identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E6654F682FFF8CFC979118E841FD3A.text	03E6654F682FFF8CFC979118E841FD3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptarma Shahdadi, Fratini & Schubart 2020	<div><p>Genus Leptarma Shahdadi, Fratini &amp; Schubart, 2020</p> <p>Type species. — Sesarma leptosoma Hilgendorf, 1869, by original designation.</p> </div>	https://treatment.plazi.org/id/03E6654F682FFF8CFC979118E841FD3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Shahdadi, Adnan;Mendoza, Jose Christopher E.	Shahdadi, Adnan, Mendoza, Jose Christopher E. (2023): Two new species of the mangrove crab genus Leptarma (Crustacea: Brachyura: Sesarmidae) from the Western Pacific islands. Raffles Bulletin of Zoology 71: 207-223, DOI: 10.26107/RBZ-2023-0016
03E6654F682FFF8BFC1A91BBEDB9FDFA.text	03E6654F682FFF8BFC1A91BBEDB9FDFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptarma schubarti Shahdadi & Mendoza 2023	<div><p>Leptarma schubarti, new species</p> <p>(Figs. 1–4)</p> <p>Material examined. Holotype: male (11.0 × 9.5 mm) (ZRC 2019.1347), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=167.1&amp;materialsCitation.latitude=-15.583333" title="Search Plazi for locations around (long 167.1/lat -15.583333)">Rata Stream</a>, flow to the Belmoul Lagoon, stn VM07, Espiritu Santo Island, Vanuatu, 15 ° 35′S, 167 ° 06′E, coll. 13 October 2006, SANTO 2006 Expedition. Paratypes: 3 males (8.6 × 7.3 mm, 11.4 × 10.0 mm, 11.7 × 10.1 mm), 1 female (10.8 × 8.6 mm), 1 ovig. female (10.4 × 8.6 mm) (ZRC 2017.0191), same data as the holotype. Other material: 1 male (9.3 × mm), 1 ovig. female (10.7 × 9.2 mm) (ZRC 2019.1343), stn. PAL 19-07, stream beside yam plantation, entry from bridge along main road, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.57526&amp;materialsCitation.latitude=7.60875" title="Search Plazi for locations around (long 134.57526/lat 7.60875)">Ngetbong</a>, Babeldaob Island, Palau, 7 ° 36′31.5″N, 134 ° 34′30.9″E, coll. 9 January 2019, B. Y. Lee &amp; I. Iesa; 1 female (10.7 × 9.0 mm) (ZRC 2019.1344), stn. PAL 19-11, mangrove on islet along highway on <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.65405&amp;materialsCitation.latitude=7.531611" title="Search Plazi for locations around (long 134.65405/lat 7.531611)">eastern Babeldaob Island</a>, Palau, 7 ° 31′53.8″N, 134 ° 39′14.6″E, coll. 10 January 2019, B. Y. Lee &amp; I. Iesa.</p> <p>Comparative material. Leptarma aurifrons (Li, Shih &amp; Ng, 2019): paratypes, 1 male (10.1 × 8.2 mm) (ZRC 2019.0819); 9 males (9.8 × 8.2 mm – 11.1 × 9.5 mm), 4 females (10.7 × 8.4 mm – 11.5 × 9.1 mm) (ZRC 2019.0822); 1 male (12.4 × 11.1 mm) (ZRC 2019.0823), 1 male (14.2 × 10.9 mm) (ZRC 2019.1075), Gankou River estuary, Hengchun, Pingtung, Taiwan. Leptarma gecko (Li, Rahayu &amp; Ng, 2018): holotype male (16.5 × 14.3 mm) (RUMF-ZC-4710), Awase, Okinawa Island, Japan; paratypes 2 males (17.4 × 14.7 mm, 15.5 × 13.0 mm), 1 female (16.5 × 14.3 mm) (RUMF-ZC-2930), Yaeyama Islands, Iriomote Island, Japan; 1 male (11.1 × 9.6 mm) (ZRC 2019.1455), stn. VM-04, intertidal, between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=167.03&amp;materialsCitation.latitude=-15.581667" title="Search Plazi for locations around (long 167.03/lat -15.581667)">Rose Point</a> &amp; Nasouli River, Espiritu Santo Island, Vanuatu, 15°34.9′S, 167°01.8′E, coll. SANTO 2006 Expedition, 11 September 2006; 1 male (17.6 × 15.9 mm) (ZRC 2019.1448), stn. LBY 2019-16, sandy-muddy mangrove, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-179.87128&amp;materialsCitation.latitude=-16.824612" title="Search Plazi for locations around (long -179.87128/lat -16.824612)">Taveuni Island</a>, Fiji, 16°49′28.6″S 179°52′16.6″W, coll. 25 July 2019, B. Y. Lee et al. Leptarma lenzii (De Man, 1895): lectotype male (13.3 × 11.3 mm) (NNM-ZMA Crus. D. 102653), Aceh, Sumatra, Indonesia. Leptarma obliquifrons (Rathbun, 1924): holotype male (12.7 × 11.1 mm) (USNM 45913), Pago Pago, American Samoa. Leptarma reunionense Shahdadi &amp; Schubart, 2020: holotype male (12.2 × 10.1 mm) (MNHN- IU-2012-660), Mozambique Channel, Europa Island.</p> <p>Diagnosis. Carapace slightly broader than long, front deflexed, slightly sinuous in dorsal view, median postfrontal lobes almost as wide as lateral ones, lateral carapace margin slightly concave; eye with peduncle short, stout, wider than long, cornea wider than peduncle; male chela large (homochelous), robust, upper surface of palm with two oblique pectinated crests, dorsal surface of dactylus with 15 low, blunt, transversely oval tubercles, distinct to tip, proximal tubercles positioned at almost median part of upper face of palm; ambulatory legs flattened dorsoventrally and relatively long, third pair (P4) longest, total length (ischium–dactylus) ca. 1.9 times carapace width; male pleon triangular, somite 2 medially longer than lateral edges; G1 robust, long, straight, apical corneous process long, tip truncated, bent at an angle of about 50° to vertical axis, aperture terminal.</p> <p>Description. (Morphometric measurements are for the holotype) Carapace subrectangular, slightly broader than long (cw/cl = 1.16), greatest width between exorbital angles; carapace surface smooth; front ca. 0.61 times carapace width, markedly deflexed, slightly sinuous in dorsal view; post-frontal lobes prominent, median lobes almost as wide as lateral ones, separated by shallow furrow; dorsal carapace regions well indicated, gastric region demarcated, cardiac region separated from intestinal region, lateral branchial ridges prominent; anterolateral margin with sharp exorbital angle, lateral margin slightly concave, lined with row of short setae. Eye with peduncle short, stout, wider than long, cornea wider than peduncle (Figs. 1, 2A, 3A, 4A).</p> <p>Male chela large (palm length/cw = 0.68), robust (palm width/ length = 0.59) (Figs. 1, 2D, 3A, E); merus with granulate dorsal border, no distinct subdistal spine, ventral border granulate, anterior border granulate, with distinct subdistal spine; upper surface of palm with 2 oblique pectinated crests, distal one composed of 18 well developed chitinous teeth, proximal one with 9–12 teeth, both crests terminate at inner end in 2 granules and granular rows on outer end (Fig. 2E, 3F); outer surface of palm with fine granules except for smooth, punctate fixed finger (Figs. 2D, 3E); inner surface of palm finely granular except area facing carpus and distal half of finger; no median vertical ridge (Figs. 2E, 3F); both sides of palm with scattered tufts of setae, cutting edge ca. 0.40 times palm length; dactylus ca. 0.60 time palm length, slightly curved downwards and inwards, dorsal surface with 15 (n = 5) low but distinct, transversely oval tubercles, distinct to tip, all tubercles positioned almost at median part of upper face of dactylus upper-inner side of dactylus scattered with small pointed granules (Figs. 2E, 3F); scattered tufts of setae on inner side, cutting edge of both fingers with a series of different sized teeth (Figs. 2D, 3E).</p> <p>Ambulatory legs flattened dorso-ventrally, relatively long, third pair (P4) longest (Figs. 1A, B, 3A, 4A), total length (ischium–dactylus) ca. 1.9 times carapace width, tufts of long setae scattered along the legs, more on propodi and dactyli, merus of P4 ca. 2.41 times as long as wide, anterior margin crenulated, propodus ca. 3.27 times as long as wide, dactylus ca. 0.82 times length of propodus.</p> <p>Male pleon triangular (Figs. 1B, 2B), with rounded telson, almost as wide as long, almost as long as somite 6; somite 6 longer than others; somites 4 and 5 trapezoidal, somite 3 widest, laterally convex, somite 2 medially longer than lateral edges (Fig. 2C).</p> <p>G1 relatively long, stout, straight; apical corneous process long, narrow, tip truncated, bent at an angle of about 50° to vertical axis, aperture terminal (Fig. 3B, C, D).</p> <p>Female (Fig. 4) with proportionately smaller chelipeds; pectinate crests reduced to rows of granules (Fig. 4C); except for few proximal ones, dactylar tubercles indistinct (Fig. 4B). Pleon broad, rounded, or even laterally slightly ovate, broadest at somite 4, fringed with long setae (Fig. 4D). Vulva in depression on anterior edge of sternite 6, touching posterior margin of sternite 5; operculum broadly oval, perpendicular to sternite lines, positioned at inner side of gonopore, with distinct posterior sternal cover (Fig. 4E).</p> <p>Habitat and distribution. Most type material was collected from streams near the coast and also from mangroves, all on muddy substrates. The new species is thus far only known from Vanuatu and Palau.</p> <p>Etymology. The new species is named in honour of Prof. Dr. Christoph D. Schubart (Universität Regensburg, Germany) for his significant contributions to brachyuran systematics, especially the family Sesarmidae.</p> <p>Remarks. Besides the two new species described in the present study, two other species of Leptarma (i.e., L. gecko and L. obliquifrons) have previously been recorded from oceanic islands in the Western Pacific (Rathbun, 1924; Li et al., 2018). Leptarma schubarti, new species, differs most significantly from L. gecko in having proportionally shorter ambulatory legs (P4/cw = 1.9 in L. schubarti vs. P4/ cw = 2.2 in L. gecko); and the apical corneous process of the G1 is uniformly narrow throughout its length and has a truncate tip (Fig. 3B, C, D) (vs. widening distally with the tip rounded in L. gecko, Li et al., 2018: fig. 5D‒G). Li et al. (2018) mentioned that in L. gecko the number of dactylar tubercles on the male chela ranges from 10 to 15 (average 11), whereas L. schubarti has 15 dactylar tubercles consistently (n = 5) in the male chela. The shape of the proximal tubercles differs between these two species: they are low and blunt in L. schubarti (Figs. 2D, 3E), but steep and sharp in L. gecko (Li et al., 2018, fig. 6D‒F). Leptarma gecko is a widely distributed species in the West Pacific, from Japan to New Caledonia and Fiji, and reportedly with intraspecific morphological variations (Li et al., 2018). In the present study we also examined male specimens from Vanuatu and Fiji (see Comparative material), and both sufficiently resemble the types from Japan for them to be considered here as conspecific. However, to properly account for its wide distribution and morphological variations, a separate phylogeographic study using an appropriate genetic marker would clarify the true identities of material from its entire distribution range.</p> <p>Leptarma schubarti differs from L. obliquifrons in possessing 15 (n = 5) small and broadly oval dactylar tubercles in the male chela (Figs. 2D, E, 3E, F) (vs. only 5–6 (n = 5) large and round tubercles in male L. obliquifrons, Li et al., 2019: fig. 5D, E), and in the G1 apical corneous process being proportionately longer (Fig. 3B) than in L. obliquifrons (Li et al., 2019: fig. 6A‒D).</p> <p>The new species differs from L. lenzii s. str. in having a truncate G1 apical corneous process (Fig. 3B, C, D) (vs. tip widened and rounded in L. lenzii, Shahdadi et al., 2019: fig. 1F‒I), and the operculum on the vulva is oval (Fig. 4E) (vs. triangular in L. lenzii, Shahdadi et al., 2019: fig. 2).</p> <p>Leptarma schubarti resembles the three Southeast Asian congeners, L. gracilipes (Li, Rahayu &amp; Ng, 2018), L. macaco (Li, Rahayu &amp; Ng, 2018), and L. tarantula (Li, Rahayu &amp; Ng, 2018), in having a long and narrow apical corneous process of the G1 (Li et al., 2018). The tip of the process, however, is rounded in these three species, while it is truncated in L. schubarti new species. The new species differs also from these species in the number of dactylar tubercles in the male chela (15 in L. schubarti (n = 5) vs. 7 or 8 in L. gracilipes (n = 2), 6‒8 (average 7) in L. macaco and 10‒12 in L. tarantula (n = 5) (Li et al., 2018)).</p> <p>According to the molecular results (see below; Fig. 12), L. schubarti is genetically closest to L. aurifrons (Li, Shih &amp; Ng, 2019). These two species, however, are quite different from each other in key morphological characters. Most significantly, the G1 has a short and wide apical process in L. aurifrons (cf. Li et al., 2019: figs. 3E, F, 4E), while it is nearly three times as long and narrower in L. schubarti (Fig. 3B, C). Leptarma aurifrons also has proportionally shorter ambulatory legs (P4/cw = 1.5 in L. aurifrons; Li et al., 2019: figs. 2A, 3A) compared to the new species (vs. P4/cw = 1.9 in L. schubarti; Figs. 1A, 3A, 4A), the differences in length observed mainly in the ambulatory propodi. Furthermore, there are 12 subcircular dactylar tubercles in the male chela of L. aurifrons (Li et al., 2019: fig. 4C), whereas there are 15 transversely oval dactylar tubercles in the male chela of L. schubarti (Figs. 2D, E, 3E, F).</p> </div>	https://treatment.plazi.org/id/03E6654F682FFF8BFC1A91BBEDB9FDFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Shahdadi, Adnan;Mendoza, Jose Christopher E.	Shahdadi, Adnan, Mendoza, Jose Christopher E. (2023): Two new species of the mangrove crab genus Leptarma (Crustacea: Brachyura: Sesarmidae) from the Western Pacific islands. Raffles Bulletin of Zoology 71: 207-223, DOI: 10.26107/RBZ-2023-0016
03E6654F6828FF87FEBC917BE88EF7B2.text	03E6654F6828FF87FEBC917BE88EF7B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptarma bertrandi Shahdadi & Mendoza 2023	<div><p>Leptarma bertrandi, new species</p> <p>(Figs. 5–8, 9A‒D, 10A‒D, 12A‒C)</p> <p>Parasesarma lenzii — Li et al., 2019: 19, fig. 12A, B. Not Leptarma lenzii (De Man, 1895).</p> <p>Material examined. Holotype: 1 male (21.4 × 19.3 mm) (MNHN-IU-2022-384), mangrove, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=166.56667&amp;materialsCitation.latitude=-20.408333" title="Search Plazi for locations around (long 166.56667/lat -20.408333)">Ouvéa Island</a>, Loyalty Islands, New Caledonia, 20 ° 24.5′S, 166 ° 34′E, coll. 10 December 2009, KIWA 2 Expedition. Paratypes: 2 males (18.0 × 15.2 mm, 16.5 × 14.2 mm), 1 female (18.6 × 16.0 mm) (ZRC 2019.1330), same data as the holotype. Other material: 1 male (14.5 × 12.5 mm) (ZRC 2019.1331), intertidal, sand and muddy sand, station VM09, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=167.10167&amp;materialsCitation.latitude=-15.596666" title="Search Plazi for locations around (long 167.10167/lat -15.596666)">Belmoul Lagoon</a>, Espiritu Santo Island, Vanuatu, 15 ° 35.8′S, 167 ° 06.1′E, coll. 13 October 2006, SANTO 2006 Expedition; 6 males (8.8 × 7.6 mm – 17.8 × 15.7 mm), 2 females (14.8 × 13.0 mm, 18.3 × 16.2 mm) (ZRC 2019.1079), Tanzih fish port, Pingtung, Taiwan, coll. 29 September 2018, J.-J. Li.</p> <p>Comparative material. Leptarma lenzii (De Man, 1895): lectotype, male (13.3 × 11.3 mm) (NNM-ZMA Crus. D. 102653), Aceh, Sumatra, Indonesia; 1 male (16.4 × 14.2 mm) (ZRC 2018.1374), Cocos (Keeling) Islands; 1 male (11.8 × 9.9 mm), 2 females (15.1 × 13.4 mm, 8.3 × 7.3 mm) (ZRC 2018.1373), Cocos (Keeling) Islands.</p> <p>Diagnosis. Carapace slightly broader than long, front deflexed, slightly sinuous in dorsal view, median postfrontal lobes wider than lateral ones, carapace lateral margin almost straight; eye with peduncle short, stout, wider than long, cornea wider than peduncle; male chela large, robust, upper surface of palm with two oblique pectinated crests, dorsal surface of dactylus with 8‒10 transversely oval tubercles, distinct to tip, proximal tubercles positioned at almost median part of upper face of palm; ambulatory legs flattened dorsoventrally and relatively long, third pair (P4) longest, total length (ischium–dactylus) ca. 1.9 times carapace width; male pleon triangular, somite 2 medially longer than lateral edges; G1 robust, long, straight, apical corneous process long, stout, tip rounded, bent at an angle of about 50° to vertical axis, aperture subterminal.</p> <p>Description. (Morphometric measurements are for the holotype) Carapace subrectangular, slightly broader than long (cw/cl = 1.10); carapace surface smooth; front ca. 0.60 times carapace width, markedly deflexed, slightly sinuous in dorsal view; post-frontal lobes prominent, median lobes wider than lateral ones, separated by shallow furrow; dorsal carapace regions well indicated, gastric region demarcated, cardiac region separated from intestinal region, lateral branchial ridges prominent, in the largest male (holotype) first ridge results in an epibranchial (anterolateral) prominence (posterior to exorbital angle); anterolateral margin with sharp exorbital angle, lateral margin almost straight, lined with row of short setae. Eye with peduncle short, stout, wider than long, cornea wider than peduncle (Figs. 5A, 6A, B, 7A, 9A‒D).</p> <p>Male chela large (palm length/cw = 0.76), robust (palm width/ length = 0.56) (Figs. 5, 6C, 10A–D); merus with granulate dorsal border, no distinct subdistal spine, ventral border granulate, anterior border granulate, with distinct subdistal spine; upper surface of palm with 2 oblique pectinated crests, distal one composed of 10–12 (10 in holotype) well-developed chitinous teeth, proximal one with 6–10 (10 in holotype) teeth, both crests terminate at inner end in 2 granules and granular row on outer side; outer surface of palm with coarse granules except for smooth, punctate fixed finger; inner surface of palm finely granular except area facing carpus and distal half of finger; no median vertical ridge; both sides of palm with scattered tufts of setae, cutting edge ca. 0.41 times palm length, with a series of different sized teeth; dactylus ca. 0.57 times palm length, slightly curved downwards and inwards, dorsal surface with 8‒10 (n = 5) (nine in holotype) distinct, transversely oval tubercles, distinct to tip, all tubercles including the proximal ones positioned almost at median part of upper face of palm; small pointed granules scattered on inner and outer side; cutting edge with two large and a series of smaller teeth (Figs. 5A, 6C‒E, 10A‒D).</p> <p>Ambulatory legs flattened dorso-ventrally, relatively long, third pair (P4) longest (Figs. 5, 7A, B, 9A‒D), total length (ischium–dactylus) 1.9 times carapace width, tufts of long setae scattered along the legs, more on propodi and dactyli, merus of P4 ca. 2.40 times as long as wide, anterior margin crenulated, propodus ca. 3.6 times as long as wide, dactylus ca. 0.66 times length of propodus.</p> <p>Male pleon triangular, with rounded telson (Figs. 5B, 6F), almost as wide as long, almost as long as somite 6; somite 6 longer than others; somites 4 and 5 trapezoidal, somite 3 widest, laterally convex, somite 2 medially longer than lateral edges (Fig. 6G).</p> <p>G1 relatively long, stout, straight; apical corneous process long, tip rounded, distally flared, bent at angle of about 50° to vertical axis, aperture sub-terminal (Figs. 6H‒J, 11A‒C).</p> <p>Female (Fig. 7) with proportionately smaller chelipeds; pectinate crests reduced to rows of granules (Fig. 7C, D); except for few proximal ones, dactylar tubercles indistinct (Fig. 7C, D). Pleon broad, rounded, or even laterally slightly ovate, broadest at somite 4, fringed with long setae (Fig. 7B). Vulva in depression on anterior edge of sternite 6, touching posterior margin of sternite 5; operculum triangular, perpendicular, positioned anterior to gonopore (Fig. 7E).</p> <p>Habitat and distribution. Specimens of this species were collected from mangroves and estuaries, in the intertidal zone, on sand or muddy sand substrates. The new species is known thus far from the Loyalty Islands (New Caledonia), Vanuatu (present study), and Taiwan (Li et al., 2019; present study).</p> <p>Etymology. This new species is named in honour of Dr. Bertrand Richer de Forges for his tireless efforts in collecting and documenting the brachyuran/crustacean fauna of the Indo-West Pacific region, particularly through expeditions held under various French-led marine biodiversity programs.</p> <p>Remarks. Leptarma bertrandi, new species, most closely resembles L. lenzii s. str. in general appearance. According to our molecular results, these two species also have a close phylogenetic association (see below). These two species, however, differ in several morphological characters. The dorsal carapace regions are more demarcated in L. bertrandi (Fig. 9A–D) than in L. lenzii (Fig. 9E, F). The lateral frontal lobes are distinctly narrower than the median ones in L. bertrandi (Figs. 6A, B, 9A–D), while they are subequal in L. lenzii (Fig. 9E, F). These two species also differ in the pattern of the dactylar tuberculation in the male chela, the new species only having 8‒10 (n =5) distinct tubercles (Fig. 10A–D), while L. lenzii has 11–13 (n = 3) small tubercles (Fig. 10E, F). The new species (Fig. 9A–D) has proportionally longer ambulatory legs than L. lenzii (Fig. 9E, F), especially the propodi: i.e., length of P4 (ischium–dactylus) = 1.9 times carapace width, merus ca. 2.4 times as long as wide, propodus ca. 3.6 times as long as wide, dactylus ca. 0.66 times length of propodus in L. bertrandi; while in L. lenzii the corresponding values are P4 = 1.8 times cw, merus ca. 2.2 times as long as wide, propodus ca. 2.9 times as long as wide, dactylus ca. 0.60 times length of propodus. The two species are similar in the general form of the G1, but the apical corneous process of L. bertrandi (Fig. 11A–C) is distinctly shorter and stouter than that of L. lenzii (Fig. 11D, E). It bears noting here that the material from Taiwan previously reported as Parasesarma lenzii (viz. Li et al., 2019: 19), is here treated as conspecific with L. bertrandi, primarily due to it possessing the diagnostic morphology of the ambulatory legs, male chela, and G1.</p> <p>Furthermore, the live colouration of the two species may be useful for differential diagnosis, based on the information gleaned from available colour photographs of these crabs. The L. bertrandi from Taiwan has orange chelae with deeper reddish-orange fingers and a predominantly whitish carapace with black marbling (Li et al., 2019, fig. 12A, B); the colour patterns on the specimen from Vanuatu (Fig. 8) are similar, but the specimen was photographed only after it had been in the freezer overnight and the colour may have already faded. On the other hand, L. lenzii from the Cocos (Keeling) Islands has greyish chelae with orange-tipped fingers (Shahdadi et al., 2019, fig. 3D) and a predominantly black carapace with small yellow spots and a distinct yellow, transverse, submarginal band on the frontal region (Shahdadi et al., 2019, fig. 3C, D). This will need further investigation as more fresh material is observed.</p> </div>	https://treatment.plazi.org/id/03E6654F6828FF87FEBC917BE88EF7B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Shahdadi, Adnan;Mendoza, Jose Christopher E.	Shahdadi, Adnan, Mendoza, Jose Christopher E. (2023): Two new species of the mangrove crab genus Leptarma (Crustacea: Brachyura: Sesarmidae) from the Western Pacific islands. Raffles Bulletin of Zoology 71: 207-223, DOI: 10.26107/RBZ-2023-0016
