taxonID	type	description	language	source
03E487F3FFF3FFD14F05031EFAFCF81C.taxon	description	(Figs 1 A, B; 6 A, B; 7 B)	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF3FFD14F05031EFAFCF81C.taxon	materials_examined	TYPE MATERIAL. — Not located. According to Vogel & van Rooijen (2011 a), the specimen examined by Wall was from the collection of St. Joseph’s College, Darjeeling, India. TYPE LOCALITY. — “ Darjeeling, ” State of West Bengal, India, by inference; unknown collector. MATERIAL EXAMINED. — India • 1 ♀; Assam, Dibrugarh; BMNH 1908.6.23.27 • 1 ♀; Mizoram, Aizawl District, Durtlang; 23 ° 14 ’ 0 ” N, 92 ° 43 ’ 31 ” E; 1215 m a. s. l.; 6. VI. 2017; Lal Biakzuala leg.; GenBank: MT 635656; MZMU 951 • 1 ♂; MZMU 1066; Mizoram, Siaha District, Phura; 22 ° 13 ’ 57 ” N, 92 ° 54 ’ 45 ” E; 226 m a. s. l.; 7. VIII. 2016; M. Vabeiryureilai leg. • 1 ♀; Mizoram, Mamit District, Reiek-kai; 23 ° 42 ’ 43 ” N, 92 ° 39 ’ 49 ” E; 150 m a. s. l.; 16. III. 2013; H. T. Lalremsanga leg.; MZMU 1077 • 1 ♂; Mizoram; Aizawl District, Sailam; 23 ° 21 ’ 10 ” N, 92 ° 47 ’ 53 ” E; 1312 m a. s. l.; 15. V. 2014; H. T. Lalremsanga leg.; MZMU 1078 • 1 ♂; Mizoram, Serchhip District, Buangpui; 23 ° 42 ’ 15 ” N, 92 ° 45 ’ 0 ” E; 710 m a. s. l.; 4. VIII. 2017; Vanlal Hrima leg.; MZMU 1079 • 1 ♂; Mizoram, Aizawl District, Sateek; 23 ° 32 ’ 49 ” N, 92 ° 42 ’ 12 ” E; 909 m a. s. l.; 18. VIII. 2018; Lal Muansanga leg.; MZMU 1080 • 1 ♀; Mizoram, Mamit District, Reiek Community Reserved Forest; 23 ° 41 ’ 48 ” N, 92 ° 36 ’ 19 ” E; 1166 m a. s. l.; 21. V. 2017; G. Z. Hmar leg.; MZMU 1101 • 1 ♂; Mizoram, Aizawl District, Tlangnuam; 23 ° 42 ’ 23 ” N, 92 ° 42 ’ 54 ” E; 1040 m a. s. l.; 12. V. 2018; Lal Biakzuala leg.; GenBank: MT 647129; MZMU 1218 • 1 ♂; Mizoram, Lunglei District, Thenhlum; 23 ° 11 ’ 51 ” N, 92 ° 34 ’ 59 ” E; 687 m a. s. l.; 17. VIII. 2014; H. T. Lalremsanga leg.; MZMU 1327; GenBank: MT 311195 • 1 ♂; Mizoram, Aizawl District, MZU campus; 23 ° 44 ’ 15 ” N, 92 ° 39 ’ 45 ” E; 789 m a. s. l.; 21. V. 2019; Lalrinsanga leg.; GenBank: MT 647128, MT 711166; MZMU 1370 • 1 ♀; Mizoram, Aizawl District, Maubuang; 23 ° 29 ’ 15 ” N, 92 ° 42 ’ 43 ” E; 1012 m a. s. l.; Denzil Lalneihsanga leg.; GenBank: MT 635657; MZMU 1432 • 1 ♂; Mizoram, Khawzawl District, Khawzawl; 23 ° 32 ’ 34 ” N, 93 ° 10 ’ 51 ” E; 1225 m a. s. l.; 7. VII. 2018; H. T. Lalremsanga leg.; MZMU 1475 • 1 ♂; Mizoram, Aizawl District, Kepran; 23 ° 56 ’ 40 ” N, 92 ° 55 ’ 50 ” E; 1360 m a. s. l.; 9. III. 2018; H. T. Lalremsanga leg.; MZMU 1507 • 1 ♀; Mizoram, Saitual District, Tamdil National Wetland; 23 ° 44 ’ 24 ” N, 92 ° 57 ’ 5 ” E; 774 m a. s. l.; H. T. Lalremsanga leg.; GenBank: MT 634738; MZMU 1509 • 1 ♀, 1 ♂; Mizoram, Serchhip District, Leng; 23 ° 9 ’ 20 ” N, 93 ° 7 ’ 14 ” E; 1230 m a. s. l.; H. T. Lalremsanga leg.; MZMU 1510, MZMU 1529 • 1 ♂; Mizoram, Serchhip District, Khawlailung; 23 ° 13 ’ 12 ” N, 92 ° 59 ’ 33 ” E; 1035 m a. s. l.; H. T. Lalremsanga leg.; MZMU 1516 • 1 ♂; Mizoram, Aizawl District, Sesawng; 23 ° 45 ’ 34 ” N, 92 ° 51 ’ 14 ” E; 853 m a. s. l.; 15. VIII. 2015; Lalrinsanga leg.; MZMU 1566 • 1 ♂; Mizoram, Aizawl District, Daifim resort; 23 ° 47 ’ 59 ” N, 92 ° 37 ’ 29 ” E; 70 m a. s. l.; 11. VI. 2019; H. Laltlanchhuaha leg.; MZMU 1598 • 1 ♂; Mizoram, Aizawl District, Tanhril; 23 ° 44 ’ 12 ” N, 92 ° 40 ’ 33 ” E; 958 m a. s. l.; 16. IX. 2019; Lalrinsanga leg.; MZMU 1602 • 1 ♂; Mizoram, Mamit District, Dampa Tiger Reserve; 23 ° 43 ’ 28 ” N, 92 ° 23 ’ 40 ” E; 255 m a. s. l.; 10. VI. 2016; Ht. Decemson leg.; MZMU 1603 • 1 ♀; Mizoram, Aizawl District, Sialsuk; 23 ° 23 ’ 50 ” N, 92 ° 44 ’ 46 ” E; 1255 m a. s. l.; 7. X. 2019; Angshuman Das leg.; GenBank: MT 274567; MZMU 1629. Myanmar • 1 ♂; Sagaing Division; CAS 210338; GenBank: KX 660183, KX 660456 (Figueroa et al. 2016) • 1 ♀; Ayeyarwady Division; GenBank: KX 660182, KX 660455; CAS 222690 (Figueroa et al. 2016). Laos • 2 ♀; no locality information; MNHN 1896.650; MNHN 1896.651 • 1 ♀; Phongsaly Province, Buon Tai; MNHN 2004.0253 • 1 ♀; Phongsaly Province, Long Nai; MNHN 2004.0254; • 2 ♂; Phongsaly Province; MNHN 2004.0263; MNHN 2004.0251. Vietnam • 1 ♀; Tonkin; MNHN 1948.82.	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF3FFD14F05031EFAFCF81C.taxon	diagnosis	DIAGNOSIS (BASED ON NEW MATERIAL). — From a total of 34 Dendrelaphis specimens collected from the Mizoram State, 22 of them were morphologically diagnosed as D. proarchos based on the combination of: 1) bronze-coloured body with bluish interstitial skin and a black lateral stripe separating cream dorsolateral stripe from whitish underside; 2) enlarged vertebral scales; 3) Ve 180 - 190 in males, 186 - 195 in females; 4) paired Sc 142 - 163 in males, 138 - 146 in females; 5) undivided As in 36.4 % of the specimens; 6) Ls single; 7) PrO 1; 8) PoO 2 - 3; 9) Te 2 + 2 or 2 + 3, rarely 3 + 2 or 1 + 2; 10) 4 th – 6 th SLE; 11) IF 9 - 10, rarely 11; 12) SL 8 - 9; 13) DSR 15: 15: 9 in males, 15: 15: 11 in females, rarely nine rows in females; and 14) average relative RTaL 0.35 (0.31 - 0.37) in males, 0.33 (0.33 - 0.34) in females. The hemipenis is not bilobate, and extends up to seven caudal plates; a prominent sulcus spermaticus runs up to the tip; a sinuous fold is present at about the middle of the organ, and the distal area is covered by somewhat evenly distributed small spines. The new materials divulged a new upper limit of Sc range i. e, 138 - 163 (sex pooled) vs 138 - 156 (Vogel & van Rooijen 2011 a). However, other morphological characters of the new materials are consistent with the specimens from Assam (Northeast India), Myanmar, Laos and Vietnam (also see Vogel & van Rooijen 2011 a).	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF3FFD14F05031EFAFCF81C.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Northeast India, Bangladesh, Myanmar, northern Laos, and northern Vietnam (Vogel & van Rooijen 2011 a). In this study, we documented D. proarchos from 22 different localities at the elevation range between 60 - 1360 m a. s. l. from Mizoram, Northeast India (Fig. 5; Appendix 1).	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF1FFD34F07003CFC64FA02.taxon	description	(Fig. 2 A, B)	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF1FFD34F07003CFC64FA02.taxon	materials_examined	TYPE MATERIAL. — Holotype. India • BMNH (no voucher number), a 690 - 700 mm specimen (native, June, 1907) (fide Wallach et al. 2014). TYPE LOCALITY. — India, Sadiya on the Bramaputra 70 odd miles north-east of Dibrugarh (Wall 1908), now Sadiya, Assam State, NE India, 27 ° 50 ’ N, 95 ° 40 ’ E, elevation 125 m a. s. l. (Wallach et al. 2014). MATERIAL EXAMINED. — India • 1 ♀; Mizoram, Kolasib District, Khuangpuilam; 24 ° 12 ’ 13 ” N, 92 ° 41 ’ 10 ” E; 665 m a. s. l.; 7. X. 2019; Jacob Lalhriatrenga leg.; GenBank: MT 711167, MT 814708, MT 731341, MT 731342; MZMU 1659 • 1 ♂; Mizoram, Aizawl District, MZU campus; 23 ° 44 ’ 14 ” N, 92 ° 39 ’ 47 ” E; 800 m a. s. l.; 6. VIII. 2020; Michael Vanlalchhuana leg.; MZMU 1812 • 1 ♂; Mizoram, Champhai District, Champhai Jail veng; 23 ° 28 ’ 4 ” N, 93 ° 19 ’ 0 ” E; 1420 m a. s. l.; 20. I. 2021; Tbc. Lalpekthara leg.; MZMU 2051.	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF1FFD34F07003CFC64FA02.taxon	diagnosis	DIAGNOSIS (BASED ON NEW MATERIAL). — Specimens of D. biloreatus (N = 4) are diagnosed by the combination of: 1) brownish dorsum with a black stripe on each side of head extending up to the neck; 2) upper and lower lips and chin yellowish; 3) Ve 203 in females, 190 - 198 in male; 4) paired Sc 137 - 151 in males, 102 in damaged tail female; 5) divided As; 6) Ls 1 - 2; 7) PrO 1; 8) PoO 2; 9) Te 1 + 2, 2 + 2, 2 + 3; 10) 4 th – 5 th SLE; 11) IF 10; 12) SL 8; 13) DSR 13: 13: 11; and 14) average RTaL 0.32 (0.26 - 0.36) in males, and 0.28 in the damaged tail female. The new materials from this study updated the known morphological characters as follows: upper limit of Ve 190 - 203 vs 192 (Wall 1908) and 187 - 199 as D. gorei (Smith 1943); lower limit of Sc 137 - 151 vs 147 (Wall 1908) and 139 - 154 as D. gorei (Smith 1943); Te 1 + 2, 2 + 2, 2 + 3 vs 1 + 1 (Wall 1908; Smith 1943). In addition, we provide brief comparative morphological data between D. cyanochloris, D. proarchos, and D. biloreatus from other zoological collections, along with the new Dendrelaphis specimens collected from Mizoram State (Table 1). See Appendix 1 for the detailed collection records, and Appendix 2 for the morphometry and scalation data of Dendrelaphis species collected from Mizoram State, India.	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF1FFD34F07003CFC64FA02.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — This species inhabits the Assam Valley with the adjoining Himalayan foothills, and northern parts of Myanmar near the border with Assam (Vogel & van Rooijen 2011 b). In this work, D. biloreatus is documented from four localities in Mizoram state at the elevation range of 443 - 1,420 m a. s. l. (Fig. 5; Appendix 1)	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF1FFD34EF207BAFE1AF8EB.taxon	description	(Figs 1 C; 7 A)	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF1FFD34EF207BAFE1AF8EB.taxon	materials_examined	TYPE MATERIAL EXAMINED. — Lectotype (as designated by Vogel & van Rooijen 2007). India • 1 ♂; BMNH 1940.3.4.27. TYPE LOCALITY. — Based on the designation of a lectotype by Vogel & van Rooijen (2007): “ Darjeeling ”, West Bengal State, India. The original description of D. cyanochloris was partly based on specimens from India (Eastern Himalaya and Assam) and “ Burma ”, now Myanmar, as well as specimens of D. humayuni Tiwari & Biswas, 1973. M ATERIAL EXAMINED. — India • 1 ♀; Mizoram, Saitual District, Keifang; 23 ° 39 ’ 31 ” N, 92 ° 57 ’ 38 ” E; 1035 m a. s. l.; 3. VII. 2009; H. T. Lalremsanga leg. MZMU 153 • 1 ♂; Mizoram, Aizawl District, Durtlang; 23 ° 47 ’ 35 ” N, 92 ° 44 ’ 1 ” E; 1104 m a. s. l.; 1. X. 2019; Lal Biakzuala leg.; GenBank: MT 711165; MZMU 985; • 1 ♂; Mizoram, Siaha District, Palak National Wetland; 22 ° 12 ’ 11 ” N, 92 ° 53 ’ 6 ” E; 290 m a. s. l.; 3. V. 2017; M. Vabeiryureilai leg.; MZMU 1102 • 1 ♂; Mizoram, Mamit District, Dampa Tiger Reserve; 23 ° 41 ’ 16 ” N, 92 ° 26 ’ 43 ” E; 454 m a. s. l.; 9. V. 2019; Ht. Decemson leg.; MZMU 1472 • 1 ♂; Mizoram, Mamit District, Lengpui; 23 ° 50 ’ 11 ” N, 92 ° 38 ’ 5 ” E; 210 m a. s. l.; 5. IV. 2016; Vanlal Hrima leg.; MZMU 1473 • 1 ♀; Mizoram, Mamit District, Reiek Community Reserved Forest; 23 ° 41 ’ 22 ” N, 92 ° 36 ’ 16 ” E; 1300 m a. s. l.; 23. VI. 2018; Lal Muansanga leg.; MZMU 1476 • 1 ♀; Mizoram, Aizawl District, MZU campus; 23 ° 44 ’ 17 ” N, 92 ° 40 ’ 6 ” E; 873 m a. s. l.; 11. VIII. 2019; Angshuman Das leg.; MZMU 1560 • 1 ♂; Mizoram, Aizawl District, Sawleng; 23 ° 58 ’ 52 ” N, 92 ° 56 ’ 0 ” E; 1107 m a. s. l.; 4. VII. 2009; Tbc. Lalbiaknunmawia leg.; MZMU 1593. Myanmar • 1 ♂; Kachin State, Putao District; CAS 221428; GenBank: KX 660454 (Figueroa et al. 2016).	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF1FFD34EF207BAFE1AF8EB.taxon	diagnosis	DIAGNOSIS (BASED ON NEW MATERIAL). — The specimens of D. cyanochloris (N = 8) are diagnosed by the combination of: 1) bronzecoloured body with conspicuous bluish interstitial skin, large black postocular streak from eye to nape, absence of lateral stripes along the body, and greenish underside; 2) Ve 184 - 207 in males, 183 - 210 in females; 3) paired Sc 138 - 153 in males, 138 - 144 in females; 4) divided As; 5) Ls single; 6) PrO 1; 7) PoO 2; 8) Te 2 + 2 or 3 + 2, rarely 1 + 2; 9) 4 th- 6 th SLE; 10) IF 9 - 10; 11) SL 9; 12) DSR 15: 15: 11, rarely nine rows; and 13) average RTaL 0.32 (0.30 - 0.36) in males, 0.29 (0.25 - 0.33) in females. The new materials from Mizoram, Northeast India enlarged the following morphological attributes (values for the lectotype are given in parentheses): Ve 184 - 207 in males and 183 - 210 in females (201); DSR at one head length before the vent 11, and rarely nine (11); SLE 4 th- 6 th (5 th- 6 th).	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
03E487F3FFF1FFD34EF207BAFE1AF8EB.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — India (Darjeeling and Mizoram), the range of distribution is also discussed. In this study, specimens of D. cyanochloris were documented from 10 different localities at elevations between 100 - 1300 m a. s. l. from Mizoram, Northeast India (Fig. 5; Appendix 1).	en	Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Tariang, Angshuman Das, Vabeiryureilai, Mathipi, Muansanga, Lal, Hrima, Vanlal, Kumar, Vikas, Kundu, Shantanu, Purkayastha, Jayaditya, Vogel, Gernot (2022): Contributions to the taxonomic status and molecular phylogeny of Asian Bronzeback Snakes (Colubridae, Ahaetuliinae, Dendrelaphis Boulenger, 1890), from Mizoram State, Northeast India. Zoosystema 44 (7): 177-196, DOI: 10.5252/zoosystema2022v44a7
