taxonID	type	description	language	source
03E487DEFFF9F843FF65FBBB05A9F81C.taxon	description	(Fig. 1)	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFF9F843FF65FBBB05A9F81C.taxon	materials_examined	Type material examined. Holotype of Geodromicus palpalis ♂: ‘ Dampelek | b. Jumla’ <printed>, ‘ Gebiet von Jumla | Westnepal, lg. H. Franz’ <printed>, ‘ HOLOTYPE’ <red, printed>, ‘ Geodromicus | palpalis | det. H. Coiffait 1976 [printed] ’ <handwritten> (MHNH). Paratypes: 1 ♀: same data as the holotype except of red printed label: ‘ PARATYPE’ (MHNH); 1 ♂ (dissected; Fig. 1): ‘ Dampelek | b. Jumla’ <printed>, ‘ Gebiet von Jumla | Westnepal, lg. H. Franz’ <printed>, ‘ TYPE’ <red, printed>, ‘ Geodromicus | palpalis | H. COIFFAIT det. 1976 [printed] ’ <handwritten>, ‘ Hygrodromicus | coriaceus | (Cameron, 1924) | Shavrin A. V. det. 2023 ’ (NMW); 2 ♀♀: ‘ Umg. Maharigaon | 3000 – 3500 m’ <printed>, ‘ Gebiet von Jumla | Westnepal, lg. H. Franz’ <printed>, ‘ TYPE’ <red, printed>, ‘ PARATYPE’ <red, printed>, ‘ Hygrodromicus | coriaceus | (Cameron, 1924) | Shavrin A. V. det. 2023 ’ (NMW). Material examined. INDIA: UTTARAKHAND: 1 ♀: ‘ Chakrata Distr., Chuli Khud 8000`’, ‘ Dr. Cameron 14. V. [19] 22. ’ (MHNG); NEPAL: KARNALI: 1 ♂: N Khari Lagna, 29 ° 22,14´N 82 ° 09,17´E. 3280 m a. s. l. 21.06.1999. M. Hartmann leg. (cSh); 2 ♀♀: same District, Talphi, NE, Maharigaun. 3300 m a. s. l. 20.06.1997. A. Weigel leg. (NME).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFF9F843FF65FBBB05A9F81C.taxon	discussion	Remarks. Geodromicus palpalis was originally described from “ Dampelek près Jumla, Népal occidental … ”. Based on the study of the type material from MHNH and NMW, I found that based on external and internal morphological features it is conspecific with H. coriaceus. Thus, G. palpalis was synonymized with the latter species. Hygrodromicus coriaceus is widely distributed in the Himalayan Region from Kashmir to Nepal (Shavrin 2021).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFAF841FF65F98C05A9FF22.taxon	materials_examined	Material examined. NEPAL: KARNALI: 1 ♂: Humla District, Maharigaon N, 29 ° 20,24´N 82 ° 23,21´E. 3250 m a. s. l. 08 – 09.07.1999. M. Hartmann leg. (NME); 1 ♀: same District, 18 km NW Simikot Chumsa Khola (Bridge), 30 ° 02´25 ´´ N 81 ° 39´06 ´´ E. 2950 – 3000 m a. s. l., river valley. 20 – 22.06.2001. E. Grill leg. (NME); 1 ♀: same District, 16 km N Simikot, 3 km NW Sankha La, 29 ° 57´19 ´´ N 81 ° 39´30 ´´ E. 4250 – 4600 m a. s. l., border of snow fields. M. Hartmann leg. (cSh); 1 ♀: Simikot, ~ 12 km SE, 29 ° 54´23 ´´ N 81 ° 55´7 ´´ E. Pasture and edge of disturbed mixed forest, nigth collecting. 17 – 18.06.2022. D. Telnov leg. (BMNH); 1 ♀: same District, N Khari Lagna, 29 ° 22,14´N 82 ° 09,17´E. 21.06.1999. M. Hartmann leg. (NME); 1 ♀: Dolpo District, Pahada W. Tal, 29 ° 04,33´N 82 ° 42,4´E. 3000 – 3500 m a. s. l. M. Hartmann leg. (NME).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFAF841FF65F98C05A9FF22.taxon	discussion	Remarks. Hygrodromicus himalayicus was recently described from India (Uttarakhand) and Nepal (Shavrin 2021).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFBF847FF65FEFF0762FD06.taxon	description	(Figs 2, 6 – 7)	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFBF847FF65FEFF0762FD06.taxon	materials_examined	Type material examined. Cotype ♀: ‘ COTYPUS | Geodromicus | klapperichi | O. Scheerpeltz’ <pink, printed>, ‘ ♀ ’ <printed>, ‘ J. Klapperich | Achmende | Dewane’ <printed>, ‘ 2700 m, Bashgul tal, | Nuristan, 23.7.52. - | Afghanistan’ <printed>, [yellow label without letters], ‘ Hygrodromicus | klapperichi | H. Coiffait det. 1982 [printed] ’ <handwritten> (MHNH). Material examined. AFGHANISTAN: 1 ♂: ‘ Vt. N. Salang 1450 m Nhenjan, bord torrent 11.8.1975. Leg. Ledoux’ (cZ); 1 ♂: ‘ Afghanistan Umgeb. Kabul leg. J. Klapperich’ (cB).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFBF847FF65FEFF0762FD06.taxon	description	Redescription. Measurements (n = 3): HW: 0.75 – 0.85; HL: 0.46 – 0.52; OL: 0.22 – 0.25; LT: 0.07; AL (averaged): 2.36; PL: 0.67 – 0.69; PWmax: 0.98 – 1.02; PWmin: 0.78 – 0.80; ESL: 1.47 – 1.70; EW: 1.60 – 1.70; MTbL (averaged): 1.02; MTrL (averaged): 0.44 (MTrL 1 – 4: 0.20; MTrL 5: 0.22); AW: 1.58 – 1.65; AedL: 0.92 – 0.97; BL: 4.15 – 4.35. Habitus as in Fig. 2. Body reddish-brown, with paler intersegmental membranes between abdominal tergites III – VII; mouthparts, antennae and legs yellow-brown to brown; apical segment of maxillary palpi and tarsi yellowish. Body glossy; head with fine microreticulation, slightly coarser between bases of antennae, distinctly finer in middle portion; neck with large mesh-like sculpture; pronotum with very dense and coarse isodiametric microsculpture, finer in mediobasal portion; scutellum with fine transverse meshes; abdomen with dense transverse microreticulation. Head 1.6 times as broad as long, with distinctly elevated portion between bases of antennae and slightly elevated infraorbital portions; supra-antennal proturberances narrow, strongly elevated, diagonally stretching from base of antennae to about level of apical margins of eyes; medioapical depression deep and wide, narrowing posteriad toward relatively shallow or deep subrectangular interocellar depression; anteocellar foveae distinct, very narrow, diagonally stretching anteriad toward level of middle or anterior third of eyes; temples short, convex, more than three times as long as longitudinal length of eyes. Eyes moderately large, very convex. Ocelli large, located below level of posterior margins of eyes, distance between ocelli about as long as distance between ocellus and posterior margin of eyes, or distinctly longer. Punctation irregular, moderately fine, sparser and finer in middle and larger and denser on infraorbital portions. Apical segment of maxillary palpi about 1.7 times as long as preapical segment. Antennae long, exceeding middle or apical third length of elytra when reclined; basal antennomere moderately wide, about three times as long as broad, antennomere 2 slightly narrower and distinctly shorter than basal antennomere, 3 – 4 slightly longer than 2, 5 – 6 slightly longer than 4, 7 – 9 slightly longer than 6, 10 slightly shorter than 9, apical antennomere about 1.4 times as long as 10, from apical third gradually narrowed toward subacute apex. Pronotum moderately small, convex, 1.4 times as broad as long, 1.2 – 1.3 times as broad as head, widest in anterior portion, gradually narrowed posteriad toward obtuse posterior angles; anterior margin rounded, about as long as straight posterior margin. Punctation dense, about as that in middle portion of head or distinctly larger, but finer and sparser in mediobasal portion, interspaces between punctures in middle about as broad of one-two diameters of nearest punctures; narrow longitudinal band of pronotal disc without punctures. Scutellum without or with several very fine punctures. Elytra somewhat flattened, about as broad as long or slightly narrower, distinctly more than twice as long as pronotum, slightly broadened posteriad, with widely rounded apical margins. Punctation dense, about as that on pronotum or slightly deeper, finer around scutellum and along suture. Wings fully developed. Metatarsus more than twice shorter than metatibiae; apical antennomere slightly longer than preceding four metatarsomeres. Abdomen distinctly narrower than elytra, with two large transverse tomentose spots in the middle of abdominal tergite IV; apical margin of abdominal tergite VII with narrow palisade fringe. Punctation regular, very dense and fine. Male. Protarsomeres 1 – 4 wide. Apical margin of abdominal tergite VIII rounded. Apical margin of abdominal sternite VIII concave. Aedeagus narrow, long, with small basal portion slightly narrowed toward apical third of median lobe, with slightly broadened apical portion, gradually narrowed toward rounded apex; parameres narrow, reaching apex of median lobe, with four moderately short apical setae; internal sac without sclerotized structures, with very long flagellum, spirally folded in basal portion (Fig. 6). Lateral aspect of the aedeagus as in Fig. 7. Female. Protarsomeres 1 – 4 narrow. Apical margins of abdominal tergite VIII and sternite VIII rounded. Comparative notes. Based on the general shape of the body and the aedeagus, H. klapperichi is similar to H. reitteri (Scheerpeltz, 1933), from which it can be distinguished by the slightly shorter body, the slightly broader apical part of the aedeagus, and the absence of sclerotized elongate structures in the middle portion of the internal sac.	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFBF847FF65FEFF0762FD06.taxon	distribution	Distribution. Hygrodromicus klapperichi is known from three localities in north-eastern part of Afghanistan. Bionomics. The type specimens were collected at an elevation of 2700 m a. s. l. The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFBF847FF65FEFF0762FD06.taxon	discussion	Remarks. Hygrodromicus klapperichi was originally described based on four specimens from “ Afghanistan, Nouristan, Bashgultal … ”. Before the original description, these specimens were labelled by O. Scheerpeltz as “ Geodromicus klapperichi ”. During my study of Omaliinae in MHNH in 2011, I found one female which could belong to the type serie (see above).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFDF844FF65FC9B0685FC1A.taxon	description	(Figs 3, 8 – 9)	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFDF844FF65FC9B0685FC1A.taxon	materials_examined	Type material examined. Holotype ♂ (Fig. 3; dissected): ‘ NEPAL Rasuwa Dis. | Langtang Kh [ola]. Vall. | Forest Camp 1900 m | 13. IV. [19] 85 A. Smetana’ <printed>, ‘ HOLOTYPE | Hygrodromicus | latilobatus sp. n. | Shavrin A. V. des. 2024 ’ <red, printed> (CNC). Paratypes: 3 ♂♂ (dissected; one specimen without left antennomeres 9 – 11 and right antennomeres 4 – 11; one specimen without right apical antennomere), 1 ♀: same data as the holotype, with additional red printed label: ‘ PARATYPE | Hygrodromicus | latilobatus sp. n. | Shavrin A. V. des. 2024 ’ (2 ♂♂: cSh; 1 ♂, 1 ♀: CNC).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFDF844FF65FC9B0685FC1A.taxon	description	Description. Measurements (n = 5): HW: 0.77 – 0.88; HL: 0.47 – 0.58; OL: 0.16 – 0.22; LT: 0.08 – 0.12; AL (holotype): 3.48; PL: 0.63 – 0.88; PWmax: 0.95 – 1.11; PWmin: 0.84 – 0.88; ESL: 1.40 – 1.51; EW: 1.65 – 1.78; MTbL (holotype): 1.22; MTrL (holotype): 0.50 (MTrL 1 – 4: 0.23; MTrL 5: 0.27); AW: 1.50 – 1.82; AedL: 0.83 – 0.94; BL: 4.20 – 4.70 (holotype: 4.45). Habitus as in Fig. 3. Body yellow-brown to reddish-brown; mouthparts, antennae and legs yellow-brown. Head with regular, dense and moderately fine isodiametric microreticulation; neck with large and coarse isodiametric sculpture; pronotum with dense isodiametric microsculpture similar to that in head, slightly transverse in lateroapical portions and finer in middle in two paratypes; scutellum with dense transverse microsculpture; abdomen with dense transverse microreticulation. Head 1.5 – 1.6 times as broad as long, with markedly elevated portions between bases of antennae and slightly convex infraorbital portions; supra-antennal proturberances moderately wide and strongly elevated; medioapical depression deep and wide, narrowing posteriad toward wide and deep subrectangular interocellar depression; anteocellar foveae narrow, long and deep, reaching level of anterior third of eyes; temples convex, about twice as long as longitudinal length of eyes. Eyes moderately small, convex. Ocelli small, but well visible, located slightly below level of posterior margins of eyes, distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Punctation dense, moderately large and deep, sparser in middle and distinctly denser on infraorbital portions. Apical segment of maxillary palpi short, distinctly twice shorter than preapical segment. Antenna very long, almost reaching apical margins of elytra when reclined; basal antrennomere wide, distinctly more than three times as long as broad, antennomere 2 narrower and shorter than basal antennomere, 3 distinctly longer than 2, 4 – 8 distinctly longer than 3, 9 – 10 slightly broader than 8, apical antennomere 1.3 – 1.4 times as long as 10. Pronotum 1.2 – 1.5 times as broad as long, 1.2 times as broad as head, widest in anterior portion, strongly narrowed posteriad toward obtuse or subacute posterior angles, with slightly and narrowly elevated longitudinal band in middle; posterior margin somewhat rounded, slightly narrower than straight posterior margin. Punctation dense, about as that in head, sparser in middle and / or mediobasal third; narrow longitudinal band without punctures. Scutellum with fine and very sparse punctation. Elytra flattened, slightly broader than long, about twice as long as pronotum or slightly shorter, significantly broadened posteriad, with widely rounded apical margins. Punctation dense, slightly larger, deeper and sparser than that on pronotum, finer around scutellum and along suture. Wings fully developed. Metatarsus more than twice shorter than metatibiae; apical antennomere slightly longer than preceding four metatarsomeres. Abdomen narrower than elytra, with two moderately small and transverse tomentose spots in the middle of abodminal tergite IV; apical margin of abdominal tergite VII with very narrow indistinct palisade fringe. Punctation dense and fine. Male. Protarsomeres 1 – 4 wide. Apical margin of abdominal tergite VIII straight or somewhat rounded. Aedeagus with very wide median lobe, strongly narrowed toward narrow apical part, and gradually narrowed toward rounded apex; parameres narrow, significantly longer than apex of median lobe, with four moderately short apical setae; internal sac without sclerotized structures, with very long flagellum, spirally folded in basal portion (Fig. 8). Lateral aspect of the aedeagus asi n Fig. 9. Female. Protarsomeres 1 – 4 narrow. Apical margins of abdominal tergite VIII and sternite VIII rounded. Comparative notes. Based on the general shape of the body and the aedeagus, H. latilobatus sp. n. is similar to H. amnicola (Champion, 1920), known from Pakistan and India (Shavrin 2021), from which it can be distinguished by the narrower and longer elytra, significantly narrower apical portion of the median lobe, and longer parameres.	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFDF844FF65FC9B0685FC1A.taxon	distribution	Distribution. Hygrodromicus latilobatus sp. n. is known only from the type locality in Rasuwa District, central Nepal. Bionomics. The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFDF844FF65FC9B0685FC1A.taxon	etymology	Etymology. The specific epithet is a combination of Latin adjective latus, - a, - um (broad) and lobatus, - a, - um (lobed). It refers to the distinctly broadened median lobe of the aedeagus.	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFEF844FF65FBE705A9FA0E.taxon	materials_examined	Material examined. CHINA: SICHUAN: 1 ♂: Gongga Shan, Hailuogou, above Camp 3, 29 ° 35´N 102 ° 00´E. 2800 – 3300 m a. s. l. 06 – 08.07.1998. J. Farkač leg. (NHMB); 6 ♂♂, 7 ♀♀: Jiuzhaigou. 3100 m a. s. l. 13 – 15.06.1992. M. Bok leg. (SMNS); YUNNAN: 1 ♀: Yulong Mts., 27 ° 01´N 100 ° 12´E. 3200 m a. s. l. 24 – 26.09.1993. Bolm leg. (NHMB); 1 ♂, 1 ♀: above Dali. 2000 – 2200 m a. s. l. 04 – 17.04.1999. W. Schawaller leg. (SMNS); 7 ♂♂, 5 ♀♀: same data and collector. 2500 – 2700 m a. s. l. 08 – 18.04.1999 (SMNS).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFEF844FF65FBE705A9FA0E.taxon	discussion	Remarks. Hygrodromicus maderi is widely distributed species in central and south-eastern China (Shavrin 2022).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFEF844FF65F993006CF811.taxon	materials_examined	Material examined. NEPAL: KARNALI: 1 ♂, 1 ♀: Humla District, 20 km W Simikot, 2 km S Chala Kairang Khola, NN, 29 ° 59´27 ´´ N 81 ° 37´30 ´´ E. Riverbank. 26.06.2001. M. Hartmann (cSh, NME); 1 ♀: same District, Jumla, NN. 2300 m a. s. l. 29.04.1995. J. Weipert leg. (NME); 1 ♀: same District, Mugu Rara Lake, southern lake-side, 29 ° 30,59´N 82 ° 04,35´E. 18.06.2011. J. Küssner leg. (NME); GANDAKI: 1 ♂: Kali Gandaki Vall., Kagbeni. 2800 m a. s. l. 25.05.2002. J. Schmidt leg. (NME); 1 ♂: Dhaulagiri SE slope, upp. Ruhagat Khola, between Chima Khola and Dwari, NN. 1750 m a. s. l. 10.05.2002. J. Schmidt leg. (NME)	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFEF844FF65F993006CF811.taxon	discussion	Remarks. Hygrodromicus recticollis is known from India (Uttarakhand) and Nepal (Shavrin 2021).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF845FF65FF4E06E5FCBA.taxon	description	(Fig. 4)	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF845FF65FF4E06E5FCBA.taxon	materials_examined	Material examined. KAZAKHSTAN: JAMBYL: 5 ♂, 4 ♀♀: Chylik R., Sarybastau Pass. 14.06.1988. V. Kashcheev leg. (cSh, ZIN); 5 ♂♂, 3 ♀: same data, 15.08.1988 (cSh, ZIN); 4 ♂, 2 ♀: Ile Alatau. 20.06.1988. V. Kashcheev leg. (cSh, ZIN); 3 ♂♂, 2 ♀♀: Kyrgyz Mnt., Merke R., near Merke. 12.06.1988. V. Kashcheev leg. (cSh); NARYN: 3 ♂♂, 1 ♀: Bayankoy R., near Narynkoi. 15.09.1988. V. Kashcheev leg. (ZIN); KYRGYZSTAN: 1 ♂: Issyk-Kul, Terskey Alatau, Uruk Valley, 42 ° 36´17 ´´ N 79 ° 18´57 ´´ E. 30.07 – 01.08.2005. 2600 m a. s. l. L. Schmidt leg. (cSh); UZBEKISTAN: 1 ♀: West Gissar, Yakkabadskiy District, Kalta-Kol. 1400 m a. s. l. 07.09.1990. Gromenko leg. (cSh); 1 ♀: Chatkal Mts., Okhangaron District, 6 km NW Irtash, Iertash [Yertoshsoy] River (right confluent of Okhangaron River), 41 ° 10´46.7 ´´ N 70 ° 17´28.7 ´´ E. 1900 m a. s. l. 03.08.2021. A. V. Shavrin leg. (cSh).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF845FF65FF4E06E5FCBA.taxon	discussion	Remarks. Geodromicus lestevoides Reitter, 1900 was originally described from “ Bokhara, Turkestan ”. Scheerpeltz (1933) replaced this name with G. reitteri Sheerpeltz, 1933 because lestevoides was previously occupied by other taxon (see details in Scheerpeltz 1933 and Zerche 1992). Luze (1903) and Zerche (1992) redescribed the species, and Zerche also designated the lectotype and provided new faunistic records. Hygrodromicus reitteri (Fig. 4) is here recorded from Kazakhstan and Kyrgyzstan for the first time.	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF845FF65FC4601D8FAEA.taxon	description	(Figs. 5, 15)	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF845FF65FC4601D8FAEA.taxon	materials_examined	Material examined. TAJIKISTAN: 2 ♀♀: Pamir-Alai, Hissar Mts., Warsob Valley. 2000 m a. s. l. 29.06.1990. M. Schülke & D. Wrase leg. (cSh, cSch); 1 ♀: Varzob District, 48 km N Dushanbe, Khodja Obi Gharm. 1900 m a. s. l. 03 – 04.07.2012. U. Valainis leg. (cSh); 1 ♀: Sarymai River, 3 km lower Gaza vill. 1300 m a. s. l. 25.04.1992. V. Grebennikov leg. (cSh).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF845FF65FC4601D8FAEA.taxon	discussion	Remarks. Hygrodromicus wrasei was originally described from: “ Asia centr., Duschanbe, Tadschikistan, Warsob-Tal / Hissar-Gebirge ”. This species (Fig. 5) is known only from Tajikistan (Fig. 15).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF84BFF65FAB401FAFCAA.taxon	description	(Figs. 10 – 12, 15)	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF84BFF65FAB401FAFCAA.taxon	materials_examined	Type material examined. Holotype ♂ (Fig. 12; dissected): ‘ NE-Afghanistan, 2008 | Shiwa Lake, 2800 – 2900 m | 37 ° 14 ’ / 70 ° 58 ’ E | 19. – 21. VII., C. Reuter’ <printed>, ‘ HOLOTYPE | Paratrichodromeus | angustilobatus sp. n. | Shavrin A. V. des. 2024 ’ <red, printed> (cF, to be eventually deposited in NME).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF84BFF65FAB401FAFCAA.taxon	description	Redescription. Measurements: HW: 0.85; HL: 0.65; OL: 0.17; TL: 0.15; AL: 2.64; PL: 0.77; PWmax: 0.97; PWmin: 0.81; ESL: 1.32; EW: 1.52; MTbL: 1.10; MTrL: 0.61 (MTrL 1 – 4: 0.31; MTrL 5: 0.30); AW: 1.49; AedL: 1.22; BL: 5.75. Body narrow, elongate and moderately convex (Fig. 12). Body reddish-brown, with distinctly paler apical portion of elytra and abdominal tergites III – V; antennae brown, with paler basal parts of antennomeres 1 – 5; mouthparts and legs yellow-brown (tarsi slightly paler). Body glossy; head with fine microsculpture, fine and dense in middle of vertex, very dense and irregular in middle, and dense and isodiametric on infraorbital portions; neck with dense and coarse isodiametric microsculpture; pronotum with fine and dense transverse microreticulation, invisible in middle; scutellum with very fine isodiametric meshes in middle, lateral portions of visible parts of scutellum without any sculpture; abdomen with dense transverse microreticulation. Head 1.3 times as broad as long, with distinctly elevated medioapical portions between bases of antennae and infraorbital portions; supra-antennal protuberances moderately wide, strongly elevated; medioapical depression wide and very deep, narrowing posteriad and connected with moderately deep and narrow subrectangular interocellar depression; anteocellar foveae narrow, subdiagonally stretching anteriad toward level of anterior third of eyes; temples convex, long, slightly shorter than longitudinal length of eyes. Eyes moderately small, convex. Ocelli reduced. Punctation irregular, finer and sparser in middle and larger and denser on infraorbital portions; basal portions of vertex without sculpture. Preapical segment of maxillary palpi about as long as preceding segment, from middle gradually narrowed toward acute apex. Antennae reaching about middle length of elytra when reclined; basal antennomere about three times as long as wide, antennomere 2 slightly narrower and distinctly shorter than basal antennomere, 3 slightly shorter and narrower than 2, 4 slightly shorter than 3, 5 – 6 slightly longer than 4, 7 slightly longer and indistinctly broader than 6, 8 – 10 slightly shorter and broader than 7, apical antennomere slightly shorter than previous two antennomeres, from apical third gradually narrowing toward subacute apex. Pronotum 1.2 times as broad as long, slightly broader than head, from widest anterior portion gradually narrowed posteriad toward obtuse posterior angles; laterobasal margins indistinctly concave; anterior margin rounded, about as long as slightly concave posterior margin; mediobasal portion with indistinct, small and oval impression. Punctation dense and fine, distinctly sparser in middle and meddiobasal portion. Scutellum without punctation. Elytra slightly broader than long, 1.7 times as long as pronotum, distinctly broadened posteriad, with very wide apical margins. Punctation about as that on pronotum, denser and deeper in basal and significantly finer and sparser in middle portion. Wings fully developed. Metatarsus slightly less than twice as long as metatibia; apical metatarsomere slightly shorter the preceding four metatarsomeres. Abdomen slightly narrower than elytra, with two wide and transverse tomentose spots in the middle of abdominal tergite IV and indistinct two very small and transverse spots in the middle of tergite V; apical margin of abdominal tergite VII without distinct palisade fringe. Punctation regular, fine and dense, distinctly finer on abdominal tergite IV. Male. Apical margin of abdominal tergite VIII rounded. Apical margin of abdominal sternite VIII sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe with moderately wide apical part gradually narrowed toward small rounded apex; parameres narrow, distinctly exceeding apex of median lobe, with small two apical and two preapical setae; internal sac with elongate sclerotized structures in middle and moderately long flagellum (Fig. 10). Lateral aspect of the aedeagus as in Fig. 11. Female unknown. Comparative notes. Paratrichodromeus angustilobatus sp. n. can be reliably distinguished from P. punctulatus (Luze, 1904) and P. sukranensis Zerche, 1992 only by the shape of the narrower apical portion of the aedeagus and different structure of the internal sac. For illustrations of the aedeagi of the compared species see Figs 9, 10 in Zerche (1992).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF84BFF65FAB401FAFCAA.taxon	distribution	Distribution. The species is known only from the type locality in north-eastern Afghaninstan (Fig. 15). Bionomics. The specimen was collected at elevation 2800 – 2900 m a. s. l. The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFFFF84BFF65FAB401FAFCAA.taxon	etymology	Etymology. The specific epithet is a combination of Latin adjective angustus, - a, - um (narrow) and lobatus, - a, - um (lobed). It refers to the moderately narrow apical part of the median lobe of the aedeagus.	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFF1F84BFF65F913001BF80C.taxon	description	(Figs. 13, 15)	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFF1F84BFF65F913001BF80C.taxon	materials_examined	Material examined. I studied eight specimens (cSch, cSh) from Tajikistan (“ … Hissar-Gebirge, Ansob-Pass N Duschanbe … ”), labels data of which were published by Zerche (1992).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFF1F84BFF65F913001BF80C.taxon	discussion	Remarks. Paratrichodromeus punctulatus (Fig. 13) is known from Tajikistan and Uzbekistan (Fig. 15).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFF2F848FF65FF4E06FCFE31.taxon	description	(Figs. 14 – 15)	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFF2F848FF65FF4E06FCFE31.taxon	materials_examined	Material examined. KYRGYZSTAN: 1 ♂: Zaalayskiy Mts., Berk-Suu R., 20 km W Daraut-Kurgan. 13.07.1995. S. V. Ovchinnikov leg. (cSh).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
03E487DEFFF2F848FF65FF4E06FCFE31.taxon	discussion	Remarks. Paratrichodromeus sukranensis (Fig. 14) was originally described from two localities in Tajikistan. It is known also from Kyrgyzstan (Schülke & Smetana 2015) (Fig. 15).	en	Shavrin, Alexey V. (2024): New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5446 (2): 236-246, DOI: 10.11646/zootaxa.5446.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5446.2.5
