & Günther (1995) could not be located in the ZMB collections, the allocation of the name P. benguelensis to a specific taxonomic unit within the various lineages of Angolan Pedioplanis remains problematic. This situation is exacerbated by the lack of details provided by Bocage (1867 a, 1867 b) in the original descriptions, and the differences in subsequent interpretations by Boulenger (1887, 1918, 1921), Bocage (1895), Laurent (1964) and Conradie et al. (2012 b). For the purpose of taxonomic and nomenclatural stability we consider it necessary to designate a neotype for this species. Our results indicate that at least two species of Pedioplanis occur near Benguela, and Bocage’s (1867 b) vague description could apply to either of them. The only known surviving specimen that was presumably examined by Bocage (MHNCUP / REP 0231) has a single transparent scale on the lower eyelid, agreeing with Boulenger’s (1918, 1921) diagnosis of P. benguelensis. However, while the material cited by Bocage (1867 a) from Benguela was shipped by Anchieta in September 1866, the first batch of specimens from Capangombe was only shipped in July 1867 and reptiles from this locality were only cited in Bocage (1873). Therefore, this specimen could not have been part of the original type series, and was never unambiguously referred to P. benguelensis. Furthermore, it is likely that Bocage’s (1867 a) original material comprised more than one taxon, and Boulenger’s (1918, 1921) decision to allocate the name P. benguelensis to his specimens with a single transparent scale on the lower eyelid was based on his own interpretation of Bocage’s vague description, as we found no evidence suggesting that he ever examined Bocage’s original material. Instead of being consistent with Boulenger’s (1918, 1921) interpretation, we believe Bocage’s (1867 b) original description of coloration — “ ligne médiane d’une raie longitudinale noiratre (…) De chaque côté de cette raie dorsale une autre raie plus large noire, bordée en dessou de blanc ” — is the most informative portion of his description and is more consistent with the coastal populations with two transparent scales on the lower eyelid, treated by Boulenger (1921) as Eremias undata. While these specimens display three distinct and well-defined dark dorsal stripes (see Diagnosis below), those with a single transparent scale on the lower eyelid most often show an indistinct vertebral stripe and broken dorsolateral stripes (see description below). Furthermore, we considered the proximity to the type locality as a main factor for the name allocation decision. Even though both species are sympatric in the vicinity of Benguela, what we designate as P. benguelensis sensu stricto has a coastal distribution centered in Benguela, while P. benguelensis sensu Boulenger (1918, 1921) is widely distributed in inland Benguela and Namibe provinces, with records from the coastal areas being scarce. Considering that Anchieta’s first exploration of Benguela was mostly limited to coastal areas (Banha de Andrade 1985), we believe that he would be more likely to have collected the former rather than the latter. A designation of a neotype for Pedioplanis benguelensis (Bocage 1867) and an updated diagnosis for the species are provided below.	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFBCCD35FF17F89622A328F3.taxon	diagnosis	Diagnosis. Pedioplanis benguelensis is a boldly striped and medium-sized Sand Lizard, with an average SVL of 48 mm (max 58 mm) and a tail roughly two and a half times the SVL (Fig. 2). It can be distinguished from other species of the genus in Angola and Namibia by the following combination of characters: (1) lower eyelid with two enlarged transparent scales (rarely one — PEM R 24110, or three — CAS 266239), with two to four smaller ones below; (2) five (rarely four or six) supralabials anterior to the subocular and usually three posteriorly; (3) two (sometimes one or three) rows of small granules between the supraoculars and supraciliaries; (4) a group of 9 – 26 (> 13 in 75 % of specimens) small granules preceding the supraoculars; (5) ventral scales in ten longitudinal rows; (6) presence of three bold black stripes extending from the back of the head to the base of the tail. The background coloration of the dorsum is brownish, with black stripes intercalated with thinner yellowish lines. The vertebral stripe splits at the neck, and in rare cases may be entirely divided into two thinner lines. Dorsolateral stripes are bold and welldefined. On the flanks there is a dark lateral stripe that starts behind the eye and extends to the hindlimb insertion, usually faint and reticulated, and often with a series of yellow spots along its lower edge. There may also be a thin and irregular line or series of streaks of reddish to black coloration, starting behind the labials and extending to the hindlimb insertion. The hindlimbs and the tail are greyish to reddish brown, with the legs often covered above by more or less distinct pale circles surrounded by dark pigmentation. The underparts are white, sometimes reddish at the base of the tail and hindlimbs. Comparison with other Pedioplanis species. Pedioplanis benguelensis is readily distinguished from P. burchelli, P. laticeps, P. breviceps (Sternfeld, 1911), P. namaquensis and P. husabensis by the presence of two enlarged transparent scales on the lower eyelid (versus eight or more opaque to semi-transparent scales in the remaining species); from P. lineoocellata (Duméril & Bibron, 1839) by the presence of an enlarged tympanic shield (versus no enlarged tympanic shield in P. lineoocellata); from P. inornata, P. rubens, P. gaerdesi and P. branchi by the presence of distinct dorsal stripes (versus no stripes in P. inornata, P. rubens, P. gaerdesi and P. branchi); and from P. undata by a higher number of granules anterior to the supraoculars (usually> 13 in P. benguelensis versus <14 in P. undata). Pedioplanis benguelensis is identical in most morphological characters to P. mayeri, from which it can be distinguished based on a greater maximum number of granules anterior to the supraocular (9 – 26 in P. benguelensis versus 9 – 16 in P. mayeri) and geographic location (P. benguelensis restricted to Angola versus P. mayeri restricted to Namibia). Regarding Angolan congeners, P. benguelensis is distinguished from P. haackei by dorsal color pattern (dark stripes distinct all the way to the tail in P. benguelensis versus mostly fading posteriorly in P. haackei); and from P. huntleyi by a higher number of granules anterior to the supraoculars (usually> 13 in P. benguelensis versus <12 in P. huntleyi), usually two rows of granules between supraoculars and supraciliaries (versus usually one in P. huntleyi) and dorsal color pattern (dark stripes distinct all the way to the tail in P. benguelensis versus faded posteriorly in P. huntleyi). It is distinguished from an undescribed species by the presence of two enlarged transparent scales on the lower eyelid (versus one, see description below). Neotype. CAS 266242 (field number AMB 9984; Fig. 3), adult male from Chimalavera Nature Reserve, vicinity of Main Camp [- 12.83377 °, 13.16991 °, 293 m], Benguela Province. Collected by Luis M. P. Ceríaco, David C. Blackburn and Aaron M. Bauer on 11 December 2015. This specimen was selected as neotype because it was collected in the same biogeographic region and in the vicinity of the original type locality (<35 km distant), has associated genetic data, and its morphological characters closely match the original description. Additional material. 49 specimens: Benguela Province: 5 km inland from Meva beach [- 13.47778 °, 12.59222 °, 244 m] (PEM R 24225); Chimalavera Nature Reserve, vicinity of Main Camp [- 12.83377 °, 13.16991 °, 293 m] (CAS 266237 * – 266241 *, 266243 * – 266246 *); Dombe Grande-Cuio road [- 12.97805 °, 13.07391 °, 130 m] (CAS 266247 *); pass into Meva Bay [- 13.42278 °, 12.57083 °, 103 m] (PEM R 24101, 24102); near Benguela [- 12.63556 °, 13.31500 °, 64 m] (PEM R 22070); road to Benguela, S of Dombe Grande [- 13.02000 °, 13.08806 °, 280 m] (PEM R 24741, 24742); small canyon near Meva fishing village [- 13.41556 °, 12.57639 °, 30 m] (PEM R 24110 *, 24111, 27735, 27736); 10 km N of Lobito [- 12.33478 °, 13.65085 °, 301 m] (TM 46575); 35 km S of Dombe Grande [- 13.27569 °, 12.94866 °, 362 m] (TM 41254); Cimo, Equimina River [- 13.333333 °, 12.933333 °, 268 m] (TM 41245); Cuanza-Sul Province: Sumbe [- 11.20026 °, 13.84186 °, 11 m] (PEM R 25195 *, 25196 *); Namibe Province: 10.4 km S of Makonga River, on tar road to Bentiaba [- 14.84194 °, 12.427778 °, 370 m] (PEM R 24209, 24210); 10 km SE of Lucira [- 13.93139 °, 12.59444 °, 546 m] (PEM R 24076 *); road to Bentiaba, 25 km N of junction with Lubango-Namibe road [- 14.85292 °, 12.42481 °, 430 m] (PEM R 21665 *, 21666 *); sand road towards Chapeu Armando [- 14.51972 °, 12.44028 °, 471 m] (PEM R 24147); 2 km S of Chapéu Armado [- 14.46917 °, 12.33750 °, 23 m] (PEM R 24138); Mucungo farm [- 14.79528 °, 12.49500 °, 342 m] (PEM R 24142 *), [- 14.77890 °, 12.48745 °, 305 m] (CAS 266250); Bentiaba Fort [- 14.27329 °, 12.38505 °, 51 m] (CAS 266254); dirt tracks N of Moçâmedes [- 14.93503 °, 12.26406 °, 167 m] (PEM R 21648 * – 21650 *); small ridges N of Moçâmedes [- 14.92408 °, 12.37192 °, 314 m] (PEM R 21651 *, 21652 *); 31.5 km E of Namibe [- 15.01686 °, 12.39003 °, 310 m] (PEM R 18540 *); 12 km S of Bentiaba [- 14.34642 °, 12.41838 °, 248 m] (TM 41148, 41149); Bentiaba [- 14.26667 °, 12.38333 °, 11 m] (TM 25464); Lucira [- 13.86667 °, 12.53333 °, 51 m] (TM 41191); Lucira road, 5 km S of Catara River [- 13.60194 °, 12.62888 °, 331 m] (TM 41198, 41201, 41229); coastal tracks 11 km N of Namibe, 1 km before old bridge across Giraul River [- 15.08906 °, 12.17014 °, 58 m] (PEM R 21658 *). The following specimens were tentatively assigned to this species based on a combination of morphological characters and geographic location (11 specimens): Benguela Province: Cimo, Equimina River [- 13.33333 °, 12.93333 °, 268 m] (TM 41244); Namibe Province: 10 km SE of Lucira [- 13.93139 °, 12.59444 °, 546 m] (PEM R 24077); road to Bentiaba, 5 km S of Makonga River [- 14.79750 °, 12.43389 °, 394 m] (PEM R 24216, 24217); 2 km S of Chapéu Armado [- 14.46917 °, 12.33750 °, 23 m] (PEM R 24139); 2 km inland from Mucuio Bay [- 14.89611 °, 12.23722 °, 110 m] (PEM R 24155); Bentiaba [- 14.26667 °, 12.38333 °, 11 m] (TM 25466, 43965); Lucira road, 5 km S of Catara River [- 13.60194 °, 12.62888 °, 331 m] (TM 41228, 41232, 41233).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFBCCD35FF17F89622A328F3.taxon	description	Description of the Neotype. The neotype is an adult male with a complete original tail (Fig. 3). SVL 57.4 mm; TL 144.1 mm. Body relatively stout (SVL / HL 4.35), with hindlimbs longer than forelimbs and tail length two and a half times the SVL (TL / SVL 2.51). Moderately sized head (HL / SVL 0.23), distinct from the neck. Other relevant measurements are presented in Table 5. Rostral wider than high, visible from above. Nostril pierced between three scales; supranasals slightly swollen and in broad contact with each other behind rostral; infranasal in contact with rostral, first supralabial and anterior loreal; postnasal small and subquadrangular, placed between the supranasal, infranasal, anterior loreal and frontonasal. Frontonasal slightly longer than wide, rounded anteriorly and slightly acuminate posteriorly. Prefrontals in broad contact with each other, the loreals, frontonasal and frontal. Two loreals, posterior largest. Frontal longer than wide, narrower posteriorly, in contact with prefrontals anteriorly, supraoculars laterally and frontoparietals posteriorly. Paired frontoparietals in broad median contact, touching the frontal and posterior supraocular anteriorly, and the parietals and interparietal posteriorly. Interparietal pentagonal, longer than broad. Occipital small, wider than long, in broad contact with interparietal, its posterior margin slightly wider than the anterior. Parietals longer than wide, in contact with frontoparietals, interparietal and occipital. Two rounded supraoculars in contact with each other and the frontal, preceded by a group of 14 (right side) and 13 (left side) small granules, those in contact with the frontal and prefrontal largest. Two rows of small granules between supraoculars and supraciliaries. Supraciliaries six (right side) and seven (left side), first longest. Temporal scales small and granular. One narrow and elongated tympanic shield at the anterodorsal edge of ear opening. Subocular bordering lip, upper margin wider than lower. Five supralabials anterior to subocular and three posteriorly. Lower eyelid with two black edged, enlarged transparent scales, with two smaller ones below. Infralabials six. Mental wider than long, in contact with first infralabials and first pair of chin shields. Four pairs of chin shields, first three in median contact and fourth largest. Gular scales 31, in a straight line between the symphysis of the chin shields and median collar plate. Collar free, comprising seven enlarged plates. Ventral scales smooth, in ten longitudinal and 28 transverse rows; outer scales smaller than others; those on the first complete transverse row posterior to collar notably longer than others. Precloacal scales irregular and subequal, central ones largest. Fifteen femoral pores on each leg. Lamellae under fourth toe 27. Dorsal scales small and granular, larger towards ventral scales. Upper forelimbs covered above by large hexagonal plates; forearm covered above by slightly imbricate and keeled scales, larger than dorsal scales, and below by enlarged plates. Hindlimbs covered above by slightly imbricate and keeled scales, larger than dorsal scales, and below by enlarged plates. Scales on tail diagonally keeled, except for those on ventral side of basal portion, which are smooth. Coloration (in preservative). Head and limbs brownish; hindlimbs covered above with pale circles surrounded by black speckling. Dorsum with three bold and well-defined dark stripes intercalated with four thinner white to yellowish lines, extending from the back of the head to the base of the tail. The black vertebral stripe splits at the nape towards parietals. Flanks with a reticulated black stripe extending from the back of the eye to the hindlimb insertion, with a series of pale spots running along its lower edge. Tail brown above, with some black keeled scales. Moderate greyish speckling on the labials and flanks. Venter white.	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFBCCD35FF17F89622A328F3.taxon	distribution	Distribution and habitat. Known records of P. benguelensis are restricted to lower elevation areas (below 546 m) in coastal Angola, as far as 30 km inland from the coastline, from Moçâmedes, Namibe Province, northwards to Sumbe, Cuanza-Sul Province (Fig. 4). It is sympatric with an undescribed species in most of coastal Benguela Province and northern Namibe Province, and P. haackei in the littoral of northern Namibe Province. This species inhabits the xeric coastal areas of southwestern Angola, which are characterized by sublittoral steppes with bushes and herbage, mostly belonging to the plant genera Senegalia, Commiphora, Colophospermum, Aristida, Schmidtia and Setaria (Grandvaux-Barbosa 1970; Fig. 5).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB5CD31FF17F93A24492867.taxon	description	(Figs. 6 – 7)	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB5CD31FF17F93A24492867.taxon	description	This species was formally described by Conradie et al. (2012 b) based on specimens collected between Lake Arco and Espinheira, in Namibe Province. It is possible that Bocage’s (1895) records of Eremias namaquensis from “ Capangombe ” and “ l’intérieur de Mossamedes ” included representatives of this species, but the loss of this material precludes any confident conclusion. Specimens from Moçâmedes and Munhino in Namibe Province, and Quilengues in Huíla Province, collected by António de Barros Machado in 1949, were later reported by Laurent (1964) as Eremias undata undata and are still deposited in the collections of the Museu Regional do Dundo. Additional specimens were collected in Namibe Province by Wulf Haacke in the 1970 s, deposited in the collections of the Ditsong National Museum of Natural History and later reported as Pedioplanis undata (Haacke 2008). New material reported by Ceríaco et al. (2016) and in this paper considerably expand the known distribution area of Pedioplanis haackei. A specimen collected by Conradie et al. (2012 b) near the bottom of the Leba escarpment (PEM R 18460 *, field number MBUR 02132) was provisionally treated by the authors as a separate lineage, Pedioplanis sp. 2. Our analysis, however, shows that this specimen belongs to a northern clade of P. haackei, and we interpret this as an indication of intraspecific genetic structure rather than a distinct taxon.	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB5CD31FF17F93A24492867.taxon	diagnosis	Diagnosis: A small Pedioplanis with an average SVL of 45 mm (max 59 mm), with a tail length roughly two and a half times the SVL (Fig. 6). It can be distinguished from other species of the genus in Angola and neighboring regions by the following combination of characters: (1) lower eyelid with two enlarged transparent scales, often with a row of smaller ones below; (2) five (rarely four or six) supralabials anterior to the subocular and two to three posteriorly; (3) two (rarely one or three) rows of small granules between the supraoculars and supraciliaries; (4) a group of 7 – 28 (> 12 in 75 % of specimens) small granules anterior to the supraoculars; (5) ventral scales in ten (rarely nine or 11) longitudinal rows; (6) variable color pattern, usually with three dark dorsal stripes faded posteriorly. Among Angolan Pedioplanis, this species seems to be the most variable in terms of color pattern. The dorsum is greyish brown anteriorly and pale brown to brick red posteriorly, usually with dark stripes starting at the back of the head and fading between midbody and two-thirds along the back. The vertebral stripe splits or widens at the neck, or may be entirely split into two thinner stripes, with a greyish interspace. Dorsolateral stripes are usually wider and more distinct than the vertebral one. On the flanks there is a dark lateral stripe that starts behind the eye and usually fades posteriorly, often faint and reticulated, with a series of yellow to blueish circles running along its lower edge. However, the extension and distinctiveness of dorsal stripes is quite variable, being completely absent in some specimens, and extending nearly to the base of the tail in others. Hind limbs and tail reddish-brown to brick red, with the hindlimbs sometimes covered above by faint pale circles. White ventrally, sometimes reddish at the base of the tail. Comparison with other Pedioplanis species. Pedioplanis haackei is readily distinguished from P. burchelli, P. laticeps, P. breviceps, P. namaquensis and P. husabensis by the presence of two enlarged transparent scales on the lower eyelid (versus eight or more opaque to semi-transparent scales in other species); from P. lineoocellata by the presence of an enlarged tympanic shield (versus no enlarged tympanic shield in P. lineoocellata); from P. inornata, P. rubens, P. gaerdesi and P. branchi by the presence of dorsal stripes (versus no stripes in P. inornata, P. rubens, P. gaerdesi and P. branchi); from P. undata by color pattern (bold anterior dorsal striping that fades posteriorly in P. haackei versus dorsal striping bold or not, may be reduced with pale longitudinal elements or even a single middorsal stripe restricted to the nape in P. undata) and a greater maximum number of granules anterior to the supraoculars (7 – 28 in P. haackei versus 8 – 13 in P. undata); and from P. mayeri by color pattern (dorsal stripes usually faded posteriorly in P. haackei versus bold and distinct all the way to the tail in P. mayeri), and a greater maximum number of granules anterior to the supraocular (7 – 28 in P. haackei versus 9 – 16 in P. mayeri). It can be further distinguished from P. undata and P. mayeri by geographic location (P. haackei restricted to Angola versus P. mayeri and P. undata restricted to Namibia). With respect to Angolan congeners, P. haackei is distinguished from P. benguelensis by color pattern (posterior dorsum usually reddish with faded stripes in P. haackei versus bold stripes all the way to the tail in P. benguelensis); it is distinguished from P. huntleyi by a smaller average SVL (mean 45 mm in P. haackei versus 54 mm in P. huntleyi), a higher number of granules anterior to the supraoculars (usually> 11 in P. haackei versus <12 in P. huntleyi) and two rows of granules between supraoculars and supraciliaries (versus one in P. huntleyi). It is distinguished from an undescribed species by the presence of two transparent scales on the lower eyelid (versus one, see description below).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB5CD31FF17F93A24492867.taxon	materials_examined	Holotype: PEM R 18465 *, an adult male collected along the road to Tambor, Namibe Province [- 15.87606 °, 12.20583 °, 196 m], by William R. Branch, G. John Measey, Werner Conradie and Krystal A. Tolley on 19 January 2009. Additional material. 121 specimens: Benguela Province: 2 km from Lucira-Benguela road, towards Cape Santa Maria [- 13.51167 °, 12.62833 °, 285 m] (PEM R 24087); Huíla Province: 13 km N of Quilengues, on road to Benguela [- 13.97247 °, 14.04717 °, 936 m] (PEM R 21661 *); Namibe Province: 10 km S of Lake Arco [- 15.83044 °, 12.14125 °, 121 m] (PEM R 18470, 18471 *, 18472 *); on the road to Benguela, 2 km N of turning to Lucira [- 13.90528 °, 12.54417 °, 312 m] (AG 98 *); road to Bentiaba, 52 km N of junction with Lubango-Namibe road [- 14.65806 °, 12.52717 °, 586 m] (PEM R 21662 *, 21663 *); 5 km along fence leading to Baptista farm [- 16.05847 °, 12.42597 °, 343 m] (PEM R 18468); Humpata-Namibe road, 7.8 km from bottom of Leba pass [- 15.04467 °, 13.15947 °, 642 m] (PEM R 18460 *, 21664 *); along road to Tambor [- 15.88778 °, 12.36417 °, 300 m] (PEM R 18464 *), [- 15.87606 °, 12.20583 °, 196 m] (PEM R 18466 – 18467); base of Serra da Neve, on dirt road to Quilengues [- 13.81594 °, 13.32644 °, 892 m] (MHNC-UP / REP 645); base of Serra da Neve, on dirt road to the top of the mountain [- 13.83424 °, 13.27669 °, 803 m] (MHNC-UP / REP 634 - 635, CAS 266123); Catara River valley [- 13.62417 °, 12.83556 °, 454 m] (PEM R 24057 *, 24058, 24059, 24060, 24061 *, 24062 *); Caraculo [- 15.01651 °, 12.64251 °, 487 m] (CAS 264769 *); Chipumpo-Virulundo [- 16.19125 °, 12.85595 °, 539 m] (CAS 264805 *); Curoca-Omahua [- 15.81918 °, 12.14273 °, 100 m] (CAS 264774 *, 264775 *, 264776), [- 15.95026 °, 12.49531 °, 394 m] (CAS 264777 * – 264779 *); dry river valley, approximately 10 km S of turning to Lucira [- 13.96389 °, 12.53222 °, 137 m] (PEM R 24223, 24224); Lucira Dam [- 13.87889 °, 12.56556 °, 339 m] (PEM R 24103 *, 24104); Mucungo farm [- 14.78111 °, 12.48983 °, 296 m] (MHNCUP / REP 633), [- 14.77890 °, 12.48745 °, 305 m] (CAS 266248); Munhino [- 14.92000 °, 13.00000 °, 393 m] (MD 1918 b); near Virulundo [- 16.28523 °, 12.94192 °, 718 m] (CAS 264806 * – 264808 *, 264810 *); near Curoca River [- 16.24468 °, 12.34068 °, 265 m] (CAS 264781 *); off road 20 km N of Omauha Lodge [- 16.07414 °, 12.43328 °, 352 m] (PEM R 18474 *); Omauha-Virei [- 16.14506 °, 12.59460 °, 538 m] (CAS 264791 *); Omauha Lodge [- 16.19858 °, 12.40073 °, 340 m] (CAS 264780 *, 264784 * – 264790 *, MHNC-UP / REP 644, INBAC / AMB 10416 *), [- 16.20033 °, 12.40003 °, 343 m] (CAS 254835 *); Omauha Lodge, N of entrance to Iona National Park [- 15.99694 °, 12.40694 °, 301 m] (PEM R 18473); Red Canyons near Lake Arco [- 15.74597 °, 12.13989 °, 81 m] (PEM R 18461 *, 18462, 18463); road from Lake Arco to Espinheira [- 15.91356 °, 12.39519 °, 319 m] (PEM R 18475 *); road to Tambor [- 15.88119 °, 12.22267 °, 192 m] (PEM R 18469 *); sandstone hill near Piambo [- 14.65833 °, 12.36944 °, 365 m] (PEM R 22041); vicinity of N’Dolondolo [- 13.80866 °, 13.13521 °, 731 m] (CAS 264757 *, 264765 *); Virei-Virulundo [- 16.31018 °, 12.79597 °, 471 m] (CAS 264794 *, 264801 *, 264803 *); 10 km E of Munhino [- 14.87821 °, 13.13195 °, 566 m] (TM 46726); 12 km N of Furnas [- 15.02513 °, 12.18828 °, 69 m] (TM 40570); 12 km N of Tambor [- 15.96780 °, 12.39581 °, 262 m] (TM 40496); 23 km W of Virei [- 15.66886 °, 12.74089 °, 397 m] (TM 41025); 30 km N of Tambor [- 15.89287 °, 12.37609 °, 312 m] (TM 40489 – 40491); 35 km S of Moçâmedes [- 15.48000 °, 12.21000 °, 150 m] (MD 1946); 3 km E of Chicambi village [- 13.89556 °, 12.78222 °, 612 m] (AG 69 * – 71 *); 5 km (by road) NW of Pico Azevedo [- 15.47547 °, 12.46219 °, 408 m] (CAS 254763 *, 254764 *), [- 15.47600 °, 12.46150 °, 399 m] (CAS 254767 *); 60 km (by road) E of Tômbua [- 15.87760 °, 12.21668 °, 189 m] (TM 40451 – 40453); Assunção [- 14.86667 °, 13.10000 °, 505 m] (TM 40181); 17 km W of Chicamba village [- 13.91722 °, 12.68111 °, 531 m] (AG 84 *); Inamangando farm [- 14.05114 °, 12.42733 °, 24 m] (TM 41157, 41158); Iona National Park, N of Tambor [- 15.99539 °, 12.40647 °, 306 m] (CAS 254840 *); Iona National Park, Curoca river crossing [- 16.30000 °, 12.43333 °, 198 m] (TM 40580); Lungo [- 14.66667 °, 13.25000 °, 679 m] (TM 24397, 24398, 24400); Moçâmedes [- 15.16667 °, 12.16667 °, 6 m] (TM 22844 – 22845); Pediva Hot Springs, S side of the river [- 16.29381 °, 12.56033 °, 235 m] (CAS 254860 *); Pico Azevedo [- 15.53400 °, 12.49197 °, 359 m] (CAS 254939); Namibe Nature Reserve [- 15.77292 °, 12.33269 °, 262 m] (CAS 254951 *, 254953 *); Saiona River [- 15.40000 °, 13.20000 °, 534 m] (TM 40983 – 40985); Tambor [- 16.13556 °, 12.42972 °, 379 m] (TM 40501, 40502), [- 16.06710 °, 12.42970 °, 348 m] (INBAC / AMB 10393 *). The following specimens were tentatively assigned to this species based on a combination of morphological characters and geographic location (10 specimens): Benguela Province: 2 km turning on Cape Santa Maria from Lucira-Benguela coastal road [- 13.51167 °, 12.62833 °, 285 m] (PEM R 24086, 24088); Namibe Province: 13.6 km S of Mucungo Farm [- 14.85528 °, 12.42889 °, 400 m] (PEM R 24153); 1 km SE of Mucungo Farm [- 14.78667 °, 12.49611 °, 303 m] (PEM R 24133 – 24135); 8.8 km S of Mucungo Farm [- 14.80167 °, 12.41917 °, 579 m] (PEM R 24160); Catara River valley [- 13.62278 °, 12.80750 °, 454 m] (PEM R 27734); Lucira [- 13.86667 °, 12.53333 °, 51 m] (TM 24430, 41190).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB5CD31FF17F93A24492867.taxon	distribution	Distribution and habitat. Pedioplanis haackei occurs over most of Namibe Province north of - 16 º latitude, extending marginally into southern Benguela Province, and northwestern Huíla Province below 1000 m (Fig. 7). It is absent from the more xeric areas of southwestern Namibe Province, as well as southeastern Namibe Province, where it is replaced by P. huntleyi, with both species occurring in sympatry near Virulundo and Pediva Hot Springs. Pedioplanis haackei is sympatric with P. benguelensis in coastal areas, and with the undescribed species in several localities along most of its range. This species inhabits a vast area covering different habitat types, from more xeric areas to areas dominated by Mopane woodlands (Grandvaux-Barbosa 1970; Fig. 5).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB0CD32FF17FF0422812D07.taxon	description	(Figs. 8 – 9)	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB0CD32FF17FF0422812D07.taxon	diagnosis	Diagnosis: A rather large Pedioplanis, with an average SVL of 55 mm (max 59 mm) and a tail length two and a half times the SVL (Fig. 8). It can be distinguished from other species of the genus in Angola and neighboring regions by the following combination of characters: (1) lower eyelid with two enlarged transparent scales, usually with a row of smaller ones below; (2) five (rarely four or six) supralabials anterior to the subocular and three posteriorly; (3) one (sometimes two) row of small granules between the supraoculars and supraciliaries; (4) supraoculars preceded by a group of 6 – 13 (<12 in 75 % of specimens) small granules; (5) ventral scales in ten longitudinal rows; (6) three dark dorsal stripes faded posteriorly. Coloration similar to P. haackei, but less variable. Dorsum greyish brown anteriorly and reddish-brown to brick red posteriorly, with three dark stripes starting behind the head and fading between midbody and two-thirds along the back. The vertebral stripe splits or widens at the neck and is often more faded than the dorsolateral stripes. On the flanks there is a dark lateral stripe that starts behind the eye and usually fades posteriorly, often faint and reticulated, with a series of yellow to blueish circles running along its lower edge. Hind limbs and tail reddish-brown to brick red, with the hindlimbs sometimes covered above by faint pale circles. White ventrally, sometimes reddish at the base of the tail. Comparison with other Pedioplanis species. Pedioplanis huntleyi is readily distinguished from P. burchelli, P. laticeps, P. breviceps, P. namaquensis and P. husabensis by the presence two enlarged transparent scales on the lower eyelid (versus eight or more opaque to semi-transparent scales in other species); from P. lineoocellata by the presence of an enlarged tympanic shield (versus no enlarged tympanic shield in P. lineoocellata); from P. inornata, P. rubens, P. gaerdesi and P. branchi by the presence of dorsal stripes (versus no stripes in P. inornata, P. rubens, P. gaerdesi and P. branchi). P. huntleyi is identical in most morphological characters to P. undata and P. mayeri, from which it can be distinguished based on color pattern (dorsal stripes always faded posteriorly in P. huntleyi versus usually bold and distinct all the way to the tail in P. mayeri and bold or faded in P. undata) and geographic location (P. huntleyi restricted to Angola versus P. mayeri and P. undata restricted to Namibia). With respect to other Angolan species, P. huntleyi can be distinguished from P. benguelensis by a lower number of granules anterior to the supraoculars (usually <12 in P. huntleyi versus> 13 in P. benguelensis), one row of granules between the supraoculars and supraciliaries (versus two in P. benguelensis) and color pattern (dorsal stripes faded posteriorly in P. huntleyi versus distinct all the way to the tail in P. benguelensis); it is distinguished from P. haackei by a larger SVL (average 54 mm in P. huntleyi versus 45 mm in P. haackei), a lower number of granules anterior to the supraoculars (usually <11 in P. huntleyi versus> 12 in P. haackei) and one row of granules between the supraoculars and supraciliaries (versus two in P. haackei). It is distinguished from an undescribed species by the presence of two transparent scales on the lower eyelid (versus one, see description below).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB0CD32FF17FF0422812D07.taxon	materials_examined	Holotype: PEM R 18479 *, an adult male collected on the road to Oncocua, 7 km from Iona, Namibe Province [- 16.85831 °, 12.61275 °, 803 m], by Werner Conradie, Eduardo Traguedo, Krystal A. Tolley and William R. Branch on 21 January 2009. Additional material. 26 specimens: Cunene Province: 26 km SE of Oncocua [- 16.86889 °, 13.52750 °, 846 m] (PEM R 18483, 18489, 18490 *); NE of Elola village [- 16.76251 °, 13.24230 °, 892 m] (MHNC-UP / REP 643); Namibe Province: 14 km W of Moimba [- 16.67944 °, 12.97389 °, 684 m] (PEM R 18481, 18487 *); 23 km W of Moimba [- 16.67444 °, 12.88500 °, 602 m] (PEM R 18482, 18488); 16 km E of Iona [- 16.79806 °, 12.68056 °, 785 m] (PEM R 18480, 18485 *); 26 km E of Iona [- 16.70056 °, 12.82417 °, 623 m] (PEM R 18486); 8 km NE of Iona [- 16.82917 °, 12.62111 °, 743 m] (PEM R 18484 *); Iona National Park [- 16.43150 °, 12.48930 °, 431 m] (MHNC- UP / REP 640 – 642), [- 16.85972 °, 12.61111 °, 811 m] (TM 40739); Iona National Park, 50 km S of Curoca River [- 16.73367 °, 12.40757 °, 472 m] (TM 40596); near Virulundo [- 16.28523 °, 12.94192 °, 718 m] (INBAC / AMB 10632 *), [- 16.29485 °, 12.94075 °, 759 m] (CAS 264811 * – 264813 *, INBAC / AMB 10645 *); road to Espinheira, 40 km S of Omauha Lodge [- 16.51164 °, 12.44761 °, 371 m] (PEM R 18476); road to Oncocua, 7 km from Iona [- 16.85834 °, 12.61275 °, 803 m] (PEM R 18477, 18478 *); Pediva Hot Springs, south side of the river [- 16.29000 °, 12.56219 °, 270 m] (CAS 254864 *).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB0CD32FF17FF0422812D07.taxon	distribution	Distribution and habitat. Known records of Pedioplanis huntleyi are restricted to southwestern Angola south of - 16 ° latitude, from eastern Iona National Park in Namibe Province eastwards to the Oncocua region in Cunene Province, between 270 and 892 m (Fig. 9). This species is sympatric with P. haackei in some localities along the northern limits of its range. This species inhabits xeric areas dominated by Mopane forests (Colophospermum mopane) (Grandvaux-Barbosa 1970; Fig. 5).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB2CD08FF17FB5D23E62E3D.taxon	description	(Figs. 10 – 13)	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB2CD08FF17FB5D23E62E3D.taxon	description	The first records of this species were probably those provided by Bocage (1895) as Eremias namaquensis from “ Capangombe par M. d’Anchieta ” and “ l’intérieur de Mossamedes par MM. Capello et Ivens ”. However, earlier records as Eremias benguelensis from “ Benguella ” (Bocage 1867 a, 1867 b) could have also represented this taxon. Nevertheless, the loss of these specimens precludes any further inferences regarding their taxonomic identity. The only exception is a specimen collected by Anchieta in Capangombe (MHNCUP / REP 0231, Fig. 11) and still deposited in the collections of MHNC-UP, which represents the first confirmed record of this species. Boulenger’s (1918, 1921) diagnosis of “ benguelensis ” was characterized by the presence of a single transparent scale on the lower eyelid, which was present in his specimens from “ Huxe ” [= Uche, Benguela Province] and “ Ponang Kuma ” [= Dongoena, Cunene Province], collected by Ansorge. Boulenger’s (1918, 1921) diagnosis was followed by most authors (e. g., Monard 1937; Szczerbak 1975; Mayer 1989; Branch 1998) and Parker (1936) reported a specimen (BMNH 1936.8.1.514) from “ Catengue ”, Benguela Province, collected during the Karl Jordan’s expedition to South-West Africa and Angola and deposited in the collections of the Natural History Museum, London. Laurent (1964) reported additional Angolan material under the name Eremias undata undata, including among his records two specimens with a single transparent scale on the lower eyelid, collected by Barros Machado in Munhino, Namibe Province (MD 1918 a), and Quilengues, Huíla Province (MD 1970 - 1). Following Boulenger’s (1918, 1921) diagnosis and the lack of topotypic comparative material, Conradie et al. (2012 b) conservatively assigned specimens collected in central and southeastern Namibe Province to P. benguelensis, providing the first molecular data for this species. Three additional specimens conforming to Boulenger’s (1918, 1921) diagnosis of P. benguelensis were collected by Ceríaco et al. (2016) near Caraculo, Namibe Province. The collection of true P. benguelensis from the coastal areas of Benguela Province allowed for a reassessment of Angolan populations assigned to this species. Our analysis revealed species-level phylogenetic distance as well as consistent morphological differences from all named species of Pedioplanis. Based on the evidence produced in this study, we here describe a new species of Angolan Pedioplanis characterized by the presence of a single transparent scale on the lower eyelid (see P. benguelensis account for discussion on the name allocation decision).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB2CD08FF17FB5D23E62E3D.taxon	diagnosis	Diagnosis: Pedioplanis serodioi sp. nov. is a rather small Pedioplanis, with an average SVL of 44 mm (max 50 mm) and a tail length roughly twice the SVL (Fig. 10). It can be distinguished from other Pedioplanis species in Angola and neighboring regions by the following combination of characters: (1) lower eyelid with a single blackedged, enlarged transparent scale; (2) four (rarely three, five or six) supralabials anterior to the subocular and two (rarely three) posteriorly; (3) One (sometimes two) row of small granules between the supraoculars and supraciliaries; (4) a group of 6 – 24 (<15 in 75 % of specimens) small granules preceding the supraoculars; (5) ventral scales in ten longitudinal rows; (6) a pair of irregularly edged dorsolateral stripes or series of spots and a faint or absent vertebral stripe. The dorsum is greyish brown to pale grey, often with extensive black speckling throughout. There may be a thin, dark vertebral stripe, but this is usually faint and indistinct, or reduced to irregular black streaks. A pair of irregularly edged, black dorsolateral stripes extend from the back of the head to the base of the tail, often broken into series of black spots on a reddish-brown background. A pale line separates the dorsolateral stripes from a lateral stripe that starts behind the eye and extends to the base of the tail, often faint and reticulated, with a series of yellow spots running along its lower edge. There is often a reddish stripe extending from the posterior labials to the hindlimb insertion, up to four scales wide, continuous or broken into series of streaks. The hindlimbs are often covered above by pale circles surrounded by black pigmentation. Venter white. Comparison with other Pedioplanis species. Pedioplanis serodioi sp. nov. is distinguished from P. gaerdesi by the number of supralabials anterior to the subocular (usually four in P. serodioi sp. nov. versus five to six in P. gaerdesi) and color pattern (dark dorsolateral stripes or series of spots in P. serodioi sp. nov. versus no dorsal stripes in P. gaerdesi); and from all the remaining species of the genus by consistently possessing a single transparent scale on the lower eyelid (versus two or more scales in other species). Regarding Angolan congeners, it can be further distinguished from P. benguelensis by a lower number of supralabials anterior to the subocular (usually four in P. serodioi sp. nov. versus five in P. benguelensis), one row of granules between supraoculars and supraciliaries (versus two in P. benguelensis), fewer granules anterior to the supraoculars (usually <15 in P. serodioi sp. nov. versus> 13 in P. benguelensis) and color pattern (dorsolateral stripes often broken and vertebral stripe indistinct in P. serodioi sp. nov. versus all stripes distinct and well-defined in P. benguelensis); from P. haackei by the presence of one row of granules between supraoculars and supraciliaries (versus two in P. haackei), fewer granules anterior to the supraoculars (usually <15 in P. serodioi sp. nov. versus> 11 in P. haackei) and color pattern (dorsolateral stripes often broken and vertebral stripe indistinct in P. serodioi sp. nov. versus dorsal stripes continuous and faded posteriorly in P. haackei); and from P. huntleyi by a smaller SVL (average 43 mm in P. serodioi sp. nov. versus 54 mm in P. huntleyi), a lower number of supralabials anterior to the subocular (four in P. serodioi sp. nov. versus five in P. huntleyi) and color pattern (dorsolateral stripes often broken and vertebral stripe indistinct in P. serodioi sp. nov. versus dorsal stripes continuous and faded posteriorly in P. huntleyi).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB2CD08FF17FB5D23E62E3D.taxon	materials_examined	Holotype. CAS 254906 (field number JVV 8598, Fig. 12), adult male collected on the north side of the road from Namibe to Lubango, road marker 59, 1.8 km West (by road) of Caraculo [- 15.01888 °, 12.64014 °, 491 m], Namibe Province, by Luis M. P. Ceríaco, E. L. Stanley, Arianna L. Kuhn, Jens V. Vindum, Sango de Sá, Suzana Bandeira and Hilária Valério on 6 December 2013. Paratypes. 11 specimens: CAS 254909 * (field number JVV 8601), with the same collecting data as the holotype. CAS 264752 * (field number AMB 10181) from Bibala Crossroad [- 14.74398 °, 13.32852 °, 863 m], collected by Luis M. P. Ceríaco, Suzanna Bandeira and Ishan Agarwal on 18 November 2016. CAS 264767 * and CAS 264768 * (field numbers AMB 10364 and 10365, respectively) from Bibala Crossroad [- 14.74398 °, 13.32852 °, 863 m], collected by Luis M. P. Ceríaco, Suzanna Bandeira and Ishan Agarwal on 23 November 2016. CAS 264754 * (field number AMB 10183) from Camacuio [- 14.11365 °, 13.24320 °, 670 m], collected by Luis M. P. Ceríaco, Suzanna Bandeira and Ishan Agarwal on 18 November 2016. CAS 264766 * (field number AMB 10334) from N’Dolondolo [- 13.81328 °, 13.13618 °, 681 m], collected by Luis M. P. Ceríaco, Suzanna Bandeira and Ishan Agarwal on 22 November 2016. MHNCUP / REP 0231, collected by José d’Anchieta in Capangombe [- 14.93309 °, 12.96477 °, 422 m]. PEM R 18449 * and 18450 * (field numbers MBUR 02146 and 02147, respectively), collected by Werner Conradie, William R. Branch, Krystal A. Tolley, G. John Measey among rocks and boulders by roadside 50 km E of Namibe [- 15.01661 °, 12.55650 °, 516 m] on 18 January 2009. PEM R 18453 * (field number MBUR 02165), collected by Dirk Bellstedt among rocks and boulders by roadside 50 km E of Namibe [- 15.01958 °, 12.52458 °, 550 m] on 18 January 2009. PEM R 18459 * (field number WC 09 - 33) from east of Moimba [- 16.71833 °, 13.12194 °, 834 m], collected by Werner Conradie on 24 January 2009. Additional material. 70 specimens: Benguela Province: Chimalavera Nature Reserve, vicinity of Main Camp [- 12.83377 °, 13.16991 °, 293 m] (INBAC / AMB 9967 *); Catengue [- 13.06222 °, 13.75611 °, 586 m] (PEM R 27699, 27700); 7 km east of Catengue, on road to Cubal [- 12.99942 °, 13.79856 °, 814 m] (PEM R 21659 *, 21660); 30 km SSW of Benguela [- 12.74389 °, 13.15167 °, 55 m] (PEM R 22071, 22072); 2 km turning on Cape Santa Maria from Lucira-Benguela coastal road [- 13.51167 °, 12.62833 °, 285 m] (PEM R 24089); Maconjo, near Benguela-Catengue road [- 12.85472 °, 13.39472 °, 399 m] (PEM R 27701); 15 km E of Lobito [- 12.37056 °, 13.72833 °, 252 m] (PEM R 27702); Namibe Province: Capangombe [- 15.09738 °, 13.13909 °, 542 m] (CAS 264240); base of Serra da Neve, on dirt road to the top of the mountain [- 13.83424 °, 13.27669 °, 803 m] (MHNC-UP / REP 636, CAS 266122); base of Serra da Neve, on dirt road to Quilengues [- 13.81594 °, 13.32644 °, 892 m] (MHNC-UP / REP 637, 638); base of Serra da Neve, Malowe village [- 13.83549 °, 13.27547 °, 798 m] (MHNC-UP / REP 639, 646); Munhino [- 14.96667 °, 12.96667 °, 393 m] (MD 1918 a); 50 km E of Moçâmedes on main road to Leba [- 15.01558 °, 12.55503 °, 516 m] (PEM R 21653 *, 21654 *, 21655 *, 21656 *, 21657 *); Camacuio [- 14.11365 °, 13.24320 °, 670 m] (CAS 264755 *); 10 km W of Lola, road northwest to Camacuio [- 14.29028 °, 13.53056 °, 802 m] (PEM R 24006 *, 24007 *, 24008, 24009, 24010 *, 24011, 24012); 30 km W of Lola, road northwest to Camacuio [- 14.27583 °, 13.45806 °, 791 m] (PEM R 24032 * – 24035 *); just before Camacuio [- 14.14333 °, 13.27556 °, 712 m] (PEM R 24050 * – 24052 *); 50 km E of Namibe by roadside [- 15.01661 °, 12.55650 °, 516 m] (PEM R 18451, 18452); approximately 20 km S of Bentiaba [- 14.40278 °, 12.44972 °, 426 m] (PEM R 24220); approximately 20 km SW of Camacuio [- 14.21472 °, 13.40556 °, 803 m] (PEM R 27733 *); road between Namibe and Humpata 47 km E of Namibe [- 15.01942 °, 12.52881 °, 527 m] (PEM R 18454 *, 18455 – 18458); road north of Bibala towards Lola [- 14.41611 °, 13.56472 °, 920 m] (PEM R 24026 – 24027); Bibala Crossroad [- 14.74398 °, 13.32852 °, 863 m] (CAS 264753); Namibe-Lubango road, 1.8 km W (by road) of Caraculo [- 15.01888 °, 12.64014 °, 491 m] (CAS 254896 *); Caraculo [- 15.01667 °, 12.66667 °, 463 m] (TM 40211); 14 km NE of Caraculo [- 14.91304 °, 12.73356 °, 446 m] (TM 40210, 40254 – 40262); Leba Pass bottom [- 15.04108 °, 13.19029 °, 696 m] (TM 46756); Lungo [- 14.66667 °, 13.25 °, 679 m] (TM 24399); 10 km E of Munhino [- 14.87821 °, 13.13195 °, 566 m] (TM 46727 – 46728); Pico do Azevedo [- 15.55000 °, 12.51667 °, 347 m] (TM 41071, 41072); Huíla Province: 15 km N of Quilengues [- 13.94208 °, 14.04557 °, 927 m] (MD 1970 - 1); between Chingoroi and Quilengues [- 13.87278 °, 13.96528 °, 926 m] (PEM R 22068).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB2CD08FF17FB5D23E62E3D.taxon	description	Description of the Holotype. The holotype is an adult male with a complete original tail (Fig. 12). SVL 48 mm; TL 104 mm. Body relatively stout (SVL / HL 4.29), with hindlimbs longer than forelimbs and tail 2.16 times as long as the SVL. Moderately sized head (HL / SVL 0.23), distinct from the neck. Other relevant measurements are presented in Table 5. Rostral wider than high, visible from above. Nostril pierced between three scales; supranasals slightly swollen and in broad contact with one another behind rostral; infranasal in contact with rostral, first supralabial and anterior loreal; postnasal small and subquadrangular, placed between supranasal, infranasal, anterior loreal and frontonasal. Frontonasal hexagonal, as long as it is wide. Prefrontals in broad contact with each other, the loreals, frontonasal and frontal. Two loreals, posterior largest. Frontal longer than wide, narrower posteriorly, in contact with prefrontals anteriorly, supraoculars laterally and frontoparietals posteriorly. Paired frontoparietals in broad median contact, touching frontal and posterior supraocular anteriorly, and the parietals and interparietal posteriorly. Interparietal longer than broad, rounded anteriorly, its posterior margin much narrower than the anterior. Occipital small and slightly wider than it is long, posterior margin more than twice as wide as anterior, in broad contact with interparietal. Parietals longer than wide, in contact with frontoparietals, interparietal and occipital. Two rounded supraoculars in contact with each other and the frontal, preceded by a group of 14 (right side) and 13 (left side) small granules, those in contact with the frontal and prefrontal largest. One row of small granules between anterior supraocular and supraciliaries, increasing to two rows posterior to supraocular suture. Supraciliaries six, the first longest. Temporal scales small and granular. One narrow and elongated tympanic shield at anterodorsal edge of the ear opening. Subocular bordering lip, upper margin much wider than lower. Five (right side) and four (left side) supralabials anterior to subocular and two posterior to subocular. Lower eyelid with a single black-edged, enlarged transparent scale. Infralabials six. Mental wider than long, in contact with first infralabials and first pair of chin shields. Four pairs of chin shields, first three in median contact and fourth largest. Gular scales 26, in a straight line between symphysis of the chin shields and median collar plate. Collar free, comprising six enlarged plates. Ventral scales smooth, in nine longitudinal and 27 transverse rows; outer scales smaller than others; first complete transverse row posterior to collar notably longer than others. Precloacal scales irregular and subequal, with a single scale larger than the other in the center. Femoral pores 13 on each leg. Lamellae under fourth toe 23. Dorsal scales small and granular, larger towards the ventral scales. Upper forelimb covered above by large hexagonal plates; forearm covered above by slightly imbricate and keeled scales, larger than dorsal scales, and below by enlarged plates. Hindlimbs covered above by slightly imbricate and keeled scales, larger than dorsal scales, and below by enlarged plates. Scales on tail diagonally keeled, except for those on ventral side of basal portion, which are smooth. Coloration (in preservative). Dorsum pale greyish with some black speckling, especially on the limbs and tail; hindlimbs covered above by pale circles surrounded by black pigmentation. There is a barely distinct, greyish vertebral stripe that splits at the neck, as well as a pair of dorsolateral series of black spots extending to the base of the tail. Lateral stripe somewhat faint and reticulated, with a series of pale circles running along its lower edge. Tail greyish above, with some black speckling and dark keeled scales. White ventrally. Variation in the type series. Variation in scalation and measurements among the type series of Pedioplanis serodioi sp. nov. is reported in Table 5. Except for minor differences in scalation and color pattern, all paratypes generally agree with the holotype. The prefrontals are separated by a small azygous scale in MHNCUP / REP 0231, CAS 254909, CAS 264766, PEM R 18453 and PEM R 18459, while in CAS 264768 and CAS 264752 there are two longitudinally aligned scales between the prefrontals. Unlike the holotype, dorsolateral stripes are continuous, but irregularly outlined, in CAS 254909, CAS 264752, CAS 264768, CAS 264766, PEM R 18453, and indistinct in MHNCUP / REP 0231. The remaining paratypes agree with the holotype in having broken (at least partially) dorsolateral stripes.	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB2CD08FF17FB5D23E62E3D.taxon	distribution	Distribution and habitat. Pedioplanis serodioi sp. nov. has a broad distribution in the low elevation areas of southwestern Angola, from central Benguela Province to western Cunene Province, except for the more xeric areas of southwestern Namibe Province (Fig. 13). Voucher specimens from northern Namibia deposited in the Ditsong National Museum of Natural History (TM 38789 – 95 from Okjivakandu; TM 33292 and 38903 from Opuwo; TM 38868, 38870 – 71 from Otjiwise) are provisionally referred to Pedioplanis cf. serodioi sp. nov. based on the presence of a single transparent scale on the lower eyelid and dorsal coloration, but the identity of these populations is pending genetic confirmation (see Discussion). The current gaps in the distribution of P. serodioi sp. nov. are likely an artifact of incomplete sampling, and further surveys in southern Angola and adjacent northern Namibia shall improve our knowledge of the distribution range of this species. At 1121 m above sea level, the specimens collected by Ansorge in “ Ponang Kuma ” [= Dongoena, Cunene Province] (Boulenger 1921) represent the highest elevation recorded for any Angolan Pedioplanis. It occurs sympatrically with all other Angolan congeners in several localities across its range. This species inhabits a vast area covering different habitat types, from more xeric areas to areas dominated by Mopane woodlands (Grandvaux-Barbosa 1970; Fig. 5).	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
03DBD63AFFB2CD08FF17FB5D23E62E3D.taxon	etymology	Etymology. The species is named after the Angolan scholar João Manuel Serôdio de Almeida (1943 – present), professor of the Biology Department of the Faculty of Sciences of Agostinho Neto University, Luanda, Angola. João Serôdio has had a pivotal role in scientific research and biodiversity conservation in Angola, as manager of several conservation areas, Vice-Minister for the Environment (1997 – 2000) and by training several generations of Angolan biologists. Prof. Serôdio has been a strong supporter and advocate of the present herpetological research in Angola. The name is formed in the genitive masculine singular. We propose the English common name of Serôdio’s Sand Lizard, and the Portuguese common name of Lagartixa da Areia de Serôdio.	en	Parrinha, Diogo, Marques, Mariana P., Heinicke, Matthew P., Khalid, Farkhanda, Parker, Kelly L., Tolley, Krystal A., Childers, Jackie L., Conradie, Werner, Bauer, Aaron M., Ceríaco, Luis M. P. (2021): A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 1-46, DOI: 10.11646/zootaxa.5032.1.1
