taxonID	type	description	language	source
03E587F7AC14FFA49E9EFF0D39E7F7CF.taxon	materials_examined	Type: — Vietnam [Indo-China], Tonkin, route de Vinh Yen au Tam Dao, May 1935 (fl.), A. Pételot 5684 (lectotype, first step, designated by Sleumer 1955: A; second step, here designated: barcode A 00035444 (fig. 1 A); isolectotypes: A!, no barcode (fig. 1 B); HNU!; NY! 00284743; P [image!] P 00750613, P 00750614).	en	Wang, Yuer (2025): Taxonomic Notes on Helicia (Proteaceae) I: Revision for Two Species from Continental Asia with new synonym and typification. Phytotaxa 714 (2): 118-128, DOI: 10.11646/phytotaxa.714.2.2, URL: https://doi.org/10.11646/phytotaxa.714.2.2
03E587F7AC14FFA49E9EFF0D39E7F7CF.taxon	description	Amended Description: Trees 3 – 8 m, branches and leaves glabrous. Leaves alternate, simple; petiole 1 – 2.5 cm; pulvinus present at base of petiole, 2 – 5 mm in length, woody; leaf blade glabrous, lanceolate to oblong-lanceolate, 8 – 20 cm × 2 – 5.5 cm, coriaceous, base cuneate to attenuate, margin entire, sparsely serrate, to serrate, apex acuminate; secondary veins 7 – 11 on each side of midvein, slightly impressed adaxially, slightly raised abaxially, anastomosing; the first pair of secondary veins arising from leaf base and located close to leaf margin. Inflorescence single or cluster of 2 (– 4) racemes that matures acropetally, ramiflorous or cauliflorous, 8 – 21 cm in length; rachis and peduncle sparsely ferrugineo-pilose; peduncle of flower pair 0 – 2 mm long; bract of flower pair triangular, ca. 1 mm. Pedicel sparsely ferrugineo-pilose, 1 – 6 mm long; receptacle erect, 0.8 – 1.5 mm wide; floral bract lanceolate to filiform, ca. 0.8 mm; perianth tube 4 - lobed, 1.5 – 4 cm in length, sparsely white-pilose, pink to magenta, turning light purple at anthesis, apex yellowish; stamens adnate to perianth lobes; anthers dehiscing with longitudinal slit; style 1.2 – 3.8 cm long with stigma dilated; ovary sessile, glabrous; hypogynous glands 4, connate; Fruits erect to oblique, one-seeded, ellipsoid to near globose, radial or bilateral symmetrical, 2 – 4 cm long, 1 – 3 cm wide, fleshy indehiscent (rarely dehiscent along ventral suture), basally contracted into a short stipe, apically markedly beaked with style and stigma persistent; ventral suture present; pericarp 1 – 2 mm thick; exocarp brown, pitted; mesocarp fleshy; cotyledons purple to magenta. Flower April to June; fruit August to February the next year.	en	Wang, Yuer (2025): Taxonomic Notes on Helicia (Proteaceae) I: Revision for Two Species from Continental Asia with new synonym and typification. Phytotaxa 714 (2): 118-128, DOI: 10.11646/phytotaxa.714.2.2, URL: https://doi.org/10.11646/phytotaxa.714.2.2
03E587F7AC14FFA49E9EFF0D39E7F7CF.taxon	distribution	Distribution and Habitat: China (SW Guangxi) to N Vietnam, elevation below 900 m. On mountain slopes, near streams in lowland tropical monsoon rainforests and cloud forests. Assessment of Conservation Status: The synonym Helicia dongxingensis was under “ VU (Vulnerable), A 2 c ” in China’s Red List of Biodiversity: Higher Plants (2020). For an assessment of H. cauliflora in its full course of distribution, the author suggests a rank of “ VU (Vulnerable), B 1 ab ” with an estimated EOO of 15,869 km 2, based on the standards of IUCN (IUCN 2025). The lowland rainforest habitat of Helicia cauliflora is highly fragmented and has severely declined during the last 20 years, resulting from logging and deforestation of the tropical rainforest in the Sino-Vietnamese border region (Woodruff 2010). The decline of habitat and populations for this species is unlikely to be reversible. Additional Specimens examined: CHINA. Guangxi: Fangchenggang City, Fangcheng District, Nasuo Town [ãūē], Pingshuiding Reservoir [ṿAEṈAEṽ], 467 m, 21.75 ° N, 108.08 ° E, 20 January 2018 (fr.), Bo Liu et al. 221 (JJF [image!] JJF 00045230); Nasuo Town, Lanshan Erta [ÎƜ 二塔]. 440 m, 108.0378 ° E, 21.7569 ° N, 22 June 2021 (st.), Yonggang Liu et al. 1502 | 4 (PE! Sheet No. 2771802); Nasuo Town, Nasuo Village [ãūť], Meizaitian [妹仔 H], 180 m, 21.7491 ° N, 108.099996 ° E, 13 January 2021 (fr.), Yonggang Liu et al. 243 | 5; Nasuo Town, Shangyuegou [上ėŝ], Siyan Road [Ŀân], 370 m, 21.764208 ° N, 108.08525 ° E, 29 June 2021 (fl.), H. N. Qin et al. 1578 | 4 (PE! Sheet No. 2771801); Nasuo Town, near front gate of Conghuanggou Scenic Area [DZ ẽŝ], 28.05 m, 21.7441223 ° N, 108.0585607 ° E, 27 April 2025 (fl. et fr.), Z. X. Ma 2304 (BAZI! 001611 to 001618); Fangchenggang City, Mt. Shiwandashan [十万大Ɯ], Pingwang Township, Vicinity of Shaping Village [ŭ ṿť], 12 February 1944 (fr.), S. H. Chun 4837 (IBSC! 0087478); Shangsi County [上ō县], Mt. Shiwandashan [十万大Ɯ], Fenghuang Township [凤凰 乡], Mt. Lugui [ṻfflƜ], 12 February 1944 (st.), S. H. Chun 4553 (IBSC! 0206122);	en	Wang, Yuer (2025): Taxonomic Notes on Helicia (Proteaceae) I: Revision for Two Species from Continental Asia with new synonym and typification. Phytotaxa 714 (2): 118-128, DOI: 10.11646/phytotaxa.714.2.2, URL: https://doi.org/10.11646/phytotaxa.714.2.2
03E587F7AC14FFA49E9EFF0D39E7F7CF.taxon	materials_examined	VIETNAM. B ắc Ninh: B ắc Giang, SƠn Đ ộng, Tu ấn M ậu, 12 April 2013 (fl.), N. T. Cuong and D. D. Cuong VN 2357 (HN!). Phú Th ọ: locality unclear, Nov 1964 (fr.), s. col. 1704 (HNU!). Qu ảng Ninh: Tiên Yên, Kau Nga Shan, and Vicinity, 13 – 22 December 1936 (fr.), W. T. Tsang 27386 (A!; IBSC! 0205809; IBK! IBK 00386384; HITBC! 053260); Vĩnh Phúc: Tam Dao National Park, along trail in the southern outskirts of Tam Dao town, 870 m, 21 º 26 ’ 54 ” N, 105 º 38 ’ 50 ” E, 2 May 2013 (fl.), Ulf Swenson et. al. 1542, (S [image!], Herb. No. S 14 - 26136; P [image!] P 00910975). Tam Đ ảo, 30 May 1977 (fl.), Vũ xuân PHƯƠng 164 (HN!); ibid., 3 August 1976 (fr.), LX-VN 983 (HN!); ibid., s. ann. (fl.), Eberhardt 4967 (P [image!] P 02309918).	en	Wang, Yuer (2025): Taxonomic Notes on Helicia (Proteaceae) I: Revision for Two Species from Continental Asia with new synonym and typification. Phytotaxa 714 (2): 118-128, DOI: 10.11646/phytotaxa.714.2.2, URL: https://doi.org/10.11646/phytotaxa.714.2.2
03E587F7AC13FFA09E9EFF673928F7F1.taxon	description	5: 194 (2003). Fig. 3 – 5. Type: — Myanmar: Lower Burma, Martaban Hills, between Sittang and Sadween watershed, ca. 1200 m, Brandis 1408 (lectotype, designated by Sleumer 1955: 66: K [image!] K 00487566; isolectotype, CAL [image!] CAL 0000022093).	en	Wang, Yuer (2025): Taxonomic Notes on Helicia (Proteaceae) I: Revision for Two Species from Continental Asia with new synonym and typification. Phytotaxa 714 (2): 118-128, DOI: 10.11646/phytotaxa.714.2.2, URL: https://doi.org/10.11646/phytotaxa.714.2.2
03E587F7AC13FFA09E9EFF673928F7F1.taxon	distribution	Distribution and Habitat: China (S. Yunnan, Guangxi), S Myanmar, Thailand, N Vietnam, elevation 400 – 1900 m. In tropical montane rainforests, damp cloud forests, tea plantations, on the edge of forests, and on roadsides. Assessment of Conservation Status: LC, with a wide range of distribution and relatively stable populations in Northern Indo-Chinese floristic region by the definition of IUCN. However, it should be noted that the rubber-tea agriculture in tropical areas may pose a threat to the habitat of this species. Additional Specimens examined: CHINA. Guangxi: Baise City, Napo County [ã 坡县], Pingmeng Town [ṿā 区], Mengda Gongshe [āżȓfl], 9 November 1965 (st.), Guangxi Forestry Expedition 46154 (IBK! IBK 00114084); Nanning City, Wuming District [ẍạ 区], Liangjiang Town [两ȗē], Mt. Damingshan Nature Preserve [大ffiƜ保 Ễ 区], Chaoyang Station [fi ẹȅ], Chaoyanggou [fi ẹŝ], 640 m, 23.49870 ° N, 108.40031 ° E, 19 October 2022 (st.), Qiu-Yan Lu et al. DMB 1162 (PE! 02419681, 02422075); ibid., 570 m, 23.49872 ° N, 108.39643 ° E, 21 February 2023 (fr.), Min Su & Shi-Li Chang DMB 1455 (PE! 02419204, 02420923); ibid., valley next to Chaoyanggou [fi ẹ ŝ], 400 m, 15 October 2011 (fr.), Lei Wu & Jin-Cai Yang D 3211 (IBK! IBK 00208713); Mt. Damingshan [大ffiƜ], Ganlan River [dzĸñ], 565 m, 31 May 2011 (fr.), Lei Wu D 2381 (IBK! IBK 00208712). Yunnan: Honghe Prefecture, Hekou County [ñ 口], Luopudi [ḙä 地], 4 August 1959 (fl.), Wei-Xin Liu 482 (HITBC! HITBC 0071636; KUN! 0485318, 0485319; PE! 00874812); Jinping County [ae ṿ 县], Huangjia Zhai [a ẋḛ], 1500 m, 22 ° 47 ’ N, 103 ° 13 ’ E, 11 August 1951 (fl.), P. I Mao 304 (HITBC! HITBC 0071716; PE! 00874813); Jinping County, locality unclear, 27 April 1956 (fl.), Sino-Soviet Joint Yunnan Expedition 852 (KUN! 0485321); Lvchun County [ũǝ 县], Ma-yu [ṉī], 1 November 1995 (fr.), S. K. Wu et al. 3385 (KUN! 0485327); Lvchun County, Sanmeng Township [三 % 乡], across from Lagu [š 姑], Zeng-Hong Yang & Zheng-Wei Lv 90 - 1261 (KUN! 0485331). Puer City, Szemao [now Puer City], locality unclear, s. ann. (fr.), A. Henry 12808 (K [image!] K 004487470; MO!; NY! 04390724); ibid., s. ann. (fl.), A. Henry 13116 (K [image!] K 004487469, K 004487470; MO!; NY! 04390722, 04390723); Jiangcheng County [ȗ ṵ 县], Jiahe Township [嘉 ‡ 乡], 1547 m, 22.7563809 ° N, 101.9161193 ° E, 4 March 2025, J. D. Ya 25 CS 28378 (BAZI!). Xishuangbanna Prefecture, Jinghong City [ḁăṄ], Puwen Township [ḃÿ 乡], 22 ° 27 ’ N, 101 ° 4 ’ E, 3 May 1980 (fr.), Jing-Yun Cui et al. 18198 (HITBC! HITBC 0071719, HITBC 0071720); Jinio Xiang [ḋë 乡], Longpa [ẑü], 1250 m, 21 ° 59 ’ N, 101 ° 4 ’ E, 29 August 1977 (fl.), G. D. Tao 16044 (HITBC! HITBC 0071731); ibid., 1250 m, 18 March 2004 (fl.), G. D. Tao 16044 (PE!); Damenglong Town [大勐 ẑē], Mengsong Village [勐 Ṫť], 21 ° 30 ’ N, 100 ° 30 ’ E, 12 October 1978 (fr.), G. D. Tao et al. 19969 (HITBC! HITBC 0071726); ibid., 1580 m, 3 December 2009 (fr.), J. T. Yin 1293 (HITBC! HITBC 0029945); Menghai County [勐ª县], near Liusha River [ĭŭñ], 1200 m, 3 March 1957 (fr.), Sino-Soviet Joint Yunnan Expedition 5539 (KUN! 0485322); Menghai County, Mt. Nannuo [南IJƜ], 1700 – 1900 m, 29 April 1967 (fr.), Yunnan Second Team 037 (PE! 02429188); ibid., 1800 m, 25 May 1998 (fl.), T. G. Gao 1709 (PE! 02429189); ibid., 1700 m, 19 June 1998 (fl.), T. G. Gao 1483 (PE! 02429187); ibid., 1890 m, 19 August 2011 (fl.), Jian W. Li 844 (HITBC! HITBC 0071728); ibid., near submit, 1700 m, 2 April 2004 (fr.), H. Wang 6968 (HITBC! HITBC 0071718); ibid., 15 September 2006 (fr.), H. Wang 8521 (HITBC! HITBC 0071729); ibid., en route from Banpo Laozhai [半坡ƭ ḛ] to Duoyi Zhai [多依 ḛ], near scenic tower, 1651.09 m, 21.5106784 ° N, 100.5060661 ° E, 2 July 2024 (fl.), Y. E. Wang 901 (BAZI!); Menghai County, Bulang Township [ssș 乡], 900 m, 17 July 2013 (fl.), Jian W. Li 3516 (HITBC! HITBC 0071727); Mengla County [勐š县], Mt. Nangong [南ĪƜ], 1800 m, 21 ° 38 ’ N, 101 ° 30 ’ E, May 1987 (fl.), G. D. Tao 14580 (HITBC! HITBC 0071721, HITBC 0071722); ibid., s. ann., Rare Species Group 33834 (HITBC! HITBC 0071730); Mengla County, Menglun Town [勐 ṙē], Xiaola Road [小šȓn] 56 Km, 21 ° 55 ’ N, 101 ° 13 ’ E, 3 September 1975 (fr.), G. D. Tao 13771, 13772 (HITBC! HITBC 0071723, HITBC 0071724; KUN! 0485532); Mengla County, Jingpiao Village [ḁȃť], 6 December 1983 (fr.), Nature Reserve Expedition 34972 (HITBC! HITBC 0071725), ibid., 7 December 1983 (fr.), Expedition 25214 (HITBC! HITBC 0071717).	en	Wang, Yuer (2025): Taxonomic Notes on Helicia (Proteaceae) I: Revision for Two Species from Continental Asia with new synonym and typification. Phytotaxa 714 (2): 118-128, DOI: 10.11646/phytotaxa.714.2.2, URL: https://doi.org/10.11646/phytotaxa.714.2.2
03E587F7AC13FFA09E9EFF673928F7F1.taxon	materials_examined	THAILAND. Chiang Mai: Mae Wang District, 1275 m, J. F. Maxwell 04 - 799 (A!); ibid., J. F. Maxwell 91 - 244 (A!); Lampang: Nuang Bahn (Pan) District, Jae Sawn National Park below Bah Miang Village and near Mae Bahn Stream, 925 m, 30 November 1995, J. F. Maxwell 95 - 1192 (A!); Nakhon Nayok: Muang District, Khao Yai National Park, near km 1, along the road to Khao Khieo, 750 m, 14 ° 24 ’ N, 101 ° 22 ’ E, 17 June 2002 (fl.), A. Boonkongchart 122 (A!); Nan: Pua District, Doi Phu Kha National Park, 1700 m, 19 ° 10 ’ N, 101 ° 07 ’ E, 26 May 2000 (fl.), P. Srisanga 1448 (P [image!] P 02786203); ibid., 1500 m, 19 ° 13 ’ N, 101 ° 05 ’ E, 13 January 2000 (fl. et fr.), P. Srisanga 1294 (P [image!] P 02786245); VIETNAM. Lào Cai: Chapa [Sa Pa], 1400 m, July 1931, A. Pételot 2721 (HNU!); ibid., 1500 m. July 1933. A. Pételot 4791 (A!; MO! MO 1204270; NY!, two sheets; P [image!] P 02309672, P 02309674). Ibid. July 1943. A. Pételot 8496 (P 02309673, P 02309675). Vĩnh Phúc: Tam Dao, 7 February 1965 (fr.), Sino-Vietnamese Expedition 1963 (KUN! 1293256; PE! 01725431).	en	Wang, Yuer (2025): Taxonomic Notes on Helicia (Proteaceae) I: Revision for Two Species from Continental Asia with new synonym and typification. Phytotaxa 714 (2): 118-128, DOI: 10.11646/phytotaxa.714.2.2, URL: https://doi.org/10.11646/phytotaxa.714.2.2
