taxonID	type	description	language	source
03E587FA3312FFA768D82A902DCEFE13.taxon	materials_examined	Type species: Hysterodus bloetei Dlabola, 1982.	en	Gnezdilov, Vladimir M., Bieman, F. M. (2025): The smallest species of the family Issidae-a new genus for Hysterodus bloetei Dlabola, 1982 (Hemiptera: Fulgoroidea). Zootaxa 5683 (3): 443-450, DOI: 10.11646/zootaxa.5683.3.7, URL: https://doi.org/10.11646/zootaxa.5683.3.7
03E587FA3312FFA768D82A902DCEFE13.taxon	diagnosis	Diagnosis. Robust, brachypterous, fore wings usually reaching hind margin of tergite VI, sometimes only hind margin of tergite III. Metope slightly elongate, with strong median carina running from its upper margin throughout postclypeus and with weak sublateral carinae running from the same point with median carina on its upper margin, but not reaching metopoclypeal suture. Coryphe slightly transverse. Fore wings spread out, with its basal parts before anterior radial vein turned downwards nearly at right angle to the rest of the corium; without hypocostal plate; clavus almost as long as whole wing, opened. Hind wings rudimentary. Hind tibia with a single lateral spine medially. First metatarsomere larger than second one, elongate, with two latero-apical and five intermediate spines. Male pygofer vertically elongate, narrow. Phallobase horseshoe-shaped in lateral view, with two hardly sclerotized plates dorso-apically bearing teeth. Aedeagus not visible above phallobase, with pair of long ventral hooks directed downwards. Capitulum of style on long neck, narrow in dorsal view. Hind margin of female sternite VII deeply concave medially. Female anal tube elongate, with pointed apex in dorsal view, convex in lateral view. Gonoplacs convex, without carinae. Anterior connective lamina of gonapophyse VIII with one distinct tooth in apical group. Endogonocoxal process with a simple narrow apical part. Posterior connective laminae of gonapophyses IX wide in ventral and lateral views, with two widely spaced lobes of median field and small lateral fields. Distal parts of posterior connective laminae of gonapophyses IX curved at obtuse angle in ventral view.	en	Gnezdilov, Vladimir M., Bieman, F. M. (2025): The smallest species of the family Issidae-a new genus for Hysterodus bloetei Dlabola, 1982 (Hemiptera: Fulgoroidea). Zootaxa 5683 (3): 443-450, DOI: 10.11646/zootaxa.5683.3.7, URL: https://doi.org/10.11646/zootaxa.5683.3.7
03E587FA3312FFA768D82A902DCEFE13.taxon	etymology	Etymology. Generic name is derived from the Greek (μωρό) for baby referring to small size of species. Neutral in gender. Composition and distribution. Monotypical genus known from southeastern Spain.	en	Gnezdilov, Vladimir M., Bieman, F. M. (2025): The smallest species of the family Issidae-a new genus for Hysterodus bloetei Dlabola, 1982 (Hemiptera: Fulgoroidea). Zootaxa 5683 (3): 443-450, DOI: 10.11646/zootaxa.5683.3.7, URL: https://doi.org/10.11646/zootaxa.5683.3.7
03E587FA3311FFA068D82E7C2A2FFC2F.taxon	description	(Figs 1 – 20)	en	Gnezdilov, Vladimir M., Bieman, F. M. (2025): The smallest species of the family Issidae-a new genus for Hysterodus bloetei Dlabola, 1982 (Hemiptera: Fulgoroidea). Zootaxa 5683 (3): 443-450, DOI: 10.11646/zootaxa.5683.3.7, URL: https://doi.org/10.11646/zootaxa.5683.3.7
03E587FA3311FFA068D82E7C2A2FFC2F.taxon	materials_examined	Material examined. Spain: 16 ♂, 13 ♀, Ciudad Real, 6 km NNW Ruidera, along CM 3115, N 39.03122 W 2.91030, 920 m, 26. V. 2016 (ZIN and CCB); 21 ♂, 19 ♀, same locality, 8. V. 2017 (ZIN and CCB); 16 ♂, 14 ♀, same locality, 17. V. 2018 (CCB); 3 ♂, 5 ♀, Ciudad Real, 8 km NNW Ruidera, near CM 3115, N 39.04506 W 2.94186, 880 m, 9. V. 2017 (CCB); 1 ♂, Ciudad Real, 1 km SW Ruidera, along N 430, N 38.97031 W 2.89434, 870 m, 18. V. 2018 (CCB); 2 ♂, 2 ♀, Albacete, 8 km SW Ossa de Montiel, Laguna de Ruidera, N 38.89160 W 2.76808, 820 m, 16. V. 2018 (CCB); 3 ♂, same locality, 19. V. 2018 (CCB); 5 ♂, 5 ♀, Valencia, 6 km NW Villargordo del Cabriel, near Embalsa de Contreras (water reservoir), N 39.55509 W 1.48678, 650 m, 22. V. 2018 (CCB), all leg. C. F. M. den Bieman. Emended description. Structure (Figs 1 – 6). Metope slightly elongate, lateral margins convex, with strong median carina running from its upper margin throughout postclypeus and with weak sublateral carinae running from the same point with median carina on its upper margin, but not reaching metopoclypeal suture which is strongly convex (Fig. 3). Upper margin of metope obtusely angulately concaved. Pedicel globular. Rostrum reaching hind coxae, 3 rd segment narrowing apically, slightly shorter than 2 nd one. Coryphe slightly transverse, 1.5 times as wide as long medially, grooved concave; anterior margin obtusely angulately convex; posterior margin concave; lateral margins keel-shaped (Figs 1, 4, 6). Eyes large, each eye is nearly as broad as coryphe width. Pronotum slightly shorter than coryphe medially, with strongly convex keel-shaped anterior margin and notched medially posterior margin. Paradiscal fields of pronotum narrow behind eyes. Paranotal lobes of pronotum wide. Mesonotum twice longer than pronotum medially, with weak lateral carinae; concave at scutellum. Brachypterous, fore wings reaching hind margin of tergite VI (in one female — only hind margin of tergite III) (Figs 1, 2, 4 – 6). Fore wings nearly round, spread out, with its basal parts before anterior radial vein turned downwards nearly at right angle to the rest of corium; without hypocostal plate. Clavus almost as long as whole wing, opened. Forewing vein branching sequence: R 4, furcating basally; r-m 1 – 4; M 2, furcating apically; m-cua 3; CuA 2, furcating apically. Pcu and first anal vein fused before middle of clavus. All veins relief. Hind wings rudimentary. Legs not flattened neither foliated. Hind tibia with a single lateral spine medially (in one male on left leg a second lateral spine present distally to the middle one) and seven spines apically (Figs 1, 4, 6). First metatarsomere larger than second one, elongate, with two latero-apical and five intermediate spines. Second metatarsomere with only two latero-apical spines. Coloration (Figs 1 – 7). Generally orange brown. Metope and pronotum with dense dark brown to black dots. Rostrum with black 3 rd segment. Pedicel dark below. Antenna with dark brown 3 rd segment and flagellum. Fore wings with dark brown dots and spots in cells. Femora and tibiae with brown to dark brown longitudinal stripes. Apices of leg spines, third tarsomeres, and claws black. Abdominal tergites with black spots surrounding setae. In females, gonocoxa VIII dark brown near to anterior connective lamina of gonapophyse VIII. Male genitalia (Figs 8 – 13). Pygofer vertically elongate, narrow, weakly convex medially in lateral view (Fig. 8). Anal tube large, enlarged medially (in dorsal view), 1.5 times as long as wide medially, with two rounded lateral processes (Figs 8, 13). Anal column short, 1 / 6 of whole anal tube. Phallobase strongly curved, horseshoe-shaped in lateral view, weakly sclerotized apically and covered by denticles, with two hardly sclerotized plates dorso-apically bearing teeth (Fig. 9, spl). Aedeagus not visible above phallobase, with pair of long, half of phallobase length, ventral hooks pointed apically and directed basad. Ventral phallobase lobe long and narrow, weakly concave apically (Fig. 10). Style with concave hind margin (Fig. 12). Capitulum of style on long neck (Fig. 12), narrow in dorsal view (Fig. 11), with large lateral tooth and distinct apical tooth. Female terminalia (Figs 7, 14 – 20). Hind margin of sternite VII deeply concave medially (Fig. 14). Anal tube elongate, 1.6 times as long as wide medially, nearly oval, with pointed apex in dorsal view (Fig. 17); convex in lateral view (Fig. 18). Anal column short. Gonoplacs convex, without carinae (Fig. 7). Gonocoxa VIII with lobe-shaped hind margin (Fig. 19). Anterior connective lamina of gonapophyse VIII wide, with two teeth in apical group (one distinct apical tooth and one smooth tooth below) and 4 – 5 keeled teeth in lateral group (Fig. 19). Endogonocoxal process with simple narrow apical part. Posterior connective laminae of gonapophyses IX wide in ventral and lateral views, with large and convex median field covered by small spines bearing two widely spaced lobes apically and with small lateral fields covered by microsetae (Figs 15, 16). Distal parts of the laminae curved at obtuse angle in ventral view (Fig. 15). Gonospiculum bridge large, elongate in lateral view (Fig. 16). Diverticulum ductus with slightly elongate bulb (Fig. 20). Total length. Males — 2.0 – 2.5 mm. Females — 2.5 – 2.9 mm.	en	Gnezdilov, Vladimir M., Bieman, F. M. (2025): The smallest species of the family Issidae-a new genus for Hysterodus bloetei Dlabola, 1982 (Hemiptera: Fulgoroidea). Zootaxa 5683 (3): 443-450, DOI: 10.11646/zootaxa.5683.3.7, URL: https://doi.org/10.11646/zootaxa.5683.3.7
